180 lines
19 KiB
XML
180 lines
19 KiB
XML
<document id="7D451E3CD521F176655DDDD05CDC3A8F" ID-DOI="10.11646/zootaxa.3683.2.3" ID-GBIF-Dataset="bb1cc4ed-5a7d-4c04-a042-2deb834b1d21" ID-ISSN="1175-5326" ID-Zenodo-Dep="219725" ID-ZooBank="1CDD7727-E1BF-4C46-B8FD-A13522FFC698" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460644285648" checkinUser="plazi" docAuthor="Dominiak, Patrycja & Alwin, Alicja" docDate="2013" docId="081B3D0DFFFBFFC342AB33E2001A89B1" docLanguage="en" docName="zt03683p144.pdf" docOrigin="Zootaxa 3683 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dasyhelea libanensis Dominiak, sp. nov." docType="treatment" docUuid="47B17A47-831F-47A1-B864-080FE059D111" docUuidSource="ZooBank" docVersion="10" lastPageNumber="138" masterDocId="F4224575FFFEFFC5423C302F02178965" masterDocTitle="Five new species and new records of biting midges of the genus Dasyhelea Kieffer from the Near East (Diptera: Ceratopogonidae)" masterLastPageNumber="144" masterPageNumber="133" pageNumber="138" updateTime="1698336543250" updateUser="plazi">
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<mods:titleInfo id="FF1BD766B6C668225CA8E27E2183CC8A">
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<mods:title id="C24BEF1AACD62C58F63C9AB00D31B513">Five new species and new records of biting midges of the genus Dasyhelea Kieffer from the Near East (Diptera: Ceratopogonidae)</mods:title>
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<mods:name id="11A8DEF3231D711F8976AE1CF27F3C4A" type="personal">
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<mods:namePart id="C9722530160D570B3C247660E3EC747C">Dominiak, Patrycja</mods:namePart>
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<mods:roleTerm id="1D46AF26C90ABCEAB2D0574A550430DF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6EA781368D4A2FA75DFCD2DA598B05B0">Alwin, Alicja</mods:namePart>
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<mods:typeOfResource id="7A1A9CB35776017CDD4C319D8B6E4B77">text</mods:typeOfResource>
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<mods:title id="D56B4276BC4E7424E8CE7A474B2468ED">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="C20DF5FE1E8765DE5EEAC1E02F1DB966">
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<mods:date id="38FCB84F97205B5AC3064E5CD47C5384">2013</mods:date>
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<mods:detail id="2B382589A5F3F7DD889827E4B92A312E" type="volume">
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<mods:number id="F16C21D5FC86225CC624D4C9D5AAB5F1">3683</mods:number>
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<mods:detail id="51C7B7D7CEA36808230AA3A4882AB792" type="issue">
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<mods:number id="A97A9D87220DBA977DDCBF4F82A07C4A">2</mods:number>
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<mods:classification id="87E1C3831B5361E7ADC4F5102E37C434">journal article</mods:classification>
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<mods:identifier id="3B017A6FF6F271C62CDCC4456B98472A" type="DOI">10.11646/zootaxa.3683.2.3</mods:identifier>
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<mods:identifier id="92BF40435D546304EF3FC83BB2D334BE" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="2762182AFB5D156A81E931F19860485D" type="Zenodo-Dep">219725</mods:identifier>
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<mods:identifier id="F3EA23994B873D687AB8C7250C02021E" type="ZooBank">1CDD7727-E1BF-4C46-B8FD-A13522FFC698</mods:identifier>
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<treatment id="081B3D0DFFFBFFC342AB33E2001A89B1" ID-DOI="http://doi.org/10.5281/zenodo.4500008" ID-GBIF-Taxon="119571708" ID-Zenodo-Dep="4500008" LSID="urn:lsid:zoobank.org:act:47B17A47-831F-47A1-B864-080FE059D111" httpUri="http://treatment.plazi.org/id/081B3D0DFFFBFFC342AB33E2001A89B1" lastPageId="6" lastPageNumber="138" pageId="5" pageNumber="138">
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<subSubSection id="C8A8DF90FFFBFFC042AB33E2031D8D62" pageId="5" pageNumber="138" type="nomenclature">
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<paragraph id="800D8C1BFFFBFFC042AB33E2006A8A82" blockId="5.[151,637,973,1031]" box="[151,637,973,999]" pageId="5" pageNumber="138">
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<heading id="DB453B77FFFBFFC042AB33E2006A8A82" bold="true" box="[151,637,973,999]" fontSize="11" level="1" pageId="5" pageNumber="138" reason="1">
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<emphasis id="B2C65009FFFBFFC042AB33E2006A8A82" bold="true" box="[151,637,973,999]" pageId="5" pageNumber="138">
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||
<taxonomicName id="47B2F798FFFBFFC042AB33E2000E8A82" ID-CoL="9HN7J" authority="Dominiak" authorityName="Dominiak" box="[151,537,973,999]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="libanensis" status="sp. nov.">
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<emphasis id="B2C65009FFFBFFC042AB33E203818A82" bold="true" box="[151,406,973,999]" italics="true" pageId="5" pageNumber="138">Dasyhelea libanensis</emphasis>
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Dominiak
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</taxonomicName>
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<taxonomicNameLabel id="A9F5ED72FFFBFFC0401C33E2006A8A82" box="[544,637,973,999]" pageId="5" pageNumber="138" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="800D8C1BFFFBFFC042AB33C0031D8D62" blockId="5.[151,637,973,1031]" box="[151,266,1007,1031]" pageId="5" pageNumber="138">
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||
(
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<figureCitation id="1889909EFFFBFFC042A333C003148D62" box="[159,259,1007,1031]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="5" pageNumber="138">Figs 6–8</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C8A8DF90FFFBFFC042AB341701B58C42" pageId="5" pageNumber="138" type="materials_examined">
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<paragraph id="800D8C1BFFFBFFC042AB341701B58C42" blockId="5.[151,1437,1079,2009]" pageId="5" pageNumber="138">
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<emphasis id="B2C65009FFFBFFC042AB3417035E8D35" bold="true" box="[151,329,1079,1104]" pageId="5" pageNumber="138">
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<typeStatus id="5F0932B9FFFBFFC042AB341702C58D35" box="[151,210,1080,1104]" pageId="5" pageNumber="138">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="5F0932B9FFFBFFC04368341803D68D2A" box="[340,449,1079,1103]" pageId="5" pageNumber="138" type="holotype">Holotype</typeStatus>
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: adult male.
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<emphasis id="B2C65009FFFBFFC04066341700DC8D35" bold="true" box="[602,715,1080,1104]" pageId="5" pageNumber="138">
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<collectingCountry id="F8A5CC8BFFFBFFC04066341700D18D35" box="[602,710,1080,1104]" name="Lebanon" pageId="5" pageNumber="138">Lebanon</collectingCountry>
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.
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</emphasis>
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Tyre, Tyre Coast Nature Reserve, N 33°
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<geoCoordinate id="E586EADCFFFBFFC04696341807018D2A" box="[1194,1302,1079,1103]" direction="east" orientation="longitude" pageId="5" pageNumber="138" precision="55" value="15.206">15.206 E</geoCoordinate>
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35°12.794,
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<date id="F40CAADBFFFBFFC042AB3473031B8D11" box="[151,268,1116,1140]" pageId="5" pageNumber="138" value="2012-05-06">06.V.2012</date>
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, net, leg. P. Dominiak (
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<collectionCode id="E6A314DEFFFBFFC04015347300938D11" box="[553,644,1116,1140]" pageId="5" pageNumber="138">BMNH</collectionCode>
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).
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<typeStatus id="5F0932B9FFFBFFC040A73472011B8D10" box="[667,780,1117,1141]" pageId="5" pageNumber="138" type="paratype">Paratypes</typeStatus>
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:
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<emphasis id="B2C65009FFFBFFC041203473019A8D11" bold="true" box="[796,909,1116,1140]" pageId="5" pageNumber="138">
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<collectingCountry id="F8A5CC8BFFFBFFC041203473019F8D11" box="[796,904,1116,1140]" name="Lebanon" pageId="5" pageNumber="138">Lebanon</collectingCountry>
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.
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</emphasis>
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Tyre, N 33°
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<geoCoordinate id="E586EADCFFFBFFC0461F347306988D11" box="[1059,1167,1116,1140]" direction="east" orientation="longitude" pageId="5" pageNumber="138" precision="55" value="15.535">15.535 E</geoCoordinate>
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35°12.726,
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<date id="F40CAADBFFFBFFC0471E347307808D11" box="[1314,1431,1116,1140]" pageId="5" pageNumber="138" value="2012-05-06">06.V.2012</date>
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, beach, net,
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<specimenCount id="96B44792FFFBFFC0431E345003978DF2" box="[290,384,1151,1175]" pageId="5" pageNumber="138" type="male">4 males</specimenCount>
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, leg. A. Alwin & P. Dominiak (UG). Tyre, Tyre Coast Nature Reserve, N 33°
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<geoCoordinate id="E586EADCFFFBFFC047123450078B8DF2" box="[1326,1436,1151,1175]" direction="east" orientation="longitude" pageId="5" pageNumber="138" precision="55" value="15.206">15.206 E</geoCoordinate>
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35°12.794,
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<date id="F40CAADBFFFBFFC04326348B03988DD9" box="[282,399,1188,1212]" pageId="5" pageNumber="138" value="2012-05-06">06.V.2012</date>
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, net,
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<specimenCount id="96B44792FFFBFFC043F1348B00318DD9" box="[461,550,1188,1212]" pageId="5" pageNumber="138" type="male">3 males</specimenCount>
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, leg. P. Dominiak (LU & UG). Nahr al-Kalb, N 33°
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<geoCoordinate id="E586EADCFFFBFFC04652348B06C08DD9" box="[1134,1239,1188,1212]" direction="east" orientation="longitude" pageId="5" pageNumber="138" precision="55" value="55.842">55.842 E</geoCoordinate>
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35°35.326, 69 m asl,
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<date id="F40CAADBFFFBFFC042F634E803578DBA" box="[202,320,1223,1247]" pageId="5" pageNumber="138" value="2012-05-03">03.V.2012</date>
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, net,
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<specimenCount id="96B44792FFFBFFC043B434E803CB8DBA" box="[392,476,1223,1247]" pageId="5" pageNumber="138" type="male">1 male</specimenCount>
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, leg. P. Dominiak (UG). Aarbit Kozhayya, St. Antonios Kozhayya Monaster, N 34°
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<geoCoordinate id="E586EADCFFFBFFC0428034C303308C61" box="[188,295,1260,1284]" direction="east" orientation="longitude" pageId="5" pageNumber="138" precision="55" value="17.046">17.046 E</geoCoordinate>
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35°56.689,
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<date id="F40CAADBFFFBFFC0438534C300258C61" box="[441,562,1260,1284]" pageId="5" pageNumber="138" value="2012-10-28">28.X.2012</date>
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, shallow cave, net,
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<specimenCount id="96B44792FFFBFFC0412234C301678C61" box="[798,880,1260,1284]" pageId="5" pageNumber="138" type="male">1 male</specimenCount>
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, leg. P. Dominiak (UG). Fanar, N 33°
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<geoCoordinate id="E586EADCFFFBFFC0471334C3078B8C61" box="[1327,1436,1260,1284]" direction="east" orientation="longitude" pageId="5" pageNumber="138" precision="55" value="52.68">52.680 E</geoCoordinate>
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35°33.547, 138 m asl,
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<date id="F40CAADBFFFBFFC043A83520001E8C42" box="[404,521,1295,1319]" pageId="5" pageNumber="138" value="2012-05-03">03.V.2012</date>
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, net,
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<specimenCount id="96B44792FFFBFFC04078352000858C42" box="[580,658,1295,1319]" pageId="5" pageNumber="138" type="male">1 male</specimenCount>
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, leg. P. Dominiak (UG).
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</paragraph>
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</subSubSection>
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<subSubSection id="C8A8DF90FFFBFFC042FB351C03128CF1" pageId="5" pageNumber="138" type="diagnosis">
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<paragraph id="800D8C1BFFFBFFC042FB351C03128CF1" blockId="5.[151,1437,1079,2009]" pageId="5" pageNumber="138">
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<emphasis id="B2C65009FFFBFFC042FB351C03568C29" bold="true" box="[199,321,1331,1356]" pageId="5" pageNumber="138">Diagnosis.</emphasis>
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The species belongs to the
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<taxonomicName id="47B2F798FFFBFFC040AE351B01158C29" box="[658,770,1332,1356]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="fasciigera">
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<emphasis id="B2C65009FFFBFFC040AE351B01158C29" box="[658,770,1332,1356]" italics="true" pageId="5" pageNumber="138">fasciigera</emphasis>
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</taxonomicName>
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group, and differs from all other members by stout and truncated apicolateral processes of 9th tergite in male genitalia, as well as by weakly sclerotized lateral borders of aedeagus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C8A8DF90FFFBFFC042FB35B001968E08" pageId="5" pageNumber="138" type="description">
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<paragraph id="800D8C1BFFFBFFC042FB35B007328E40" blockId="5.[151,1437,1079,2009]" pageId="5" pageNumber="138">
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<emphasis id="B2C65009FFFBFFC042FB35B003B28CDD" bold="true" box="[199,421,1439,1464]" pageId="5" pageNumber="138">Description. Male.</emphasis>
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Flagellum length
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<quantity id="474A21FEFFFBFFC04045358F01378CDD" box="[633,800,1440,1464]" metricMagnitude="-4" metricUnit="m" metricValue="5.949999999999999" metricValueMax="6.3" metricValueMin="5.6" pageId="5" pageNumber="138" unit="mm" value="0.595" valueMax="0.63" valueMin="0.56">0.56–0.63 mm</quantity>
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, AR 0.81–0.92 (n=9). Distal flagellomeres bearing two rows of long setae (
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<figureCitation id="1889909EFFFBFFC043B135EB03CF8CB9" box="[397,472,1476,1500]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="5" pageNumber="138">Fig. 6</figureCitation>
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). Clypeus elongate, composed of two separate sclerites, with 7–12 strong setae. Maxillary palp as in fig. 7. Third palpal segment slender, 57–67 μm long, PR III 3.80–5.58 (n=10). Thorax. Transverse suture absent, 8–10 supraalar setae present. Scutellum entirely pale or dark, with 6–7 strong and few small setae. Wing length
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<quantity id="474A21FEFFFBFFC043F8361D007A8F2F" box="[452,621,1586,1610]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" metricValueMax="9.1" metricValueMin="7.9" pageId="5" pageNumber="138" unit="mm" value="0.85" valueMax="0.91" valueMin="0.79">0.79–0.91 mm</quantity>
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, CR 0.43–0.46 (n=11). Only one radial cell present, wing membrane covered with macrotrichia. Legs unicolored, dark,
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<collectingCountry id="F8A5CC8BFFFBFFC040E1367701168F0A" box="[733,769,1624,1647]" name="Chile" pageId="5" pageNumber="138">TR</collectingCountry>
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I 1.9–2.1 (n=9),
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<collectingCountry id="F8A5CC8BFFFBFFC04181367701F68F0A" box="[957,993,1624,1647]" name="Chile" pageId="5" pageNumber="138">TR</collectingCountry>
|
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II 2.3–2.5 (n=10),
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<collectingCountry id="F8A5CC8BFFFBFFC04693367706C48F0A" box="[1199,1235,1624,1647]" name="Chile" pageId="5" pageNumber="138">TR</collectingCountry>
|
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III 2.0–2.2 (n=11). Genitalia (
|
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<figureCitation id="1889909EFFFBFFC0432D3652034D8FF0" box="[273,346,1661,1685]" captionStart="FIGURES 1 – 8" captionStartId="2.[151,264,1792,1815]" captionTargetBox="[286,1299,222,1743]" captionTargetId="figure@2.[260,1326,195,1768]" captionTargetPageId="2" captionText="FIGURES 1 – 8. Dasyhelea antonii Dominiak sp. nov. (1 – 3), holotype male (3), paratype male (1, 2): 1. distal flagellomeres, 2. palpus, 3. ventral view of genitalia; Dasyhelea avia Dominiak sp. nov. (4 – 5), holotype male (5), paratype male (4): 4. distal flagellomeres, 5. ventral view of genitalia; Dasyhelea libanensis Dominiak sp. nov. (6 – 8), holotype male (6, 8), paratype male (7): 6. distal flagellomeres, 7. palpus, 8. ventral view of genitalia." httpUri="https://zenodo.org/record/219726/files/figure.png" pageId="5" pageNumber="138">Fig. 8</figureCitation>
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). Posterior margin of tergite 9 with a pair of stout, apically truncated apicolateral processes. Posterior margin of sternite 9 elongate, conical. Gonostylus composed of two parts. Outer gonostylus long and straight, with varying length of finger-like prolongation near its base. Inner gonostylus strongly curved, sharp. Parameres asymmetrical. Posterior process broadly fused to right basal arm, stout basally, with divided apex. Lateral projections of aedeagus bent in apical half, with weakly sclerotized membranous lateral borders.
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</paragraph>
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<paragraph id="800D8C1BFFFBFFC042FB371D00498E2C" blockId="5.[151,1437,1079,2009]" box="[199,606,1841,1866]" pageId="5" pageNumber="138">
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<emphasis id="B2C65009FFFBFFC042FB371D03F68E2F" bold="true" box="[199,481,1841,1866]" pageId="5" pageNumber="138">Female and immatures.</emphasis>
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Unknown.
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</paragraph>
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<paragraph id="800D8C1BFFFBFFC042FB377A01968E08" blockId="5.[151,1437,1079,2009]" box="[199,897,1877,1902]" pageId="5" pageNumber="138">
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<emphasis id="B2C65009FFFBFFC042FB377A03F78E0B" bold="true" box="[199,480,1877,1902]" pageId="5" pageNumber="138">Derivation of the name.</emphasis>
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The species is named after
|
||
<collectingCountry id="F8A5CC8BFFFBFFC0412B377A016A8E08" box="[791,893,1877,1901]" name="Lebanon" pageId="5" pageNumber="138">Lebanon</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8A8DF90FFFBFFC042FB375603C68EF4" box="[199,465,1913,1938]" pageId="5" pageNumber="138" type="distribution">
|
||
<paragraph id="800D8C1BFFFBFFC042FB375603C68EF4" blockId="5.[151,1437,1079,2009]" box="[199,465,1913,1938]" pageId="5" pageNumber="138">
|
||
<emphasis id="B2C65009FFFBFFC042FB375603778EF7" bold="true" box="[199,352,1913,1938]" pageId="5" pageNumber="138">Distribution.</emphasis>
|
||
<collectingCountry id="F8A5CC8BFFFBFFC0435B375603DA8EF4" box="[359,461,1913,1937]" name="Lebanon" pageId="5" pageNumber="138">Lebanon</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C8A8DF90FFFBFFC342FB37B2001A89B1" lastPageId="6" lastPageNumber="139" pageId="5" pageNumber="138" type="discussion">
|
||
<paragraph id="800D8C1BFFFBFFC342FB37B2001A89B1" blockId="5.[151,1437,1079,2009]" lastBlockId="6.[151,1436,151,212]" lastPageId="6" lastPageNumber="139" pageId="5" pageNumber="138">
|
||
<emphasis id="B2C65009FFFBFFC042FB37B203588ED3" bold="true" box="[199,335,1949,1974]" pageId="5" pageNumber="138">Comments.</emphasis>
|
||
This newly described species is similar to
|
||
<taxonomicName id="47B2F798FFFBFFC0410D37B106628ED0" authority="Kieffer" authorityName="Kieffer" box="[817,1141,1949,1974]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="fasciigera">
|
||
<emphasis id="B2C65009FFFBFFC0410D37B106098ED3" box="[817,1054,1950,1974]" italics="true" pageId="5" pageNumber="138">Dasyhelea fasciigera</emphasis>
|
||
Kieffer
|
||
</taxonomicName>
|
||
but differs in the shape of apicolateral processes of tergite 9, which are truncated, whereas they are rounded in the latter species. Other diagnostic characters for the new species are the shape of the lateral processes of aedeagus bearing membranous weakly chitinized lateral borders.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |