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<mods:title id="A14547097835703A6453BAE1681B3A9E">Five new species and new records of biting midges of the genus Dasyhelea Kieffer from the Near East (Diptera: Ceratopogonidae)</mods:title>
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<treatment id="081B3D0DFFFAFFC042AB377E01FC8B7D" ID-DOI="http://doi.org/10.5281/zenodo.4499994" ID-GBIF-Taxon="119571713" ID-Zenodo-Dep="4499994" LSID="urn:lsid:plazi:treatment:081B3D0DFFFAFFC042AB377E01FC8B7D" httpUri="http://treatment.plazi.org/id/081B3D0DFFFAFFC042AB377E01FC8B7D" lastPageId="5" lastPageNumber="138" pageId="4" pageNumber="137">
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<paragraph id="800D8C1BFFFAFFC142AB377E00908E0E" blockId="4.[151,647,1873,1899]" box="[151,647,1873,1899]" pageId="4" pageNumber="137">
<heading id="DB453B77FFFAFFC142AB377E00908E0E" bold="true" box="[151,647,1873,1899]" fontSize="11" level="1" pageId="4" pageNumber="137" reason="1">
<taxonomicName id="47B2F798FFFAFFC142AB377E00908E0E" ID-CoL="348HK" authority="Boorman &amp; Harten" authorityName="Boorman &amp; Harten" box="[151,647,1873,1899]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="137" phylum="Arthropoda" rank="species" species="deemingi">
<emphasis id="B2C65009FFFAFFC142AB377E00908E0E" bold="true" box="[151,647,1873,1899]" pageId="4" pageNumber="137">
<emphasis id="B2C65009FFFAFFC142AB377E039D8E0E" bold="true" box="[151,394,1873,1899]" italics="true" pageId="4" pageNumber="137">Dasyhelea deemingi</emphasis>
Boorman &amp; Harten
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFAFFC142AB37B701768EA9" pageId="4" pageNumber="137" type="reference_group">
<paragraph id="800D8C1BFFFAFFC142AB37B700EA8ECA" blockId="4.[151,865,1944,1997]" box="[151,765,1944,1967]" pageId="4" pageNumber="137">
<treatmentCitationGroup id="A0A2AB35FFFAFFC142AB37B700EA8ECA" box="[151,765,1944,1967]" pageId="4" pageNumber="137">
<taxonomicName id="47B2F798FFFAFFC142AB37B700EE8ECA" ID-CoL="348HK" authority="Boorman &amp; Harten, 2002: 441" authorityName="Boorman &amp; Harten" authorityPageNumber="441" authorityYear="2002" box="[151,761,1944,1967]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="137" phylum="Arthropoda" rank="species" species="deemingi">
<emphasis id="B2C65009FFFAFFC142AB37B703728ECB" box="[151,357,1944,1966]" italics="true" pageId="4" pageNumber="137">Dasyhelea deemingi</emphasis>
<treatmentCitation id="0113AA0AFFFAFFC1435737B600B68ECB" author="Boorman" box="[363,673,1945,1966]" page="441" pageId="4" pageNumber="137" year="2002">
<bibRefCitation id="E423F1EAFFFAFFC1435737B600B68ECB" author="Boorman, J. &amp; van Harten, A." box="[363,673,1945,1966]" firstAuthor="Boorman" journalOrPublisher="Fauna of Arabia" pageId="4" pageNumber="137" pagination="427 - 462" part="19" refId="ref6017" refString="Boorman, J. &amp; van Harten, A. (2002) Some Ceratopogonidae (Insecta: Diptera) from the Arabian Peninsula, with particular reference to the Republic of Yemen. Fauna of Arabia, 19, 427 - 462." title="Some Ceratopogonidae (Insecta: Diptera) from the Arabian Peninsula, with particular reference to the Republic of Yemen" type="journal article" year="2002">Boorman &amp; Harten, 2002: 441</bibRefCitation>
</treatmentCitation>
(
<collectingCountry id="F8A5CC8BFFFAFFC1409337B600E58ECA" box="[687,754,1945,1967]" name="Oman" pageId="4" pageNumber="137">Oman</collectingCountry>
)
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="800D8C1BFFFAFFC142AB379801768EA9" blockId="4.[151,865,1944,1997]" box="[151,865,1974,1997]" pageId="4" pageNumber="137">
<taxonomicName id="47B2F798FFFAFFC142AB379800B38EA9" ID-CoL="9HSFG" authority="Mazumdar &amp; Chaudhuri, 2009" authorityName="Mazumdar &amp; Chaudhuri" authorityYear="2009" box="[151,676,1975,1997]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="137" phylum="Arthropoda" rank="species" species="labinoda">
<emphasis id="B2C65009FFFAFFC142AB379803758EA8" box="[151,354,1975,1997]" italics="true" pageId="4" pageNumber="137">Dasyhelea labinoda</emphasis>
<bibRefCitation id="E423F1EAFFFAFFC14354379800B38EA9" author="Mazumdar, A. &amp; Chaudhuri, P. K." box="[360,676,1975,1997]" firstAuthor="Mazumdar" journalOrPublisher="International Journal of Dipterological Research" pageId="4" pageNumber="137" pagination="191 - 201" part="20" refId="ref7183" refString="Mazumdar, A. &amp; Chaudhuri, P. K. (2009) Indian species of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae). International Journal of Dipterological Research, 20, 191 - 201." title="Indian species of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae)" type="journal article" year="2009">Mazumdar &amp; Chaudhuri, 2009</bibRefCitation>
</taxonomicName>
: (
<collectingCountry id="F8A5CC8BFFFAFFC14089379800F98EA8" box="[693,750,1975,1997]" name="India" pageId="4" pageNumber="137">India</collectingCountry>
).
<emphasis id="B2C65009FFFAFFC140C3379901768EA9" bold="true" box="[767,865,1974,1997]" pageId="4" pageNumber="137">Syn. nov.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFBFFC042AB30B801D38879" pageId="5" pageNumber="138" type="materials_examined">
<paragraph id="800D8C1BFFFBFFC042AB30B8012E89B1" blockId="5.[151,1436,151,536]" pageId="5" pageNumber="138">
<emphasis id="B2C65009FFFBFFC042AB30B803C989D5" bold="true" box="[151,478,151,176]" pageId="5" pageNumber="138">
Material examined.
<collectingCountry id="F8A5CC8BFFFBFFC043BA30B703CE89D5" box="[390,473,152,176]" name="Yemen" pageId="5" pageNumber="138">Yemen</collectingCountry>
.
</emphasis>
Al Mukalla, N 14˚33' E 49˚08',
<date id="F40CAADBFFFBFFC0417830B801A789CA" box="[836,944,151,175]" pageId="5" pageNumber="138" value="2003-01" valueMax="2003-02-31" valueMin="2003-01-00">
I
<bibRefCitation id="E423F1EAFFFBFFC0416130B801A789CA" author="II" box="[861,944,151,175]" pageId="5" pageNumber="138" refString="Szadziewski, R. &amp; Borkent, A. (2003) New synonyms, combinations and records of biting midges (Diptera: Ceratopogonidae). Polish Journal of Entomology, 72, 249 - 260." type="journal article" year="2003">II.2003</bibRefCitation>
</date>
, light trap,
<specimenCount id="96B44792FFFBFFC0461230B8066C89CA" box="[1070,1147,151,175]" pageId="5" pageNumber="138" type="male">1 male</specimenCount>
, leg. A. van Harten (UG);
<date id="F40CAADBFFFBFFC042AB3093032989B1" box="[151,318,188,212]" pageId="5" pageNumber="138" value="2003-07" valueMax="2003-08-31" valueMin="2003-07-00">VIIVIII.2003</date>
, light trap,
<specimenCount id="96B44792FFFBFFC043813093000189B1" box="[445,534,188,212]" pageId="5" pageNumber="138" type="male">5 males</specimenCount>
, leg. A. van Harten (UG).
</paragraph>
<paragraph id="800D8C1BFFFBFFC042FB30CF01D38879" blockId="5.[151,1436,151,536]" pageId="5" pageNumber="138">
<emphasis id="B2C65009FFFBFFC042FB30CF0362899D" bold="true" box="[199,373,223,248]" pageId="5" pageNumber="138">
<typeStatus id="5F0932B9FFFBFFC042FB30CF0315899D" box="[199,258,224,248]" pageId="5" pageNumber="138">Type</typeStatus>
material.
</emphasis>
<typeStatus id="5F0932B9FFFBFFC0434130CF03C8899D" box="[381,479,224,248]" pageId="5" pageNumber="138" type="paratype">Paratype</typeStatus>
male of
<taxonomicName id="47B2F798FFFBFFC0407F30CE00DD899D" box="[579,714,224,248]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="labinoda">
<emphasis id="B2C65009FFFBFFC0407F30CE0040899D" box="[579,599,225,248]" italics="true" pageId="5" pageNumber="138">D</emphasis>
.
<emphasis id="B2C65009FFFBFFC0405A30CF00DD899D" box="[614,714,224,248]" italics="true" pageId="5" pageNumber="138">labinoda</emphasis>
</taxonomicName>
, labeled as follows: Para Adult 3, Loc. Burdwan, West Bengal,
<date id="F40CAADBFFFBFFC042AB312B03068879" box="[151,273,260,284]" pageId="5" pageNumber="138" value="1989-11-02">02.xi.1989</date>
, Coll. N. Saha,
<taxonomicName id="47B2F798FFFBFFC043FD312B01D38879" authority="Mazumdar &amp; Chaudhuri." authorityName="Mazumdar &amp; Chaudhuri." box="[449,964,260,284]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="labinoda">
<emphasis id="B2C65009FFFBFFC043FD312B00B58879" box="[449,674,260,284]" italics="true" pageId="5" pageNumber="138">Dasyhelea labinoda</emphasis>
Mazumdar &amp; Chaudhuri.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFBFFC042FB3108037488E2" pageId="5" pageNumber="138" type="distribution">
<paragraph id="800D8C1BFFFBFFC042FB3108037488E2" blockId="5.[151,1436,151,536]" pageId="5" pageNumber="138">
<emphasis id="B2C65009FFFBFFC042FB310803778825" bold="true" box="[199,352,295,320]" pageId="5" pageNumber="138">Distribution.</emphasis>
Up to now this species was mentioned from
<collectingCountry id="F8A5CC8BFFFBFFC0414B31080607885A" box="[887,1040,295,319]" name="Saudi Arabia" pageId="5" pageNumber="138">Saudi Arabia</collectingCountry>
,
<collectingCountry id="F8A5CC8BFFFBFFC046233108066C885A" box="[1055,1147,295,319]" name="Bahrain" pageId="5" pageNumber="138">Bahrain</collectingCountry>
,
<collectingCountry id="F8A5CC8BFFFBFFC046B5310806D9885A" box="[1161,1230,295,319]" name="Oman" pageId="5" pageNumber="138">Oman</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFBFFC046E3310803318801" author="Boorman, J. &amp; van Harten, A." firstAuthor="Boorman" journalOrPublisher="Fauna of Arabia" pageId="5" pageNumber="138" pagination="427 - 462" part="19" refId="ref6017" refString="Boorman, J. &amp; van Harten, A. (2002) Some Ceratopogonidae (Insecta: Diptera) from the Arabian Peninsula, with particular reference to the Republic of Yemen. Fauna of Arabia, 19, 427 - 462." title="Some Ceratopogonidae (Insecta: Diptera) from the Arabian Peninsula, with particular reference to the Republic of Yemen" type="journal article" year="2002">Boorman &amp; van Harten 2002</bibRefCitation>
), the
<collectingCountry id="F8A5CC8BFFFBFFC0435F3163004E8801" box="[355,601,332,356]" name="United Arab Emirates" pageId="5" pageNumber="138">United Arab Emirates</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFBFFC04054316301638801" author="Szadziewski, R. &amp; Gwizdalska-Kentzer, M. &amp; Gilka, W." box="[616,884,332,356]" editor="van Harten, A." etAl="et al." firstAuthor="Szadziewski" journalOrPublisher="Dar Al Ummah Printing, Publishing, Distribution &amp; Advertising, Abu Dhabi, UAE" pageId="5" pageNumber="138" pagination="636 - 653" refId="ref7546" refString="Szadziewski, R., Gwizdalska-Kentzer, M. &amp; Gilka, W. (2011) Order Diptera, family Ceratopogonidae. In: Sheikh Tahnoon Bin Zayed Al Nahyan, H. H. (patron), van Harten, A. (Ed.), Arthropod fauna of the United Arab Emirates. Volume 4. Dar Al Ummah Printing, Publishing, Distribution &amp; Advertising, Abu Dhabi, UAE, pp. 636 - 653." title="Order Diptera, family Ceratopogonidae" type="book chapter" volumeTitle="Arthropod fauna of the United Arab Emirates. Volume 4" year="2011">
Szadziewski
<emphasis id="B2C65009FFFBFFC040C0316201398801" box="[764,814,332,356]" italics="true" pageId="5" pageNumber="138">et al</emphasis>
. 2011
</bibRefCitation>
) and
<collectingCountry id="F8A5CC8BFFFBFFC0418F316301FA8801" box="[947,1005,332,356]" name="India" pageId="5" pageNumber="138">India</collectingCountry>
(
<bibRefCitation id="E423F1EAFFFBFFC041C7316307438801" author="Mazumdar, A. &amp; Chaudhuri, P. K." box="[1019,1364,332,356]" firstAuthor="Mazumdar" journalOrPublisher="International Journal of Dipterological Research" pageId="5" pageNumber="138" pagination="191 - 201" part="20" refId="ref7183" refString="Mazumdar, A. &amp; Chaudhuri, P. K. (2009) Indian species of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae). International Journal of Dipterological Research, 20, 191 - 201." title="Indian species of the genus Dasyhelea Kieffer (Diptera: Ceratopogonidae)" type="journal article" year="2009">Mazumdar &amp; Chaudhuri 2009</bibRefCitation>
). New record for
<collectingCountry id="F8A5CC8BFFFBFFC04331315F034888E2" box="[269,351,368,391]" name="Yemen" pageId="5" pageNumber="138">Yemen</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C8A8DF90FFFBFFC042FB31BC01FC8B7D" pageId="5" pageNumber="138" type="discussion">
<paragraph id="800D8C1BFFFBFFC042FB31BC01FC8B7D" blockId="5.[151,1436,151,536]" pageId="5" pageNumber="138">
<emphasis id="B2C65009FFFBFFC042FB31BC035888C9" bold="true" box="[199,335,403,428]" pageId="5" pageNumber="138">Comments.</emphasis>
After examination of the
<typeStatus id="5F0932B9FFFBFFC040A431BA00EF88C8" box="[664,760,405,429]" pageId="5" pageNumber="138" type="paratype">paratype</typeStatus>
male of
<taxonomicName id="47B2F798FFFBFFC0414F31BB064B88C9" box="[883,1116,404,428]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="labinoda">
<emphasis id="B2C65009FFFBFFC0414F31BB064B88C9" box="[883,1116,404,428]" italics="true" pageId="5" pageNumber="138">Dasyhelea labinoda</emphasis>
</taxonomicName>
we consider there are no differences in the morphological characters between this species and
<taxonomicName id="47B2F798FFFBFFC041823196065C88B5" box="[958,1099,440,464]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="deemingi">
<emphasis id="B2C65009FFFBFFC04182319601C588B5" box="[958,978,441,464]" italics="true" pageId="5" pageNumber="138">D</emphasis>
.
<emphasis id="B2C65009FFFBFFC041D83197065C88B5" box="[996,1099,440,464]" italics="true" pageId="5" pageNumber="138">deemingi</emphasis>
</taxonomicName>
. Szadziewski and co-authors (
<bibRefCitation id="E423F1EAFFFBFFC042A231F303A68891" author="Szadziewski, R. &amp; Gwizdalska-Kentzer, M. &amp; Gilka, W." box="[158,433,476,500]" editor="van Harten, A." etAl="et al." firstAuthor="Szadziewski" journalOrPublisher="Dar Al Ummah Printing, Publishing, Distribution &amp; Advertising, Abu Dhabi, UAE" pageId="5" pageNumber="138" pagination="636 - 653" refId="ref7546" refString="Szadziewski, R., Gwizdalska-Kentzer, M. &amp; Gilka, W. (2011) Order Diptera, family Ceratopogonidae. In: Sheikh Tahnoon Bin Zayed Al Nahyan, H. H. (patron), van Harten, A. (Ed.), Arthropod fauna of the United Arab Emirates. Volume 4. Dar Al Ummah Printing, Publishing, Distribution &amp; Advertising, Abu Dhabi, UAE, pp. 636 - 653." title="Order Diptera, family Ceratopogonidae" type="book chapter" volumeTitle="Arthropod fauna of the United Arab Emirates. Volume 4" year="2011">
Szadziewski
<emphasis id="B2C65009FFFBFFC0430931F2037E8891" box="[309,361,476,500]" italics="true" pageId="5" pageNumber="138">et al</emphasis>
. 2011
</bibRefCitation>
) suggested that these two names are probably conspecific, and currently we propose to treat
<taxonomicName id="47B2F798FFFBFFC042F3322E03438B7D" box="[207,340,512,536]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="labinoda">
<emphasis id="B2C65009FFFBFFC042F3322E02F48B7D" box="[207,227,513,536]" italics="true" pageId="5" pageNumber="138">D</emphasis>
.
<emphasis id="B2C65009FFFBFFC042CD322F03438B7D" box="[241,340,512,536]" italics="true" pageId="5" pageNumber="138">labinoda</emphasis>
</taxonomicName>
as a new junior synonym of
<taxonomicName id="47B2F798FFFBFFC040A6322E01358B7D" box="[666,802,512,536]" class="Insecta" family="Ceratopogonidae" genus="Dasyhelea" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="138" phylum="Arthropoda" rank="species" species="deemingi">
<emphasis id="B2C65009FFFBFFC040A6322E00B98B7D" box="[666,686,513,536]" italics="true" pageId="5" pageNumber="138">D</emphasis>
.
<emphasis id="B2C65009FFFBFFC04080322F01358B7D" box="[700,802,512,536]" italics="true" pageId="5" pageNumber="138">deemingi</emphasis>
</taxonomicName>
.
<emphasis id="B2C65009FFFBFFC0410C322F01FC8B7D" bold="true" box="[816,1003,512,536]" pageId="5" pageNumber="138">New synonymy.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>