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<mods:title id="3D83E73DD3924670DBAF241465E29EE4">Integrative description of two new species of Malagasy chirping giant pill-millipedes, genus Sphaeromimus (Diplopoda: Sphaerotheriida: Arthrosphaeridae)</mods:title>
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<mods:date id="F6654F138DF7363743D5CDABE9B3E4CE">2017</mods:date>
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<treatment id="08124C2AFFF5FF8EFE7FF95EFAA254A4" ID-DOI="http://doi.org/10.5281/zenodo.5623723" ID-GBIF-Taxon="139350446" ID-Zenodo-Dep="5623723" LSID="urn:lsid:plazi:treatment:08124C2AFFF5FF8EFE7FF95EFAA254A4" httpUri="http://treatment.plazi.org/id/08124C2AFFF5FF8EFE7FF95EFAA254A4" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" scope_class="Diplopoda" scope_family="Arthrosphaeridae" scope_order="Sphaerotheriida">
<subSubSection id="C8A1AEB7FFF5FF8CFE7FF95EFB815277" box="[480,1108,1776,1803]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="8004FD3CFFF5FF8CFE7FF95EFB815277" blockId="4.[480,1108,1776,1803]" box="[480,1108,1776,1803]" pageId="4" pageNumber="5">
<heading id="DB4C4A50FFF5FF8CFE7FF95EFB815277" box="[480,1108,1776,1803]" centered="true" fontSize="36" level="4" pageId="4" pageNumber="5" reason="4">
Genus
<taxonomicName id="47BB86BFFFF5FF8CFDADF95FFB815277" authority="de Saussure &amp; Zehntner, 1902" authorityName="de Saussure &amp; Zehntner" authorityYear="1902" box="[562,1108,1776,1803]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B2CF212EFFF5FF8CFDADF95FFD3C5277" bold="true" box="[562,745,1776,1803]" italics="true" pageId="4" pageNumber="5">Sphaeromimus</emphasis>
de Saussure &amp; Zehntner, 1902
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<paragraph id="8004FD3CFFF5FF8CFF22F89CFD385231" blockId="4.[189,749,1842,1869]" box="[189,749,1842,1869]" pageId="4" pageNumber="5">
For a recent revision, see
<bibRefCitation id="E42A80CDFFF5FF8CFE75F89CFD3D5231" author="Wesener T. &amp; Minh-Tu Le D. &amp; Loria S. F." box="[490,744,1842,1869]" journalOrPublisher="ZooKeys" pageId="4" pageNumber="5" pagination="67 - 104" part="414" publicationUrl="https://doi.org/10.3897/zookeys.414.7730" refId="ref12874" refString="Wesener T., Minh-Tu Le D. &amp; Loria S. F. 2014. Integrative revision of the giant pill-millipede genus Sphaeromimus, with the description of seven new species (Diplopoda, Sphaerotheriida, Arthrosphaeridae). ZooKeys 414: 67 - 104. https: // doi. org / 10.3897 / zookeys. 414.7730" title="Integrative revision of the giant pill-millipede genus Sphaeromimus, with the description of seven new species (Diplopoda, Sphaerotheriida, Arthrosphaeridae)" type="journal article" year="2014">
Wesener
<emphasis id="B2CF212EFFF5FF8CFDCAF89CFD455230" box="[597,656,1842,1868]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
(
<date id="F405DBFCFFF5FF8CFD00F89DFD0E5230" box="[671,731,1842,1868]" pageId="4" pageNumber="5" value="2014">2014</date>
)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C8A1AEB7FFF5FF8CFF22F8D8FD0452EA" box="[189,721,1910,1942]" pageId="4" pageNumber="5" type="multiple">
<paragraph id="8004FD3CFFF5FF8CFF22F8D8FD0452EA" blockId="4.[189,721,1910,1942]" box="[189,721,1910,1942]" pageId="4" pageNumber="5">
<heading id="DB4C4A50FFF5FF8CFF22F8D8FD0452EA" bold="true" box="[189,721,1910,1942]" fontSize="13" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="B2CF212EFFF5FF8CFF22F8D8FD0452EA" bold="true" box="[189,721,1910,1942]" pageId="4" pageNumber="5">
<emphasis id="B2CF212EFFF5FF8CFF22F8D8FE0B52E9" bold="true" box="[189,478,1910,1942]" italics="true" pageId="4" pageNumber="5">Key to the species of</emphasis>
<taxonomicName id="47BB86BFFFF5FF8CFE79F8D8FD1052EA" authorityName="de Saussure &amp; Zehntner" authorityYear="1902" box="[486,709,1911,1942]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="4" pageNumber="24" phylum="Arthropoda" rank="genus">Sphaeromimus</taxonomicName>
:
</emphasis>
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</paragraph>
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<paragraph id="8004FD3CFFF5FF8CFF22F803FAA352BA" blockId="4.[189,1399,1964,2026]" box="[189,1398,1964,1991]" pageId="4" pageNumber="5">
<keyLead id="3B4A5E09FFF5FF8CFF22F803FAA352BA" box="[189,1398,1964,1991]" pageId="4" pageNumber="5">1. Male harp on the anterior telopod with 3 stridulation ribs …………………………………………2</keyLead>
</paragraph>
<paragraph id="8004FD3CFFF5FF8CFF5EF87FFAA25296" blockId="4.[189,1399,1964,2026]" box="[193,1399,1999,2026]" pageId="4" pageNumber="5">
<keyLead id="3B4A5E09FFF5FF8CFF5EF87FFAA25296" box="[193,1399,1999,2026]" pageId="4" pageNumber="5">– Male harp on the anterior telopod with 4–6 stridulation ribs ………………………………………7</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B4FE599FFF4FF8DFF22FEA4FAA354A7" pageId="5" pageNumber="6">
<paragraph id="8004FD3CFFF4FF8DFF22FEA4FAA25412" blockId="5.[189,1406,267,2027]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF22FEA4FAA25412" pageId="5" pageNumber="6">
2. Body length &lt;
<quantity id="474350D9FFF4FF8DFE0BFEA4FE3C545A" box="[404,489,267,294]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="6" unit="mm" value="20.0">20 mm</quantity>
. Mid-body legs without a coxal lobe. Endotergum with single row of marginal bristles. Immovable finger of posterior telopod slender, apically strongly curved like a hook. Colour pink or brown ………………………………………………………………………………………3
</keyLead>
</paragraph>
<paragraph id="8004FD3CFFF4FF8DFF5EFED6FAA354A7" blockId="5.[189,1406,267,2027]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF5EFED6FAA354A7" pageId="5" pageNumber="6">
– Body length&gt;
<quantity id="474350D9FFF4FF8DFE3EFED6FE2A54EF" box="[417,511,377,403]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="6" unit="mm" value="20.0">20 mm</quantity>
. Mid-body legs at least with a weak coxal lobe. Endotergum usually with at least two, rarely only one
<emphasis id="B2CF212EFFF4FF8DFD58FE32FD0454CB" box="[711,721,413,439]" italics="true" pageId="5" pageNumber="6">(</emphasis>
<taxonomicName id="47BB86BFFFF4FF8DFD4EFE32FBC354CB" authority="Wesener, 2014" authorityName="Wesener" authorityYear="2014" box="[721,1046,413,439]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="titanus">
<emphasis id="B2CF212EFFF4FF8DFD4EFE32FC9F54CB" box="[721,842,413,439]" italics="true" pageId="5" pageNumber="6">S. titanus</emphasis>
<bibRefCitation id="E42A80CDFFF4FF8DFCC2FE32FBC354CB" author="Wesener T." box="[861,1046,413,439]" journalOrPublisher="Zootaxa" pageId="5" pageNumber="6" pagination="587 - 594" part="3790" publicationUrl="https://doi.org/10.11646/zootaxa.3790.4.7" refId="ref12228" refString="Wesener T. 2014 a. Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda). Zootaxa 3790 (4): 587 - 594. https: // doi. org / 10.11646 / zootaxa. 3790.4.7" title="Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda)" type="journal article" year="2014">Wesener, 2014</bibRefCitation>
</taxonomicName>
) row of marginal bristles. Immovable finger of posterior telopod at least basally wide. Colour black or brown …………4
</keyLead>
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</keyStep>
<keyStep id="3B4FE599FFF4FF8DFF22FDABFAA257D3" pageId="5" pageNumber="6">
<paragraph id="8004FD3CFFF4FF8DFF22FDABFAA2571B" blockId="5.[189,1406,267,2027]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF22FDABFAA2571B" pageId="5" pageNumber="6">
3. Colour pink, surface shiny. Process of telopoditomere 2 of anterior telopods in anterior view visible laterally. Littoral forest of Mandena and rainforest of Enato …………………………… ……………………………………………………………
<taxonomicName id="47BB86BFFFF4FF8DFC12FDE2FAA2571B" authority="Wesener &amp; Sierwald, 2005" authorityName="Wesener &amp; Sierwald" authorityYear="2005" box="[909,1399,588,615]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="inexpectatus">
<emphasis id="B2CF212EFFF4FF8DFC12FDE2FB95571B" bold="true" box="[909,1088,589,615]" italics="true" pageId="5" pageNumber="6">S. inexpectatus</emphasis>
<bibRefCitation id="E42A80CDFFF4FF8DFBD9FDE2FAA2571B" author="Wesener T. &amp; Sierwald P." box="[1094,1399,588,615]" journalOrPublisher="Proceedings of the California Academy of Science" pageId="5" pageNumber="6" pagination="557 - 599" part="56" refId="ref12487" refString="Wesener T. &amp; Sierwald P. 2005. The giant pill-millipedes of Madagascar - Revision of the genus Sphaeromimus. Proceedings of the California Academy of Science 56: 557 - 599." title="The giant pill-millipedes of Madagascar - Revision of the genus Sphaeromimus" type="journal article" year="2005">Wesener &amp; Sierwald, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8004FD3CFFF4FF8DFF22FDDEFAA257D3" blockId="5.[189,1406,267,2027]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF22FDDEFAA257D3" pageId="5" pageNumber="6">
– Colour light brown, surface dull. Process of telopoditomere 2 of anterior telopods in anterior view not visible. Littoral forest of Sainte Luce, fragment S8 ………………
<taxonomicName id="47BB86BFFFF4FF8DFB84FD3AFAA257D3" authority="Wesener, 2014" authorityName="Wesener" authorityYear="2014" box="[1051,1399,661,688]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="saintelucei">
<emphasis id="B2CF212EFFF4FF8DFB84FD3AFB6957CC" bold="true" box="[1051,1212,661,688]" italics="true" pageId="5" pageNumber="6">S. saintelucei</emphasis>
<bibRefCitation id="E42A80CDFFF4FF8DFB55FD39FAA257D3" author="Wesener T." box="[1226,1399,661,688]" journalOrPublisher="Zootaxa" pageId="5" pageNumber="6" pagination="587 - 594" part="3790" publicationUrl="https://doi.org/10.11646/zootaxa.3790.4.7" refId="ref12228" refString="Wesener T. 2014 a. Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda). Zootaxa 3790 (4): 587 - 594. https: // doi. org / 10.11646 / zootaxa. 3790.4.7" title="Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda)" type="journal article" year="2014">Wesener, 2014</bibRefCitation>
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="3B4FE599FFF4FF8DFF22FD77FAA3561C" pageId="5" pageNumber="6">
<paragraph id="8004FD3CFFF4FF8DFF22FD77FAA2566A" blockId="5.[189,1406,267,2027]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF22FD77FAA2566A" pageId="5" pageNumber="6">
4. Body length&gt;
<quantity id="474350D9FFF4FF8DFE0EFD77FE31578E" box="[401,484,728,754]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="6" unit="mm" value="30.0">30mm</quantity>
, light brown. Endotergum with single row of marginal bristles. Movable finger of posterior telopod straight (
<figureCitation id="1880E1B9FFF4FF8DFDDBFD53FD74566A" box="[580,673,764,790]" captionStart="Fig.2" captionStartId="8.[189,232,1930,1956]" captionTargetBox="[189,1398,1079,1886]" captionTargetId="figure@8.[74,1398,1079,1887]" captionTargetPageId="8" captionText="Fig.2.Sphaeromimus kalambatritrasp.nov.,SEM ofleft antenna,♂, holotype (CASENT9068297-A). A. Overview. B.Antennal disc, detail. C. Margin of antennal disc, detail.Abbreviations: aD = antennal disc; SB = sensilla basiconica; SC = sensorycone; I–VI = antennomeres." httpUri="https://zenodo.org/record/1118114/files/figure.png" pageId="5" pageNumber="6">Fig. 2B</figureCitation>
). Lowland forest of Manombo ……
<taxonomicName id="47BB86BFFFF4FF8DFBD0FD53FAA2566A" authority="Wesener, 2014" authorityName="Wesener" authorityYear="2014" box="[1103,1399,764,791]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="titanus">
<emphasis id="B2CF212EFFF4FF8DFBD0FD53FB14566A" bold="true" box="[1103,1217,764,790]" italics="true" pageId="5" pageNumber="6">S. titanus</emphasis>
<bibRefCitation id="E42A80CDFFF4FF8DFB56FD52FAA2566A" author="Wesener T." box="[1225,1399,764,791]" journalOrPublisher="Zootaxa" pageId="5" pageNumber="6" pagination="587 - 594" part="3790" publicationUrl="https://doi.org/10.11646/zootaxa.3790.4.7" refId="ref12228" refString="Wesener T. 2014 a. Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda). Zootaxa 3790 (4): 587 - 594. https: // doi. org / 10.11646 / zootaxa. 3790.4.7" title="Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda)" type="journal article" year="2014">Wesener, 2014</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8004FD3CFFF4FF8DFF5EFC8EFAA3561C" blockId="5.[189,1406,267,2027]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF5EFC8EFAA3561C" pageId="5" pageNumber="6">
– Body length
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, black or dark brown. Endotergum with two rows of marginal bristles. Movable finger of posterior telopod curved ………………………………………………………5
</keyLead>
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</keyStep>
<keyStep id="3B4FE599FFF4FF8DFF22FC28FAA35696" pageId="5" pageNumber="6">
<paragraph id="8004FD3CFFF4FF8DFF22FC28FAA256BB" blockId="5.[189,1406,267,2027]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF22FC28FAA256BB" pageId="5" pageNumber="6">
5. Mid-body legs with strongly developed coxal process. Tergite surface shiny. Littoral forest of Sainte Luce, fragment S9 …………………………………………
<taxonomicName id="47BB86BFFFF4FF8DFC05FC03FAA256BB" authority="Wesener &amp; Sierwald, 2005" authorityName="Wesener &amp; Sierwald" authorityYear="2005" box="[922,1399,940,967]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="splendidus">
<emphasis id="B2CF212EFFF4FF8DFC05FC03FBED56BB" bold="true" box="[922,1080,940,967]" italics="true" pageId="5" pageNumber="6">S. splendidus</emphasis>
<bibRefCitation id="E42A80CDFFF4FF8DFBDDFC02FAA256BB" author="Wesener T. &amp; Sierwald P." box="[1090,1399,940,967]" journalOrPublisher="Proceedings of the California Academy of Science" pageId="5" pageNumber="6" pagination="557 - 599" part="56" refId="ref12487" refString="Wesener T. &amp; Sierwald P. 2005. The giant pill-millipedes of Madagascar - Revision of the genus Sphaeromimus. Proceedings of the California Academy of Science 56: 557 - 599." title="The giant pill-millipedes of Madagascar - Revision of the genus Sphaeromimus" type="journal article" year="2005">Wesener &amp; Sierwald, 2005</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8004FD3CFFF4FF8DFF5EFC7EFAA35696" blockId="5.[189,1406,267,2027]" box="[193,1398,976,1003]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF5EFC7EFAA35696" box="[193,1398,976,1003]" pageId="5" pageNumber="6">– Mid-body legs with barely developed coxal process. Tergite surface dull …………………………6</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B4FE599FFF4FF8DFF22FBBCFAA2519F" pageId="5" pageNumber="6">
<paragraph id="8004FD3CFFF4FF8DFF22FBBCFAA2510A" blockId="5.[189,1406,267,2027]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF22FBBCFAA2510A" pageId="5" pageNumber="6">
6. Endotergum with strongly developed cuticular patterns. Movable finger of posterior telopod without small pits, carrying 20–22 crenulated teeth. Andohahela mountain chain, Manantantely, Malio and Isaka-Ivondro ……………………………………………
<taxonomicName id="47BB86BFFFF4FF8DFB91FBF3FAA2510A" authority="Wesener, 2014" authorityName="Wesener" authorityYear="2014" box="[1038,1399,1115,1142]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="andohahela">
<emphasis id="B2CF212EFFF4FF8DFB91FBF3FB6B510A" bold="true" box="[1038,1214,1115,1142]" italics="true" pageId="5" pageNumber="6">S. andohahela</emphasis>
<bibRefCitation id="E42A80CDFFF4FF8DFB57FBF3FAA2510A" author="Wesener T." box="[1224,1399,1116,1142]" journalOrPublisher="Zootaxa" pageId="5" pageNumber="6" pagination="587 - 594" part="3790" publicationUrl="https://doi.org/10.11646/zootaxa.3790.4.7" refId="ref12228" refString="Wesener T. 2014 a. Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda). Zootaxa 3790 (4): 587 - 594. https: // doi. org / 10.11646 / zootaxa. 3790.4.7" title="Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda)" type="journal article" year="2014">Wesener, 2014</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8004FD3CFFF4FF8DFF5EFB2EFAA2519F" blockId="5.[189,1406,267,2027]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF5EFB2EFAA2519F" pageId="5" pageNumber="6">
– Endotergum with weakly developed cuticular patterns. Movable finger of posterior telopod covered with small pits, carrying 23 or 24 crenulated teeth. Inside Grotte d’Andrahomana and in deep ravines N of Ankapaky ……………………………………………………
<taxonomicName id="47BB86BFFFF4FF8DFC6BFB66FAA2519F" authority="Wesener, 2014" authorityName="Wesener" authorityYear="2014" box="[1012,1399,1225,1252]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="andrahomana">
<emphasis id="B2CF212EFFF4FF8DFC6BFB66FB155198" bold="true" box="[1012,1216,1225,1252]" italics="true" pageId="5" pageNumber="6">S. andrahomana</emphasis>
<bibRefCitation id="E42A80CDFFF4FF8DFB57FB66FAA2519F" author="Wesener T." box="[1224,1399,1225,1251]" journalOrPublisher="Zootaxa" pageId="5" pageNumber="6" pagination="587 - 594" part="3790" publicationUrl="https://doi.org/10.11646/zootaxa.3790.4.7" refId="ref12228" refString="Wesener T. 2014 a. Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda). Zootaxa 3790 (4): 587 - 594. https: // doi. org / 10.11646 / zootaxa. 3790.4.7" title="Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda)" type="journal article" year="2014">Wesener, 2014</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B4FE599FFF4FF8DFF22FAA3FAA25012" pageId="5" pageNumber="6">
<paragraph id="8004FD3CFFF4FF8DFF22FAA3FAA35036" blockId="5.[189,1406,267,2027]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF22FAA3FAA35036" pageId="5" pageNumber="6">
7. Unique black pattern on orange-reddish basic colour. Harp with 5 ribs. Endotergum with three rows of marginal bristles. Widespread in the SW spiny forest ……
<taxonomicName id="47BB86BFFFF4FF8DFC3BFA9FFAA35036" authority="(Saussure &amp; Zehntner, 1897)" authorityName="" authorityYear="1897" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" box="[932,1398,1328,1354]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="musicus">
<emphasis id="B2CF212EFFF4FF8DFC3BFA9FFBF15036" bold="true" box="[932,1060,1328,1354]" italics="true" pageId="5" pageNumber="6">
S.
<taxonomicName id="47BB86BFFFF4FF8DFC5EFA9FFBF15036" box="[961,1060,1328,1354]" pageId="5" pageNumber="6" rank="species" species="musicus">musicus</taxonomicName>
</emphasis>
(Saussure &amp; Zehntner, 1897)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8004FD3CFFF4FF8DFF5EFAFAFAA25012" blockId="5.[189,1406,267,2027]" box="[193,1399,1364,1391]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF5EFAFAFAA25012" box="[193,1399,1364,1391]" pageId="5" pageNumber="6">– Colour different, either uniformly black or brown. Rainforest species ……………………………8</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B4FE599FFF4FF8DFF22FA38FAA35362" pageId="5" pageNumber="6">
<paragraph id="8004FD3CFFF4FF8DFF22FA38FAA25086" blockId="5.[189,1406,267,2027]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF22FA38FAA25086" pageId="5" pageNumber="6">
8. Harp with 4 stridulation ribs. Endotergum with two rows of marginal bristles, not reaching tergite margin. Operculum well-rounded. Dark brown to black species. Lavasoa Mountain ……… ……………………………………………………………………………
<taxonomicName id="47BB86BFFFF4FF8DFBA2FA4FFAA25086" authority="Wesener, 2014" authorityName="Wesener" authorityYear="2014" box="[1085,1399,1503,1530]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lavasoa">
<emphasis id="B2CF212EFFF4FF8DFBA2FA4FFB6D5086" bold="true" box="[1085,1208,1503,1530]" italics="true" pageId="5" pageNumber="6">S. lavasoa</emphasis>
<bibRefCitation id="E42A80CDFFF4FF8DFB56FA4FFAA25086" author="Wesener T." box="[1225,1399,1504,1530]" journalOrPublisher="Zootaxa" pageId="5" pageNumber="6" pagination="587 - 594" part="3790" publicationUrl="https://doi.org/10.11646/zootaxa.3790.4.7" refId="ref12228" refString="Wesener T. 2014 a. Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda). Zootaxa 3790 (4): 587 - 594. https: // doi. org / 10.11646 / zootaxa. 3790.4.7" title="Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda)" type="journal article" year="2014">Wesener, 2014</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8004FD3CFFF4FF8DFF5EF9AAFAA35362" blockId="5.[189,1406,267,2027]" box="[193,1398,1540,1567]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF5EF9AAFAA35362" box="[193,1398,1540,1567]" pageId="5" pageNumber="6">– Harp with 5 or 6 stridulation ribs. Endotergum with one to three rows of marginal bristles …9</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B4FE599FFF4FF8DFF22F9E8FAA353B2" pageId="5" pageNumber="6">
<paragraph id="8004FD3CFFF4FF8DFF22F9E8FAA353F9" blockId="5.[189,1406,267,2027]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF22F9E8FAA353F9" pageId="5" pageNumber="6">
9. Harp with 5 stridulation ribs (
<figureCitation id="1880E1B9FFF4FF8DFDF9F9E8FD1C531D" box="[614,713,1606,1633]" captionStart="Fig. 5" captionStartId="11.[189,232,1788,1814]" captionTargetBox="[193,1396,571,1748]" captionTargetId="figure@11.[192,1396,569,1748]" captionTargetPageId="11" captionText="Fig. 5. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♂, holotype (CASENT 9068297-A). A.1 st leftcoxaeandstigmacarryingplate.B.2 nd leftcoxaandstigmacarryingplate.C.9 th leftleg.D.Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Left anterior telopod, lateral view. G. Left posterior telopod, anterior view. H. Left posterior telopod, posterior view.Abbreviations: as = anterior spine; cl = claw; ct = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; lb = lobe; pof = postfemur; pref =prefemur; sp = spine;sr =stridulation rib; st =stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scalebars =1 mm." httpUri="https://zenodo.org/record/1118120/files/figure.png" pageId="5" pageNumber="6">Fig. 5D</figureCitation>
). Movable finger of posterior telopod with one or two membranous lobes. Marginal bristles of endotergum protruding beyond tergite margin (
<figureCitation id="1880E1B9FFF4FF8DFB49F9C4FAFB53F9" box="[1238,1326,1643,1669]" captionStart="Fig.4" captionStartId="10.[189,232,1928,1954]" captionTargetBox="[189,1401,1078,1882]" captionTargetId="figure@10.[189,1401,1078,1883]" captionTargetPageId="10" captionText="Fig.4.Sphaeromimuskalambatritra sp. nov., SEM of endotergum, ♂, holotype (CASENT9068297-A). A.Overview,endotergum.B. Marginofendotergum,detail.Abbreviations:(1)=innerpart; (2)=middle part; (3) =outer part." httpUri="https://zenodo.org/record/1118118/files/figure.png" pageId="5" pageNumber="6">Fig. 4B</figureCitation>
) …10
</keyLead>
</paragraph>
<paragraph id="8004FD3CFFF4FF8DFF5EF93FFAA353B2" blockId="5.[189,1406,267,2027]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF5EF93FFAA353B2" pageId="5" pageNumber="6">
– Harp with 6 stridulation ribs (
<figureCitation id="1880E1B9FFF4FF8DFDD3F920FD7F53D6" box="[588,682,1679,1706]" captionStart="Fig. 9" captionStartId="15.[189,232,1789,1815]" captionTargetBox="[189,1399,522,1754]" captionTargetId="figure@15.[189,1399,518,1754]" captionTargetPageId="15" captionText="Fig. 9. Sphaeromimus midongy  sp.  nov., schematic drawing,  ♂,  holotype (FMNH-INS 3119888). A. 1st left coxae and stima carrying plate. B. 2nd left coxae and stigma carrying plate. C. 9th left leg. D. Anterior view of right anterior telopod. E. Posterior view of right anterior telopod. F. Lateral view of right anterior telopod. G. Left posterior telopod, anterior view. H. Posterior view of left posterior telopod.Abbreviations:as =anterior spine; cl = claw;ct =crenulated teeth; cx= coxa; fe =femur; gp = gonopore; lb = lobe; pof = postfemur; pref = prefemur; sp = spine; sr = stridulation rib; st = stigmatic plate; syn =syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scale bars = 1 mm." httpUri="https://zenodo.org/record/1118128/files/figure.png" pageId="5" pageNumber="6">Fig.9D</figureCitation>
). Movable finger of posterior telopod with two membranous lobes (
<figureCitation id="1880E1B9FFF4FF8DFEA3F91BFE4953B2" box="[316,412,1716,1742]" captionStart="Fig. 9" captionStartId="15.[189,232,1789,1815]" captionTargetBox="[189,1399,522,1754]" captionTargetId="figure@15.[189,1399,518,1754]" captionTargetPageId="15" captionText="Fig. 9. Sphaeromimus midongy  sp.  nov., schematic drawing,  ♂,  holotype (FMNH-INS 3119888). A. 1st left coxae and stima carrying plate. B. 2nd left coxae and stigma carrying plate. C. 9th left leg. D. Anterior view of right anterior telopod. E. Posterior view of right anterior telopod. F. Lateral view of right anterior telopod. G. Left posterior telopod, anterior view. H. Posterior view of left posterior telopod.Abbreviations:as =anterior spine; cl = claw;ct =crenulated teeth; cx= coxa; fe =femur; gp = gonopore; lb = lobe; pof = postfemur; pref = prefemur; sp = spine; sr = stridulation rib; st = stigmatic plate; syn =syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scale bars = 1 mm." httpUri="https://zenodo.org/record/1118128/files/figure.png" pageId="5" pageNumber="6">Fig. 9G</figureCitation>
) ……………………………………………………………………………………11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3B4FE599FFF4FF8DFF22F958FAA25297" pageId="5" pageNumber="6">
<paragraph id="8004FD3CFFF4FF8DFF22F958FAA25225" blockId="5.[189,1406,267,2027]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF22F958FAA25225" pageId="5" pageNumber="6">
10. Operculum apically recessed. Medium-sized species. Movable finger of posterior telopod with single membranous lobe and 26 crenulated teeth. Eyes with&gt;60 ocelli. Rudimentary lateral palpi consisting of 3 sensilla. Vevembe-Vatovavy area …
<taxonomicName id="47BB86BFFFF4FF8DFBBEF890FAA25225" authority="Wesener, 2014" authorityName="Wesener" authorityYear="2014" box="[1057,1399,1855,1882]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vatovavy">
<emphasis id="B2CF212EFFF4FF8DFBBEF890FB645226" bold="true" box="[1057,1201,1855,1882]" italics="true" pageId="5" pageNumber="6">S. vatovavy</emphasis>
<bibRefCitation id="E42A80CDFFF4FF8DFB5EF8EFFAA25225" author="Wesener T." box="[1217,1399,1855,1882]" journalOrPublisher="Zootaxa" pageId="5" pageNumber="6" pagination="587 - 594" part="3790" publicationUrl="https://doi.org/10.11646/zootaxa.3790.4.7" refId="ref12228" refString="Wesener T. 2014 a. Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda). Zootaxa 3790 (4): 587 - 594. https: // doi. org / 10.11646 / zootaxa. 3790.4.7" title="Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda)" type="journal article" year="2014">Wesener, 2014</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8004FD3CFFF4FF8DFF5EF8CBFAA25297" blockId="5.[189,1406,267,2027]" pageId="5" pageNumber="6">
<keyLead id="3B4A5E09FFF4FF8DFF5EF8CBFAA25297" pageId="5" pageNumber="6">
– Operculum well-rounded, typical for genus (
<figureCitation id="1880E1B9FFF4FF8DFC81F8CCFC505202" box="[798,901,1891,1918]" captionStart="Fig. 6" captionStartId="12.[189,232,758,784]" captionTargetBox="[189,1396,263,730]" captionTargetId="figure@12.[189,1396,263,730]" captionTargetPageId="12" captionText="Fig. 6. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♀, paratype (CASENT 9058301). A. Left vulva (2 nd coxa). B. Subanal plate. Abbreviations: cx = coxa; ep = exterior plate; ip = interior plate; op = operculum; pref =prefemur; sr = stridulation ribs. Scale bars =1 mm." httpUri="https://zenodo.org/record/1118122/files/figure.png" pageId="5" pageNumber="6">Fig. 6A</figureCitation>
). Gigantic species (&gt;
<quantity id="474350D9FFF4FF8DFB0DF8CCFB265202" box="[1170,1267,1891,1918]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="6" unit="mm" value="50.0">50 mm</quantity>
). Movable finger of posterior telopod with two membranous lobes and 30 crenulated teeth (
<figureCitation id="1880E1B9FFF4FF8DFB5BF827FAF152DF" box="[1220,1316,1928,1955]" captionStart="Fig. 5" captionStartId="11.[189,232,1788,1814]" captionTargetBox="[193,1396,571,1748]" captionTargetId="figure@11.[192,1396,569,1748]" captionTargetPageId="11" captionText="Fig. 5. Sphaeromimus kalambatritra sp. nov., schematic drawing, ♂, holotype (CASENT 9068297-A). A.1 st leftcoxaeandstigmacarryingplate.B.2 nd leftcoxaandstigmacarryingplate.C.9 th leftleg.D.Left anterior telopod, anterior view. E. Left anterior telopod, posterior view. F. Left anterior telopod, lateral view. G. Left posterior telopod, anterior view. H. Left posterior telopod, posterior view.Abbreviations: as = anterior spine; cl = claw; ct = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; lb = lobe; pof = postfemur; pref =prefemur; sp = spine;sr =stridulation rib; st =stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scalebars =1 mm." httpUri="https://zenodo.org/record/1118120/files/figure.png" pageId="5" pageNumber="6">Fig.5H</figureCitation>
). Eyes with&gt;80 ocelli. Rudimentary lateral palpi consisting of 4 sensilla (
<figureCitation id="1880E1B9FFF4FF8DFBCEF803FB6352BA" box="[1105,1206,1964,1990]" captionStart="Fig. 3" captionStartId="9.[189,232,1825,1851]" captionTargetBox="[189,1399,438,1783]" captionTargetId="figure@9.[189,1399,438,1783]" captionTargetPageId="9" captionText="Fig. 3.Sphaeromimus kalambatritra sp. nov., SEM of mouthparts, ♂, holotype (CASENT9068297-A). A. Gnathochilarium overview, ventral. B. Gnathochilarium overview, frontodorsal. C. Lateral palpus, detail.D.Innerpalpus, detail.E. Centralpad, detail.F.Rightmandible,overview.G.Pectinatelamellae, detail.Abbreviations:C=condyles;cP =centralpad;eT =exteriorteeth; Hyp=hypopharynx;iP =inner palpus; iT = inner teeth; LL = lamella lingualis; LP = lateral palpus; Me = mentum; mp = molar plate; pl = pectinate lamellae; ST = stipes of gnathochilarium." httpUri="https://zenodo.org/record/1118116/files/figure.png" pageId="5" pageNumber="6">Fig. 3C</figureCitation>
). Kalambatritra Rainforest …………………………………………………………………
<taxonomicName id="47BB86BFFFF4FF8DFBD4F87EFAC45297" authority="Moritz &amp; Wesener, 2017" authorityName="Moritz &amp; Wesener" authorityYear="2017" box="[1099,1297,2000,2027]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="kalambatritra" status="sp. nov.">
<emphasis id="B2CF212EFFF4FF8DFBD4F87EFAC45297" bold="true" box="[1099,1297,2000,2027]" italics="true" pageId="5" pageNumber="6">S. kalambatritra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9FC9C55FFF4FF8DFA84F87EFAA25297" box="[1307,1399,2001,2027]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<keyStep id="3B4FE599FFF7FF8EFF22FEA4FAA254A4" pageId="6" pageNumber="7">
<paragraph id="8004FD3CFFF7FF8EFF22FEA4FAA25411" blockId="6.[189,1399,267,472]" pageId="6" pageNumber="7">
<keyLead id="3B4A5E09FFF7FF8EFF22FEA4FAA25411" pageId="6" pageNumber="7">
11. Small, black species, red legs. Endotergum with single sparse row of very short marginal bristles. Posterior telopod with four spines and 36 crenulated teeth. Ivohibe Mountain ………… ……………………………………………………………………………
<taxonomicName id="47BB86BFFFF7FF8EFBA2FEFDFAA25411" authority="Wesener, 2014" authorityName="Wesener" authorityYear="2014" box="[1085,1399,338,365]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="ivohibe">
<emphasis id="B2CF212EFFF7FF8EFBA2FEFDFB6C5411" bold="true" box="[1085,1209,338,365]" italics="true" pageId="6" pageNumber="7">S. ivohibe</emphasis>
<bibRefCitation id="E42A80CDFFF7FF8EFB5AFEFCFAA25411" author="Wesener T." box="[1221,1399,339,365]" journalOrPublisher="Zootaxa" pageId="6" pageNumber="7" pagination="587 - 594" part="3790" publicationUrl="https://doi.org/10.11646/zootaxa.3790.4.7" refId="ref12228" refString="Wesener T. 2014 a. Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda). Zootaxa 3790 (4): 587 - 594. https: // doi. org / 10.11646 / zootaxa. 3790.4.7" title="Redescription of Polyzonium malagassum, a new synonym of Rhinotus purpureus (Pocock, 1894), with notes about the occurrence of the order Polyzoniida on Madagascar (Diplopoda)" type="journal article" year="2014">Wesener, 2014</bibRefCitation>
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</paragraph>
<paragraph id="8004FD3CFFF7FF8EFF5EFED8FAA254A4" blockId="6.[189,1399,267,472]" pageId="6" pageNumber="7">
<keyLead id="3B4A5E09FFF7FF8EFF5EFED8FAA254A4" pageId="6" pageNumber="7">
– Gigantic (&gt;
<quantity id="474350D9FFF7FF8EFEEBFED9FE0554ED" box="[372,464,374,401]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="6" pageNumber="7" unit="mm" value="50.0">50 mm</quantity>
), brown species. Endotergum with two dense rows of long marginal bristles (
<figureCitation id="1880E1B9FFF7FF8EFF6CFE35FE9B54C8" box="[243,334,410,436]" captionStart="Fig. 9" captionStartId="15.[189,232,1789,1815]" captionTargetBox="[189,1399,522,1754]" captionTargetId="figure@15.[189,1399,518,1754]" captionTargetPageId="15" captionText="Fig. 9. Sphaeromimus midongy  sp.  nov., schematic drawing,  ♂,  holotype (FMNH-INS 3119888). A. 1st left coxae and stima carrying plate. B. 2nd left coxae and stigma carrying plate. C. 9th left leg. D. Anterior view of right anterior telopod. E. Posterior view of right anterior telopod. F. Lateral view of right anterior telopod. G. Left posterior telopod, anterior view. H. Posterior view of left posterior telopod.Abbreviations:as =anterior spine; cl = claw;ct =crenulated teeth; cx= coxa; fe =femur; gp = gonopore; lb = lobe; pof = postfemur; pref = prefemur; sp = spine; sr = stridulation rib; st = stigmatic plate; syn =syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scale bars = 1 mm." httpUri="https://zenodo.org/record/1118128/files/figure.png" pageId="6" pageNumber="7">Fig. 9B</figureCitation>
). Posterior telopod with two spines and 32 crenulated teeth (
<figureCitation id="1880E1B9FFF7FF8EFC64FE35FB8254C8" box="[1019,1111,410,436]" captionStart="Fig. 9" captionStartId="15.[189,232,1789,1815]" captionTargetBox="[189,1399,522,1754]" captionTargetId="figure@15.[189,1399,518,1754]" captionTargetPageId="15" captionText="Fig. 9. Sphaeromimus midongy  sp.  nov., schematic drawing,  ♂,  holotype (FMNH-INS 3119888). A. 1st left coxae and stima carrying plate. B. 2nd left coxae and stigma carrying plate. C. 9th left leg. D. Anterior view of right anterior telopod. E. Posterior view of right anterior telopod. F. Lateral view of right anterior telopod. G. Left posterior telopod, anterior view. H. Posterior view of left posterior telopod.Abbreviations:as =anterior spine; cl = claw;ct =crenulated teeth; cx= coxa; fe =femur; gp = gonopore; lb = lobe; pof = postfemur; pref = prefemur; sp = spine; sr = stridulation rib; st = stigmatic plate; syn =syncoxite; ta = tarsus; ti = tibia; vs = ventral spine; 1–4 = podomeres. Scale bars = 1 mm." httpUri="https://zenodo.org/record/1118128/files/figure.png" pageId="6" pageNumber="7">Fig. 9H</figureCitation>
). Midongy Rainforest … …………………………………………………………………………………
<taxonomicName id="47BB86BFFFF7FF8EFBE7FE12FAD354A4" authority="Moritz &amp; Wesener, 2017" authorityName="Moritz &amp; Wesener" authorityYear="2017" box="[1144,1286,445,472]" class="Diplopoda" family="Sphaerotheriidae" genus="Sphaeromimus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sphaerotheriida" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="midongy" status="sp. nov.">
<emphasis id="B2CF212EFFF7FF8EFBE7FE12FAD354A4" bold="true" box="[1144,1286,445,472]" italics="true" pageId="6" pageNumber="7">S. midongy</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A9FC9C55FFF7FF8EFA8AFE11FAA254A4" box="[1301,1399,446,472]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
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</treatment>
</document>
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