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<document id="55367942430BB7A0188E55B9DA76AD4D" ID-CLB-Dataset="26691" ID-DOI="http://dx.doi.org/10.3897/evolsyst.2.30778" ID-GBIF-Dataset="00106674-20d9-4d70-bb0f-8f000fb311f2" ID-Pensoft-Pub="2535-0730-2-169" ID-ZooBank="C0E89FEC8BE54DE9803D784FF6727BA0" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="2535-0730-2-169" ModsDocOrigin="Evolutionary Systematics 2 (2)" ModsDocTitle="The wolf spider genus Artoria in New South Wales and the Australian Capital Territory, Australia (Araneae, Lycosidae, Artoriinae)" checkinTime="1558621757476" checkinUser="pensoft" docAuthor="Framenau, Volker W. &amp; Baehr, Barbara C." docDate="2018" docId="06E7300E55DCD2F17314FCE353EB5AE4" docLanguage="en" docName="EvolutSyst 2(2): 169-241" docOrigin="Evolutionary Systematics 2 (2)" docSource="http://dx.doi.org/10.3897/evolsyst.2.30778" docTitle="Artoria helensmithae Framenau &amp; Baehr, 2018, sp. n." docType="treatment" docUuid="2E554E14-1913-49EF-8853-E0CDF2ABD877" docUuidSource="ZooBank" docVersion="6" lastPageNumber="169" masterDocId="FFE8FFB72743554D943BFFB83D324B46" masterDocTitle="The wolf spider genus Artoria in New South Wales and the Australian Capital Territory, Australia (Araneae, Lycosidae, Artoriinae)" masterLastPageNumber="241" masterPageNumber="169" pageNumber="169" updateTime="1701381204780" updateUser="ExternalLinkService">
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<mods:title id="E4CFAAB2B6C850006A1109A3993CDE78">The wolf spider genus Artoria in New South Wales and the Australian Capital Territory, Australia (Araneae, Lycosidae, Artoriinae)</mods:title>
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<mods:namePart id="E6B0E969D8C0284AA415ED2A30445539">Framenau, Volker W.</mods:namePart>
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<mods:namePart id="E20C80935EDD4DBE976327499B077529">Baehr, Barbara C.</mods:namePart>
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<mods:title id="9AAC49251DFF4A92E409364F49F911C5">Evolutionary Systematics</mods:title>
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<mods:date id="F857D57EF5FBE5F96795CB32E069CB99">2018</mods:date>
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<treatment id="06E7300E55DCD2F17314FCE353EB5AE4" ID-GBIF-Taxon="157251919" LSID="urn:lsid:zoobank.org:act:2E554E14-1913-49EF-8853-E0CDF2ABD877" httpUri="http://treatment.plazi.org/id/06E7300E55DCD2F17314FCE353EB5AE4" lastPageNumber="169" pageId="0" pageNumber="169">
<subSubSection id="C69032A62E4E422BBECA223F4DD31C62" pageId="0" pageNumber="169" type="nomenclature">
<paragraph id="0C4E113FF12D7F96BBDD85D0422D3FCA" pageId="0" pageNumber="169">
<taxonomicName id="46D09F8B9566A04845BDD84F35D12D6B" ID-CoL="H2BS" LSID="http://zoobank.org/2E554E14-1913-49EF-8853-E0CDF2ABD877" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria helensmithae" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="helensmithae">Artoria helensmithae</taxonomicName>
<taxonomicNameLabel id="F17F85F5DA07BAFA826B1B8BCCBC0A7C" pageId="0" pageNumber="169">sp. n.</taxonomicNameLabel>
Figs 23, 24
<normalizedToken id="9D39842CBFFA569BF329CF9E57AC24E0" originalValue="AH">A-H</normalizedToken>
, 47K
<normalizedToken id="399643240D7ABA6029C96A47689B78A1" originalValue="Helens">Helen's</normalizedToken>
Forest Runner
</paragraph>
</subSubSection>
<subSubSection id="93F623DF568CED46D33A34F5CE6C1877" pageId="0" pageNumber="169" type="materials_examined">
<paragraph id="DD702892FACB921B0B950A8653F9C10D" pageId="0" pageNumber="169">Material examined.</paragraph>
<paragraph id="08481B98CB5A3DD291E883285022462E" pageId="0" pageNumber="169">
Holotype, Booti Booti National Park, S of Forster, southern end of park (
<geoCoordinate id="D9F329668880939C49DCBE2965F0F519" direction="south" orientation="latitude" precision="15" value="-32.27972">
32°16
<normalizedToken id="1EF134415EF546031B0276C0F9FFB223" originalValue="47”">'47&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="BDFC339D9EAE708FF911135A50AE0858" direction="east" orientation="longitude" precision="15" value="152.52444">
152°31
<normalizedToken id="FA6934890B53BAC5A92B668E2220FF78" originalValue="28”">'28&quot;</normalizedToken>
E
</geoCoordinate>
, New South Wales, AUSTRALIA), 30
<normalizedToken id="332A7F602DF0FF7138AA42089E5B325A" originalValue="September">September-</normalizedToken>
9 October 1997, L. Wilkie, pitfall trap (AM KS128076). Paratypes: 4 males, same data as holotype (AM KS61893); 1 female, Munmorah State Recreation Area,
<geoCoordinate id="C52CC21904D5CFC7EAEA00E6040F9A0E" direction="south" orientation="latitude" precision="15" value="-33.207222">
33°12
<normalizedToken id="A75ACEAC5AA06A149762F88E9F48F092" originalValue="26”">'26&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="13243D29157C8857BD1B51AE2650E0E3" direction="east" orientation="longitude" precision="15" value="151.57695">
151°34
<normalizedToken id="D34D1250AB33425FE0D75E8728D51B7C" originalValue="37”">'37&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS61928); 7 females, Booti Booti National Park (
<geoCoordinate id="EA74C3F0076856124187DA553FAC09A7" direction="south" orientation="latitude" precision="15" value="-32.24111">
32°14
<normalizedToken id="57DD3BFE6B66C98DF40AF641EB535B9C" originalValue="28”">'28&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="2DCC212CEDA425A7F5711587D602D9A3" direction="east" orientation="longitude" precision="15" value="152.54723">
152°32
<normalizedToken id="14FDDC54E84954E15000E099BA78FD4A" originalValue="50”">'50&quot;</normalizedToken>
E
</geoCoordinate>
, New South Wales, AUSTRALIA), 25 November 1997, pitfall trap, L. Wilkie, BBC02/03 (AM KS61914); 1 female, same data (ZMH A0002168); 33 males, Bungonia Caves area, near Information Centre (
<geoCoordinate id="D264B64D07F4E858DC59AD1CF36BB6EE" direction="south" orientation="latitude" precision="15" value="-34.800556">
34°48
<normalizedToken id="4D838E59281E47C26C07927B07528B6E" originalValue="02”">'02&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="F89599DB68CFFC3E138FCD0BBCBDC3E7" direction="east" orientation="longitude" precision="15" value="150.01584">
150°00
<normalizedToken id="891E6E438FA7FA5BE0A82C3C0231D552" originalValue="57”">'57&quot;</normalizedToken>
E
</geoCoordinate>
, New South Wales, AUSTRALIA),
<normalizedToken id="29851899B9E7D9D9F64F8FBD49864C87" originalValue="mid-Octobermid-November">mid-October-mid-November</normalizedToken>
1989, G. Hunt, pitfall trap (AM KS22556); 1 male, same data (ZSMH A0002169).
</paragraph>
<caption id="07E25CE369F53C38D485BCA3FBC61E54" pageId="0" pageNumber="169">
<paragraph id="2E91B2D7A072979285AF7EF2615226BD" pageId="0" pageNumber="169">
Figure 23. Distribution records of
<taxonomicName id="7E930DBE0F9ECB6B92BC09925FBE510C" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria grahammilledgei" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="grahammilledgei">Artoria grahammilledgei</taxonomicName>
sp. n. (open circles),
<taxonomicName id="A9CB2510DA3DE7609D219227F5D9D406" lsidName="A. helensmithae" pageId="0" pageNumber="169" rank="species" species="helensmithae">A. helensmithae</taxonomicName>
sp. n. (full circles) and
<taxonomicName id="90840FA9AC775AD1018993B0134F8864" lsidName="A. howquaensis" pageId="0" pageNumber="169" rank="species" species="howquaensis">A. howquaensis</taxonomicName>
Framenau, 2002 (grey squares) in NSW. IBRA bioregions with spider records: BBS - Brigalow Belt South; NAN - Nandewar; NET - New England Tablelands; NNC - NSW North Coast; SEQ - South East Queensland; SEH - South Eastern Highlands; SEC - South East Corner; SYB - Sydney Basin.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="04B88DF3AA2717645A5046733FF4FEA6" pageId="0" pageNumber="169" type="materials_examined">
<paragraph id="02F19F791AE1DD51632050102B262488" pageId="0" pageNumber="169">Other material examined.</paragraph>
<paragraph id="FEDD1EBD51BA971643E93542C5F19620" pageId="0" pageNumber="169">
175 males, 100 females (2 with spiderlings) and 1 juvenile 134 records. AUSTRALIA: New South Wales: 2 males, 4 km NE of Mt Wog Wog, 17 km SE Bombala,
<geoCoordinate id="9FA6663C35BB7629B37F5D37FD0FFAB7" direction="south" orientation="latitude" precision="15" value="-37.075">
37°04
<normalizedToken id="366403BDC1A41711E43ABECAA68698FD" originalValue="30”">'30&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="29D275C00726DA0DD3AC7F05CB2875EB" direction="east" orientation="longitude" precision="15" value="149.46666">
149°28
<normalizedToken id="2B3700AFA4ECF216D5E414E7696747E0" originalValue="00”">'00&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS98791, KS98810); 1 male, Barrington Cave Pigna Barney PB1-20,
<geoCoordinate id="257A890E2CB30F08C65EEBD5ADE15322" direction="south" orientation="latitude" precision="15" value="-31.773333">
31°46
<normalizedToken id="452AFA45AC137B2BC1D1EBD1742347DB" originalValue="24”">'24&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="109514D9DAF99BF7E1091140E486DCCF" direction="east" orientation="longitude" precision="15" value="151.52138">
151°31
<normalizedToken id="B970A49560B25D46BB53EEF84A0B2F57" originalValue="17”">'17&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS52386); 3 males, 1 female, Bondi State Forest, S of Bombala,
<geoCoordinate id="0B46B3F287B479A466650834B81BE80F" direction="south" orientation="latitude" precision="925" value="-37.116665">37°07'S</geoCoordinate>
,
<geoCoordinate id="AEB87AACF96D0784673AC3E1FAA29391" direction="east" orientation="longitude" precision="925" value="149.13333">149°08'E</geoCoordinate>
(AM KS18203-5, KS70234); 47 males, 21 females, Booti Booti National Park,
<geoCoordinate id="7D1B2890CF99F00FAEDF15771950D498" direction="south" orientation="latitude" precision="15" value="-32.24111">
32°14
<normalizedToken id="150FE2F3878AA31ACDB31C1C13E24BD2" originalValue="28”">'28&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="9FE77567882309A2106F61BE66B57F09" direction="east" orientation="longitude" precision="15" value="152.54723">
152°32
<normalizedToken id="3D7970E15B148B5B17ED077F71EBBD50" originalValue="50”">'50&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS50502-13, KS52069-70, KS61861, KS61868, KS61872, KS61875, KS61879, KS61881, KS61889, KS61892-4, KS61897-8, KS61909, KS61911-2, KS61919, KS61923, KS61927, KS61935-6, KS61978, KS64040-48); 2 males, Cherry Tree north State Forest,
<geoCoordinate id="E942F69F477380FEA9B3258081DAB591" direction="south" orientation="latitude" precision="925" value="-28.966667">28°58'S</geoCoordinate>
,
<geoCoordinate id="4DA959F5E7E4DA57F0BDA5BFFF52EEE1" direction="east" orientation="longitude" precision="925" value="152.25">152°15'E</geoCoordinate>
(AM KS52392); 1 female, Dover Heights, Rodney Reserve,
<geoCoordinate id="4AC3F28415AE95FBA13868CEB940FCFA" direction="south" orientation="latitude" precision="15" value="-33.873886">
33°52
<normalizedToken id="10AA105F0EDFE1802B9DCE225ED5709B" originalValue="26”">'26&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="F06CD51B7C498583DE6C55306B4FC390" direction="east" orientation="longitude" precision="15" value="151.28305">
151°16
<normalizedToken id="4B67BB563D88F209AF5DACB1DC2AC7B3" originalValue="59”">'59&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS110534); 2 males, 6 females, 1 juv., Harrington, 3-5 km NE,
<geoCoordinate id="80F400EC4A3D1EA61EF893537FA28A7E" direction="south" orientation="latitude" precision="925" value="-31.866667">31°52'S</geoCoordinate>
,
<geoCoordinate id="DD8F519190695894B73D8F1F31B11954" direction="east" orientation="longitude" precision="925" value="152.7">152°42'E</geoCoordinate>
(AM KS23427); 1 male, 1 female, Munmorah State Conservation Area, 30 m off Birdie Beach Drive and 200m north of Freemans,
<geoCoordinate id="33FE376141C1E0EC177EB6516C2408EE" direction="south" orientation="latitude" precision="15" value="-33.204723">
33°12
<normalizedToken id="7779A4F1ADADDF1D139B87E35BCB6686" originalValue="17”">'17&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="A268731BD87CB54312B5D0B70695F82C" direction="east" orientation="longitude" precision="15" value="151.60251">
151°36
<normalizedToken id="40027B8C2CB7BCA5B8385743DC4CC05B" originalValue="9”">'9&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS122654, KS122908); 10 males, Munmorah State Recreation Area,
<geoCoordinate id="C66851E7E5B76E41C31D0F32485590B5" direction="south" orientation="latitude" precision="15" value="-33.20667">
33°12
<normalizedToken id="26F74432043019A81DC4A2CE8C1BC597" originalValue="24”">'24&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="6C5226A6108DC94EE1692E98D51888D1" direction="east" orientation="longitude" precision="15" value="151.58305">
151°34
<normalizedToken id="188F7CAF3CAA96E9674B54DFA5724539" originalValue="59”">'59&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS61862, KS61864); 2 males, Munmorah State Recreation Area,
<geoCoordinate id="D3A5FD9A49A03DDAA3BC4B62689C0F59" direction="south" orientation="latitude" precision="15" value="-33.207222">
33°12
<normalizedToken id="4ED591F8323B619CD1AF0EA61E90ACE2" originalValue="26”">'26&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="3DB8C760381EAB317945433106434EB3" direction="east" orientation="longitude" precision="15" value="151.57695">
151°34
<normalizedToken id="58599675F202D90CE05635F352DDB49D" originalValue="37”">'37&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS61852-3); 1 female, Munmorah State Recreation Area,
<geoCoordinate id="BB99783F252C6BE9379FC12981A0F3F0" direction="south" orientation="latitude" precision="15" value="-33.2075">
33°12
<normalizedToken id="DB9FD0AF83338976138885187CE816D7" originalValue="27”">'27&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="701BD8CFEAD55BE08ED8521B84F6E6EE" direction="east" orientation="longitude" precision="15" value="151.57695">
151°34
<normalizedToken id="444035593223861C23576D7F4CC482B5" originalValue="37”">'37&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS61907); 15 males, 9 females, Munmorah State Recreation Area,
<geoCoordinate id="BB2404E7AB619DD2D9331D887AB2A09C" direction="south" orientation="latitude" precision="15" value="-33.209446">
33°12
<normalizedToken id="40B214E461900C229E337EEE77FCC883" originalValue="34”">'34&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="031FD5F156252D819B9D2074FED0CC49" direction="east" orientation="longitude" precision="15" value="151.58305">
151°34
<normalizedToken id="83B5CF4B1D90D9CEA6C6BD55CB1320F4" originalValue="59”">'59&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS61848, KS61856, KS61876, KS61883-4, KS61896, KS61908, KS61930-31, KS61933-4, KS86380); 18 males, 1 female, Munmorah State Recreation Area,
<geoCoordinate id="11AFB5833EC88B6C4BD5070FBD3A2107" direction="south" orientation="latitude" precision="15" value="-33.219166">
33°13
<normalizedToken id="A3680775DD54DB35DB19CD565F46DA2A" originalValue="09”">'09&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="389195534BF90F2731470234E21AA02C" direction="east" orientation="longitude" precision="15" value="151.57083">
151°34
<normalizedToken id="25F53C9A43EAAB7F79F3E18F84DCD531" originalValue="15”">'15&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS61859, KS61871, KS61885, KS61891, KS61903, KS64049); 1 male, 1 female, Myall Lakes National Park,
<geoCoordinate id="B3B309808298AD36759D35B8811DA5D8" direction="south" orientation="latitude" precision="15" value="-32.489445">
32°29
<normalizedToken id="A03EC947F62F1E002D9D527008DBA20F" originalValue="22”">'22&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="FD79C22AF25B3D8BBA079C8E98AF789D" direction="east" orientation="longitude" precision="15" value="152.39806">
152°23
<normalizedToken id="5468CDECBEDCFD0F8514D429CCA566C3" originalValue="53”">'53&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS61887, KS64051); 2 males, Myall Lakes National Park,
<geoCoordinate id="0A44E16348DD8F90C62B7FC296284530" direction="south" orientation="latitude" precision="15" value="-32.50722">
32°30
<normalizedToken id="A9A7FDA9EECFBBD047945E4CAE36F210" originalValue="26”">'26&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="A01BEA6F9024D8C725DBDD457FEAB338" direction="east" orientation="longitude" precision="15" value="152.36528">
152°21
<normalizedToken id="FACEBC2B27E964EB8C1FFC35F196BB63" originalValue="55”">'55&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS61899); 2 males, 1 female, Myall Lakes National Park,
<geoCoordinate id="3F9A475FDDA24AFCCA6C5B8C2A7C8027" direction="south" orientation="latitude" precision="15" value="-32.579166">
32°34
<normalizedToken id="5C06A7D5DE62E6DAFAB0C0413B4BDEC7" originalValue="45”">'45&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="5306A1F7F69EDF0E04D3A7081D8DCD2C" direction="east" orientation="longitude" precision="15" value="152.29085">
152°17
<normalizedToken id="8CC54D4B15904AC90C2C36D840B741EF" originalValue="27”">'27&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS61900, KS64052); 5 male, 2 females, Myall Lakes National Park,
<geoCoordinate id="989F8CF12F1DA14E6D67566C0DF70D3F" direction="south" orientation="latitude" precision="15" value="-32.579166">
32°34
<normalizedToken id="8AF301421D24A3065E01F654936F4F54" originalValue="45”">'45&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="FD1E3A8F08350C698E60B2CC11ABC031" direction="east" orientation="longitude" precision="15" value="152.2075">
152°12
<normalizedToken id="7C441DAF53D7BE0D6E7828B12C86F467" originalValue="27”">'27&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS61849, KS61865, KS61873, KS61888, KS61890, KS61916); 9 male, 8 females, Myall Lakes National Park,
<geoCoordinate id="5B723D7D52D93FF06607008BF29C67E3" direction="south" orientation="latitude" precision="15" value="-32.63222">
32°37
<normalizedToken id="E2C51F42348EEB2156C22C159CC5CFD5" originalValue="56”">'56&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="D6A49114DC1D1F6E159A262445946483" direction="east" orientation="longitude" precision="15" value="152.2075">
152°12
<normalizedToken id="51A3603E32F15D7C57AEA73306E3BCE5" originalValue="27”">'27&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS61850-51, KS61855, KS61869, KS61878, KS61886, KS61906, KS61915, KS61921-2, KS61926); 4 males, 5 females, Nullica Beach,
<geoCoordinate id="C2F865E1E7E0A71551E5CDBE2A01A4E2" direction="south" orientation="latitude" precision="925" value="-37.1">37°06'S</geoCoordinate>
,
<geoCoordinate id="53406F8FFEFD02FC6A8B5E9ED7BC57DE" direction="east" orientation="longitude" precision="925" value="149.88333">149°53'E</geoCoordinate>
(AM KS83713, KS83718, KS83723); 2 females, Nullica River Bridge, S of river between bridge and coast,
<geoCoordinate id="7F61B117D5E135F5FD0E6B625FB50BE3" direction="south" orientation="latitude" precision="925" value="-37.066666">37°04'S</geoCoordinate>
,
<geoCoordinate id="D90E044F509A5CF4A87B721B44A7E978" direction="east" orientation="longitude" precision="925" value="149.8">149°48'E</geoCoordinate>
(WAM T56211); 1 male, Seven Mile Beach National Park, picnic area.,
<geoCoordinate id="43B6D9C7E95FDC0956D60B34D0ACF5E7" direction="south" orientation="latitude" precision="15" value="-34.791668">
34°47
<normalizedToken id="8E1B1BE88E312526D88D44608DECA119" originalValue="30”">'30&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="C4A5EDFFFE8101F2F136E42B4CD58AF7" direction="east" orientation="longitude" precision="15" value="150.77888">
150°46
<normalizedToken id="DED9588907576BCE5B49E9E261069386" originalValue="44”">'44&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS119173); 2 females, Tamarama, Marks Park,
<geoCoordinate id="E05EF0E9CCDBEF419397232A7FA970BB" direction="south" orientation="latitude" precision="15" value="-33.89889">
33°53
<normalizedToken id="FF587D811B4D0064E7D005279039B92C" originalValue="56”">'56&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="3A215CC71BAECE0C955B106317A64671" direction="east" orientation="longitude" precision="15" value="151.275">
151°16
<normalizedToken id="B751432C9893E34701673B5C16AE67B6" originalValue="30”">'30&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS110537); 1 female, Wallaroo State Forest, Flaggy Creek,
<geoCoordinate id="6AAEF959DEF9A743FB852251E6A82779" direction="south" orientation="latitude" precision="15" value="-32.60083">
32°36
<normalizedToken id="275F3C2D12D3E406E7765CD8E881B368" originalValue="03”">'03&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="4330C85F553A628629A3C9BC134CB95F" direction="east" orientation="longitude" precision="15" value="151.80194">
151°48
<normalizedToken id="A62B8633FEEB697BE8B03D70AA5D0D6E" originalValue="07”">'07&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS39707); 9 males, 20 females, Wyrrabalong National Park,
<geoCoordinate id="F16D587365AFBDA0E6709AA3A5460FFC" direction="south" orientation="latitude" precision="15" value="-33.27889">
33°16
<normalizedToken id="8ED7710C1F69C18519F67994D4174834" originalValue="44”">'44&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="BA19BB2E4402488C092AAEE911AC14A0" direction="east" orientation="longitude" precision="15" value="151.5475">
151°32
<normalizedToken id="B595591246760548C5EB0543E336AEDC" originalValue="51”">'51&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS61854, KS61874, KS61895, KS61904, KS61917, KS61920, KS61924-5, KS64053-5); 3 male, 3 females, Wyrrabalong National Park,
<geoCoordinate id="3E659BAD9E6DB31264945FE3AC305D5F" direction="south" orientation="latitude" precision="15" value="-33.27972">
33°16
<normalizedToken id="728FB3D1383E49C9497C504D05798291" originalValue="47”">'47&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="81C53260DB53F8F3C4BBB3B32251A168" direction="east" orientation="longitude" precision="15" value="151.54445">
151°32
<normalizedToken id="015F41ECC0A613DB8A5BAF488D5E1B23" originalValue="40”">'40&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS61866, KS61870, KS61880, KS61905, KS61913, KS61929); 3 males, 2 females, Wyrrabalong National Park,
<geoCoordinate id="24E413332B4739B3145F1B7765863774" direction="south" orientation="latitude" precision="15" value="-33.28">
33°16
<normalizedToken id="0491C2B32C26ECFBF9F344B4BF227BDE" originalValue="48”">'48&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="8ACF1305749248EBFF1B5F52A94F4B45" direction="east" orientation="longitude" precision="15" value="151.54584">
151°32
<normalizedToken id="28C708E8BA9C524989FFB866131502D6" originalValue="45”">'45&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS61867, KS61877, KS64057, KS64059-60); 1 male, Wyrrabalong National Park,
<geoCoordinate id="DD50C17C5D20443AA53CD5415A4AB73F" direction="south" orientation="latitude" precision="15" value="-33.280834">
33°16
<normalizedToken id="47E2FAF0A2BC5610400A80CEDB9F52E9" originalValue="51”">'51&quot;</normalizedToken>
S
</geoCoordinate>
,
<geoCoordinate id="25863425F7EABBBE364F3FCA3330AF10" direction="east" orientation="longitude" precision="15" value="151.54362">
151°32
<normalizedToken id="EF09CA080614EF2F28816F91E919A38D" originalValue="37”">'37&quot;</normalizedToken>
E
</geoCoordinate>
(AM KS64058).
</paragraph>
</subSubSection>
<subSubSection id="4306D29E3FC319BAEC9724BAA106D932" pageId="0" pageNumber="169" type="etymology">
<paragraph id="581D13EF2E9B011C556D297EFC95C536" pageId="0" pageNumber="169">Etymology.</paragraph>
<paragraph id="58C7F168E88294EA132150ED6935E3A5" pageId="0" pageNumber="169">The specific epithet is a matronym honouring Dr Helen Smith of the Australian Museum, Sydney, for her ongoing support of our spider research.</paragraph>
</subSubSection>
<subSubSection id="67BC0CDB13AFCC9EEF8ECC3F4458CE51" pageId="0" pageNumber="169" type="diagnosis">
<paragraph id="02511D6356FD671CAC2C2423FB2C51C9" pageId="0" pageNumber="169">Diagnosis.</paragraph>
<paragraph id="0E63A1FA7DE37DE7686EF96454B8A542" pageId="0" pageNumber="169">
<taxonomicName id="6E6CA764D1E48CEAA0D56F59ABDF9581" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria helensmithae" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="helensmithae">Artoria helensmithae</taxonomicName>
sp. n. males and females are very similar to those of
<taxonomicName id="B44DB86B41269E11585240004F73863A" lsidName="A. beaury" pageId="0" pageNumber="169" rank="species" species="beaury">A. beaury</taxonomicName>
sp. n. (see diagnosis above).
</paragraph>
</subSubSection>
<subSubSection id="26EC19A4547CD190919A1ECE91F168CA" pageId="0" pageNumber="169" type="description">
<paragraph id="77D7F8712B1832ADF60D2A75809070A7" pageId="0" pageNumber="169">Description.</paragraph>
<paragraph id="1A5216D6804766513284D79E0C373539" pageId="0" pageNumber="169">Male (based on holotype, AM KS128076).</paragraph>
<paragraph id="15967016A34EC28A66A91FD1698840D4" pageId="0" pageNumber="169">Total length 3.9.</paragraph>
<paragraph id="57AE9B7467C0456BA0A8D03DBED0061B" pageId="0" pageNumber="169">Prosoma. Length 2.2, width 1.5; carapace light yellow-brown with dark radial pattern and black V-shaped pattern between cephalic and thoracic region; indistinct and irregular lighter central and marginal band (Fig. 24A); sternum light, margin dusted dark grey (Fig. 24B).</paragraph>
<paragraph id="37CBA0BE94F07928D609BA4EB78896D1" pageId="0" pageNumber="169">Eyes. Diameter of AME: 0.10; ALE: 0.07; PME: 0.28; PLE: 0.20.</paragraph>
<paragraph id="7AD63C2246C9E8D7F12D6A1043D0E9BA" pageId="0" pageNumber="169">Anterior eye row. Slightly procurved, evenly spaced.</paragraph>
<paragraph id="CF240E64852772B2888A7F6668DE9C8F" pageId="0" pageNumber="169">Chelicerae. Dark brown darker apically.</paragraph>
<paragraph id="F4717659969C05001497071C475A5F08" pageId="0" pageNumber="169">Labium. Dark brown, with lighter anterior rim (Fig. 24B)</paragraph>
<paragraph id="050310C3D3AB9281B96970C1BCBB73FE" pageId="0" pageNumber="169">Pedipalp coxae. Dark brown, with lighter anterior rim (Fig. 24B).</paragraph>
<paragraph id="95031B7CFF784B7BF90CFE644488F598" pageId="0" pageNumber="169">Legs. Femora, tibiae and metatarsi of leg I very dark to almost black; other legs brown, with darker annulations; tarsi and metatarsi lighter brown (Fig. 24A).</paragraph>
<paragraph id="9D0B38F06AED6854E389CB12FE9CC991" pageId="0" pageNumber="169">Opisthosoma. Length 1.7, width 1.3; dark grey with light yellow-brown anterior cardiac mark and light irregular pattern (Fig. 24A). Venter pale with darker pattern anteriorly (Fig. 24B); spinnerets dark grey.</paragraph>
<paragraph id="64EE50F7674E12C299A3C53FFF547296" pageId="0" pageNumber="169">Pedipalps. Tibia as long as broad; cymbium tip with 8-10 macrosetae (Fig. 24E, F); dorsal scopula patch absent; tegular apophysis distally widely scooped, basally narrowed to 1/2, retrolateral tip strongly hooked and reaching beyond margin of cymbium (Fig. 24E, F); palea about 1 1/2 times as long as wide, basoembolic apophysis large, triangular; embolus broad, widely semicircular; terminal apophysis triangular, tip scooped (Fig. 47K).</paragraph>
<paragraph id="E3A78BF5CD453A8D374439ED4EB62BB0" pageId="0" pageNumber="169">Female (based on AM KS61928).</paragraph>
<paragraph id="4988A5589D6562EB7AA9CDB53B87FAC6" pageId="0" pageNumber="169">Total length 3.8.</paragraph>
<paragraph id="A9951268FF18B568BD3100A1E75EA735" pageId="0" pageNumber="169">Prosoma. Length 2.2, width 1.6; carapace and sternum colouration as male (Fig. 24C, D).</paragraph>
<paragraph id="F71BBA66230AE76812F099D6309EC178" pageId="0" pageNumber="169">Eyes. Diameter of AME 0.08, ALE 0.08, PME 0.26, PLE 0.19.</paragraph>
<paragraph id="CC703D08CE02BF38A5ED98175F7328E6" pageId="0" pageNumber="169">Anterior eye row. Slightly procurved, evenly spaced.</paragraph>
<paragraph id="AFF3043E913445B9A21584173AB5BC86" pageId="0" pageNumber="169">Opisthosoma. Length 1.6, width 1.5; otherwise as male, but legs I not darker and opisthosoma pattern more obscure (Fig. 24C, D).</paragraph>
<paragraph id="CC399E57F502F19C46E741A669C923F0" pageId="0" pageNumber="169">Epigyne about slightly longer than wide, strongly sclerotized at posterior tips (Fig. 24G); spermathecal heads globular around 1 diameter apart, spermathecal stalks attached laterally slightly s-shaped (Fig. 24H).</paragraph>
<caption id="506B1D6D1B35592EF3CCBCAA4872DD4D" pageId="0" pageNumber="169">
<paragraph id="EDE379329DA537BF442A0EF19194FD5E" pageId="0" pageNumber="169">
Figure 24. A
<taxonomicName id="27F23007AE5DD39B9028FBD885F04EA6" lsidName="helensmithae" pageId="0" pageNumber="169" rank="species" species="helensmithae">helensmithae</taxonomicName>
sp. n., male holotype (AM KS128076), female paratype (AM KS61928): A, male habitus, dorsal view; B, male habitus, ventral view; C, female habitus, dorsal view; D, female habitus, ventral view; E, male pedipalp, ventral view; F, male pedipalp, retrolateral view; G, epigyne, ventral view; H, epigyne, dorsal view. Scale bars: habitus 1.0 mm; pedipalp, epigyne 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="1F15F3C52EA9C315D8E8695CC4A637E1" pageId="0" pageNumber="169" type="life history and habitat preferences">
<paragraph id="2D99436BD4B6FBFAA27ACED6F5A980D2" pageId="0" pageNumber="169">Life history and habitat preferences.</paragraph>
<paragraph id="AA8FC11C506A7CAD93BF38BD205484C1" pageId="0" pageNumber="169">
Habitat descriptions found on the collection labels of
<taxonomicName id="9B18D3BA7F737E1EC2BFA2D3F7EC3B98" lsidName="A. helensmithae" pageId="0" pageNumber="169" rank="species" species="helensmithae">A. helensmithae</taxonomicName>
sp. n. include 'under tall
<normalizedToken id="2B7043A16824211A1A17846CE36F3219" originalValue="shrubs">shrubs'</normalizedToken>
, 'sand
<normalizedToken id="0FCAB7440BF7A2E62BBC5CFB07ED8AA8" originalValue="dunes">dunes'</normalizedToken>
, open forest,
<normalizedToken id="757A099B2E881FDCA03680EA354C7041" originalValue="litter">litter'</normalizedToken>
,
<normalizedToken id="94BA7117915E9F92C6A383379680396D" originalValue="rainforest">'rainforest'</normalizedToken>
and 'dry
<normalizedToken id="DE2A0A46C5D0B6FBD5B0C1BE0B72BEFE" originalValue="forest">forest'</normalizedToken>
. A single record is from the dark zone of a cave. Therefore,
<taxonomicName id="5B7803B8C03A48B2A302A3469979A180" lsidName="A. helensmithae" pageId="0" pageNumber="169" rank="species" species="helensmithae">A. helensmithae</taxonomicName>
sp. n. seems to show some variation in its habitat preferences. Most records also state
<normalizedToken id="7B8FE5EAF9457BCC4D78DC8420AF4A3A" originalValue="litter">'litter'</normalizedToken>
.
</paragraph>
<paragraph id="F8EFC62CAE959545B1630DF3EB01CD07" pageId="0" pageNumber="169">
<taxonomicName id="5A364B27D93D1E50149C83EC1AB245D9" class="Arachnida" family="Lycosidae" genus="Artoria" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Artoria helensmithae" order="Araneae" pageId="0" pageNumber="169" phylum="Arthropoda" rank="species" species="helensmithae">Artoria helensmithae</taxonomicName>
sp. n. appears to have three peaks of reproductive activity, the first around April/May, then in July and again from October to December. Females with spiderlings were found in December. This is an unusual phenology for wolf spiders, i.e. this species appears to be reproductively active in autumn, winter and summer.
</paragraph>
</subSubSection>
<subSubSection id="EE4906A815820E294ECDD26DEA3B3C70" pageId="0" pageNumber="169" type="distribution">
<paragraph id="D3738506EEB94A539903BE60B44AA04F" pageId="0" pageNumber="169">Distribution.</paragraph>
<paragraph id="72B3A27AA5EB5F2993461C79C41643A3" pageId="0" pageNumber="169">
In NSW,
<taxonomicName id="837C91C07308F52104FAA426A12EF5E4" lsidName="A. helensmithae" pageId="0" pageNumber="169" rank="species" species="helensmithae">A. helensmithae</taxonomicName>
sp. n. has generally been found east of the Great Dividing Range, with records from the NSW North Coast (NNC), Sydney Basin (SYB), South East Corner (SEC) and South East Highlands (SHE) IBRA regions (Fig. 23). The species has also been found in eastern and central Victoria (V.W. Framenau, unpublished data).
</paragraph>
</subSubSection>
</treatment>
</document>