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<document id="AE333410F0AC246E874570A2421EA7DD" ID-DOI="10.1111/j.1096-3642.2007.00281.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5429392" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630420803834" checkinUser="felipe" docAuthor="Stiassny, Melanie L. J. &amp; Getahun, Abebe" docDate="2007" docId="065F9C4AFFB5FFD8DEE3FD88FC08F908" docLanguage="en" docName="j.1096-3642.2007.00281.x.pdf" docOrigin="Zoological Journal of the Linnean Society 150 (1)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2007.00281.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Garra regressus Stiassny &amp; Getahun 2007, SP. NOV." docType="treatment" docVersion="3" lastPageNumber="55" masterDocId="FA66E432FFB9FFD6DE4FFFFEFF86FFDE" masterDocTitle="An overview of labeonin relationships and the phylogenetic placement of the Afro-Asian genus Garra Hamilton, 1922 (Teleostei: Cyprinidae), with the description of five new species of Garra from Ethiopia, and a key to all African species" masterLastPageNumber="83" masterPageNumber="41" pageNumber="53" updateTime="1699264050007" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:titleInfo id="AE906633ABD02F3FC21BF1A31CD44DA2">
<mods:title id="B26303005BCA85686B25EA1D1C55BEB7">An overview of labeonin relationships and the phylogenetic placement of the Afro-Asian genus Garra Hamilton, 1922 (Teleostei: Cyprinidae), with the description of five new species of Garra from Ethiopia, and a key to all African species</mods:title>
</mods:titleInfo>
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<mods:namePart id="7C0A6CA213B229D83F88237C55D49B1C">Stiassny, Melanie L. J.</mods:namePart>
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<mods:namePart id="115716D1CC1919C93C8FDE68A8F19B8A">Getahun, Abebe</mods:namePart>
</mods:name>
<mods:typeOfResource id="65ED4EB1DAACF4A72B09038AB3CC736C">text</mods:typeOfResource>
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<mods:title id="7F5849AA9DD4BE2EC530AB9EAB54B7AA">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="16115FB0FFA88E5A5AC4AD9D03490895">
<mods:date id="EF598E5BBFFD22CACF2D059CFE911665">2007</mods:date>
<mods:detail id="9085224703DAB12C4B075E6F9D00333E" type="pubDate">
<mods:number id="F336C4CC5EDC13E6B84F91648F84DFB7">2007-05-31</mods:number>
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<mods:detail id="B023BD9153E7E7BEDFE42A8D61C63153" type="volume">
<mods:number id="27A09A32990733B7127AE0DAC1EF9B00">150</mods:number>
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<mods:number id="7FB8A3AFB89ED8A9CC5F9B800E5DCE0E">1</mods:number>
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<mods:url id="7F5C39D9F6471B7D1DABCF3781274B41">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2007.00281.x</mods:url>
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<mods:classification id="6899E11E155BEC80F008A5B4FEB2C577">journal article</mods:classification>
<mods:identifier id="988262C870E3506CAA2C764C23C1120C" type="DOI">10.1111/j.1096-3642.2007.00281.x</mods:identifier>
<mods:identifier id="29E31309FA4422E25BB9EA36BEB81DE1" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="84CD35E140B5C75F64088A067D45F162" type="Zenodo-Dep">5429392</mods:identifier>
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<treatment id="065F9C4AFFB5FFD8DEE3FD88FC08F908" LSID="urn:lsid:plazi:treatment:065F9C4AFFB5FFD8DEE3FD88FC08F908" httpUri="http://treatment.plazi.org/id/065F9C4AFFB5FFD8DEE3FD88FC08F908" lastPageId="14" lastPageNumber="55" pageId="12" pageNumber="53">
<subSubSection id="C6EC7ED7FFB5FFDADEE3FD88FD5EFD50" box="[172,728,630,654]" pageId="12" pageNumber="53" type="nomenclature">
<paragraph id="8E492D5CFFB5FFDADEE3FD88FD5EFD50" blockId="12.[172,728,630,654]" box="[172,728,630,654]" pageId="12" pageNumber="53">
<heading id="D5019A30FFB5FFDADEE3FD88FD5EFD50" box="[172,728,630,654]" centered="true" fontSize="9" level="2" pageId="12" pageNumber="53" reason="2">
<taxonomicName id="49F656DFFFB5FFDADEE3FD88FE16FD53" ID-CoL="3FBM7" authority="Stiassny &amp; Getahun, 2007" authorityName="Stiassny &amp; Getahun" authorityYear="2007" box="[172,400,630,654]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="53" phylum="Chordata" rank="species" species="regressus" status="sp. nov.">GARRA REGRESSUS</taxonomicName>
<emphasis id="BC82F14EFFB5FFDADFD6FD84FE7BFD50" bold="true" box="[409,509,630,654]" pageId="12" pageNumber="53">
<taxonomicNameLabel id="A7B14C35FFB5FFDADFD6FD84FE7BFD50" box="[409,509,630,654]" pageId="12" pageNumber="53" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="16CD31D9FFB5FFDADC41FD88FDE5FD50" box="[526,611,630,654]" captionStart="Figure 11" captionStartId="14.[142,220,1015,1034]" captionTargetBox="[302,1251,202,982]" captionTargetId="figure-437@14.[302,1262,194,982]" captionTargetPageId="14" captionText="Figure 11. Garra regressus, holotype, AMNH 228487, 121.8 mm standard length." figureDoi="http://doi.org/10.5281/zenodo.5429414" httpUri="https://zenodo.org/record/5429414/files/figure.png" pageId="12" pageNumber="53">FIG. 11</figureCitation>
;
<tableCitation id="C37418E7FFB5FFDADC3FFD88FD56FD50" box="[624,720,630,654]" captionStart="Table 2" captionStartId="12.[142,206,863,882]" captionText="Table 2. Morphometric and meristic data for Garra regressus sp. nov. Proportional measurements (mm) in either the percentage of standard length (SL) or the percentage of head length (HL), unless noted otherwise. When range of values, meristics of holotype are given in parenthesis" pageId="12" pageNumber="53">TABLE 2</tableCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C6EC7ED7FFB5FFDADEC1FD61FDE4FCCF" pageId="12" pageNumber="53" type="reference_group">
<paragraph id="8E492D5CFFB5FFDADEC1FD61FDE4FCCF" blockId="12.[142,758,671,785]" pageId="12" pageNumber="53">
<taxonomicName id="49F656DFFFB5FFDADEC1FD61FDB7FD6A" ID-CoL="3FBKX" authority="Getahun, 2000" authorityName="Getahun" authorityYear="2000" box="[142,561,671,693]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="53" phylum="Chordata" rank="species" species="microstoma">
<emphasis id="BC82F14EFFB5FFDADEC1FD61FEEFFD6A" box="[142,361,671,692]" italics="true" pageId="12" pageNumber="53">Garra microstoma</emphasis>
<bibRefCitation id="EA6750ADFFB5FFDADF33FD61FDB7FD6A" author="Getahun A" box="[380,561,671,693]" pageId="12" pageNumber="53" refId="ref28792" refString="Getahun A. 2000. Systematic studies of the African species of the genus Garra (Pisces: Cyprinidae) PhD Thesis, City University of New York." type="book" year="2000">Getahun, 2000</bibRefCitation>
</taxonomicName>
,
<emphasis id="BC82F14EFFB5FFDADC09FD61FD73FD6A" box="[582,757,671,692]" italics="true" pageId="12" pageNumber="53">nomen nudum</emphasis>
[p. 92, unpubl. PhD thesis, preoccupied by
<taxonomicName id="49F656DFFFB5FFDADCFFFD40FE29FD2C" ID-CoL="3FBKX" authority="Mai (1978)" authorityName="Mai" authorityYear="1978" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="53" phylum="Chordata" rank="species" species="microstoma">
<emphasis id="BC82F14EFFB5FFDADCFFFD40FE94FD2C" italics="true" pageId="12" pageNumber="53">Garra microstoma</emphasis>
<bibRefCitation id="EA6750ADFFB5FFDADF68FD22FE29FD2C" author="Mai DY" box="[295,431,732,754]" pageId="12" pageNumber="53" refId="ref29610" refString="Mai DY. 1978. Identification of the Fresh-Water Fishes of North Viet Nam. Ha Noi: Scientific &amp; Technology Publisher." type="book" year="1978">Mai (1978)</bibRefCitation>
</taxonomicName>
and valid as
<taxonomicName id="49F656DFFFB5FFDADC33FD22FDDAFCCF" ID-CoL="36TQB" authority="(Mai, 1978) (Kottelat, 2001)" authorityName="Kottelat" authorityYear="2001" baseAuthorityName="Mai" baseAuthorityYear="1978" class="Actinopterygii" family="Cyprinidae" genus="Discogobio" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="53" phylum="Chordata" rank="species" species="microstoma">
<emphasis id="BC82F14EFFB5FFDADC33FD22FE94FCCE" italics="true" pageId="12" pageNumber="53">Discogobio microstoma</emphasis>
(
<bibRefCitation id="EA6750ADFFB5FFDADF6CFD05FE10FCCF" author="Mai DY" box="[291,406,763,785]" pageId="12" pageNumber="53" refId="ref29610" refString="Mai DY. 1978. Identification of the Fresh-Water Fishes of North Viet Nam. Ha Noi: Scientific &amp; Technology Publisher." type="book" year="1978">Mai, 1978</bibRefCitation>
) (
<bibRefCitation id="EA6750ADFFB5FFDADFE1FD05FDD2FCCF" author="Kottelat M" box="[430,596,762,785]" pageId="12" pageNumber="53" pagination="1 - 18" refId="ref29308" refString="Kottelat M. 2001. Freshwater fishes of northern Vietnam. A preliminary check-list of the fishes known or expected to occur in northern Vietnam with comments on systematics and nomenclature. Environment Social Development Unit, East Asia and Pacific Region. The World Bank, i - iii, 1 - 123, 1 - 18." type="journal article" year="2001">Kottelat, 2001</bibRefCitation>
)
</taxonomicName>
]
</paragraph>
</subSubSection>
<subSubSection id="C6EC7ED7FFB5FFD8DD69FF3AFC08F908" lastPageId="14" lastPageNumber="55" pageId="12" pageNumber="53" type="diagnosis">
<paragraph id="8E492D5CFFB5FFDADD69FF3AFA00FEC9" blockId="12.[806,1421,195,279]" pageId="12" pageNumber="53">
<emphasis id="BC82F14EFFB5FFDADD69FF3AFC1BFF07" box="[806,925,196,217]" italics="true" pageId="12" pageNumber="53">Diagnosis:</emphasis>
Unique among African
<taxonomicName id="49F656DFFFB5FFDADA89FF3AFA8DFF07" authorityName="Hamilton" authorityYear="1822" box="[1222,1291,196,217]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="BC82F14EFFB5FFDADA89FF3AFA8DFF07" box="[1222,1291,196,217]" italics="true" pageId="12" pageNumber="53">Garra</emphasis>
</taxonomicName>
in possession of a small, acutely pointed head, narrow mouth, regressed rostral cap, and a fully exposed upper jaw.
</paragraph>
<paragraph id="8E492D5CFFB5FFD8DD69FED1FE35F976" blockId="12.[806,1422,303,785]" lastBlockId="14.[142,758,1100,1704]" lastPageId="14" lastPageNumber="55" pageId="12" pageNumber="53">
<emphasis id="BC82F14EFFB5FFDADD69FED1FC28FE9A" box="[806,942,303,324]" italics="true" pageId="12" pageNumber="53">Description:</emphasis>
Morphometric and meristic data for
<typeStatus id="514D93FEFFB5FFDADD69FEB3FC01FEBD" box="[806,903,333,355]" pageId="12" pageNumber="53" type="holotype">holotype</typeStatus>
and
<specimenCount id="98F0E6D5FFB5FFDADD86FEB0FBE8FEBD" box="[969,1134,334,355]" pageId="12" pageNumber="53" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
given in
<tableCitation id="C37418E7FFB5FFDADAA8FEB3FABAFEBC" box="[1255,1340,333,355]" captionStart="Table 2" captionStartId="12.[142,206,863,882]" captionText="Table 2. Morphometric and meristic data for Garra regressus sp. nov. Proportional measurements (mm) in either the percentage of standard length (SL) or the percentage of head length (HL), unless noted otherwise. When range of values, meristics of holotype are given in parenthesis" pageId="12" pageNumber="53">Table 2</tableCitation>
. Maximum size:
<quantity id="490E80B9FFB5FFDADDE2FE92FB98FE5F" box="[941,1054,364,385]" metricMagnitude="-1" metricUnit="m" metricValue="1.218" pageId="12" pageNumber="53" unit="mm" value="121.8">121.8 mm</quantity>
SL (
<typeStatus id="514D93FEFFB5FFDADA10FE92FB45FE5C" box="[1119,1219,364,386]" pageId="12" pageNumber="53" type="holotype">holotype</typeStatus>
, AMNH 228487). Currently known only from type series. All specimens relatively slender (body depth 19.8–20.6, m. 20.1% SL) and elongate, greatest body depth located at level of dorsal-fin origin. Head relatively short (19.7–21.0, m. 20.5% SL), snout acutely pointed and often studded with small tubercles, mouth narrow. Eye positioned a little posterior on head. Dorsal head profile rising steeply over snout, dorsum of head more-orless flattened, dorsal body profile smoothly convex to dorsal-fin origin. Disc type A, weakly developed and inconspicuous, without free posterior margin. Rostral cap thin and regressed leaving upper jaw exposed. Two pairs of small barbels, maxillary pair usually longest. Thirty-nine pored scales along lateral line; either eight or nine predorsal scales anterior to dorsal fin; either three or four scales from lateral line to origin of pelvic fin; either four or five scales from lateral line to origin of dorsal fin. Chest asquamate, belly with a few deeply embedded scales, and postpelvic region fully scaled. Dorsal-fin rays iii, seven, inserted well anterior to pelvic fin, with short subacuminate tip, first and second branched rays longest, extending slightly beyond last ray; posterior margin straight, posteriormost ray not extending to tip of pelvic fin. Predorsal length (distance between dorsal-fin origin and tip of snout) 44.0–46.3% SL. Anal-fin rays iii, five, with short subacuminate tip; posterior margin straight. Pectoral-fin rays iv, 11; pectoral length 17.5–21.0% SL. Vent located relatively close to anal fin (vent distance 19.3–23.8, mean 21.9%). Intestine very short (SL 108–116% Int.L). Gas bladder well developed and large (posterior chamber 21.7–24.9% SL).
</paragraph>
<caption id="DA897DD4FFB5FFDADEC1FCA1FDF7FC72" pageId="12" pageNumber="53" startId="12.[142,206,863,882]">
<paragraph id="8E492D5CFFB5FFDADEC1FCA1FDF7FC72" blockId="12.[142,1422,862,941]" pageId="12" pageNumber="53">
<emphasis id="BC82F14EFFB5FFDADEC1FCA1FF6EFCAF" bold="true" box="[142,232,862,882]" pageId="12" pageNumber="53">Table 2.</emphasis>
Morphometric and meristic data for
<emphasis id="BC82F14EFFB5FFDADC38FCA0FC15FCAC" bold="true" box="[631,915,862,882]" italics="true" pageId="12" pageNumber="53">
<taxonomicName id="49F656DFFFB5FFDADC38FCA0FCB3FCAC" authority="Stiassny &amp; Getahun, 2007" authorityName="Stiassny &amp; Getahun" authorityYear="2007" box="[631,821,862,882]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="12" pageNumber="53" phylum="Chordata" rank="species" species="regressus" status="sp. nov.">Garra regressus</taxonomicName>
<taxonomicNameLabel id="A7B14C35FFB5FFDADD72FCA1FC15FCAC" box="[829,915,863,882]" pageId="12" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Proportional measurements (mm) in either the percentage of standard length (SL) or the percentage of head length (HL), unless noted otherwise. When range of values, meristics of holotype are given in parenthesis
</paragraph>
</caption>
<paragraph id="8E492D5CFFB5FFDADEC1FC2AFCE2F889" pageId="12" pageNumber="53">
<table id="FCF6DFFCFFB50029DEC1FC2AFA0AF889" box="[142,1420,980,1879]" gridcols="6" gridrows="30" pageId="12" pageNumber="53">
<tr id="30C62F1EFFB50029DEC1FC2AFA0AFC39" box="[142,1420,980,999]" gridrow="0" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FC2AFE6DFC39" box="[142,491,980,999]" gridcol="0" gridrow="0" pageId="12" pageNumber="53">Character</th>
<th id="73174662FFB50029DC2EFC2AFDE9FC39" box="[609,623,980,999]" gridcol="1" gridrow="0" pageId="12" pageNumber="53">
<emphasis id="BC82F14EFFB5FFDADC2EFC2AFDE9FC38" box="[609,623,980,998]" italics="true" pageId="12" pageNumber="53">n</emphasis>
</th>
<th id="73174662FFB50029DCACFC2AFCE3FC39" box="[739,869,980,999]" gridcol="2" gridrow="0" pageId="12" pageNumber="53">holotype</th>
<th id="73174662FFB50029DD8EFC2AFBB4FC39" box="[961,1074,980,999]" gridcol="3" gridrow="0" pageId="12" pageNumber="53">range</th>
<th id="73174662FFB50029DAE9FC2AFB66FC39" box="[1190,1248,980,999]" gridcol="4" gridrow="0" pageId="12" pageNumber="53">mean</th>
<th id="73174662FFB50029DB1CFC2AFA0AFC39" box="[1363,1420,980,999]" gridcol="5" gridrow="0" pageId="12" pageNumber="53">SD</th>
</tr>
<tr id="30C62F1EFFB50029DEC1FBF0FA0AFBFC" box="[142,1420,1038,1058]" gridrow="1" pageId="12" pageNumber="53" rowspan-4="1" rowspan-5="1">
<th id="73174662FFB50029DEC1FBF0FE6DFBFC" box="[142,491,1038,1058]" gridcol="0" gridrow="1" pageId="12" pageNumber="53">Standard length (SL)</th>
<td id="73174662FFB50029DC2EFBF0FDE9FBFC" box="[609,623,1038,1058]" gridcol="1" gridrow="1" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACFBF0FCE3FBFC" box="[739,869,1038,1058]" gridcol="2" gridrow="1" pageId="12" pageNumber="53">121.8</td>
<td id="73174662FFB50029DD8EFBF0FBB4FBFC" box="[961,1074,1038,1058]" gridcol="3" gridrow="1" pageId="12" pageNumber="53">94.7–121.8</td>
</tr>
<tr id="30C62F1EFFB50029DEC1FBD2FA0AFBE1" box="[142,1420,1068,1087]" gridrow="2" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FBD2FA0AFBE1" box="[142,1420,1068,1087]" colspan="6" colspanRight="5" gridcol="0" gridrow="2" pageId="12" pageNumber="53">% SL</th>
</tr>
<tr id="30C62F1EFFB50029DEC1FBB7FA0AFB82" box="[142,1420,1097,1116]" gridrow="3" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FBB7FE6DFB82" box="[142,491,1097,1116]" gridcol="0" gridrow="3" pageId="12" pageNumber="53">Body depth</th>
<td id="73174662FFB50029DC2EFBB7FDE9FB82" box="[609,623,1097,1116]" gridcol="1" gridrow="3" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACFBB7FCE3FB82" box="[739,869,1097,1116]" gridcol="2" gridrow="3" pageId="12" pageNumber="53">20.1</td>
<td id="73174662FFB50029DD8EFBB7FBB4FB82" box="[961,1074,1097,1116]" gridcol="3" gridrow="3" pageId="12" pageNumber="53">19.8–20.6</td>
<td id="73174662FFB50029DAE9FBB7FB66FB82" box="[1190,1248,1097,1116]" gridcol="4" gridrow="3" pageId="12" pageNumber="53">20.1</td>
<td id="73174662FFB50029DB1CFBB7FA0AFB82" box="[1363,1420,1097,1116]" gridcol="5" gridrow="3" pageId="12" pageNumber="53">0.44</td>
</tr>
<tr id="30C62F1EFFB50029DEC1FB98FA0AFBA4" box="[142,1420,1126,1146]" gridrow="4" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FB98FE6DFBA4" box="[142,491,1126,1146]" gridcol="0" gridrow="4" pageId="12" pageNumber="53">Head length</th>
<td id="73174662FFB50029DC2EFB98FDE9FBA4" box="[609,623,1126,1146]" gridcol="1" gridrow="4" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACFB98FCE3FBA4" box="[739,869,1126,1146]" gridcol="2" gridrow="4" pageId="12" pageNumber="53">19.6</td>
<td id="73174662FFB50029DD8EFB98FBB4FBA4" box="[961,1074,1126,1146]" gridcol="3" gridrow="4" pageId="12" pageNumber="53">19.7–21.0</td>
<td id="73174662FFB50029DAE9FB98FB66FBA4" box="[1190,1248,1126,1146]" gridcol="4" gridrow="4" pageId="12" pageNumber="53">20.5</td>
<td id="73174662FFB50029DB1CFB98FA0AFBA4" box="[1363,1420,1126,1146]" gridcol="5" gridrow="4" pageId="12" pageNumber="53">0.72</td>
</tr>
<tr id="30C62F1EFFB50029DEC1FB7AFA0AFB49" box="[142,1420,1156,1175]" gridrow="5" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FB7AFE6DFB49" box="[142,491,1156,1175]" gridcol="0" gridrow="5" pageId="12" pageNumber="53">Predorsal length</th>
<td id="73174662FFB50029DC2EFB7AFDE9FB49" box="[609,623,1156,1175]" gridcol="1" gridrow="5" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACFB7AFCE3FB49" box="[739,869,1156,1175]" gridcol="2" gridrow="5" pageId="12" pageNumber="53">46.2</td>
<td id="73174662FFB50029DD8EFB7AFBB4FB49" box="[961,1074,1156,1175]" gridcol="3" gridrow="5" pageId="12" pageNumber="53">44.0–46.3</td>
<td id="73174662FFB50029DAE9FB7AFB66FB49" box="[1190,1248,1156,1175]" gridcol="4" gridrow="5" pageId="12" pageNumber="53">45.0</td>
<td id="73174662FFB50029DB1CFB7AFA0AFB49" box="[1363,1420,1156,1175]" gridcol="5" gridrow="5" pageId="12" pageNumber="53">1.18</td>
</tr>
<tr id="30C62F1EFFB50029DEC1FB5FFA0AFB6A" box="[142,1420,1185,1204]" gridrow="6" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FB5FFE6DFB6A" box="[142,491,1185,1204]" gridcol="0" gridrow="6" pageId="12" pageNumber="53">Pectoral fin length</th>
<td id="73174662FFB50029DC2EFB5FFDE9FB6A" box="[609,623,1185,1204]" gridcol="1" gridrow="6" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACFB5FFCE3FB6A" box="[739,869,1185,1204]" gridcol="2" gridrow="6" pageId="12" pageNumber="53">19.6</td>
<td id="73174662FFB50029DD8EFB5FFBB4FB6A" box="[961,1074,1185,1204]" gridcol="3" gridrow="6" pageId="12" pageNumber="53">17.5–21.0</td>
<td id="73174662FFB50029DAE9FB5FFB66FB6A" box="[1190,1248,1185,1204]" gridcol="4" gridrow="6" pageId="12" pageNumber="53">19.6</td>
<td id="73174662FFB50029DB1CFB5FFA0AFB6A" box="[1363,1420,1185,1204]" gridcol="5" gridrow="6" pageId="12" pageNumber="53">1.83</td>
</tr>
<tr id="30C62F1EFFB50029DEC1FB40FA0AFB0C" box="[142,1420,1214,1234]" gridrow="7" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FB40FE6DFB0C" box="[142,491,1214,1234]" gridcol="0" gridrow="7" pageId="12" pageNumber="53">Caudal peduncle length</th>
<td id="73174662FFB50029DC2EFB40FDE9FB0C" box="[609,623,1214,1234]" gridcol="1" gridrow="7" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACFB40FCE3FB0C" box="[739,869,1214,1234]" gridcol="2" gridrow="7" pageId="12" pageNumber="53">18.1</td>
<td id="73174662FFB50029DD8EFB40FBB4FB0C" box="[961,1074,1214,1234]" gridcol="3" gridrow="7" pageId="12" pageNumber="53">18.2–18.9</td>
<td id="73174662FFB50029DAE9FB40FB66FB0C" box="[1190,1248,1214,1234]" gridcol="4" gridrow="7" pageId="12" pageNumber="53">18.6</td>
<td id="73174662FFB50029DB1CFB40FA0AFB0C" box="[1363,1420,1214,1234]" gridcol="5" gridrow="7" pageId="12" pageNumber="53">0.35</td>
</tr>
<tr id="30C62F1EFFB50029DEC1FB22FA0AFB31" box="[142,1420,1244,1263]" gridrow="8" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FB22FE6DFB31" box="[142,491,1244,1263]" gridcol="0" gridrow="8" pageId="12" pageNumber="53">Caudal peduncle depth</th>
<td id="73174662FFB50029DC2EFB22FDE9FB31" box="[609,623,1244,1263]" gridcol="1" gridrow="8" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACFB22FCE3FB31" box="[739,869,1244,1263]" gridcol="2" gridrow="8" pageId="12" pageNumber="53">10.2</td>
<td id="73174662FFB50029DD8EFB22FBB4FB31" box="[961,1074,1244,1263]" gridcol="3" gridrow="8" pageId="12" pageNumber="53">9.9–11.1</td>
<td id="73174662FFB50029DAE9FB22FB66FB31" box="[1190,1248,1244,1263]" gridcol="4" gridrow="8" pageId="12" pageNumber="53">10.7</td>
<td id="73174662FFB50029DB1CFB22FA0AFB31" box="[1363,1420,1244,1263]" gridcol="5" gridrow="8" pageId="12" pageNumber="53">0.67</td>
</tr>
<tr id="30C62F1EFFB50029DEC1FB07FA0AFAD2" box="[142,1420,1273,1292]" gridrow="9" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FB07FE6DFAD2" box="[142,491,1273,1292]" gridcol="0" gridrow="9" pageId="12" pageNumber="53">Dorsal fin length</th>
<td id="73174662FFB50029DC2EFB07FDE9FAD2" box="[609,623,1273,1292]" gridcol="1" gridrow="9" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACFB07FCE3FAD2" box="[739,869,1273,1292]" gridcol="2" gridrow="9" pageId="12" pageNumber="53">18.7</td>
<td id="73174662FFB50029DD8EFB07FBB4FAD2" box="[961,1074,1273,1292]" gridcol="3" gridrow="9" pageId="12" pageNumber="53">18.7–24.1</td>
<td id="73174662FFB50029DAE9FB07FB66FAD2" box="[1190,1248,1273,1292]" gridcol="4" gridrow="9" pageId="12" pageNumber="53">20.7</td>
<td id="73174662FFB50029DB1CFB07FA0AFAD2" box="[1363,1420,1273,1292]" gridcol="5" gridrow="9" pageId="12" pageNumber="53">2.90</td>
</tr>
<tr id="30C62F1EFFB50029DEC1FAE8FA0AFAF4" box="[142,1420,1302,1322]" gridrow="10" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FAE8FE6DFAF4" box="[142,491,1302,1322]" gridcol="0" gridrow="10" pageId="12" pageNumber="53">Anal fin length</th>
<td id="73174662FFB50029DC2EFAE8FDE9FAF4" box="[609,623,1302,1322]" gridcol="1" gridrow="10" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACFAE8FCE3FAF4" box="[739,869,1302,1322]" gridcol="2" gridrow="10" pageId="12" pageNumber="53">16.9</td>
<td id="73174662FFB50029DD8EFAE8FBB4FAF4" box="[961,1074,1302,1322]" gridcol="3" gridrow="10" pageId="12" pageNumber="53">16.8–18.7</td>
<td id="73174662FFB50029DAE9FAE8FB66FAF4" box="[1190,1248,1302,1322]" gridcol="4" gridrow="10" pageId="12" pageNumber="53">17.5</td>
<td id="73174662FFB50029DB1CFAE8FA0AFAF4" box="[1363,1420,1302,1322]" gridcol="5" gridrow="10" pageId="12" pageNumber="53">1.08</td>
</tr>
<tr id="30C62F1EFFB50029DEC1FACAFA0AFA99" box="[142,1420,1332,1351]" gridrow="11" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FACAFE6DFA99" box="[142,491,1332,1351]" gridcol="0" gridrow="11" pageId="12" pageNumber="53">Pelvic fin length</th>
<td id="73174662FFB50029DC2EFACAFDE9FA99" box="[609,623,1332,1351]" gridcol="1" gridrow="11" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACFACAFCE3FA99" box="[739,869,1332,1351]" gridcol="2" gridrow="11" pageId="12" pageNumber="53">16.8</td>
<td id="73174662FFB50029DD8EFACAFBB4FA99" box="[961,1074,1332,1351]" gridcol="3" gridrow="11" pageId="12" pageNumber="53">16.8–20.8</td>
<td id="73174662FFB50029DAE9FACAFB66FA99" box="[1190,1248,1332,1351]" gridcol="4" gridrow="11" pageId="12" pageNumber="53">19.0</td>
<td id="73174662FFB50029DB1CFACAFA0AFA99" box="[1363,1420,1332,1351]" gridcol="5" gridrow="11" pageId="12" pageNumber="53">2.04</td>
</tr>
<tr id="30C62F1EFFB50029DEC1FAAFFA0AFABB" box="[142,1420,1361,1381]" gridrow="12" pageId="12" pageNumber="53" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="73174662FFB50029DEC1FAAFFE6DFABB" box="[142,491,1361,1381]" gridcol="0" gridrow="12" pageId="12" pageNumber="53">% HL</th>
</tr>
<tr id="30C62F1EFFB50029DEC1FA90FA0AFA5C" box="[142,1420,1390,1410]" gridrow="13" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FA90FE6DFA5C" box="[142,491,1390,1410]" gridcol="0" gridrow="13" pageId="12" pageNumber="53">Head width</th>
<td id="73174662FFB50029DC2EFA90FDE9FA5C" box="[609,623,1390,1410]" gridcol="1" gridrow="13" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACFA90FCE3FA5C" box="[739,869,1390,1410]" gridcol="2" gridrow="13" pageId="12" pageNumber="53">64.6</td>
<td id="73174662FFB50029DD8EFA90FBB4FA5C" box="[961,1074,1390,1410]" gridcol="3" gridrow="13" pageId="12" pageNumber="53">62.7–71.2</td>
<td id="73174662FFB50029DAE9FA90FB66FA5C" box="[1190,1248,1390,1410]" gridcol="4" gridrow="13" pageId="12" pageNumber="53">66.8</td>
<td id="73174662FFB50029DB1CFA90FA0AFA5C" box="[1363,1420,1390,1410]" gridcol="5" gridrow="13" pageId="12" pageNumber="53">4.26</td>
</tr>
<tr id="30C62F1EFFB50029DEC1FA72FA0AFA41" box="[142,1420,1420,1439]" gridrow="14" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FA72FE6DFA41" box="[142,491,1420,1439]" gridcol="0" gridrow="14" pageId="12" pageNumber="53">Head depth</th>
<td id="73174662FFB50029DC2EFA72FDE9FA41" box="[609,623,1420,1439]" gridcol="1" gridrow="14" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACFA72FCE3FA41" box="[739,869,1420,1439]" gridcol="2" gridrow="14" pageId="12" pageNumber="53">59.4</td>
<td id="73174662FFB50029DD8EFA72FBB4FA41" box="[961,1074,1420,1439]" gridcol="3" gridrow="14" pageId="12" pageNumber="53">57.5–61.0</td>
<td id="73174662FFB50029DAE9FA72FB66FA41" box="[1190,1248,1420,1439]" gridcol="4" gridrow="14" pageId="12" pageNumber="53">59.8</td>
<td id="73174662FFB50029DB1CFA72FA0AFA41" box="[1363,1420,1420,1439]" gridcol="5" gridrow="14" pageId="12" pageNumber="53">1.97</td>
</tr>
<tr id="30C62F1EFFB50029DEC1FA57FA0AFA62" box="[142,1420,1449,1468]" gridrow="15" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FA57FE6DFA62" box="[142,491,1449,1468]" gridcol="0" gridrow="15" pageId="12" pageNumber="53">Snout length</th>
<td id="73174662FFB50029DC2EFA57FDE9FA62" box="[609,623,1449,1468]" gridcol="1" gridrow="15" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACFA57FCE3FA62" box="[739,869,1449,1468]" gridcol="2" gridrow="15" pageId="12" pageNumber="53">44.5</td>
<td id="73174662FFB50029DD8EFA57FBB4FA62" box="[961,1074,1449,1468]" gridcol="3" gridrow="15" pageId="12" pageNumber="53">38.9–44.7</td>
<td id="73174662FFB50029DAE9FA57FB66FA62" box="[1190,1248,1449,1468]" gridcol="4" gridrow="15" pageId="12" pageNumber="53">41.5</td>
<td id="73174662FFB50029DB1CFA57FA0AFA62" box="[1363,1420,1449,1468]" gridcol="5" gridrow="15" pageId="12" pageNumber="53">2.96</td>
</tr>
<tr id="30C62F1EFFB50029DEC1FA38FA0AFA04" box="[142,1420,1478,1498]" gridrow="16" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FA38FE6DFA04" box="[142,491,1478,1498]" gridcol="0" gridrow="16" pageId="12" pageNumber="53">Orbit diameter</th>
<td id="73174662FFB50029DC2EFA38FDE9FA04" box="[609,623,1478,1498]" gridcol="1" gridrow="16" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACFA38FCE3FA04" box="[739,869,1478,1498]" gridcol="2" gridrow="16" pageId="12" pageNumber="53">16.5</td>
<td id="73174662FFB50029DD8EFA38FBB4FA04" box="[961,1074,1478,1498]" gridcol="3" gridrow="16" pageId="12" pageNumber="53">16.3–21.7</td>
<td id="73174662FFB50029DAE9FA38FB66FA04" box="[1190,1248,1478,1498]" gridcol="4" gridrow="16" pageId="12" pageNumber="53">19.3</td>
<td id="73174662FFB50029DB1CFA38FA0AFA04" box="[1363,1420,1478,1498]" gridcol="5" gridrow="16" pageId="12" pageNumber="53">2.75</td>
</tr>
<tr id="30C62F1EFFB50029DEC1FA1AFA0AFA29" box="[142,1420,1508,1527]" gridrow="17" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1FA1AFE6DFA29" box="[142,491,1508,1527]" gridcol="0" gridrow="17" pageId="12" pageNumber="53">Interorbital width</th>
<td id="73174662FFB50029DC2EFA1AFDE9FA29" box="[609,623,1508,1527]" gridcol="1" gridrow="17" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACFA1AFCE3FA29" box="[739,869,1508,1527]" gridcol="2" gridrow="17" pageId="12" pageNumber="53">44.3</td>
<td id="73174662FFB50029DD8EFA1AFBB4FA29" box="[961,1074,1508,1527]" gridcol="3" gridrow="17" pageId="12" pageNumber="53">41.0–44.3</td>
<td id="73174662FFB50029DAE9FA1AFB66FA29" box="[1190,1248,1508,1527]" gridcol="4" gridrow="17" pageId="12" pageNumber="53">42.7</td>
<td id="73174662FFB50029DB1CFA1AFA0AFA29" box="[1363,1420,1508,1527]" gridcol="5" gridrow="17" pageId="12" pageNumber="53">1.65</td>
</tr>
<tr id="30C62F1EFFB50029DEC1F9FFFA0AF9CA" box="[142,1420,1537,1556]" gridrow="18" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1F9FFFA0AF9CA" box="[142,1420,1537,1556]" colspan="6" colspanRight="5" gridcol="0" gridrow="18" pageId="12" pageNumber="53">% Orbit diameter</th>
</tr>
<tr id="30C62F1EFFB50029DEC1F9E0FA0AF9EC" box="[142,1420,1566,1586]" gridrow="19" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1F9E0FE6DF9EC" box="[142,491,1566,1586]" gridcol="0" gridrow="19" pageId="12" pageNumber="53">Rostral barbel length</th>
<td id="73174662FFB50029DC2EF9E0FDE9F9EC" box="[609,623,1566,1586]" gridcol="1" gridrow="19" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACF9E0FCE3F9EC" box="[739,869,1566,1586]" gridcol="2" gridrow="19" pageId="12" pageNumber="53">56.2</td>
<td id="73174662FFB50029DD8EF9E0FBB4F9EC" box="[961,1074,1566,1586]" gridcol="3" gridrow="19" pageId="12" pageNumber="53">44.0–66.0</td>
<td id="73174662FFB50029DAE9F9E0FB66F9EC" box="[1190,1248,1566,1586]" gridcol="4" gridrow="19" pageId="12" pageNumber="53">54.0</td>
<td id="73174662FFB50029DB1CF9E0FA0AF9EC" box="[1363,1420,1566,1586]" gridcol="5" gridrow="19" pageId="12" pageNumber="53">11.14</td>
</tr>
<tr id="30C62F1EFFB50029DEC1F9C2FA0AF991" box="[142,1420,1596,1615]" gridrow="20" pageId="12" pageNumber="53">
<th id="73174662FFB50029DEC1F9C2FE6DF991" box="[142,491,1596,1615]" gridcol="0" gridrow="20" pageId="12" pageNumber="53">Maxillary barbel length</th>
<td id="73174662FFB50029DC2EF9C2FDE9F991" box="[609,623,1596,1615]" gridcol="1" gridrow="20" pageId="12" pageNumber="53">3</td>
<td id="73174662FFB50029DCACF9C2FCE3F991" box="[739,869,1596,1615]" gridcol="2" gridrow="20" pageId="12" pageNumber="53">57.1</td>
<td id="73174662FFB50029DD8EF9C2FBB4F991" box="[961,1074,1596,1615]" gridcol="3" gridrow="20" pageId="12" pageNumber="53">35.6–68.4</td>
<td id="73174662FFB50029DAE9F9C2FB66F991" box="[1190,1248,1596,1615]" gridcol="4" gridrow="20" pageId="12" pageNumber="53">54.0</td>
<td id="73174662FFB50029DB1CF9C2FA0AF991" box="[1363,1420,1596,1615]" gridcol="5" gridrow="20" pageId="12" pageNumber="53">16.74</td>
</tr>
<tr id="30C62F1EFFB50029DEC1F9A6FA0AF9B3" box="[142,1420,1624,1645]" gridrow="21" pageId="12" pageNumber="53" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="73174662FFB50029DEC1F9A6FE6DF9B3" box="[142,491,1624,1645]" gridcol="0" gridrow="21" pageId="12" pageNumber="53">Dorsal fin rays</th>
<td id="73174662FFB50029DCACF9A6FCE3F9B3" box="[739,869,1624,1645]" gridcol="2" gridrow="21" pageId="12" pageNumber="53">III + 7</td>
</tr>
<tr id="30C62F1EFFB50029DEC1F98BFA0AF954" box="[142,1420,1653,1674]" gridrow="22" pageId="12" pageNumber="53" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="73174662FFB50029DEC1F98BFE6DF954" box="[142,491,1653,1674]" gridcol="0" gridrow="22" pageId="12" pageNumber="53">Anal fin rays</th>
<td id="73174662FFB50029DCACF98BFCE3F954" box="[739,869,1653,1674]" gridcol="2" gridrow="22" pageId="12" pageNumber="53">III + 5</td>
</tr>
<tr id="30C62F1EFFB50029DEC1F96DFA0AF979" box="[142,1420,1683,1703]" gridrow="23" pageId="12" pageNumber="53" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="73174662FFB50029DEC1F96DFE6DF979" box="[142,491,1683,1703]" gridcol="0" gridrow="23" pageId="12" pageNumber="53">Pectoral fin rays</th>
<td id="73174662FFB50029DCACF96DFCE3F979" box="[739,869,1683,1703]" gridcol="2" gridrow="23" pageId="12" pageNumber="53">IV + 11</td>
</tr>
<tr id="30C62F1EFFB50029DEC1F94EFA0AF91B" box="[142,1420,1712,1733]" gridrow="24" pageId="12" pageNumber="53" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="73174662FFB50029DEC1F94EFE6DF91B" box="[142,491,1712,1733]" gridcol="0" gridrow="24" pageId="12" pageNumber="53">Pelvic fin rays</th>
<td id="73174662FFB50029DCACF94EFCE3F91B" box="[739,869,1712,1733]" gridcol="2" gridrow="24" pageId="12" pageNumber="53">II + 6</td>
</tr>
<tr id="30C62F1EFFB50029DEC1F930FA0AF93C" box="[142,1420,1742,1762]" gridrow="25" pageId="12" pageNumber="53" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="73174662FFB50029DEC1F930FE6DF93C" box="[142,491,1742,1762]" gridcol="0" gridrow="25" pageId="12" pageNumber="53">Lateral line scales</th>
<td id="73174662FFB50029DCACF930FCE3F93C" box="[739,869,1742,1762]" gridcol="2" gridrow="25" pageId="12" pageNumber="53">39</td>
</tr>
<tr id="30C62F1EFFB50029DEC1F912FA0AF921" box="[142,1420,1772,1791]" gridrow="26" pageId="12" pageNumber="53" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="73174662FFB50029DEC1F912FE6DF921" box="[142,491,1772,1791]" gridcol="0" gridrow="26" pageId="12" pageNumber="53">Scale rows lateral line–dorsal fin</th>
<td id="73174662FFB50029DCACF912FCE3F921" box="[739,869,1772,1791]" gridcol="2" gridrow="26" pageId="12" pageNumber="53">4–5 (H, 5)</td>
</tr>
<tr id="30C62F1EFFB50029DEC1F8F7FA0AF8C3" box="[142,1420,1801,1821]" gridrow="27" pageId="12" pageNumber="53" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="73174662FFB50029DEC1F8F7FE6DF8C3" box="[142,491,1801,1821]" gridcol="0" gridrow="27" pageId="12" pageNumber="53">Scale rows lateral line–pelvic fin</th>
<td id="73174662FFB50029DCACF8F7FCE3F8C3" box="[739,869,1801,1821]" gridcol="2" gridrow="27" pageId="12" pageNumber="53">3–4 (H, 4)</td>
</tr>
<tr id="30C62F1EFFB50029DEC1F8D8FA0AF8E4" box="[142,1420,1830,1850]" gridrow="28" pageId="12" pageNumber="53" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="73174662FFB50029DEC1F8D8FE6DF8E4" box="[142,491,1830,1850]" gridcol="0" gridrow="28" pageId="12" pageNumber="53">Scale rows lateral line–anal fin</th>
<td id="73174662FFB50029DCACF8D8FCE3F8E4" box="[739,869,1830,1850]" gridcol="2" gridrow="28" pageId="12" pageNumber="53">4.5</td>
</tr>
<tr id="30C62F1EFFB50029DEC1F8BAFA0AF889" box="[142,1420,1860,1879]" gridrow="29" pageId="12" pageNumber="53" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="73174662FFB50029DEC1F8BAFE6DF889" box="[142,491,1860,1879]" gridcol="0" gridrow="29" pageId="12" pageNumber="53">Predorsal scales</th>
<td id="73174662FFB50029DCACF8BAFCE3F889" box="[739,869,1860,1879]" gridcol="2" gridrow="29" pageId="12" pageNumber="53">8–9 (H, 8)</td>
</tr>
</table>
</paragraph>
<table id="FCF6DFFCFFB40029DEEEFF3CFA22F8CF" box="[161,1444,194,1809]" pageId="13" pageNumber="54">
<subSection id="E77934B7FFB4FFDBDFAAFF2CFA14F93F" pageId="13" pageNumber="54" type="multiple">
<paragraph id="8E492D5CFFB4FFDBDFAAFF2CFBE6FF37" blockId="13.[485,1120,210,234]" box="[485,1120,210,234]" pageId="13" pageNumber="54">
<heading id="D5019A30FFB4FFDBDFAAFF2CFBE6FF37" box="[485,1120,210,234]" centered="true" fontSize="9" level="2" pageId="13" pageNumber="54" reason="2">
ARTIFICIAL KEY TO
<taxonomicName id="49F656DFFFB4FFDBDC88FF28FCEAFF37" authority="OF THE" authorityName="OF THE" box="[711,876,214,233]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="genus">
<emphasis id="BC82F14EFFB4FFDBDC88FF28FC97FF37" box="[711,785,214,233]" italics="true" pageId="13" pageNumber="54">GARRA</emphasis>
OF THE
</taxonomicName>
AFRICAN CONTINENT
</heading>
</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDFF05FA14FEF5" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">
1. Two pairs of barbels (maxillary and rostral) .................................................................................................................. 2 Single pair of (maxillary) barbels.........................................................................................
<taxonomicName id="49F656DFFFB4FFDBDA3FFEE6FA14FEF5" authority="(Ethiopia)" authorityName="Stiassny &amp; Getahun" authorityYear="2007" baseAuthorityName="Ethiopia" box="[1136,1426,280,299]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="duobarbis">
<emphasis id="BC82F14EFFB4FFDBDA3FFEE6FAA7FEF5" box="[1136,1313,280,299]" italics="true" pageId="13" pageNumber="54">Garra duobarbis</emphasis>
(
<collectingCountry id="F6E16DCCFFB4FFDBDB7FFEE6FA0DFEF5" box="[1328,1419,280,299]" name="Ethiopia" pageId="13" pageNumber="54">Ethiopia</collectingCountry>
)
</taxonomicName>
</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDFECBFA14FE5F" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">
2. Rostral fold ventrally expanded and covering upper jaw; mouth broad .......................................................................3 Rostral fold not ventrally expanded and upper jaw exposed; mouth narrow................................................................. .............................................................................................................................
<taxonomicName id="49F656DFFFB4FFDBDDB4FE90FB24FE5E" authorityName="Stiassny &amp; Getahun" authorityYear="2007" box="[1019,1186,366,385]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="regressus">
<emphasis id="BC82F14EFFB4FFDBDDB4FE90FB24FE5E" box="[1019,1186,366,385]" italics="true" pageId="13" pageNumber="54">Garra regressus</emphasis>
</taxonomicName>
(
<collectingCountry id="F6E16DCCFFB4FFDBDAFDFE90FA89FE5F" box="[1202,1295,366,385]" name="Ethiopia" pageId="13" pageNumber="54">Ethiopia</collectingCountry>
, Lake Tana)
</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDFE75FA14FE09" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">
3. Mental disc moderately to very well developed, with free lateral and posterior margins (
<figureCitation id="16CD31D9FFB4FFDBDACBFE74FB6BFE40" box="[1156,1261,394,414]" captionStart="Figure 1" captionStartId="3.[162,240,1287,1306]" captionTargetBox="[164,1448,194,1249]" captionTargetId="figure-312@3.[162,1448,194,1254]" captionTargetPageId="3" captionText="Figure 1. Range of variation in disc type among African Garra: (A) type-A disc, weakly developed; (B) type-B disc, intermediate development; (C) type-C disc, well-developed." figureDoi="http://doi.org/10.5281/zenodo.5429394" httpUri="https://zenodo.org/record/5429394/files/figure.png" pageId="13" pageNumber="54">Fig. 1B, C</figureCitation>
) ......................4 Mental disc reduced to a central callus, sometimes with a narrow free posterior margin, but entirely lacking free lateral margins and papillate periphery (
<figureCitation id="16CD31D9FFB4FFDBDCDFFE3AFD5DFE09" box="[656,731,452,471]" captionStart="Figure 1" captionStartId="3.[162,240,1287,1306]" captionTargetBox="[164,1448,194,1249]" captionTargetId="figure-312@3.[162,1448,194,1254]" captionTargetPageId="3" captionText="Figure 1. Range of variation in disc type among African Garra: (A) type-A disc, weakly developed; (B) type-B disc, intermediate development; (C) type-C disc, well-developed." figureDoi="http://doi.org/10.5281/zenodo.5429394" httpUri="https://zenodo.org/record/5429394/files/figure.png" pageId="13" pageNumber="54">Fig. 1A</figureCitation>
) ........................................................................................................ 14
</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDFE1FFA14FDCE" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">4. Chest with either many, or some scattered scales ..........................................................................................................5 Chest asquamate ............................................................................................................................................................... 6</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDFDE4FA14FD5D" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">
5. Lateral line scales 34–35; vent located close to anal fin (vent distance 8.6–19.1% of distance between anterior end of anal- and pelvic-fin insertions); eye median ...............................
<taxonomicName id="49F656DFFFB4FFDBDD3CFDC9FBE4FD97" baseAuthorityName="Ruppell" baseAuthorityYear="1931" box="[883,1122,566,586]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="quadrimaculata">
<emphasis id="BC82F14EFFB4FFDBDD3CFDC9FBE4FD97" box="[883,1122,566,586]" italics="true" pageId="13" pageNumber="54">Garra quadrimaculata</emphasis>
</taxonomicName>
(
<collectingCountry id="F6E16DCCFFB4FFDBDA3DFDC8FB49FD97" box="[1138,1231,566,585]" name="Ethiopia" pageId="13" pageNumber="54">Ethiopia</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDA94FDC8FAAFFD97" box="[1243,1321,566,585]" name="Eritrea" pageId="13" pageNumber="54">Eritrea</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDB7AFDC8FA0DFD97" box="[1333,1419,566,585]" name="Somalia" pageId="13" pageNumber="54">Somalia</collectingCountry>
) Lateral line scales 40–42; vent located relatively far from anal fin (vent distance 29.3–30.2% of distance between anterior end of anal- and pelvic-fin insertions); eye posterior ....
<taxonomicName id="49F656DFFFB4FFDBDD32FD8EFB85FD5D" baseAuthorityName="Boulenger" baseAuthorityYear="1905" box="[893,1027,624,643]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="hindii">
<emphasis id="BC82F14EFFB4FFDBDD32FD8EFB85FD5D" box="[893,1027,624,643]" italics="true" pageId="13" pageNumber="54">Garra hindii</emphasis>
</taxonomicName>
(
<collectingCountry id="F6E16DCCFFB4FFDBDA5DFD8EFB06FD5D" box="[1042,1152,624,643]" name="Cameroon" pageId="13" pageNumber="54">Cameroon</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDAC3FD8EFB57FD5D" box="[1164,1233,624,643]" name="Democratic Republic of the Congo" pageId="13" pageNumber="54">Congo</collectingCountry>
*,
<collectingCountry id="F6E16DCCFFB4FFDBDAA9FD8EFAAAFD5D" box="[1254,1324,624,643]" name="Kenya" pageId="13" pageNumber="54">Kenya</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDB77FD8EFA0DFD5D" box="[1336,1419,624,643]" name="Uganda" pageId="13" pageNumber="54">Uganda</collectingCountry>
)
</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDFD72FA14FD62" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">6. Post-pelvic region asquamate ........................................................................................................................................... 7 Post-pelvic region scaled ................................................................................................................................................... 9</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDFD38FA14FCF1" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">
7. Body either with distinct midlateral band, or darker dorsally than ventrally; black spots in basal membrane of dorsal fin; predorsal region only partially scaled; region between vent and anal fin scaled ............................................8 Uniformly pale body coloration; no markings in dorsal fin; fully scaled predorsal region; no scales between vent and anal fin ........................................................................................................
<taxonomicName id="49F656DFFFB4FFDBDD89FCE2FBF3FCF1" baseAuthorityName="Boulenger" baseAuthorityYear="1901" box="[966,1141,796,815]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="blanfordii">
<emphasis id="BC82F14EFFB4FFDBDD89FCE2FBF3FCF1" box="[966,1141,796,815]" italics="true" pageId="13" pageNumber="54">Garra blanfordii</emphasis>
</taxonomicName>
(
<collectingCountry id="F6E16DCCFFB4FFDBDACBFCE2FB67FCF1" box="[1156,1249,796,815]" name="Ethiopia" pageId="13" pageNumber="54">Ethiopia</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDAA2FCE2FABDFCF1" box="[1261,1339,796,815]" name="Eritrea" pageId="13" pageNumber="54">Eritrea</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDB08FCE2FA0DFCF1" box="[1351,1419,796,815]" name="Sudan" pageId="13" pageNumber="54">Sudan</collectingCountry>
)
</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDFCC7FA14FC7F" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">
8. Between five and nine scales in predorsal region; body depth 14.9–16.8, m. 16.0% SL; intestine short (SL 95.0– 102.0% Int.L) ..................................................................................................................................
<taxonomicName id="49F656DFFFB4FFDBDAE5FCABFA14FCB6" authority="(Ethiopia)" authorityName="Stiassny &amp; Getahun" authorityYear="2007" baseAuthorityName="Ethiopia" box="[1194,1426,853,872]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="geba">
<emphasis id="BC82F14EFFB4FFDBDAE5FCABFAA7FCB6" box="[1194,1313,853,872]" italics="true" pageId="13" pageNumber="54">Garra geba</emphasis>
(
<collectingCountry id="F6E16DCCFFB4FFDBDB7FFCABFA0DFCB6" box="[1328,1419,853,872]" name="Ethiopia" pageId="13" pageNumber="54">Ethiopia</collectingCountry>
)
</taxonomicName>
Between zero and three scales in predorsal region; body depth 15.6–23.9, m. 20.4% SL; intestine moderately long (SL 39.0–42.0% Int.L) .........................
<taxonomicName id="49F656DFFFB4FFDBDC0EFC71FD79FC7F" baseAuthorityName="RUPPELL" baseAuthorityYear="1836" box="[577,767,910,930]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="dembeensis">
<emphasis id="BC82F14EFFB4FFDBDC0EFC71FD79FC7F" box="[577,767,910,930]" italics="true" pageId="13" pageNumber="54">Garra dembeensis</emphasis>
</taxonomicName>
(
<collectingCountry id="F6E16DCCFFB4FFDBDD40FC71FCFBFC7C" box="[783,893,911,930]" name="Cameroon" pageId="13" pageNumber="54">Cameroon</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDDC7FC70FC44FC7F" box="[904,962,910,929]" name="Chad" pageId="13" pageNumber="54">Chad</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDD81FC71FB88FC7C" box="[974,1038,911,930]" name="Egypt" pageId="13" pageNumber="54">Egypt</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDA55FC70FBF0FC7F" box="[1050,1142,910,929]" name="Ethiopia" pageId="13" pageNumber="54">Ethiopia</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDACDFC71FB4EFC7C" box="[1154,1224,911,930]" name="Kenya" pageId="13" pageNumber="54">Kenya</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDA9BFC70FAA3FC7F" box="[1236,1317,910,929]" name="Nigeria" pageId="13" pageNumber="54">Nigeria</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDB66FC70FA0DFC7F" box="[1321,1419,910,929]" name="Tanzania" pageId="13" pageNumber="54">Tanzania</collectingCountry>
)
</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDFC55FA14FC05" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">9. Predorsal region either naked or only partially scaled ................................................................................................ 10 Predorsal region fully scaled .......................................................................................................................................... 11</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDFC1BFA14FB93" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">
10. Body and head markedly dorso-ventrally flattened (body depth 10.6–15.0, m. 11.9% SL); gas bladder extremely reduced, posterior chamber 3.1–3.8% SL.................................................................
<taxonomicName id="49F656DFFFB4FFDBDA6CFBFFFB51FBCA" authorityName="Poll" authorityYear="1959" box="[1059,1239,1025,1044]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="congoensis">
<emphasis id="BC82F14EFFB4FFDBDA6CFBFFFB51FBCA" box="[1059,1239,1025,1044]" italics="true" pageId="13" pageNumber="54">Garra congoensis</emphasis>
</taxonomicName>
(
<collectingCountry id="F6E16DCCFFB4FFDBDAA9FBFFFAB4FBCA" box="[1254,1330,1025,1044]" name="Angola" pageId="13" pageNumber="54">Angola</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDB71FBFFFA07FBCA" box="[1342,1409,1025,1044]" name="Democratic Republic of the Congo" pageId="13" pageNumber="54">Congo</collectingCountry>
*) Body depth 15.8–22.3, m. 17.8% SL; posterior chamber of gas bladder 6.2–10.0% SL ................................................. ...............................................
<taxonomicName id="49F656DFFFB4FFDBDC44FBC5FD11FB93" baseAuthorityName="Nichols &amp; Griscom" baseAuthorityYear="1917" box="[523,663,1083,1102]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="ornata">
<emphasis id="BC82F14EFFB4FFDBDC44FBC5FD11FB93" box="[523,663,1083,1102]" italics="true" pageId="13" pageNumber="54">Garra ornata</emphasis>
</taxonomicName>
(
<collectingCountry id="F6E16DCCFFB4FFDBDCE9FBC5FC92FB90" box="[678,788,1083,1102]" name="Cameroon" pageId="13" pageNumber="54">Cameroon</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDD50FBC4FBACFB93" box="[799,1066,1082,1101]" name="Central African Republic" pageId="13" pageNumber="54">Central African Republic</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDA79FBC5FBFCFB90" box="[1078,1146,1083,1102]" name="Democratic Republic of the Congo" pageId="13" pageNumber="54">Congo</collectingCountry>
*,
<collectingCountry id="F6E16DCCFFB4FFDBDADFFBC4FB50FB93" box="[1168,1238,1082,1101]" name="Gabon" pageId="13" pageNumber="54">Gabon</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDAAEFBC4FAA9FB93" box="[1249,1327,1082,1101]" name="Guinea-Bissau" pageId="13" pageNumber="54">Guinea</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDB74FBC4FA0DFB93" box="[1339,1419,1082,1101]" name="Nigeria" pageId="13" pageNumber="54">Nigeria</collectingCountry>
)
</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDFBA9FA14FB7A" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">
11. Either 29 or 30 scales on the lateral line; body depth 29.1–30.9, m. 30.0% SL; 27–28 vertebrae................................ ..............................................................................................................................................
<taxonomicName id="49F656DFFFB4FFDBDA29FB8AFA14FB59" authority="(Eritrea)" authorityName="Menon" authorityYear="1958" baseAuthorityName="Eritrea" box="[1126,1426,1140,1159]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="ethelwynnae">
<emphasis id="BC82F14EFFB4FFDBDA29FB8AFAA8FB59" box="[1126,1326,1140,1159]" italics="true" pageId="13" pageNumber="54">Garra ethelwynnae</emphasis>
(
<collectingCountry id="F6E16DCCFFB4FFDBDB71FB8AFA0DFB59" box="[1342,1419,1140,1159]" name="Eritrea" pageId="13" pageNumber="54">Eritrea</collectingCountry>
)
</taxonomicName>
More than 34 scales on the lateral line; body depth less than 28.0% SL; more than 30 vertebrae ......................... 12
</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDFB53FA14FAC8" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">
12. Disc moderately developed (
<typeStatus id="514D93FEFFB4FFDBDC43FB53FDBCFB1E" box="[524,570,1197,1216]" pageId="13" pageNumber="54">type</typeStatus>
B); dark ash-grey midlateral band to base of caudal peduncle; belly scaled; no tubercles on snout ..............................................................................................................
<taxonomicName id="49F656DFFFB4FFDBDA10FB36FA14FB03" authority="(Ethiopia)" baseAuthorityName="Ethiopia" baseAuthorityYear="1927" box="[1119,1426,1224,1245]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="aethiopica">
<emphasis id="BC82F14EFFB4FFDBDA10FB36FAA7FB03" box="[1119,1313,1224,1245]" italics="true" pageId="13" pageNumber="54">Garra aethiopica</emphasis>
(
<collectingCountry id="F6E16DCCFFB4FFDBDB7FFB34FA0DFB03" box="[1328,1419,1226,1245]" name="Ethiopia" pageId="13" pageNumber="54">Ethiopia</collectingCountry>
)
</taxonomicName>
Disc well developed (
<typeStatus id="514D93FEFFB4FFDBDFF8FB19FE63FB24" box="[439,485,1255,1274]" pageId="13" pageNumber="54">type</typeStatus>
C); body more-or-less uniformly dark on flanks; belly either asquamate or with only a few embedded scales; tubercles on snout ............................................................................................................................. 13
</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDFADEFA14FA57" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">
13. Some scales on belly; distance from vent to anal fin 10.3–16.9, m. 13.6% of distance between anal- and pelvic-fin insertions; eye either median, or posterior in large specimens .........................................
<taxonomicName id="49F656DFFFB4FFDBDA3FFAC3FA14FA91" authority="(Ethiopia)" baseAuthorityName="Ethiopia" box="[1136,1426,1340,1360]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="makiensis">
<emphasis id="BC82F14EFFB4FFDBDA3FFAC3FAA7FA91" box="[1136,1313,1340,1360]" italics="true" pageId="13" pageNumber="54">Garra makiensis</emphasis>
(
<collectingCountry id="F6E16DCCFFB4FFDBDB7FFAC2FA0DFA91" box="[1328,1419,1340,1359]" name="Ethiopia" pageId="13" pageNumber="54">Ethiopia</collectingCountry>
)
</taxonomicName>
Belly asquamate; distance from vent to anal fin 19.2–29.5, m. 18.2% of distance of anal- and pelvic-fin insertions; eye median........................................................................................................................................
<taxonomicName id="49F656DFFFB4FFDBDAC2FA88FA14FA57" authority="(Ethiopia)" baseAuthorityName="Ethiopia" baseAuthorityYear="1925" box="[1165,1426,1398,1417]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="ignestii">
<emphasis id="BC82F14EFFB4FFDBDAC2FA88FAA7FA56" box="[1165,1313,1398,1417]" italics="true" pageId="13" pageNumber="54">Garra ignestii</emphasis>
(
<collectingCountry id="F6E16DCCFFB4FFDBDB7FFA88FA0DFA57" box="[1328,1419,1398,1417]" name="Ethiopia" pageId="13" pageNumber="54">Ethiopia</collectingCountry>
)
</taxonomicName>
</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDFA6DFA14FA1C" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">
14. Post-pelvic region asquamate, Ethiopian distribution.................................................................................................. 15 Post-pelvic region scaled, found in
<collectingCountry id="F6E16DCCFFB4FFDBDC02FA51FD3EFA1C" box="[589,696,1455,1474]" name="Cameroon" pageId="13" pageNumber="54">Cameroon</collectingCountry>
and
<collectingCountry id="F6E16DCCFFB4FFDBDCA1FA51FCA9FA1C" box="[750,815,1455,1474]" name="Chad" pageId="13" pageNumber="54">Tchad</collectingCountry>
............................................................................................ 16
</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDFA32FA14F9B0" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">
15. Length of caudal peduncle 20.4–22.3, m. 20.2% SL; either 39 or 40 scales in lateral line; intestine short (SL 62.7–79% Int.L); spots in basal membrane of dorsal fin either absent or small and faint ........................................... .....................................................................................................................................
<taxonomicName id="49F656DFFFB4FFDBDA64F9FBFB25F9C6" authorityName="Stiassny &amp; Getahun" authorityYear="2007" box="[1067,1187,1541,1560]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="tana">
<emphasis id="BC82F14EFFB4FFDBDA64F9FBFB25F9C6" box="[1067,1187,1541,1560]" italics="true" pageId="13" pageNumber="54">Garra tana</emphasis>
</taxonomicName>
(
<collectingCountry id="F6E16DCCFFB4FFDBDAFDF9FBFA89F9C6" box="[1202,1295,1541,1560]" name="Ethiopia" pageId="13" pageNumber="54">Ethiopia</collectingCountry>
, Lake Tana) Length of caudal peduncle 13.3–20.7, m. 16.8% SL; 37–38 scales in lateral line; intestine longer (SL 40–56% Int.L); either four or five elongate black spots in basal membrane of dorsal fin....................................................................... ...................................................................................................
<taxonomicName id="49F656DFFFB4FFDBDD18F9A5FB81F9B0" authorityName="Stiassny &amp; Getahun" authorityYear="2007" box="[855,1031,1627,1646]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="dembecha">
<emphasis id="BC82F14EFFB4FFDBDD18F9A5FB81F9B0" box="[855,1031,1627,1646]" italics="true" pageId="13" pageNumber="54">Garra dembecha</emphasis>
</taxonomicName>
(
<collectingCountry id="F6E16DCCFFB4FFDBDA59F9A5FBF5F9B0" box="[1046,1139,1627,1646]" name="Ethiopia" pageId="13" pageNumber="54">Ethiopia</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDA31F9A5FB4AF9B0" box="[1150,1228,1627,1646]" name="Eritrea" pageId="13" pageNumber="54">Eritrea</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDA97F9A2FA98F9B1" box="[1240,1310,1628,1647]" name="Kenya" pageId="13" pageNumber="54">Kenya</collectingCountry>
,
<collectingCountry id="F6E16DCCFFB4FFDBDB66F9A5FA0DF9B0" box="[1321,1419,1627,1646]" name="Tanzania" pageId="13" pageNumber="54">Tanzania</collectingCountry>
)
</paragraph>
<paragraph id="8E492D5CFFB4FFDBDEFDF986FA14F93F" blockId="13.[178,1426,251,1790]" pageId="13" pageNumber="54">
16. Vent close to anal fin (vent distance 13.3–17.4% of distance between pelvic- and anal-fin insertions) ....................... ............................................................................................................................................
<taxonomicName id="49F656DFFFB4FFDBDA16F96BFA14F979" authority="(Tchad)" authorityName="Blache &amp; Miton" authorityYear="1960" baseAuthorityName="Tchad" box="[1113,1426,1684,1704]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="lancrenonensis">
<emphasis id="BC82F14EFFB4FFDBDA16F96BFABCF979" box="[1113,1338,1684,1704]" italics="true" pageId="13" pageNumber="54">Garra lancrenonensis</emphasis>
(
<collectingCountry id="F6E16DCCFFB4FFDBDB06F96AFA0DF979" box="[1353,1419,1684,1703]" name="Chad" pageId="13" pageNumber="54">Tchad</collectingCountry>
)
</taxonomicName>
Vent situated relatively far from anal fin (vent distance 28.5–30.6% of distance between pelvic- and anal-fin insertions) ............................................................................................................................
<taxonomicName id="49F656DFFFB4FFDBDA2BF930FA14F93F" authority="(Cameroon)" authorityName="Roberts" authorityYear="1990" baseAuthorityName="Cameroon" box="[1124,1426,1742,1761]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="13" pageNumber="54" phylum="Chordata" rank="species" species="allostoma">
<emphasis id="BC82F14EFFB4FFDBDA2BF930FA96F93F" box="[1124,1296,1742,1761]" italics="true" pageId="13" pageNumber="54">Garra allostoma</emphasis>
(
<collectingCountry id="F6E16DCCFFB4FFDBDB50F930FA0AF93F" box="[1311,1420,1742,1761]" name="Cameroon" pageId="13" pageNumber="54">Cameroon</collectingCountry>
)
</taxonomicName>
</paragraph>
</subSection>
<footnote id="EDED3152FFB4FFDBDEFDF915FE00F923" box="[178,390,1770,1790]" pageId="13" pageNumber="54">
<paragraph id="8E492D5CFFB4FFDBDEFDF915FE00F923" blockId="13.[178,1426,251,1790]" box="[178,390,1770,1790]" pageId="13" pageNumber="54">*Congo River basin.</paragraph>
</footnote>
</table>
<caption id="DA897DD4FFB7FFD8DEC1FC09FC74FBD4" ID-DOI="http://doi.org/10.5281/zenodo.5429414" ID-Zenodo-Dep="5429414" box="[142,1010,1015,1035]" httpUri="https://zenodo.org/record/5429414/files/figure.png" pageId="14" pageNumber="55" startId="14.[142,220,1015,1034]" targetBox="[302,1251,202,982]" targetPageId="14">
<paragraph id="8E492D5CFFB7FFD8DEC1FC09FC74FBD4" blockId="14.[142,1010,1015,1035]" box="[142,1010,1015,1035]" pageId="14" pageNumber="55">
<emphasis id="BC82F14EFFB7FFD8DEC1FC09FE85FBD4" bold="true" box="[142,259,1015,1034]" pageId="14" pageNumber="55">Figure 11.</emphasis>
<taxonomicName id="49F656DFFFB7FFD8DF40FC09FE30FBD4" authorityName="Stiassny &amp; Getahun" authorityYear="2007" box="[271,438,1015,1034]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="55" phylum="Chordata" rank="species" species="regressus">
<emphasis id="BC82F14EFFB7FFD8DF40FC09FE30FBD4" box="[271,438,1015,1034]" italics="true" pageId="14" pageNumber="55">Garra regressus</emphasis>
</taxonomicName>
, holotype, AMNH 228487, 121.8 mm standard length.
</paragraph>
</caption>
<paragraph id="8E492D5CFFB7FFD8DEC1F93EFC4FFB25" blockId="14.[142,758,1728,1904]" lastBlockId="14.[806,1422,1100,1275]" pageId="14" pageNumber="55">
<emphasis id="BC82F14EFFB7FFD8DEC1F93EFE26F908" box="[142,416,1728,1750]" italics="true" pageId="14" pageNumber="55">Colour in preservation:</emphasis>
Body dark brown dorsally, somewhat lighter brown below lateral line but not markedly so. In largest specimen (
<figureCitation id="16CD31D9FFB7FFD8DC51F900FDEAF8CA" box="[542,620,1790,1812]" captionStart="Figure 11" captionStartId="14.[142,220,1015,1034]" captionTargetBox="[302,1251,202,982]" captionTargetId="figure-437@14.[302,1262,194,982]" captionTargetPageId="14" captionText="Figure 11. Garra regressus, holotype, AMNH 228487, 121.8 mm standard length." figureDoi="http://doi.org/10.5281/zenodo.5429414" httpUri="https://zenodo.org/record/5429414/files/figure.png" pageId="14" pageNumber="55">Fig. 11</figureCitation>
) traces of a dark midlateral band evident. Head, snout, cheek, and opercle: dark brown. Lips, lower cheek, gular region, and branchiostegal membranes: pale yellow brown. Dorsal and anal fins dark brown distally, paler proximally, with pale, creamy subacuminate tips. Either four or five discrete black spots between rays in basal membrane of dorsal fin. Caudal fin uniformly dark brown. Pelvic and pectoral fins light brown with pale leading edges.
</paragraph>
<paragraph id="8E492D5CFFB7FFD8DD69FAEDFCE9FA7A" blockId="14.[806,1422,1299,1444]" pageId="14" pageNumber="55">
<emphasis id="BC82F14EFFB7FFD8DD69FAEDFBE6FAF6" box="[806,1120,1299,1320]" italics="true" pageId="14" pageNumber="55">Distribution and habitat:</emphasis>
Currently known from region of Gerima in southern part of Lake Tana,
<collectingCountry id="F6E16DCCFFB7FFD8DB1CFACCFCDFFAB8" name="Ethiopia" pageId="14" pageNumber="55">Ethiopia</collectingCountry>
. Specimens were collected by trawl in papyrus beds close to shore (
<quantity id="490E80B9FFB7FFD8DA6EFA91FB03FA5A" box="[1057,1157,1391,1413]" metricMagnitude="1" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="14" pageNumber="55" unit="m" value="27.5" valueMax="30.0" valueMin="25.0">25–30 m</quantity>
) in water
<emphasis id="BC82F14EFFB7FFD8DB43FA8EFA91FA5B" box="[1292,1303,1392,1413]" italics="true" pageId="14" pageNumber="55">c</emphasis>
.
<quantity id="490E80B9FFB7FFD8DB65FA91FAECFA5A" box="[1322,1386,1391,1413]" metricMagnitude="0" metricUnit="m" metricValue="2.5" pageId="14" pageNumber="55" unit="m" value="2.5">2.5 m</quantity>
in depth.
</paragraph>
<paragraph id="8E492D5CFFB7FFD8DD69FA42FBEAF9D1" blockId="14.[806,1421,1468,1551]" pageId="14" pageNumber="55">
<emphasis id="BC82F14EFFB7FFD8DD69FA42FBA7FA0F" box="[806,1057,1468,1489]" italics="true" pageId="14" pageNumber="55">
Etymology:
<taxonomicName id="49F656DFFFB7FFD8DDFEFA42FBA7FA0F" authorityName="Stiassny &amp; Getahun" authorityYear="2007" box="[945,1057,1468,1489]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="55" phylum="Chordata" rank="species" species="regressus">Regressus</taxonomicName>
</emphasis>
, from the Latin regress, meaning to go back, in reference to the regressed rostral cap characteristic of the species.
</paragraph>
<paragraph id="8E492D5CFFB7FFD8DD69F9D9FC08F908" blockId="14.[806,1422,1575,1750]" pageId="14" pageNumber="55">
<emphasis id="BC82F14EFFB7FFD8DD69F9D9FB63F9E2" box="[806,1253,1575,1597]" italics="true" pageId="14" pageNumber="55">Material examined – type material:</emphasis>
<typeStatus id="514D93FEFFB7FFD8DAB0F9D9FAE3F9E3" box="[1279,1381,1575,1597]" pageId="14" pageNumber="55" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="49F656DFFFB7FFD8DD69F9B8FB9DF985" authority=", AMNH" authorityName="AMNH" box="[806,1051,1606,1627]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="55" phylum="Chordata" rank="species" species="regressus">
<emphasis id="BC82F14EFFB7FFD8DD69F9B8FC35F985" box="[806,947,1606,1627]" italics="true" pageId="14" pageNumber="55">G. regressus,</emphasis>
AMNH
</taxonomicName>
<quantity id="490E80B9FFB7FFD8DA6DF9B8FA84F985" box="[1058,1282,1606,1627]" metricMagnitude="5" metricUnit="m" metricValue="2.284871218" pageId="14" pageNumber="55" unit="mm" value="2.284871218E8">228487, 121.8 mm</quantity>
SL, Lake
<taxonomicName id="49F656DFFFB7FFD8DD69F99BFC4CF9A4" authority=", Gerima" authorityName="Gerima" box="[806,970,1636,1658]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="55" phylum="Chordata" rank="species" species="tana">Tana, Gerima</taxonomicName>
,
<emphasis id="BC82F14EFFB7FFD8DD94F99BFC60F9A4" box="[987,998,1637,1658]" italics="true" pageId="14" pageNumber="55">c</emphasis>
.
<quantity id="490E80B9FFB7FFD8DDB7F99AFBB7F9A4" box="[1016,1073,1636,1658]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="55" unit="m" value="30.0">30 m</quantity>
off shore (
<geoCoordinate id="EBC24B9BFFB7FFD8DAF5F99AFA91F9A4" box="[1210,1303,1635,1658]" degrees="11" direction="north" minutes="35" orientation="latitude" pageId="14" pageNumber="55" precision="925" value="11.583333">11°35′N</geoCoordinate>
,
<geoCoordinate id="EBC24B9BFFB7FFD8DB69F99AFA07F9A4" box="[1318,1409,1635,1658]" degrees="37" direction="east" minutes="24" orientation="longitude" pageId="14" pageNumber="55" precision="925" value="37.4">37°24′E</geoCoordinate>
);
<typeStatus id="514D93FEFFB7FFD8DD69F97DFC11F946" box="[806,919,1667,1688]" pageId="14" pageNumber="55" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="49F656DFFFB7FFD8DD82F97DFB3AF946" authority=", AMNH" authorityName="AMNH" box="[973,1212,1667,1689]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="14" pageNumber="55" phylum="Chordata" rank="species" species="regressus">
<emphasis id="BC82F14EFFB7FFD8DD82F97DFBDCF947" box="[973,1114,1667,1689]" italics="true" pageId="14" pageNumber="55">G. regressus,</emphasis>
AMNH
</taxonomicName>
228487,
<specimenCount id="98F0E6D5FFB7FFD8DB62F97DFA0FF946" box="[1325,1417,1667,1688]" pageId="14" pageNumber="55" type="generic">two ex.</specimenCount>
,
<specimenCount id="98F0E6D5FFB7FFD8DD69F95CFCF9F969" box="[806,895,1698,1719]" pageId="14" pageNumber="55" type="generic">one ex.</specimenCount>
C &amp; S,
<quantity id="490E80B9FFB7FFD8DDA4F95FFB1FF969" box="[1003,1177,1697,1719]" metricMagnitude="-1" metricUnit="m" metricValue="1.0515" metricValueMax="1.156" metricValueMin="0.947" pageId="14" pageNumber="55" unit="mm" value="105.15" valueMax="115.6" valueMin="94.7">94.7–115.6 mm</quantity>
SL, same data as
<typeStatus id="514D93FEFFB7FFD8DD69F93EFC0CF908" box="[806,906,1728,1750]" pageId="14" pageNumber="55" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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