treatments-xml/data/06/52/43/06524372FFE0FF87E6EEA5C9B2C8357A.xml
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<document id="444399400DA9CD7C00F6FFB7A3853536" ID-ISSN="1175­5334" ID-ZooBank="C62D776F-2E8B-41B0-B296-C50782687653" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629808173969" checkinUser="felipe" docAuthor="Scherrer, Marcus V. &amp; Aguiar, Alexandre P." docDate="2012" docId="06524372FFE0FF87E6EEA5C9B2C8357A" docLanguage="en" docName="zt03469p076.pdf" docOrigin="Zootaxa 3469" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Debilos frater Scherrer &amp; Aguiar 2012, sp. nov." docType="treatment" docVersion="2" lastPageNumber="21" masterDocId="FA6B3B0AFFF3FF93E679A55EB1093464" masterDocTitle="3469" masterLastPageNumber="76" masterPageNumber="1" pageNumber="20" updateTime="1699115037135" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:namePart id="3FC5D8BB9026CD6D513DA2C6A373D915">Scherrer, Marcus V.</mods:namePart>
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<treatment id="06524372FFE0FF87E6EEA5C9B2C8357A" LSID="urn:lsid:plazi:treatment:06524372FFE0FF87E6EEA5C9B2C8357A" httpUri="http://treatment.plazi.org/id/06524372FFE0FF87E6EEA5C9B2C8357A" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<subSubSection id="C6E1A1EFFFE0FF80E6EEA5C9B31534D5" box="[151,540,151,177]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8E44F264FFE0FF80E6EEA5C9B31534D5" blockId="19.[151,540,151,210]" box="[151,540,151,177]" pageId="19" pageNumber="20">
<heading id="D50C4508FFE0FF80E6EEA5C9B31534D5" bold="true" box="[151,540,151,177]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<emphasis id="BC8F2E76FFE0FF80E6EEA5C9B31534D5" bold="true" box="[151,540,151,177]" pageId="19" pageNumber="20">
<taxonomicName id="49FB89E7FFE0FF80E6EEA5C9B0B834D5" authority="Scherrer" authorityName="Scherrer &amp; Aguiar" authorityYear="2012" box="[151,433,151,177]" class="Insecta" family="Ichneumonidae" genus="Debilos" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="frater" status="sp. nov.">
<emphasis id="BC8F2E76FFE0FF80E6EEA5C9B03434D5" bold="true" box="[151,317,151,177]" italics="true" pageId="19" pageNumber="20">Debilos frater</emphasis>
Scherrer
</taxonomicName>
,
<taxonomicNameLabel id="A7BC930DFFE0FF80E7C6A5C9B31534D5" box="[447,540,151,177]" pageId="19" pageNumber="20" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C6E1A1EFFFE0FF80E6EEA5E4B0C031AD" pageId="19" pageNumber="20" type="description">
<paragraph id="8E44F264FFE0FF80E6EEA5E4B0D534B6" blockId="19.[151,540,151,210]" box="[151,476,186,210]" pageId="19" pageNumber="20">
(
<figureCitation id="16C0EEE1FFE0FF80E6E6A5E4B1FC34B6" box="[159,245,186,210]" captionStart="FIGURES 714" captionStartId="56.[151,264,1901,1924]" captionTargetBox="[222,1364,155,1888]" captionTargetId="figure-118@56.[222,1364,155,1888]" captionTargetPageId="56" captionText="FIGURES 714. Propodeum posterior area, dorsal, ♀. 7, Debilos soror (Trentepohl), Holotype. 8, D. orthum sp. nov., Holotype. 9, D. typurum sp. nov., Holotype. 10, D. prionilix sp. nov., Holotype. 11, D. palaior sp. nov., Holotype. 12, D. trochanteratum (Szépligeti). Specimen from Brazil, RO, Candeias do Jamari, 28.I4.II.2006 (UFES). 13, D. indistinctum sp. nov., Holotype. 14, D. frater sp. nov., Holotype. Illustrations not to scale. Fig. 7 photographed by Berthil B. Longo." pageId="19" pageNumber="20">Figs 14</figureCitation>
,
<figureCitation id="16C0EEE1FFE0FF80E778A5E4B01634B6" box="[257,287,186,210]" captionStart="FIGURES 2533" captionStartId="58.[151,263,1338,1361]" captionTargetBox="[151,1435,193,1317]" captionTargetId="figure-153@58.[151,1435,193,1317]" captionTargetPageId="58" captionText="FIGURES 2533. ♀. 25, Debilos palaior sp. nov., fore wing, dorsal. Specimen from Ecuador, Morona-Santiago, Cordillera de Cutucú, 28.III.1981 (BMNH). 2628, fore and hind wings, dorsal, Holotypes. 26, D. soror (Trentepohl). 27, D. orthum sp. nov. 28, D. frater sp. nov., arrow: ramellus. 29, hind wing, dorsal, D. curtuvenatum sp. nov., Holotype. 3033, fore and hind wings, dorsal. 30, D. trochanteratum (Szépligeti). Specimen from Brazil, RO, Candeias do Jamari, 28.I4.II.2006 (UFES). 31, D. indistinctum sp. nov., Holotype. 32, D. prionilix sp. nov., Holotype. 33, D. typurum sp. nov., Holotype. Illustrations not to scale. Fig. 26 photographed by Berthil B. Longo." pageId="19" pageNumber="20">28</figureCitation>
,
<figureCitation id="16C0EEE1FFE0FF80E752A5E4B04034B6" box="[299,329,186,210]" captionStart="FIGURES 4958" captionStartId="61.[151,264,1277,1300]" captionTargetBox="[151,1435,193,1256]" captionTargetId="figure-153@61.[151,1435,193,1256]" captionTargetPageId="61" captionText="FIGURES 4958. ♀. 49, 50, propodeum, dorsal, Holotypes. 49, Debilos masipum sp. nov. 50, D. sanimum sp. nov. 5153, metapleuron apical fovea (arrow), metapleuron, frontal. 51, D. soror (Trentepohl), Holotype. 52, D. dnopheror sp. nov., Holotype. 53, D. frater sp. nov., Holotype. 5458, propodeal spiracle, frontal. 54, D. palaior sp. nov., Holotype. 55, D. dialeucor sp. nov., Holotype. 56, D. typurum sp. nov., Holotype. 57, D. indistinctum sp. nov., Holotype. 58, D. trochanteratum (Szépligeti). Specimen from Brazil, RO, Candeias do Jamari, 28.I4.II.2006 (UFES). Illustrations not to scale. Fig. 63 photographed by Berthil B. Longo." pageId="19" pageNumber="20">53</figureCitation>
,
<figureCitation id="16C0EEE1FFE0FF80E72CA5E4B07A34B6" box="[341,371,186,210]" captionStart="FIGURES 7390" captionStartId="63.[151,264,1800,1823]" captionTargetBox="[256,1339,155,1783]" captionTargetId="figure-239@63.[255,1339,155,1783]" captionTargetPageId="63" captionText="FIGURES 7390. ♀. 7384, head, frontal, Holotypes. 73, Debilos soror (Trentepohl). 74, D. dialeucor sp. nov. 75, D. dnopheror sp. nov. 76, D. orthum sp. nov. 77, D. gnampthum sp. nov. 78, D. typurum sp. nov. 79, D. prionilix sp. nov. 80, D. owinum sp. nov. 81, D. pekujum sp. nov. 82, D. crusilix sp. nov. 83, D. frater sp. nov. 84, D. indistinctum sp. nov. 85, D. trochanteratum (Szépligeti), head, frontal. Specimen from Brazil, RO, Candeias do Jamari, 28.I4.II.2006 (UFES). 86, 87, clypeus, lateral. 86, D. soror (Trentepohl). Specimen from Brazil, PA, Serra Norte, Pedreira, 1922.VI.1985 (MPEG). 87, D. palaior sp. nov., Holotype. 88, D. dnopheror sp. nov., left antenna, dorsal. Specimen from Brazil, PA, Melgaço, Floresta Nacional de Caxiuanã, 1821.ix.2003 (UFES). 89, 90, pedicel and flagellomere 1, dorsal, Holotypes. 89, D. orthum sp. nov. 90, D. gnampthum sp. nov. Illustrations not to scale. Fig. 73 photographed by Berthil B. Longo." pageId="19" pageNumber="20">83</figureCitation>
,
<figureCitation id="16C0EEE1FFE0FF80E706A5E4B09434B6" box="[383,413,186,210]" captionStart="FIGURES 96100" captionStartId="65.[151,263,1849,1872]" captionTargetBox="[198,1388,193,1828]" captionTargetId="figure-70@65.[198,1388,193,1828]" captionTargetPageId="65" captionText="FIGURES 96100. Holotypes, ♀, lateral. 96, Debilos orthum sp. nov. 97, D. gnampthum sp. nov. 98, D. frater sp. nov. 99, D. prionilix sp. nov. 100, D. crusilix sp. nov. Illustrations not to scale." pageId="19" pageNumber="20">98</figureCitation>
,
<figureCitation id="16C0EEE1FFE0FF80E7D0A5E4B0DD34B6" box="[425,468,186,210]" captionStart="FIGURES 129134" captionStartId="72.[151,264,1521,1544]" captionTargetBox="[233,1354,193,1500]" captionTargetId="figure-78@72.[233,1354,193,1500]" captionTargetPageId="72" captionText="FIGURES 129134. Diagrams with distribution records for: 129, Debilos frater sp. nov.; 130, D. crusilix sp. nov.; 131, D. prionilix sp. nov.; 132, D. indistinctum sp. nov.; 133, D. gnampthum sp. nov.; 134, D. orthum sp. nov." pageId="19" pageNumber="20">129</figureCitation>
)
</paragraph>
<paragraph id="8E44F264FFE0FF80E6EEA5A5B58835E5" blockId="19.[151,1436,251,2029]" pageId="19" pageNumber="20">
<emphasis id="BC8F2E76FFE0FF80E6EEA5A5B0213577" box="[151,296,251,276]" inLineHeading="true" pageId="19" pageNumber="20" reason="1">
<emphasis id="BC8F2E76FFE0FF80E6EEA5A5B0283570" bold="true" box="[151,289,251,276]" pageId="19" pageNumber="20">Description</emphasis>
.
</emphasis>
Female. Fore wing
<quantity id="49035F81FFE0FF80E45EA5A2B37B3577" box="[551,626,252,276]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="19" pageNumber="20" unit="mm" value="8.0">8 mm</quantity>
.
<emphasis id="BC8F2E76FFE0FF80E4FDA5A2B3C93570" box="[644,704,252,276]" italics="true" pageId="19" pageNumber="20">Head</emphasis>
(
<figureCitation id="16C0EEE1FFE0FF80E4A1A5A2B23D3570" box="[728,820,252,276]" captionStart="FIGURES 7390" captionStartId="63.[151,264,1800,1823]" captionTargetBox="[256,1339,155,1783]" captionTargetId="figure-239@63.[255,1339,155,1783]" captionTargetPageId="63" captionText="FIGURES 7390. ♀. 7384, head, frontal, Holotypes. 73, Debilos soror (Trentepohl). 74, D. dialeucor sp. nov. 75, D. dnopheror sp. nov. 76, D. orthum sp. nov. 77, D. gnampthum sp. nov. 78, D. typurum sp. nov. 79, D. prionilix sp. nov. 80, D. owinum sp. nov. 81, D. pekujum sp. nov. 82, D. crusilix sp. nov. 83, D. frater sp. nov. 84, D. indistinctum sp. nov. 85, D. trochanteratum (Szépligeti), head, frontal. Specimen from Brazil, RO, Candeias do Jamari, 28.I4.II.2006 (UFES). 86, 87, clypeus, lateral. 86, D. soror (Trentepohl). Specimen from Brazil, PA, Serra Norte, Pedreira, 1922.VI.1985 (MPEG). 87, D. palaior sp. nov., Holotype. 88, D. dnopheror sp. nov., left antenna, dorsal. Specimen from Brazil, PA, Melgaço, Floresta Nacional de Caxiuanã, 1821.ix.2003 (UFES). 89, 90, pedicel and flagellomere 1, dorsal, Holotypes. 89, D. orthum sp. nov. 90, D. gnampthum sp. nov. Illustrations not to scale. Fig. 73 photographed by Berthil B. Longo." pageId="19" pageNumber="20">Fig. 83</figureCitation>
): Clypeus smooth, sparsely punctate towards apex, moderately convex, regularly curved, CWH 1.59; MWC 0.55, MLW 2.08, MWW 0.42; malar space moderately long, MSM 1; supra-antennal area with faint, median, longitudinal carina, indistinct at middle; minimum distance from eye margin to occipital carina laterally approximately same width of mandible base.
</paragraph>
<paragraph id="8E44F264FFE0FF80E6BEA4D1B3EB37C1" blockId="19.[151,1436,251,2029]" pageId="19" pageNumber="20">
<taxonomicName id="49FB89E7FFE0FF80E6BEA4D1B03635C2" box="[199,319,399,422]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="BC8F2E76FFE0FF80E6BEA4D1B03635C2" box="[199,319,399,422]" italics="true" pageId="19" pageNumber="20">Mesosoma</emphasis>
</taxonomicName>
(
<figureCitation id="16C0EEE1FFE0FF80E72DA4D0B0A735C2" box="[340,430,398,422]" captionStart="FIGURES 96100" captionStartId="65.[151,263,1849,1872]" captionTargetBox="[198,1388,193,1828]" captionTargetId="figure-70@65.[198,1388,193,1828]" captionTargetPageId="65" captionText="FIGURES 96100. Holotypes, ♀, lateral. 96, Debilos orthum sp. nov. 97, D. gnampthum sp. nov. 98, D. frater sp. nov. 99, D. prionilix sp. nov. 100, D. crusilix sp. nov. Illustrations not to scale." pageId="19" pageNumber="20">Fig. 98</figureCitation>
). Pronotum lateral portion inconspicuously strigate to rugose, epomia faint; notaulus moderately impressed, finely corrugated, reaching about middle of mesoscutum; mesopleuron dorsal half medially faintly strigulate-rugulose, ventral half without distinct ridges or rugae, epicnemial carina restricted to ventral 0.7 of mesopleuron, posteriorly inconspicuously, densely corrugated; mesopleural groove markedly, sparsely corrugated; sternaulus entirely faintly corrugated; postpectal carina medially distinctly bent; metapleuron, subapical fovea shallow, subcircular (
<figureCitation id="16C0EEE1FFE0FF80E436A71AB3AE3638" box="[591,679,580,604]" captionStart="FIGURES 4958" captionStartId="61.[151,264,1277,1300]" captionTargetBox="[151,1435,193,1256]" captionTargetId="figure-153@61.[151,1435,193,1256]" captionTargetPageId="61" captionText="FIGURES 4958. ♀. 49, 50, propodeum, dorsal, Holotypes. 49, Debilos masipum sp. nov. 50, D. sanimum sp. nov. 5153, metapleuron apical fovea (arrow), metapleuron, frontal. 51, D. soror (Trentepohl), Holotype. 52, D. dnopheror sp. nov., Holotype. 53, D. frater sp. nov., Holotype. 5458, propodeal spiracle, frontal. 54, D. palaior sp. nov., Holotype. 55, D. dialeucor sp. nov., Holotype. 56, D. typurum sp. nov., Holotype. 57, D. indistinctum sp. nov., Holotype. 58, D. trochanteratum (Szépligeti). Specimen from Brazil, RO, Candeias do Jamari, 28.I4.II.2006 (UFES). Illustrations not to scale. Fig. 63 photographed by Berthil B. Longo." pageId="19" pageNumber="20">Fig. 53</figureCitation>
); pleural carina stout; propodeum anterior area short, PLL 0.71, anterior transverse carina distinct, complete, medially distinctly arched forwards, posterior area moderately strigate-rugose, with ridges medially slightly arched backwards (
<figureCitation id="16C0EEE1FFE0FF80E5EDA7D0B2E536C2" box="[916,1004,654,678]" captionStart="FIGURES 714" captionStartId="56.[151,264,1901,1924]" captionTargetBox="[222,1364,155,1888]" captionTargetId="figure-118@56.[222,1364,155,1888]" captionTargetPageId="56" captionText="FIGURES 714. Propodeum posterior area, dorsal, ♀. 7, Debilos soror (Trentepohl), Holotype. 8, D. orthum sp. nov., Holotype. 9, D. typurum sp. nov., Holotype. 10, D. prionilix sp. nov., Holotype. 11, D. palaior sp. nov., Holotype. 12, D. trochanteratum (Szépligeti). Specimen from Brazil, RO, Candeias do Jamari, 28.I4.II.2006 (UFES). 13, D. indistinctum sp. nov., Holotype. 14, D. frater sp. nov., Holotype. Illustrations not to scale. Fig. 7 photographed by Berthil B. Longo." pageId="19" pageNumber="20">Fig. 14</figureCitation>
), spiracle very elongate, SWL 2.50, propodeal apophyses long, thorn-shaped, AHD 2.50. Legs: Hind t4 posterior lobe about 0.5 × as long as anterior lobe. Wings (
<figureCitation id="16C0EEE1FFE0FF80E74BA789B081368B" box="[306,392,727,751]" captionStart="FIGURES 2533" captionStartId="58.[151,263,1338,1361]" captionTargetBox="[151,1435,193,1317]" captionTargetId="figure-153@58.[151,1435,193,1317]" captionTargetPageId="58" captionText="FIGURES 2533. ♀. 25, Debilos palaior sp. nov., fore wing, dorsal. Specimen from Ecuador, Morona-Santiago, Cordillera de Cutucú, 28.III.1981 (BMNH). 2628, fore and hind wings, dorsal, Holotypes. 26, D. soror (Trentepohl). 27, D. orthum sp. nov. 28, D. frater sp. nov., arrow: ramellus. 29, hind wing, dorsal, D. curtuvenatum sp. nov., Holotype. 3033, fore and hind wings, dorsal. 30, D. trochanteratum (Szépligeti). Specimen from Brazil, RO, Candeias do Jamari, 28.I4.II.2006 (UFES). 31, D. indistinctum sp. nov., Holotype. 32, D. prionilix sp. nov., Holotype. 33, D. typurum sp. nov., Holotype. Illustrations not to scale. Fig. 26 photographed by Berthil B. Longo." pageId="19" pageNumber="20">Fig. 28</figureCitation>
): Fore wing vein (Rs+M)b perfectly straight, limit with crossvein 1m-cu distinct, ramellus present; vein Rs&amp;1M uniformly slightly curved; crossvein 1cu-a posteriorly slightly convex, arising opposite vein Rs&amp;1M, angle with vein M+Cu distinctly obtuse; vein 2Cua moderately shorter than crossvein 2cu-a, FWC 0.75; areolet small, APH 0.67; vein 2Ma distinctly longer than vein 2Mb; vein 2Mb tubular; crossvein 2m-cu with bulla short, about 0.2 × as long as vein length or shorter; vein 2Rsb slightly sinuous. Hind wing vein 2-1A long, reaching about 0.8 × the distance to wing margin, HWC 2.00.
</paragraph>
<paragraph id="8E44F264FFE0FF80E6BEA6EDB40C3038" blockId="19.[151,1436,251,2029]" pageId="19" pageNumber="20">
<emphasis id="BC8F2E76FFE0FF80E6BEA6EDB03537AE" box="[199,316,947,970]" italics="true" pageId="19" pageNumber="20">Metasoma</emphasis>
(
<figureCitation id="16C0EEE1FFE0FF80E735A6ECB0A937AE" box="[332,416,946,970]" captionStart="FIGURES 96100" captionStartId="65.[151,263,1849,1872]" captionTargetBox="[198,1388,193,1828]" captionTargetId="figure-70@65.[198,1388,193,1828]" captionTargetPageId="65" captionText="FIGURES 96100. Holotypes, ♀, lateral. 96, Debilos orthum sp. nov. 97, D. gnampthum sp. nov. 98, D. frater sp. nov. 99, D. prionilix sp. nov. 100, D. crusilix sp. nov. Illustrations not to scale." pageId="19" pageNumber="20">Fig. 98</figureCitation>
). T1 petiole basal half smooth towards base, apical half, postpetiole coriarious-punctulate, except pospetiole apical parts centrally smooth, sparsely punctulate, spiracle at about basal 0.7, very conspicuously prominent, T1LW 3.04, T1WW 3.00, T1GL 0.36; T2 moderately short, apically robust, T2T1L 0.76, thyridium distinctly longer than wide, T2LW 0.94, T2WW 2.07. Ovipositor approximately straight, OST 0.47, dorsal valve with nodus absent, notch absent; ventral valve tip with inconspicuous serrations restricted to very tip.
</paragraph>
<paragraph id="8E44F264FFE0FF80E6BEA137B52431E4" blockId="19.[151,1436,251,2029]" pageId="19" pageNumber="20">
<emphasis id="BC8F2E76FFE0FF80E6BEA137B00E30E5" box="[199,263,1129,1153]" italics="true" pageId="19" pageNumber="20">Color</emphasis>
. Head black, legs and metasoma black, mesosoma dark orangish brown (127, 065, 047). Head: Flagellomere 1 dorsally white; mouth parts whitish (218, 216, 195), except mandible basal 0.25 and apex blackish (
<figureCitation id="16C0EEE1FFE0FF80E6E6A1EFB1FD30AD" box="[159,244,1201,1225]" captionStart="FIGURES 4148" captionStartId="60.[151,264,1912,1935]" captionTargetBox="[210,1377,153,1890]" captionTargetId="figure-130@60.[209,1377,153,1890]" captionTargetPageId="60" captionText="FIGURES 4148. Holotypes, ♀. 4144, mesoscutum, dorsal. 41, Debilos okarum sp. nov. 42, D. typurum sp. nov. 43, D. palaior sp. nov. 44, D. dnopheror sp. nov. 45, 46, median portion of postpectal carina (arrow), mesothoracic venter, frontal. 45, D. masipum sp. nov. 46, D. sanimum sp. nov. 47, 48, epicnemial carina, mesopleuron, frontal, arrow: carina dorsal end. 47, D. okum sp. nov. 48, D. owinum sp. nov. Illustrations not to scale." pageId="19" pageNumber="20">Fig. 45</figureCitation>
).
<taxonomicName id="49FB89E7FFE0FF80E771A1ECB0FE30AD" authority=": Ventrally" authorityName="Ventrally" box="[264,503,1201,1225]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">Mesosoma: Ventrally</taxonomicName>
darkest; propodeal apophyses white; propleuron darker than pronotum. Legs: Fore leg entirely and mid tibia and tarsus lightest; fore coxa apical margin, fore femur towards apex, fore and mid tibiae anteriorly and posteriorly, femora basal margin slightly and hind tibia basal 0.2 pale yellow (241, 224, 144) to whitish; hind t1 apical half, t2 and 3 entirely, and t4 basal half white. Fore wing with one inconspicuous dark spot, around vein 2m-cu. Metasoma: T1 apical 0.1, and T2 and 3 margin lightest; T68 apical margin centrally white; ovipositor sheath dark brown (092, 054, 024), except basal 0.65 ventrally whitish.
</paragraph>
<paragraph id="8E44F264FFE0FF80E6BEA0D3B08131C1" blockId="19.[151,1436,251,2029]" box="[199,392,1421,1445]" pageId="19" pageNumber="20">
<emphasis id="BC8F2E76FFE0FF80E6BEA0D3B00D31C1" bold="true" box="[199,260,1421,1445]" pageId="19" pageNumber="20">Male</emphasis>
. Unknown.
</paragraph>
<paragraph id="8E44F264FFE0FF80E6BEA0EFB0C031AD" blockId="19.[151,1436,251,2029]" box="[199,457,1457,1482]" pageId="19" pageNumber="20">
<emphasis id="BC8F2E76FFE0FF80E6BEA0EFB04531AD" box="[199,332,1457,1482]" inLineHeading="true" pageId="19" pageNumber="20" reason="1">
<emphasis id="BC8F2E76FFE0FF80E6BEA0EFB04C31AE" bold="true" box="[199,325,1457,1482]" pageId="19" pageNumber="20">Variability</emphasis>
.
</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C6E1A1EFFFE0FF80E6BEA08BB578331B" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8E44F264FFE0FF80E6BEA08BB0FC32C0" blockId="19.[151,1436,251,2029]" pageId="19" pageNumber="20">
<emphasis id="BC8F2E76FFE0FF80E6BEA08BB046318A" box="[199,335,1493,1518]" inLineHeading="true" pageId="19" pageNumber="20" reason="1">
<emphasis id="BC8F2E76FFE0FF80E6BEA08BB041318A" bold="true" box="[199,328,1493,1518]" pageId="19" pageNumber="20">Comments</emphasis>
.
</emphasis>
As mentioned in “Species Delimitation” of the genus, this species cannot be placed in any of the proposed species group because of its high morphological dissimilarities with all other species, as well as the presence of some unique features. According to its general body shape and size the single available specimen may resemble species of the
<taxonomicName id="49FB89E7FFE0FF80E7DBA31AB0F6323F" baseAuthorityName="Trentepohl" baseAuthorityYear="1829" box="[418,511,1604,1627]" class="Insecta" family="Ichneumonidae" genus="Debilos" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="soror">
<emphasis id="BC8F2E76FFE0FF80E7DBA31AB0BF323F" box="[418,438,1604,1627]" italics="true" pageId="19" pageNumber="20">D</emphasis>
.
<emphasis id="BC8F2E76FFE0FF80E7BDA31AB0F6323F" box="[452,511,1604,1627]" italics="true" pageId="19" pageNumber="20">soror</emphasis>
</taxonomicName>
group, but differs from those by having the mesosoma orangish brown (
<emphasis id="BC8F2E76FFE0FF80E353A31AB448323F" box="[1322,1345,1604,1627]" italics="true" pageId="19" pageNumber="20">vs</emphasis>
. black), propodeal apophyses very long, thorn-shaped (
<emphasis id="BC8F2E76FFE0FF80E4CBA337B3C032E4" box="[690,713,1641,1664]" italics="true" pageId="19" pageNumber="20">vs</emphasis>
. short, conical apophyses) and ovipositor dorsal valve without nodus (
<emphasis id="BC8F2E76FFE0FF80E693A3D3B00832C0" box="[234,257,1677,1700]" italics="true" pageId="19" pageNumber="20">vs</emphasis>
. with nodus distinct).
</paragraph>
<paragraph id="8E44F264FFE0FF80E6BEA3EFB578331B" blockId="19.[151,1436,251,2029]" pageId="19" pageNumber="20">
The morphological characterization under the diagnosis of the
<taxonomicName id="49FB89E7FFE0FF80E5C4A3ECB54A32AD" authorityName="Scherrer &amp; Aguiar" authorityYear="2012" box="[957,1091,1714,1737]" class="Insecta" family="Ichneumonidae" genus="Debilos" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="typurum">
<emphasis id="BC8F2E76FFE0FF80E5C4A3ECB2D832AD" box="[957,977,1714,1737]" italics="true" pageId="19" pageNumber="20">D</emphasis>
.
<emphasis id="BC8F2E76FFE0FF80E59FA3ECB54A32AD" box="[998,1091,1714,1737]" italics="true" pageId="19" pageNumber="20">typurum</emphasis>
</taxonomicName>
species group (see “Species Delimitation”) may fit somewhat well for
<taxonomicName id="49FB89E7FFE0FF80E409A389B3DB328A" authorityName="Scherrer &amp; Aguiar" authorityYear="2012" box="[624,722,1750,1774]" class="Insecta" family="Ichneumonidae" genus="Debilos" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="BC8F2E76FFE0FF80E409A389B38D328A" box="[624,644,1751,1774]" italics="true" pageId="19" pageNumber="20">D</emphasis>
.
<emphasis id="BC8F2E76FFE0FF80E4EBA388B3DB328A" box="[658,722,1750,1774]" italics="true" pageId="19" pageNumber="20">frater</emphasis>
</taxonomicName>
, but it should not be associated with that species group because it is largely unlike those species, mostly by having a quite large body size, with fore wing length about 8.00 mm (
<emphasis id="BC8F2E76FFE0FF80E6D9A27EB1BE3353" box="[160,183,1824,1847]" italics="true" pageId="19" pageNumber="20">vs</emphasis>
. medium size, fore wing length about
<quantity id="49035F81FFE0FF80E40BA241B2123353" box="[626,795,1823,1847]" metricMagnitude="-3" metricUnit="m" metricValue="5.21" metricValueMax="6.32" metricValueMin="4.1" pageId="19" pageNumber="20" unit="mm" value="5.21" valueMax="6.32" valueMin="4.1">4.106.32 mm</quantity>
), and metasoma mostly blackish, without white stripe at T2 apical parts (
<emphasis id="BC8F2E76FFE0FF80E735A21AB06A333F" box="[332,355,1860,1883]" italics="true" pageId="19" pageNumber="20">vs</emphasis>
. mostly dark brownish, T2 with apical white stripe). It is also remarkable that this individual has a developed ramellus on the fore wing, whereas it is absent on all other
<taxonomicName id="49FB89E7FFE0FF80E5C3A236B578331B" box="[954,1137,1895,1920]" class="Insecta" family="Ichneumonidae" genus="Debilos" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BC8F2E76FFE0FF80E5C3A236B51833E4" box="[954,1041,1896,1920]" italics="true" pageId="19" pageNumber="20">Debilos</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C6E1A1EFFFE0FF80E6BEA2D3B0353389" pageId="19" pageNumber="20" type="etymology">
<paragraph id="8E44F264FFE0FF80E6BEA2D3B0353389" blockId="19.[151,1436,251,2029]" pageId="19" pageNumber="20">
<emphasis id="BC8F2E76FFE0FF80E6BEA2D3B04433C0" box="[199,333,1933,1957]" inLineHeading="true" pageId="19" pageNumber="20" reason="1">
<emphasis id="BC8F2E76FFE0FF80E6BEA2D3B04F33C1" bold="true" box="[199,326,1933,1957]" pageId="19" pageNumber="20">Etymology</emphasis>
.
</emphasis>
From the Greek word
<taxonomicName id="49FB89E7FFE0FF80E42FA2D3B39C33C1" authorityName="Scherrer &amp; Aguiar" authorityYear="2012" box="[598,661,1933,1957]" class="Insecta" family="Ichneumonidae" genus="Debilos" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="frater">
<emphasis id="BC8F2E76FFE0FF80E42FA2D3B39C33C1" box="[598,661,1933,1957]" italics="true" pageId="19" pageNumber="20">frater</emphasis>
</taxonomicName>
, meaning “brother,” in reference to the color pattern of this species (orangish thorax and blackish legs) that seems like the inverse of the color pattern of
<taxonomicName id="49FB89E7FFE0FF80E22BA2ECB5A633AD" baseAuthorityName="Trentepohl" baseAuthorityYear="1829" box="[1106,1199,1970,1993]" class="Insecta" family="Ichneumonidae" genus="Debilos" kingdom="Animalia" order="Hymenoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="soror">
<emphasis id="BC8F2E76FFE0FF80E22BA2ECB56F33AD" box="[1106,1126,1970,1993]" italics="true" pageId="19" pageNumber="20">D</emphasis>
.
<emphasis id="BC8F2E76FFE0FF80E20DA2ECB5A633AD" box="[1140,1199,1970,1993]" italics="true" pageId="19" pageNumber="20">soror</emphasis>
</taxonomicName>
(blackish thorax and orangish legs).
</paragraph>
</subSubSection>
<subSubSection id="C6E1A1EFFFE7FF87E6BEA5C9B2C8357A" pageId="20" pageNumber="21" type="distribution">
<paragraph id="8E44F264FFE7FF87E6BEA5C9B2C8357A" blockId="20.[151,1436,151,286]" pageId="20" pageNumber="21">
<emphasis id="BC8F2E76FFE7FF87E6BEA5C9B06934CB" box="[199,352,151,176]" inLineHeading="true" pageId="20" pageNumber="21" reason="1">
<emphasis id="BC8F2E76FFE7FF87E6BEA5C9B05034D4" bold="true" box="[199,345,151,176]" pageId="20" pageNumber="21">Distribution</emphasis>
.
</emphasis>
Peru. Found at the locality of Tingo María,
<collectingRegion id="4C3F3C86FFE7FF87E537A5C9B2BA34CB" box="[846,947,151,175]" country="Peru" name="Huanuco" pageId="20" pageNumber="21">Huánuco</collectingRegion>
(around
<geoCoordinate id="EBCF94A3FFE7FF87E260A5C9B57C34CB" box="[1049,1141,151,175]" degrees="09" direction="south" minutes="17" orientation="latitude" pageId="20" pageNumber="21" precision="925" value="-9.283334">09°17S</geoCoordinate>
<geoCoordinate id="EBCF94A3FFE7FF87E202A5C9B5DC34CB" box="[1147,1237,151,175]" degrees="76" direction="west" minutes="0" orientation="longitude" pageId="20" pageNumber="21" precision="925" value="-76.0">76°0W</geoCoordinate>
) (
<figureCitation id="16C0EEE1FFE7FF87E294A5C9B45934CB" box="[1261,1360,151,175]" captionStart="FIGURES 129134" captionStartId="72.[151,264,1521,1544]" captionTargetBox="[233,1354,193,1500]" captionTargetId="figure-78@72.[233,1354,193,1500]" captionTargetPageId="72" captionText="FIGURES 129134. Diagrams with distribution records for: 129, Debilos frater sp. nov.; 130, D. crusilix sp. nov.; 131, D. prionilix sp. nov.; 132, D. indistinctum sp. nov.; 133, D. gnampthum sp. nov.; 134, D. orthum sp. nov." pageId="20" pageNumber="21">Fig. 129</figureCitation>
).
<emphasis id="BC8F2E76FFE7FF87E6BEA5E2B0BB34B1" bold="true" box="[199,434,188,213]" pageId="20" pageNumber="21">Material Examined</emphasis>
.
<typeStatus id="51404CC6FFE7FF87E7BDA5E2B32534B0" box="[452,556,188,212]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
<specimenCount id="98FD39EDFFE7FF87E44FA5E3B34234B0" box="[566,587,189,212]" pageId="20" pageNumber="21" type="female"></specimenCount>
from
<collectingCountry id="F6ECB2F4FFE7FF87E4EFA5E3B3E934B0" box="[662,736,189,212]" name="Peru" pageId="20" pageNumber="21">PERU</collectingCountry>
,
<collectingRegion id="4C3F3C86FFE7FF87E496A5E2B25E34B0" box="[751,855,188,212]" country="Peru" name="Huanuco" pageId="20" pageNumber="21">Huánuco</collectingRegion>
, Tingo María,
<date id="FA45D4A4FFE7FF87E27CA5E2B57B34B0" box="[1029,1138,188,212]" pageId="20" pageNumber="21" value="1947-01-12">12.I.1947</date>
, Alt.
<quantity id="49035F81FFE7FF87E2CEA5E2B41A34B0" box="[1207,1299,188,212]" metricMagnitude="2" metricUnit="m" metricValue="6.7056" pageId="20" pageNumber="21" unit="ft" value="2200.0">2200 ft.</quantity>
, JCPallister Coll., Donor Frank Johnson (AMNH). Mounted with a pin on its mesoscutum; antennae lacking flagellomeres 3 to tip; left hind tarsus lacking; left ovipositor sheath posterior half missing.
</paragraph>
</subSubSection>
</treatment>
</document>