195 lines
17 KiB
XML
195 lines
17 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.53.4955" ID-PMC="PMC4547023" ID-Pensoft-Pub="1314-2003-53-39" ID-Pensoft-UUID="824BD167D1535A15FC27FF98FFD3A404" ID-PubMed="26312040" ID-Zenodo-Dep="576284" ModsDocID="1314-2003-53-39" checkinTime="1451251414861" checkinUser="pensoft" docAuthor="Luong, Thien Tam, Hovenkamp, Peter H. & Sosef, Marc S. M." docDate="2015" docId="063489BAC2BA5D4DB572233F5519035F" docLanguage="en" docName="PhytoKeys 53: 39-71" docOrigin="PhytoKeys 53" docPubDate="2015-07-21" docSource="http://dx.doi.org/10.3897/phytokeys.53.4955" docTitle="Orthiopteris campylura (Kunze) Copel., Fern Fl. Philip. 1: 87. 1958." docType="treatment" docVersion="3" id="824BD167D1535A15FC27FF98FFD3A404" lastPageNumber="47" masterDocId="824BD167D1535A15FC27FF98FFD3A404" masterDocTitle="Revision of the fern genus Orthiopteris (Saccolomataceae) in Malesia and adjacent regions" masterLastPageNumber="71" masterPageNumber="39" pageNumber="46" updateTime="1668141422086" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the fern genus Orthiopteris (Saccolomataceae) in Malesia and adjacent regions</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Luong, Thien Tam</mods:namePart>
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<mods:affiliation>Department of Ecology - Evolutionary Biology, Viet Nam National University Ho Chi Minh city (VNUHCM) - University of Science. 227 Nguyen Van Cu, Ho Chi Minh City, Vietnam & Naturalis Biodiversity Center, section Botany. PO Box 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hovenkamp, Peter H.</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, section Botany. PO Box 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sosef, Marc S. M.</mods:namePart>
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<mods:affiliation>Botanic Garden Meise, Nieuwelaan 38, 1860 Meise, Belgium</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>PhytoKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2015-07-21</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>53</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>39</mods:start>
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<mods:end>71</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.53.4955</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.53.4955</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2003-53-39</mods:identifier>
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<mods:identifier type="Pensoft-UUID">824BD167D1535A15FC27FF98FFD3A404</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576284</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="182227825" LSID="urn:lsid:plazi:treatment:063489BAC2BA5D4DB572233F5519035F" httpUri="http://treatment.plazi.org/id/063489BAC2BA5D4DB572233F5519035F" lastPageId="8" lastPageNumber="47" pageId="7" pageNumber="46">
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<subSubSection pageId="7" pageNumber="46" type="nomenclature">
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<paragraph pageId="7" pageNumber="46">
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<pageBreakToken pageId="7" pageNumber="46" start="start">1</pageBreakToken>
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.
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<taxonomicName LSID="063489BA-C2BA-5D4D-B572-233F5519035F" authority="(Kunze) Copel., Fern Fl. Philip. 1: 87. 1958." authorityName="(Kunze) Copel., Fern Fl. Philip. 1: 87. 1958." class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris campylura" order="Polypodiales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="species" species="campylura">Orthiopteris campylura (Kunze) Copel., Fern Fl. Philip. 1: 87. 1958.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="46" type="reference_group">
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<paragraph pageId="7" pageNumber="46">
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<taxonomicName authorityName="Hook" authorityYear="1846" class="Polypodiopsida" family="Saccolomataceae" genus="Davallia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Davallia inaequalis subsp. var. var. minor" order="Polypodiales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="variety" species="inaequalis" subSpecies="var." variety="minor">Davallia inaequalis var. minor</taxonomicName>
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Hook., Sp. Fil. 1: 180, pl. 58A. 1846. Lectotype (selected here): PHILIPPINES. Luzon,
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<emphasis italics="true" pageId="7" pageNumber="46">Cuming 119</emphasis>
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(K, on 2 sheets: K000794851*, K000794853*, isolectotypes: K: K000794864*, MICH: MICH1259585*).
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</paragraph>
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<paragraph pageId="7" pageNumber="46">
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<taxonomicName authorityName="Kunze" authorityYear="1850" class="Polypodiopsida" family="Saccolomataceae" genus="Davallia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Davallia campylura" order="Polypodiales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="species" species="campylura">Davallia campylura</taxonomicName>
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Kunze, Bot. Zeitung (Berlin) 8: 132. 1850. Lectotype (selected here): PHILIPPINES. Luzon,
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<emphasis italics="true" pageId="7" pageNumber="46">Cuming 119</emphasis>
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(herb. Kunze, not traced, isolectotypes: K: K000794851*, K000794853*, K000794864*, MICH: MICH1259585*, see below).
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</paragraph>
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<paragraph pageId="7" pageNumber="46">
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<taxonomicName authorityName="Mett" authorityYear="1861" baseAuthorityName="Kunze" class="Polypodiopsida" family="Saccolomataceae" genus="Saccoloma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Saccoloma campylurum" order="Polypodiales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="species" species="campylurum">Saccoloma campylurum</taxonomicName>
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(Kunze) Mett. Ann. Sci. Nat. Bot.
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<normalizedToken originalValue="sér">ser</normalizedToken>
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. 4, 15: 80. 1861. Type. Based on
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<taxonomicName authorityName="Kunze" authorityYear="1850" class="Polypodiopsida" family="Saccolomataceae" genus="Davallia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Davallia campylura" order="Polypodiales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="species" species="campylura">
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<emphasis italics="true" pageId="7" pageNumber="46">Davallia campylura</emphasis>
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</taxonomicName>
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Kunze
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</paragraph>
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<paragraph pageId="7" pageNumber="46">
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Saccoloma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Saccoloma moluccanum subsp. var. var. stenolobum" order="Polypodiales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="variety" species="moluccanum" subSpecies="var." variety="stenolobum">Saccoloma moluccanum (Blume) Mett. in Kuhn var. stenolobum</taxonomicName>
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Christ, Bull. Herb. Boissier, ser. 2 6: 1005. 1906. Type. PHILIPPINES. Mabacal/Rizal, III, 1906, Loher s.n. (holo: P, on 4 sheets: P01566098*, P01566099*, P01566100*, P01566101*).
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</paragraph>
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<paragraph pageId="7" pageNumber="46">
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<taxonomicName authorityName="C. Chr" authorityYear="1929" baseAuthorityName="Hook." class="Polypodiopsida" family="Saccolomataceae" genus="Saccoloma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Saccoloma minus" order="Polypodiales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="species" species="minus">Saccoloma minus</taxonomicName>
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(Hook.) C.Chr., Gard. Bull. Straits Settlem. 4: 399. 1929. Type. Based on
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Saccoloma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="subsp. var. var. minor" order="Polypodiales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="variety" species="minus" subSpecies="var." variety="minor">var. minor</taxonomicName>
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Hook.
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</paragraph>
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<paragraph pageId="7" pageNumber="46">
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<taxonomicName authorityName="C. Chr" authorityYear="1934" baseAuthorityName="Hook." class="Polypodiopsida" family="Saccolomataceae" genus="Ithycaulon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Ithycaulon minus" order="Polypodiales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="species" species="minus">Ithycaulon minus</taxonomicName>
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(Hook.) C.Chr., Index. Fil. Suppl. 3: 116. 1934. Type. Based on
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Ithycaulon" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="subsp. var. var. minor" order="Polypodiales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="variety" species="minus" subSpecies="var." variety="minor">var. minor</taxonomicName>
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Hook.
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</paragraph>
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<paragraph pageId="7" pageNumber="46">
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<taxonomicName authorityName="Copel" authorityYear="1947" baseAuthorityName="Hook." class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris minor" order="Polypodiales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="species" species="minor">Orthiopteris minor</taxonomicName>
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(Hook.) Copel., Gen. Fil.: 50. 1947. Type. Based on
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="subsp. var. var. minor" order="Polypodiales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="variety" species="minor" subSpecies="var." variety="minor">var. minor</taxonomicName>
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Hook.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="7" pageNumber="46" type="type">
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<paragraph pageId="7" pageNumber="46">Type.</paragraph>
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<paragraph pageId="7" pageNumber="46">
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PHILIPPINES. Luzon,
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<emphasis italics="true" pageId="7" pageNumber="46">Cuming 119</emphasis>
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</paragraph>
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<paragraph pageId="7" pageNumber="46">(K000794851 [http://specimens.kew.org/herbarium/K000794851],</paragraph>
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<paragraph pageId="7" pageNumber="46">K000794853 [http://specimens.kew.org/herbarium/K000794853],</paragraph>
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<paragraph pageId="7" pageNumber="46">K000794864 [http://specimens.kew.org/herbarium/K000794864];</paragraph>
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<paragraph pageId="7" pageNumber="46">MICH, 1259585 [http://quod.lib.umich.edu/h/herb2ic/x-mich1259585/mich1259585.tif]).</paragraph>
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<paragraph pageId="7" pageNumber="46">
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non
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<taxonomicName authorityName="Mett" authorityYear="1861" baseAuthorityName="Hew." class="Polypodiopsida" family="Saccolomataceae" genus="Saccoloma" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Saccoloma alatum" order="Polypodiales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="species" species="alatum">
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<emphasis italics="true" pageId="7" pageNumber="46">Saccoloma alatum</emphasis>
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</taxonomicName>
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(Heward) (J. Sm.) Mett., Ann. Sci. Nat., Bot.
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<normalizedToken originalValue="sér">ser</normalizedToken>
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. 4, 15: 80. 1861, nec
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Microlepia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Microlepia alata" order="Polypodiales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="species" species="alata">
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<emphasis italics="true" pageId="7" pageNumber="46">Microlepia alata</emphasis>
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</taxonomicName>
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J. Sm., J. Bot. (Hooker) 3: 416. 1841, which is based on
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<taxonomicName class="Polypodiopsida" family="Dennstaedtiaceae" genus="Davallia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Davallia alata" order="Polypodiales" pageId="7" pageNumber="46" phylum="Tracheophyta" rank="species" species="alata">
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<emphasis italics="true" pageId="7" pageNumber="46">Davallia alata</emphasis>
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</taxonomicName>
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Heward, Mag. Nat. Hist. & J. Zool. II. ser. 2, 2: 465. 1838, nom. illeg, non
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<bibRefCitation author="Blume, CL" journalOrPublisher="J. W. van Leeuwen, Leiden" pageId="23" pageNumber="62" refId="B4" refString="Blume, CL, 1828. Enumeratio plantarum Javae et insularum adjacentium : minus cognitarum vel novarum ex herbariis Reinwardtii, Kuhlii, Hasseltii et Blumii curavit Carol. Ludov. Blume. J.W. van Leeuwen, Leiden" title="Enumeratio plantarum Javae et insularum adjacentium: minus cognitarum vel novarum ex herbariis Reinwardtii, Kuhlii, Hasseltii et Blumii curavit Carol. Ludov. Blume" year="1828">Blume (1828)</bibRefCitation>
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.
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</paragraph>
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<paragraph pageId="7" pageNumber="46">
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Type. Jamaica,
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<emphasis italics="true" pageId="7" pageNumber="46">Heward</emphasis>
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. John
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<bibRefCitation author="Smith, J" journalOrPublisher="Enumeration Filicum Philippinarum. Journal of Botany, (Being a Second Series of the Botanical Miscellany), Containing Figures and Descriptions London" pageId="25" pageNumber="64" pagination="393 - 422" refId="B51" refString="Smith, J, 1841. . Enumeration Filicum Philippinarum. Journal of Botany, (Being a Second Series of the Botanical Miscellany), Containing Figures and Descriptions London III: 393 - 422" volume="III" year="1841">Smith (1841)</bibRefCitation>
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applied this name to the Asiatic specimen (
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<emphasis italics="true" pageId="7" pageNumber="46">Cuming 119</emphasis>
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, Luzon, Philippines) but as
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<normalizedToken originalValue="Smith’s">Smith's</normalizedToken>
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the name is validated by
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<normalizedToken originalValue="Heward’s">Heward's</normalizedToken>
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description it is typified by the Heward specimen from Jamaica.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="8" lastPageNumber="47" pageId="7" pageNumber="46" type="description">
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<paragraph pageId="7" pageNumber="46">Description.</paragraph>
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<paragraph lastPageId="8" lastPageNumber="47" pageId="7" pageNumber="46">
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Rhizome erect, rising at least 20 cm above ground, diameter 1.5 cm. Rhizome scales pseudopeltate to peltate, 4-7
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<normalizedToken originalValue="×">x</normalizedToken>
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1.0-1.7 mm, narrowly lanceolate, attenuate toward apex. Fronds 100-130
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<normalizedToken originalValue="×">x</normalizedToken>
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40-50 cm; stipes slender, 37-40 cm long, 0.5 -1 cm across (at base), dark brown; lamina deltoid, tripinnate, 70-100
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<normalizedToken originalValue="×">x</normalizedToken>
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40-50 cm, rather thick herbaceous, rigid, brown-green when dry, scaly; pinnae at 40-45° to rachis, largest at base, slightly overlapping, stalked 1 cm, including stalk up to 23-25
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<normalizedToken originalValue="×">x</normalizedToken>
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9-10 cm, lanceolate, first basiscopic pinnules of lowest pinnae enlarged; ultimate
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<pageBreakToken pageId="8" pageNumber="47" start="start">segments</pageBreakToken>
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0.9-2.0
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<normalizedToken originalValue="×">x</normalizedToken>
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0.3-0.6 cm, sessile or very short-stalked, oblong trapeziform, apex obtuse to acute, margin with weak cartilaginous border; distance from vein to base of sinuses ca. 1.0 mm; crenate-dentate, shallowly to deeply incised, lobes obtuse to acute, veins in lobes 3-15 forks, light to dark brown, not strongly contrasting with lamina, percurrent. Scales on rachis mostly present at base of pinnae or pinnules, irregularly triangular, basifix, 0.7-0.9
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<normalizedToken originalValue="×">x</normalizedToken>
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0.3-0.4 mm.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="47" type="distribution">
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<paragraph pageId="8" pageNumber="47">Distribution.</paragraph>
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<paragraph pageId="8" pageNumber="47">
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Sumatra, Peninsular Malaysia (Pahang, Perak, Selangor), Java, Borneo (Sabah, Kalimantan Timur), Philippines (Leyte, Luzon, Mindanao, Negros), Moluccas (Seram), New Guinea, Vanuatu (Aneityum), Fiji (Vanua Levu, Taveuni) (Figure
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Distribution of Orthiopteris campylura and its varieties." figureDoi="10.3897/phytokeys.53.4955.figure8" httpUri="https://binary.pensoft.net/fig/47610" pageId="8" pageNumber="47">8</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="47" type="ecology">
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<paragraph pageId="8" pageNumber="47">Ecology.</paragraph>
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<paragraph pageId="8" pageNumber="47">Terrestrial, mostly in shaded moist habitats, river banks, moist slopes, ravines etc., in primary or disturbed forest, at 0-2000 m altitude.</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="47" type="discussion">
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<paragraph pageId="8" pageNumber="47">Discussion.</paragraph>
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<paragraph pageId="8" pageNumber="47">
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<taxonomicName class="Polypodiopsida" family="Saccolomataceae" genus="Orthiopteris" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Orthiopteris campylura" order="Polypodiales" pageId="8" pageNumber="47" phylum="Tracheophyta" rank="species" species="campylura">
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<emphasis italics="true" pageId="8" pageNumber="47">Orthiopteris campylura</emphasis>
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</taxonomicName>
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is the most variable species within the genus. It is widely distributed from Western Malesia to Papua New Guinea and the Pacific, existing as different entities in each sub-geographical region, which in many cases have been recognized as individual species. However, we have regularly observed samples that were intermediate between the forms, and accordingly treat them here as varieties rather than as subspecies. Characters of the sori provide the best criteria to distinguish these varieties.
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</paragraph>
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<paragraph pageId="8" pageNumber="47">There are no data that suggest that any of the varieties is distinguished by a different ecology.</paragraph>
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</subSubSection>
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</treatment>
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</document> |