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<document ID-DOI="10.11646/zootaxa.5256.5.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7758872" ID-ZooBank="E5B9708E-D626-4923-825A-A9FD59AE4024" approvalRequired="3" approvalRequired_for_textStreams="2" approvalRequired_for_treatments="1" checkinTime="1679468565563" checkinUser="plazi" docAuthor="Matalin, Andrey V. &amp; Wiesner, Jürgen" docDate="2023" docId="050B87D78D472466E1B81A80FB567460" docLanguage="en" docName="zootaxa.5256.5.1.pdf" docOrigin="Zootaxa 5256 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.5.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Therates pseudovitalisi subsp. tibetana Matalin et Wiesner 2023, ssp. n." docType="treatment" docVersion="2" lastPageNumber="409" masterDocId="F932FFAF8D40246EE12F1976FFC07601" masterDocTitle="Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae)" masterLastPageNumber="433" masterPageNumber="401" pageNumber="408" updateTime="1679469078842" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title>Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae)</mods:title>
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<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Matalin, Andrey V.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-7790-8709</mods:nameIdentifier>
<mods:affiliation>Moscow State Pedagogical University, Department of Zoology &amp; Ecology, Kibalchicha str. 6, build. 3, Moscow 129164, Russia. andrei-matalin @ yandex. ru; https: // orcid. org / 0000 - 0002 - 7790 - 8709 &amp; Pirogov National Research Medical University, Department of Biology, Ostrovitianova str. 1, Moscow 117997, Russia.</mods:affiliation>
<mods:nameIdentifier type="email">andrei-matalin@yandex.ru</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wiesner, Jürgen</mods:namePart>
<mods:affiliation>Dresdener Ring 11, D- 38444 Wolfsburg, Germany.</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-03-22</mods:number>
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<subSubSection box="[151,871,1014,1041]" pageId="7" pageNumber="408" type="nomenclature">
<paragraph blockId="7.[151,871,1014,1077]" box="[151,871,1014,1041]" pageId="7" pageNumber="408">
<heading bold="true" box="[151,871,1014,1041]" fontSize="11" level="1" pageId="7" pageNumber="408" reason="1">
<emphasis bold="true" box="[151,871,1014,1041]" pageId="7" pageNumber="408">
<taxonomicName authority="Matalin et Wiesner" authorityName="Matalin et Wiesner" authorityYear="2023" box="[151,781,1014,1041]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="subSpecies" species="pseudovitalisi" status="ssp. n." subSpecies="tibetana">
<emphasis bold="true" box="[151,534,1014,1041]" italics="true" pageId="7" pageNumber="408">Therates pseudovitalisi tibetana</emphasis>
Matalin et Wiesner
</taxonomicName>
,
<taxonomicNameLabel box="[794,871,1015,1041]" pageId="7" pageNumber="408" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,549,1050,1077]" pageId="7" pageNumber="408" type="description">
<paragraph blockId="7.[151,871,1014,1077]" box="[151,549,1050,1077]" pageId="7" pageNumber="408">
<figureCitation box="[151,235,1051,1077]" captionStart="FIGURES 114" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 114. Therates spp., habitus: 16—T. fruhstorferi fruhstorferi (13—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 913— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 314—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="7" pageNumber="408">Figs 11</figureCitation>
,
<figureCitation box="[247,319,1051,1077]" captionStart="FIGURES 2949" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 2949. Therates spp., left antennomeres 611: 2932—T. fruhstorferi fruhstorferi (29, 30 Vietnam, Na Hang; 31, 32 Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="7" pageNumber="408">38, 39</figureCitation>
,
<figureCitation box="[331,403,1051,1077]" captionStart="FIGURES 5864" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 5864. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="7" pageNumber="408">60, 61</figureCitation>
,
<figureCitation box="[415,487,1050,1077]" captionStart="FIGURES 8198" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 8198. Therates spp., left elytron: 8187—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 9193, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="7" pageNumber="408">94, 95</figureCitation>
,
<figureCitation box="[499,544,1050,1077]" captionStart="FIGURES 115135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115135. Therates spp., aedeagus: 115125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="7" pageNumber="408">135</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[151,1066,1119,1143]" pageId="7" pageNumber="408" type="reference_group">
<paragraph blockId="7.[151,1066,1119,1143]" box="[151,1066,1119,1143]" pageId="7" pageNumber="408">
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<taxonomicName ID-CoL="8MKSW" authority="W. Horn, 1913" authorityName="W. Horn" authorityYear="1913" box="[151,592,1119,1143]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="vitalisi">
<emphasis box="[151,438,1119,1143]" italics="true" pageId="7" pageNumber="408">Therates fruhstorferi vitalisi</emphasis>
W.
<bibRefCitation author="Horn, W." box="[478,592,1119,1143]" pageId="7" pageNumber="408" pagination="362 - 366" refId="ref25772" refString="Horn, W. (1913) Materiaux pour servir a l'etude de la faune entomologique de l'Indochine. Cicindelinae. Annales de la Societe Entomologique de Belgique, 57, 362 - 366. https: // doi. org / 10.5962 / bhl. part. 4597" type="journal article" year="1913">Horn, 1913</bibRefCitation>
</taxonomicName>
<treatmentCitation author="Sawada, H. &amp; Wiesner, J." box="[618,919,1119,1143]" page="463" pageId="7" pageNumber="408" year="2006">
<bibRefCitation author="Sawada, H. &amp; Wiesner, J." box="[618,919,1119,1143]" pageId="7" pageNumber="408" pagination="462 - 464" refId="ref26874" refString="Sawada, H. &amp; Wiesner, J. (2006) New records of tiger beetles from Xizang (China) (Coleoptera: Cicindelidae). Lambillionea, 106 (3), 462 - 464." type="journal article" year="2006">Sawada &amp;Wiesner 2006: 463</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation author="Wu, X. - Q." box="[929,1062,1119,1143]" page="31" pageId="7" pageNumber="408" year="2011">
<bibRefCitation author="Wu, X. - Q." box="[929,1062,1119,1143]" pageId="7" pageNumber="408" pagination="21 - 43" refId="ref27836" refString="Wu, X. - Q. (2011) A revised list of the tiger beetles of China (Coleoptera: Cicindelidae). Cicindela, 43 (2), 21 - 43." type="journal article" year="2011">Wu 2011: 31</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="408" type="materials_examined">
<paragraph blockId="7.[151,1437,1189,2007]" pageId="7" pageNumber="408">
<emphasis bold="true" box="[151,326,1189,1215]" pageId="7" pageNumber="408">Type material.</emphasis>
<materialsCitation collectingDate="2004-07-28" collectionCode="JW" country="China" elevation="2250" location="County" municipality="Cha Yu" pageId="7" pageNumber="408" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<typeStatus box="[333,451,1189,1215]" pageId="7" pageNumber="408">HOLOTYPE</typeStatus>
, 1Ô—
<collectingCountry box="[526,598,1191,1218]" name="China" pageId="7" pageNumber="408">China</collectingCountry>
,
<collectingRegion box="[610,671,1189,1215]" country="China" name="Xizang" pageId="7" pageNumber="408">Tibet</collectingRegion>
,
<collectingMunicipality box="[682,765,1189,1216]" pageId="7" pageNumber="408">Cha Yu</collectingMunicipality>
[
<emphasis box="[782,864,1190,1215]" italics="true" pageId="7" pageNumber="408">= Zayü</emphasis>
]
<location LSID="urn:lsid:plazi:treatment:050B87D78D472466E1B81A80FB567460:887D601A8D472469E25E1DD3FC0572BE" box="[881,965,1189,1215]" country="China" municipality="Cha Yu" name="County" pageId="7" pageNumber="408" stateProvince="Xizang">County</location>
, Xia-Cha-Yu,
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<elevation box="[1126,1284,1189,1216]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.6" metricValueMin="1.9" pageId="7" pageNumber="408" unit="m" value="2250.0" valueMax="2600.0" valueMin="1900.0">19002600 m</elevation>
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, 1
<date box="[1319,1436,1189,1215]" pageId="7" pageNumber="408" value="2004-07-28">
<collectingDate box="[1319,1436,1189,1215]" pageId="7" pageNumber="408" value="2004-07-28">28.7.2004</collectingDate>
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(
<collectionCode box="[159,202,1225,1251]" pageId="7" pageNumber="408">JW</collectionCode>
)
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;
<materialsCitation box="[222,793,1225,1252]" collectionCode="JW" country="China" pageId="7" pageNumber="408" specimenCount="1" specimenCount-female="1" stateProvince="Xizang" typeStatus="paratype">
<typeStatus box="[222,332,1226,1252]" pageId="7" pageNumber="408">PARATYPE</typeStatus>
,
<specimenCount box="[342,365,1225,1251]" pageId="7" pageNumber="408" type="female">1♀</specimenCount>
—same labeled as the
<typeStatus box="[629,727,1225,1251]" pageId="7" pageNumber="408">holotype</typeStatus>
(
<collectionCode box="[742,785,1225,1251]" pageId="7" pageNumber="408">JW</collectionCode>
)
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.
</paragraph>
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<subSubSection pageId="7" pageNumber="408" type="diagnosis">
<paragraph blockId="7.[151,1437,1189,2007]" pageId="7" pageNumber="408">
<emphasis bold="true" box="[199,322,1261,1287]" pageId="7" pageNumber="408">Diagnosis.</emphasis>
This new subspecies is best distinguished from the nominotypical form by the shape of antennomeres 911 (
<figureCitation box="[221,347,1297,1323]" captionStart="FIGURES 2949" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 2949. Therates spp., left antennomeres 611: 2932—T. fruhstorferi fruhstorferi (29, 30 Vietnam, Na Hang; 31, 32 Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="7" pageNumber="408">Figs 38, 39</figureCitation>
<emphasis box="[354,384,1298,1323]" italics="true" pageId="7" pageNumber="408">vs.</emphasis>
<figureCitation box="[391,518,1297,1323]" captionStart="FIGURES 2949" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 2949. Therates spp., left antennomeres 611: 2932—T. fruhstorferi fruhstorferi (29, 30 Vietnam, Na Hang; 31, 32 Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="7" pageNumber="408">Figs 36, 37</figureCitation>
); by the proportions of hind tibiae and tarsi—mean HTbL/HTaL = 1.25 vs. HTbL/ HTaL = 1.32; by virtually transverse, clearly dilated in lateral portion central dot (
<figureCitation box="[1092,1180,1333,1359]" captionStart="FIGURES 114" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 114. Therates spp., habitus: 16—T. fruhstorferi fruhstorferi (13—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 913— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 314—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="7" pageNumber="408">Figs 11</figureCitation>
,
<figureCitation box="[1194,1266,1333,1360]" captionStart="FIGURES 8198" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 8198. Therates spp., left elytron: 8187—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 9193, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="7" pageNumber="408">94, 95</figureCitation>
<emphasis box="[1275,1305,1334,1359]" italics="true" pageId="7" pageNumber="408">vs.</emphasis>
<figureCitation box="[1314,1431,1333,1359]" captionStart="FIGURES 114" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 114. Therates spp., habitus: 16—T. fruhstorferi fruhstorferi (13—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 913— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 314—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="7" pageNumber="408">Figs 9, 10</figureCitation>
,
<figureCitation box="[151,222,1369,1395]" captionStart="FIGURES 8198" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 8198. Therates spp., left elytron: 8187—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 9193, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="7" pageNumber="408">9093</figureCitation>
) and by shorter elytra—mean EL/AL = 2.48
<emphasis box="[728,759,1370,1395]" italics="true" pageId="7" pageNumber="408">vs.</emphasis>
mean EL/AL = 2.61.
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="409" pageId="7" pageNumber="408" type="description">
<paragraph blockId="7.[151,1437,1189,2007]" box="[199,836,1405,1432]" pageId="7" pageNumber="408">
<emphasis bold="true" box="[199,345,1405,1431]" pageId="7" pageNumber="408">Description.</emphasis>
TL =
<quantity box="[414,514,1405,1432]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="7" pageNumber="408" unit="mm" value="10.2">10.2 mm</quantity>
in male,
<quantity box="[618,717,1405,1432]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="7" pageNumber="408" unit="mm" value="11.6">11.6 mm</quantity>
in female.
</paragraph>
<paragraph blockId="7.[151,1437,1189,2007]" pageId="7" pageNumber="408">
Head shining black-blue with light violet reflection; orbital plates slightly striated in posterior third; frons semi-circular (see above), slightly downward anteriorly (see at the side), smooth, frontal sulci deep and indistinctly convergent in anterior 2/3, shallow and sharply divergent in posterior third; occiput flat or slightly concave in anterior margin. Mandibles in male brown underside, pale-yellow topside with brown teeth and apical molar; in female dark brown except yellow-brown basal half of topside. Two basal labial palpomeres light brown, palpomere 3 pale-yellow with brown apex, apical palpomere dark brown; palpomeres 1 and 2 of maxillary palpi brown, palpomeres 3 and 4 brown-black. Antennae barely extend shoulders, antennomeres
<quantity box="[1086,1170,1657,1683]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="2.794" metricValueMin="2.286" pageId="7" pageNumber="408" unit="in" value="10.0" valueMax="11.0" valueMin="9.0">911 in</quantity>
males distinctly dilated and slightly club-shaped, antennomeres 9 and 10 with protruding downward anterior lower margins, antennomere 11 with sharply cut straight anterior margin (
<figureCitation box="[667,753,1729,1755]" captionStart="FIGURES 2949" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 2949. Therates spp., left antennomeres 611: 2932—T. fruhstorferi fruhstorferi (29, 30 Vietnam, Na Hang; 31, 32 Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="7" pageNumber="408">Fig. 39</figureCitation>
); antennomeres 9 and
<quantity box="[1013,1072,1729,1755]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="7" pageNumber="408" unit="in" value="10.0">10 in</quantity>
female only slightly protruding downward, antennomere 11 oval with protruding apex (
<figureCitation box="[781,867,1765,1791]" captionStart="FIGURES 2949" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 2949. Therates spp., left antennomeres 611: 2932—T. fruhstorferi fruhstorferi (29, 30 Vietnam, Na Hang; 31, 32 Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="7" pageNumber="408">Fig. 38</figureCitation>
). Labrum as long as width (LL = LW—
<quantity box="[1323,1407,1765,1792]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="408" unit="mm" value="1.2">1.2 mm</quantity>
in male;
<quantity box="[220,305,1801,1828]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="408" unit="mm" value="1.5">1.5 mm</quantity>
in female), in male black-brown with medium-sized yellow-brown apical spot clearly separated from central apical teeth (
<figureCitation box="[378,461,1837,1863]" captionStart="FIGURES 5864" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 5864. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="7" pageNumber="408">Fig. 60</figureCitation>
), in female with small barely visible brown spot in anterior third (
<figureCitation box="[1200,1283,1837,1863]" captionStart="FIGURES 5864" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 5864. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="7" pageNumber="408">Fig. 61</figureCitation>
).
</paragraph>
<paragraph blockId="7.[151,1437,1189,2007]" pageId="7" pageNumber="408">Pronotum shining black-blue; indistinctly transverse, PW/PL = 1.051.06 (mean = 1.05, n = 2), apical lobe indistinctly wider in central portion; thorax shining blue with light golden reflection.</paragraph>
<paragraph blockId="7.[151,1437,1189,2007]" lastBlockId="8.[151,1437,151,609]" lastPageId="8" lastPageNumber="409" pageId="7" pageNumber="408">Fore femora yellow-brown with blue-tinged posterior side; middle femora in male black-brown with yellowbrown base and light blue-tinged anterior margin, in female entirely brown with blue tinge; hind femora pale-yellow in basal third and black with blue tinge in apical 2/3; fore and middle tibiae in male light brown in basal third and dark-brown blue-tinged in apical third; hind tibia entirely black with violet reflection; in female all tibiae black with blue lustre; tarsi black with indistinct blue tinge; HTbL/HTaL = 1.241.25 (mean = 1.25, n = 2).</paragraph>
<paragraph blockId="8.[151,1437,151,609]" pageId="8" pageNumber="409">
Elytra shining blue with light purple-violet reflection; indistinctly divergent toward apex, EL/EW = 1.951.97 (mean = 1.96, n = 2); punctuation deep and regular between base of humeral hump and central dot, shallower and sparse in the middle and practically absent in apical quarter; scutellum black with blue tinge; apical margin clearly cut. Elytral pattern consists of pale-yellow very small basal portion of humeral lunule (see in front) and elongate practically transverse central dot clearly dilated in lateral portion; basal dot virtually invisible in male, entirely absent in female (
<figureCitation box="[350,435,439,465]" captionStart="FIGURES 114" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 114. Therates spp., habitus: 16—T. fruhstorferi fruhstorferi (13—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 913— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 314—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="8" pageNumber="409">Figs 11</figureCitation>
,
<figureCitation box="[447,521,438,465]" captionStart="FIGURES 8198" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 8198. Therates spp., left elytron: 8187—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 9193, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="8" pageNumber="409">94, 95</figureCitation>
).
</paragraph>
<paragraph blockId="8.[151,1437,151,609]" pageId="8" pageNumber="409">
Aedeagus gradually sinuate with elongate rounded apex lacking lateral carinae (
<figureCitation box="[1083,1180,474,501]" captionStart="FIGURES 115135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115135. Therates spp., aedeagus: 115125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="8" pageNumber="409">Fig. 135</figureCitation>
); AL =
<quantity box="[1261,1348,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="409" unit="mm" value="2.6">2.6 mm</quantity>
, EL/AL = 2.48.
</paragraph>
</subSubSection>
<subSubSection box="[199,1276,547,573]" pageId="8" pageNumber="409" type="etymology">
<paragraph blockId="8.[151,1437,151,609]" box="[199,1276,547,573]" pageId="8" pageNumber="409">
<emphasis bold="true" box="[199,334,547,573]" pageId="8" pageNumber="409">Etymology.</emphasis>
This new species is named after the
<typeStatus box="[745,793,547,573]" pageId="8" pageNumber="409">type</typeStatus>
area—Chinese
<collectingRegion box="[972,1272,547,573]" country="China" name="Xizang" pageId="8" pageNumber="409">Tibet Autonomous Region</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection box="[199,1174,582,609]" pageId="8" pageNumber="409" type="distribution">
<paragraph blockId="8.[151,1437,151,609]" box="[199,1174,582,609]" pageId="8" pageNumber="409">
<emphasis bold="true" box="[199,353,582,608]" pageId="8" pageNumber="409">Distribution.</emphasis>
<collectingCountry box="[359,437,583,609]" name="China" pageId="8" pageNumber="409">CHINA</collectingCountry>
:
<collectingRegion box="[447,743,583,609]" country="China" name="Xizang" pageId="8" pageNumber="409">Tibet Autonomous Region</collectingRegion>
(Cha Yu [
<emphasis box="[858,939,583,608]" italics="true" pageId="8" pageNumber="409">= Zayü</emphasis>
] County) (
<figureCitation box="[1063,1161,583,609]" captionStart="FIGURES 184" captionStartId="28.[152,270,1579,1603]" captionTargetBox="[151,1435,597,1556]" captionTargetId="figure-226@28.[151,1435,597,1556]" captionTargetPageId="28" captionText="FIGURES 184. Distribution of Therates spp.: dark blue solid circle—T. vitalisi vitalisi; dark blue half-solid circle—T. vitalisi ida; lilac solid circle—T. hunnanensis sp. n.; orange solid circle—T. biserratus; pink solid circle—T. motoensis; red open circle—T. sauteri; red solid quadrat—T. fruhstorferi fruhstorferi; red half-solid quadrat—T. fruhstorferi australis ssp. n.; blue solid quadrat—T. ziyardamensis sp. n.; green solid quadrat—T. pseudovitalisi sp. n. (nominotypical subspecies); green halfsolid quadrat—T. pseudovitalisi tibetana ssp. n.; L in the symbols—according only to the literature sources. Map source— https://commons.wikimedia.org/wiki/File:East_Asia_topographic_map.png." figureDoi="http://doi.org/10.5281/zenodo.7758916" httpUri="https://zenodo.org/record/7758916/files/figure.png" pageId="8" pageNumber="409">Fig. 184</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>