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<mods:title id="FF67F91BBBFD955DBE19FD3C8DC3D18C">Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae)</mods:title>
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<mods:namePart id="F1200A61BA1749FAEFC55454E630764F">Matalin, Andrey V.</mods:namePart>
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<mods:affiliation id="9205A858038F296864EE85296D3CCB6A">Moscow State Pedagogical University, Department of Zoology &amp; Ecology, Kibalchicha str. 6, build. 3, Moscow 129164, Russia. andrei-matalin @ yandex. ru; https: // orcid. org / 0000 - 0002 - 7790 - 8709 &amp; Pirogov National Research Medical University, Department of Biology, Ostrovitianova str. 1, Moscow 117997, Russia.</mods:affiliation>
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<mods:namePart id="B90CDD150B4D21978D96ED015B3756DF">Wiesner, Jürgen</mods:namePart>
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<taxonomicName id="4AA24D428D462468E1B81A10FD647581" authority="Matalin et Wiesner" authorityName="Matalin et Wiesner" authorityYear="2023" box="[151,676,870,897]" class="Insecta" family="Cicindelidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="pseudovitalisi" status="sp. nov.">
<emphasis id="BFD6EAD38D462468E1B81A10FE6D7580" bold="true" box="[151,429,870,897]" italics="true" pageId="6" pageNumber="407">Therates pseudovitalisi</emphasis>
Matalin et Wiesner
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, 
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<paragraph id="8D1D36C18D462468E1B81AFDFDBA75A4" blockId="6.[151,755,870,933]" box="[151,634,906,933]" pageId="6" pageNumber="407">
<figureCitation id="15992A448D462468E1B81AFDFEC875A4" box="[151,264,907,933]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="6" pageNumber="407">Figs 9, 10</figureCitation>
, 
<figureCitation id="15992A448D462468E03B1AFDFE9C75A4" box="[276,348,907,933]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="6" pageNumber="407">36, 37</figureCitation>
, 
<figureCitation id="15992A448D462468E0471AFCFE7075A4" box="[360,432,906,933]" captionStart="FIGURES 58–64" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="6" pageNumber="407">58, 59</figureCitation>
, 
<figureCitation id="15992A448D462468E0931AFDFDC575A4" box="[444,517,907,933]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="6" pageNumber="407">90–93</figureCitation>
, 
<figureCitation id="15992A448D462468E33F1AFDFDB675A4" box="[528,630,907,933]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="6" pageNumber="407">132–134</figureCitation>
.
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<emphasis id="BFD6EAD38D462468E1B81AB9FE7675E6" box="[151,438,975,999]" italics="true" pageId="6" pageNumber="407">Therates fruhstorferi vitalisi</emphasis>
W. 
<bibRefCitation id="E9334B308D462468E0F11AA6FD9075E6" author="Horn, W." box="[478,592,975,999]" pageId="6" pageNumber="407" pagination="362 - 366" refId="ref25772" refString="Horn, W. (1913) Materiaux pour servir a l'etude de la faune entomologique de l'Indochine. Cicindelinae. Annales de la Societe Entomologique de Belgique, 57, 362 - 366. https: // doi. org / 10.5962 / bhl. part. 4597" type="journal article" year="1913">Horn, 1913</bibRefCitation>
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<bibRefCitation id="E9334B308D462468E3451AB9FC7575E6" author="Sawada, H. &amp; Wiesner, J." box="[618,949,975,999]" pageId="6" pageNumber="407" pagination="59 - 68" refId="ref26747" refString="Sawada, H. &amp; Wiesner, J. (2000) New records of tiger beetles collected in Laos (Coleoptera: Cicindelidae). Entomological Revue of Japan, 55 (1), 59 - 68." type="journal article" year="2000">Sawada &amp; Wiesner 2000: 63–64</bibRefCitation>
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<bibRefCitation id="E9334B308D462468E2EF1AB9FB9C75E6" author="Sawada, H. &amp; Wiesner, J." box="[960,1116,975,999]" pageId="6" pageNumber="407" pagination="255 - 274" refId="ref26828" refString="Sawada, H. &amp; Wiesner, J. (2004) Enumeration of the tiger beetles from Laos with description of the three new species (Coleoptera: Cicindelidae). Entomological Revue of Japan, 59 (2), 255 - 274." type="journal article" year="2004">2004: 263–264</bibRefCitation>
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<bibRefCitation id="E9334B308D462468E54A1AB9FABE75E6" author="Wiesner, J. &amp; Geiser, M." box="[1125,1406,975,999]" pageId="6" pageNumber="407" pagination="61 - 117" refId="ref27796" refString="Wiesner, J. &amp; Geiser, M. (2016) Faunistic survey of the Tiger Beetles (Coleoptera, Carabidae, Cicindelinae) of Laos. Entomologica Basiliensia et Collectionis Frey, 35, 61 - 117." type="journal article" year="2016">Wiesner &amp; Geiser 2016: 88</bibRefCitation>
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<emphasis id="BFD6EAD38D462468E1B81D63FE88722E" bold="true" box="[151,328,1045,1071]" pageId="6" pageNumber="407">Type material.</emphasis>
<materialsCitation id="3DCA3C9C8D462468E07E1D63FCB87252" ID-GBIF-Occurrence="4063374319" collectingDate="1999-05-16" collectionCode="JW" collectorName="Vit Kuban &amp; C. Holzschuh" country="Laos" elevation="1200" latitude="20.33" location="Ban Song Cha" longitude="102.23333" pageId="6" pageNumber="407" specimenCount="1" stateProvince="Luang Prabang Prov." typeStatus="holotype">
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</geoCoordinate>
, ± 
<quantity id="4A5A9B248D462468E1B81D4FFF327255" box="[151,242,1081,1108]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="407" unit="m" value="1200.0">
<elevation id="068FD1F28D462468E1B81D4FFF327255" box="[151,242,1081,1108]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="407" unit="m" value="1200.0">1200 m</elevation>
</quantity>
, 24.4– 
<date id="F91C10018D462468E06E1D4FFE727255" box="[321,434,1081,1108]" pageId="6" pageNumber="407" value="1999-05-16">
<collectingDate id="E958E9E98D462468E06E1D4FFE727255" box="[321,434,1081,1108]" pageId="6" pageNumber="407" value="1999-05-16">16.5.1999</collectingDate>
</date>
, leg. 
<collectorName id="205753178D462468E0DB1D4FFDAC7252" box="[500,620,1081,1107]" pageId="6" pageNumber="407">Vit Kubaň</collectorName>
&amp; 
<collectorName id="205753178D462468E3B91D4FFCF27252" box="[662,818,1081,1107]" pageId="6" pageNumber="407">C. Holzschuh</collectorName>
(
<collectionCode id="EBB3AE048D462468E26A1D4FFCB07252" box="[837,880,1081,1107]" pageId="6" pageNumber="407">JW</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3DCA3C9C8D462468E2A81D4FFF127276" ID-GBIF-Occurrence="4063374332" collectionCode="JW" country="Laos" pageId="6" pageNumber="407" specimenCount="1" specimenCount-female="1" stateProvince="Louangphabang" typeStatus="paratype">
<typeStatus id="521988638D462468E2A81D4FFBC17252" box="[903,1025,1081,1107]" pageId="6" pageNumber="407" type="paratype">PARATYPES</typeStatus>
: 
<specimenCount id="9BA4FD488D462468E5211D4FFBE57252" box="[1038,1061,1081,1107]" pageId="6" pageNumber="407" type="female">1♀</specimenCount>
—same labeled as the holotype (
<collectionCode id="EBB3AE048D462468E1B01D2BFF0A7276" box="[159,202,1117,1143]" collectionName="Jan Wieringa" pageId="6" pageNumber="407">JW</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3DCA3C9C8D462468E1CF1D29FDF0729F" ID-GBIF-Occurrence="4063374377" collectingDate="1998-05-01" collectionCode="JW, MSPU" collectorName="Vit Kuban" country="Laos" elevation="1100" latitude="20.75" location="Oudom Xai" longLatPrecision="1266" longitude="102.15" pageId="6" pageNumber="407" specimenCount="19" specimenCount-female="13" typeStatus="paratype">
<specimenCount id="9BA4FD488D462468E1CF1D29FED9727B" box="[224,281,1119,1146]" pageId="6" pageNumber="407">3♂♂</specimenCount>
<specimenCount id="9BA4FD488D462468E0351D29FE86727B" box="[282,326,1119,1146]" pageId="6" pageNumber="407" type="female">6♀♀</specimenCount>
— 
<collectingCountry id="F5B576518D462468E0481D29FE64727B" box="[359,420,1119,1146]" name="Laos" pageId="6" pageNumber="407">Laos</collectingCountry>
, 
<location id="887D601A8D462468E09D1D2BFDF67276" LSID="urn:lsid:plazi:treatment:050B87D78D462469E1B81A10FB2875A4:887D601A8D462468E09D1D2BFDF67276" box="[434,566,1117,1143]" country="Laos" latitude="20.75" longLatPrecision="1266" longitude="102.15" name="Oudom Xai" pageId="6" pageNumber="407">Oudom Xai</location>
(
<quantity id="4A5A9B248D462468E3671D2BFD517276" box="[584,657,1117,1143]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="407" unit="km" value="17.0">
<locationDeviation id="9CAF2FE38D462468E3671D2BFD517276" box="[584,657,1117,1143]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="6" pageNumber="407" unit="km" value="17.0">17 km</locationDeviation>
</quantity>
NEE), 
<geoCoordinate id="E89650068D462468E3C61D2BFC877279" box="[745,839,1117,1144]" degrees="20" direction="north" minutes="45" orientation="latitude" pageId="6" pageNumber="407" precision="925" value="20.75">
20 
<superScript id="7AD79B898D462468E22A1D2BFCCD726A" attach="none" box="[773,781,1117,1131]" fontSize="6" pageId="6" pageNumber="407">o</superScript>
45’N
</geoCoordinate>
<geoCoordinate id="E89650068D462468E27F1D2BFC7B7276" box="[848,955,1117,1143]" degrees="102" direction="east" minutes="09" orientation="longitude" pageId="6" pageNumber="407" precision="925" value="102.15">
102 
<superScript id="7AD79B898D462468E2551D2BFC42726A" attach="none" box="[890,898,1117,1131]" fontSize="6" pageId="6" pageNumber="407">o</superScript>
09’E
</geoCoordinate>
, ~ 
<quantity id="4A5A9B248D462468E2CF1D2BFBF77279" box="[992,1079,1117,1144]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="407" unit="m" value="1100.0">
<elevation id="068FD1F28D462468E2CF1D2BFBF77279" box="[992,1079,1117,1144]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="407" unit="m" value="1100.0">1100 m</elevation>
</quantity>
, 18.4– 
<date id="F91C10018D462468E5AB1D2BFB287279" box="[1156,1256,1117,1144]" pageId="6" pageNumber="407" value="1998-05-01">
<collectingDate id="E958E9E98D462468E5AB1D2BFB287279" box="[1156,1256,1117,1144]" pageId="6" pageNumber="407" value="1998-05-01">1.5.1998</collectingDate>
</date>
, leg. 
<collectorName id="205753178D462468E4081D2BFA5D7276" box="[1319,1437,1117,1143]" pageId="6" pageNumber="407">Vit Kubaň</collectorName>
(
<specimenCount id="9BA4FD488D462468E18F1DF5FF1B729F" box="[160,219,1155,1182]" pageId="6" pageNumber="407">2♂♂</specimenCount>
<specimenCount id="9BA4FD488D462468E1F41DF5FEC6729F" box="[219,262,1155,1182]" pageId="6" pageNumber="407" type="female">5♀♀</specimenCount>
— 
<collectionCode id="EBB3AE048D462468E0081DF5FE92729F" box="[295,338,1155,1182]" pageId="6" pageNumber="407">JW</collectionCode>
, 
<specimenCount id="9BA4FD488D462468E0701DF5FEBF729F" box="[351,383,1155,1182]" pageId="6" pageNumber="407">1♂</specimenCount>
<specimenCount id="9BA4FD488D462468E0501DF5FE6C729F" box="[383,428,1155,1182]" pageId="6" pageNumber="407" type="female">2♀♀</specimenCount>
— 
<collectionCode id="EBB3AE048D462468E0E21DF5FDE6729F" box="[461,550,1155,1182]" pageId="6" pageNumber="407">MSPU</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3DCA3C9C8D462468E3121DF5FAD9729A" ID-GBIF-Occurrence="4063374406" box="[573,1305,1153,1182]" collectingDate="2011-07" collectionCode="MSPU" country="Laos" location="Phou Pane Mt" pageId="6" pageNumber="407" specimenCount="1" specimenCount-male="1" stateProvince="Houaphanh Prov." typeStatus="paratype">
<specimenCount id="9BA4FD488D462468E3121DF5FD9B729F" box="[573,603,1155,1182]" pageId="6" pageNumber="407" type="male">1♂</specimenCount>
— 
<collectingCountry id="F5B576518D462468E3551DF5FD75729F" box="[634,693,1155,1182]" name="Laos" pageId="6" pageNumber="407">Laos</collectingCountry>
, 
<collectingRegion id="4F66F8238D462468E3ED1DF7FC4A729D" box="[706,906,1153,1180]" country="Laos" name="Houaphan" pageId="6" pageNumber="407">Houaphanh Prov.</collectingRegion>
, 
<location id="887D601A8D462468E2B81DF7FBFC729D" LSID="urn:lsid:plazi:treatment:050B87D78D462469E1B81A10FB2875A4:887D601A8D462468E2B81DF7FBFC729D" box="[919,1084,1153,1180]" country="Laos" name="Phou Pane Mt" pageId="6" pageNumber="407" stateProvince="Houaphanh Prov.">Phou Pane Mt</location>
, 
<date id="F91C10018D462468E5661DF7FB6E729A" box="[1097,1198,1153,1179]" pageId="6" pageNumber="407" value="2011-07">
<collectingDate id="E958E9E98D462468E5661DF7FB6E729A" box="[1097,1198,1153,1179]" pageId="6" pageNumber="407" value="2011-07">VII.2011</collectingDate>
</date>
(
<collectionCode id="EBB3AE048D462468E5901DF7FAD2729A" box="[1215,1298,1153,1179]" collectionName="MSPU" pageId="6" pageNumber="407">MSPU</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3DCA3C9C8D462468E4081DF7FDA872BE" ID-GBIF-Occurrence="4063374382" collectingDate="2002-05-17" collectionCode="JW" country="Laos" location="Neua" municipality="Sam" pageId="6" pageNumber="407" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9BA4FD488D462468E4081DF7FAFE729A" box="[1319,1342,1153,1179]" pageId="6" pageNumber="407" type="female">1♀</specimenCount>
— 
<collectingCountry id="F5B576518D462468E4701DF7FA5C729A" box="[1375,1436,1153,1179]" name="Laos" pageId="6" pageNumber="407">Laos</collectingCountry>
NE, 
<collectingMunicipality id="6D79ACBB8D462468E1E41DD3FF3D72BE" box="[203,253,1189,1215]" pageId="6" pageNumber="407">Sam</collectingMunicipality>
[
<emphasis id="BFD6EAD38D462468E0221DD0FE9A72BE" box="[269,346,1190,1215]" italics="true" pageId="6" pageNumber="407">= Xam</emphasis>
] 
<location id="887D601A8D462468E0451DD0FE6872C1" LSID="urn:lsid:plazi:treatment:050B87D78D462469E1B81A10FB2875A4:887D601A8D462468E0451DD0FE6872C1" box="[362,424,1190,1216]" country="Laos" municipality="Sam" name="Neua" pageId="6" pageNumber="407">Neua</location>
, 
<date id="F91C10018D462468E09B1DD3FDE672BE" box="[436,550,1189,1215]" pageId="6" pageNumber="407" value="2002-05-17">
<collectingDate id="E958E9E98D462468E09B1DD3FDE672BE" box="[436,550,1189,1215]" pageId="6" pageNumber="407" value="2002-05-17">17.V.2002</collectingDate>
</date>
(
<collectionCode id="EBB3AE048D462468E3191DD3FDA172BE" box="[566,609,1189,1215]" collectionName="Jan Wieringa" pageId="6" pageNumber="407">JW</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3DCA3C9C8D462468E35B1DD1FCBA72E2" ID-GBIF-Occurrence="4063374330" collectingDate="2003-06-20" collectionCode="JW" collectorName="P. Pacholatko" country="Laos" elevation="1100" latitude="21.633333" location="Ban Bun Neua" longLatPrecision="1263" longitude="101.95" pageId="6" pageNumber="407" specimenCount="1" specimenCount-male="1" stateProvince="Phongsaly prov." typeStatus="paratype">
<specimenCount id="9BA4FD488D462468E35B1DD1FD5172C3" box="[628,657,1191,1218]" pageId="6" pageNumber="407" type="male">1♂</specimenCount>
—Laos-N, 
<collectingRegion id="4F66F8238D462468E23A1DD3FC0D72C1" box="[789,973,1189,1216]" country="Laos" name="Phongsali" pageId="6" pageNumber="407">Phongsaly prov.</collectingRegion>
, 
<location id="887D601A8D462468E2F71DD0FBBD72C1" LSID="urn:lsid:plazi:treatment:050B87D78D462469E1B81A10FB2875A4:887D601A8D462468E2F71DD0FBBD72C1" box="[984,1149,1190,1216]" country="Laos" latitude="21.633333" longLatPrecision="1263" longitude="101.95" name="Ban Bun Neua" pageId="6" pageNumber="407" stateProvince="Phongsaly prov.">Ban Bun Neua</location>
[= 
<emphasis id="BFD6EAD38D462468E58B1DD0FAE172BE" box="[1188,1313,1190,1215]" italics="true" pageId="6" pageNumber="407">Boun Neua</emphasis>
] (
<locationDeviation id="9CAF2FE38D462468E4151DD3FA4F72BE" box="[1338,1423,1189,1215]" pageId="6" pageNumber="407">
<quantity id="4A5A9B248D462468E4151DD3FAB472BE" box="[1338,1396,1189,1215]" metricMagnitude="3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="407" unit="km" value="4.0">4 km</quantity>
E
</locationDeviation>
), 
<date id="F91C10018D462468E1B81DBFFEDA72E2" box="[151,282,1225,1251]" pageId="6" pageNumber="407" value="2003-06-20">
<collectingDate id="E958E9E98D462468E1B81DBFFEDA72E2" box="[151,282,1225,1251]" pageId="6" pageNumber="407" value="2003-06-20">20.VI.2003</collectingDate>
</date>
, ~ 
<quantity id="4A5A9B248D462468E01C1DBFFE4A72E5" box="[307,394,1225,1252]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="407" unit="m" value="1100.0">
<elevation id="068FD1F28D462468E01C1DBFFE4A72E5" box="[307,394,1225,1252]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="407" unit="m" value="1100.0">1100 m</elevation>
</quantity>
, 
<geoCoordinate id="E89650068D462468E0B91DBFFE3372E2" box="[406,499,1225,1251]" degrees="21" direction="north" minutes="38" orientation="latitude" pageId="6" pageNumber="407" precision="925" value="21.633333">
21 
<superScript id="7AD79B898D462468E09D1DBFFE7A72D6" attach="none" box="[434,442,1225,1239]" fontSize="6" pageId="6" pageNumber="407">o</superScript>
38’N
</geoCoordinate>
<geoCoordinate id="E89650068D462468E0D51DBFFDA572E5" box="[506,613,1225,1252]" degrees="101" direction="east" minutes="57" orientation="longitude" pageId="6" pageNumber="407" precision="925" value="101.95">
101 
<superScript id="7AD79B898D462468E30B1DBFFDEC72D6" attach="none" box="[548,556,1225,1239]" fontSize="6" pageId="6" pageNumber="407">o</superScript>
57’E
</geoCoordinate>
, 
<collectorName id="205753178D462468E35E1DBCFCC772E2" box="[625,775,1225,1252]" pageId="6" pageNumber="407">P. Pacholátko</collectorName>
leg. (
<collectionCode id="EBB3AE048D462468E2681DBFFCB272E2" box="[839,882,1225,1251]" collectionName="Jan Wieringa" pageId="6" pageNumber="407">JW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C5B8654A8D462468E1E81D9BFB9B734E" pageId="6" pageNumber="407" type="reference_group">
<paragraph id="8D1D36C18D462468E1E81D9BFB9B734E" blockId="6.[151,1437,1045,2008]" pageId="6" pageNumber="407">
<emphasis id="BFD6EAD38D462468E1E81D9BFE907306" bold="true" box="[199,336,1261,1287]" pageId="6" pageNumber="407">References.</emphasis>
<materialsCitation id="3DCA3C9C8D462468E0781D9BFAE07309" ID-GBIF-Occurrence="4063374324" box="[343,1312,1261,1290]" collectingDate="2002-07-15" collectingDateMax="2003-06-21" collectingDateMin="2002-07-15" collectorName="Xam Nua Distr &amp; Phu Pan" country="Laos" elevation="1350" location="Ban Saleui" municipality="Xam Nua Distr" pageId="6" pageNumber="407" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Houaphanh Prov.">
<collectingCountry id="F5B576518D462468E0781D9BFE517306" box="[343,401,1261,1287]" name="Laos" pageId="6" pageNumber="407">LAOS</collectingCountry>
, 
<emphasis id="BFD6EAD38D462468E0B01D9BFDA67306" box="[415,614,1261,1287]" italics="true" pageId="6" pageNumber="407">
<collectingRegion id="4F66F8238D462468E0B01D9BFDA67306" box="[415,614,1261,1287]" country="Laos" name="Houaphan" pageId="6" pageNumber="407">Houaphanh Prov.</collectingRegion>
</emphasis>
: 
<specimenCount id="9BA4FD488D462468E35B1D99FD53730B" box="[628,659,1263,1290]" pageId="6" pageNumber="407" type="male">1♂</specimenCount>
<specimenCount id="9BA4FD488D462468E3BC1D99FD01730B" box="[659,705,1263,1290]" pageId="6" pageNumber="407" type="female">2♀♀</specimenCount>
— 
<collectingMunicipality id="6D79ACBB8D462468E3CD1D99FC587306" box="[738,920,1261,1290]" pageId="6" pageNumber="407">Xam Nua Distr</collectingMunicipality>
, 
<location id="887D601A8D462468E28C1D98FBE17306" LSID="urn:lsid:plazi:treatment:050B87D78D462469E1B81A10FB2875A4:887D601A8D462468E28C1D98FBE17306" box="[931,1057,1261,1288]" country="Laos" municipality="Xam Nua Distr" name="Ban Saleui" pageId="6" pageNumber="407" stateProvince="Houaphanh Prov.">Ban Saleui</location>
, 
<quantity id="4A5A9B248D462468E5031D9BFB437309" box="[1068,1155,1261,1288]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="407" unit="m" value="1350.0">
<elevation id="068FD1F28D462468E5031D9BFB437309" box="[1068,1155,1261,1288]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="6" pageNumber="407" unit="m" value="1350.0">1350 m</elevation>
</quantity>
, 
<date id="F91C10018D462468E5A01D9BFADB7309" box="[1167,1307,1261,1288]" pageId="6" pageNumber="407" value="2002-07-15">
<collectingDate id="E958E9E98D462468E5A01D9BFADB7309" box="[1167,1307,1261,1288]" pageId="6" pageNumber="407" value="2002-07-15">15.VII.2002</collectingDate>
</date>
;
</materialsCitation>
<materialsCitation id="3DCA3C9C8D462468E4081D99FC3A732A" ID-GBIF-Occurrence="4063374302" collectingDate="2003-06-17" collectingDateMax="2003-06-21" collectingDateMin="2003-06-17" collectorName="Xam Nua Distr &amp; Phu Pan" country="Laos" elevation="1800" location="Phu Pan" municipality="Xam Nua Distr" pageId="6" pageNumber="407" specimenCount="1" specimenCount-female="2" specimenCount-male="1" stateProvince="Xam Nua Distr">
<specimenCount id="9BA4FD488D462468E4081D99FA83730B" box="[1319,1347,1263,1290]" pageId="6" pageNumber="407" type="male">1♂</specimenCount>
— 
<collectingRegion id="4F66F8238D462468E44D1D99FEC7732A" pageId="6" pageNumber="407">Xam Nua Distr</collectingRegion>
, 
<location id="887D601A8D462468E03C1C67FEB3732D" LSID="urn:lsid:plazi:treatment:050B87D78D462469E1B81A10FB2875A4:887D601A8D462468E03C1C67FEB3732D" box="[275,371,1297,1324]" country="Laos" municipality="Xam Nua Distr" name="Phu Pan" pageId="6" pageNumber="407" stateProvince="Xam Nua Distr">Phu Pan</location>
, 
<quantity id="4A5A9B248D462468E0511C67FE16732D" box="[382,470,1297,1324]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="407" unit="m" value="1800.0">
<elevation id="068FD1F28D462468E0511C67FE16732D" box="[382,470,1297,1324]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="6" pageNumber="407" unit="m" value="1800.0">1800 m</elevation>
</quantity>
, 
<date id="F91C10018D462468E0CD1C67FD4C732A" box="[482,652,1297,1323]" pageId="6" pageNumber="407" value="2003-06-17" valueMax="2003-06-21" valueMin="2003-06-17">
<collectingDate id="E958E9E98D462468E0CD1C67FD4C732A" box="[482,652,1297,1323]" pageId="6" pageNumber="407" value="2003-06-17" valueMax="2003-06-21" valueMin="2003-06-17">17–21.VI.2003</collectingDate>
</date>
(both 
<bibRefCitation id="E9334B308D462468E3F91C67FC32732A" author="Sawada, H. &amp; Wiesner, J." box="[726,1010,1297,1323]" pageId="6" pageNumber="407" pagination="255 - 274" refId="ref26828" refString="Sawada, H. &amp; Wiesner, J. (2004) Enumeration of the tiger beetles from Laos with description of the three new species (Coleoptera: Cicindelidae). Entomological Revue of Japan, 59 (2), 255 - 274." type="journal article" year="2004">Sawada &amp; Wiesner 2004</bibRefCitation>
)
</materialsCitation>
; 
<materialsCitation id="3DCA3C9C8D462468E5291C67FB97734E" ID-GBIF-Occurrence="4063374305" collectingDate="2009-06-01" collectingDateMax="2009-06-16" collectingDateMin="2009-06-01" collectorName="Z. Kraus" country="Laos" elevation="1515" location="Phou Pane Mt." longitude="103.99833" pageId="6" pageNumber="407" specimenCount="1" specimenCount-female="1" stateProvince="Houaphanh Prov.">
<specimenCount id="9BA4FD488D462468E5291C67FBDD732A" box="[1030,1053,1297,1323]" pageId="6" pageNumber="407" type="female">1♀</specimenCount>
— 
<location id="887D601A8D462468E5121C67FB2E732D" LSID="urn:lsid:plazi:treatment:050B87D78D462469E1B81A10FB2875A4:887D601A8D462468E5121C67FB2E732D" box="[1085,1262,1297,1324]" country="Laos" longitude="103.99833" name="Phou Pane Mt." pageId="6" pageNumber="407" stateProvince="Houaphanh Prov.">Phou Pane Mt.</location>
, 
<quantity id="4A5A9B248D462468E5D51C67FA58732D" box="[1274,1432,1297,1324]" metricMagnitude="3" metricUnit="m" metricValue="1.515" metricValueMax="1.55" metricValueMin="1.48" pageId="6" pageNumber="407" unit="m" value="1515.0" valueMax="1550.0" valueMin="1480.0">
<elevation id="068FD1F28D462468E5D51C67FA58732D" box="[1274,1432,1297,1324]" metricMagnitude="3" metricUnit="m" metricValue="1.515" metricValueMax="1.55" metricValueMin="1.48" pageId="6" pageNumber="407" unit="m" value="1515.0" valueMax="1550.0" valueMin="1480.0">1480–1550 m</elevation>
</quantity>
, 20 
<superScript id="7AD79B898D462468E19C1C43FF7B7342" attach="none" box="[179,187,1333,1347]" fontSize="6" pageId="6" pageNumber="407">o</superScript>
13’09–19’’N 103º59’54’’E, 
<date id="F91C10018D462468E0D91C43FD55734E" box="[502,661,1333,1359]" pageId="6" pageNumber="407" value="2009-06-01" valueMax="2009-06-16" valueMin="2009-06-01">
<collectingDate id="E958E9E98D462468E0D91C43FD55734E" box="[502,661,1333,1359]" pageId="6" pageNumber="407" value="2009-06-01" valueMax="2009-06-16" valueMin="2009-06-01">1–16.VI.2009</collectingDate>
</date>
, leg. 
<collectorName id="205753178D462468E3FF1C40FCF27351" box="[720,818,1334,1360]" pageId="6" pageNumber="407">Z. Kraus</collectorName>
(
<bibRefCitation id="E9334B308D462468E26F1C43FB8F734E" author="Wiesner, J. &amp; Geiser, M." box="[832,1103,1333,1359]" pageId="6" pageNumber="407" pagination="61 - 117" refId="ref27796" refString="Wiesner, J. &amp; Geiser, M. (2016) Faunistic survey of the Tiger Beetles (Coleoptera, Carabidae, Cicindelinae) of Laos. Entomologica Basiliensia et Collectionis Frey, 35, 61 - 117." type="journal article" year="2016">Wiesner &amp; Geiser 2016</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C5B8654A8D462468E1E81C2FFA9B706E" pageId="6" pageNumber="407" type="diagnosis">
<paragraph id="8D1D36C18D462468E1E81C2FFA9B706E" blockId="6.[151,1437,1045,2008]" pageId="6" pageNumber="407">
<emphasis id="BFD6EAD38D462468E1E81C2FFE827372" bold="true" box="[199,322,1369,1395]" pageId="6" pageNumber="407">Diagnosis.</emphasis>
This new species is readily distinguished from 
<taxonomicName id="4AA24D428D462468E25F1C2CFC1B7372" baseAuthorityName="Wiesner &amp; Bi" baseAuthorityYear="2014" box="[880,987,1369,1395]" class="Insecta" family="Cicindelidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="BFD6EAD38D462468E25F1C2CFC1B7372" box="[880,987,1369,1395]" italics="true" pageId="6" pageNumber="407">T. vitalisi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4AA24D428D462468E2C31C2CFBBD7372" authorityName="Tan, Mo, Liang" authorityYear="1991" box="[1004,1149,1369,1395]" class="Insecta" family="Cicindelidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="biserratus">
<emphasis id="BFD6EAD38D462468E2C31C2CFBBD7372" box="[1004,1149,1369,1395]" italics="true" pageId="6" pageNumber="407">T. biserratus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4AA24D428D462468E5A21C2CFAEA7372" authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[1165,1322,1369,1395]" class="Insecta" family="Cicindelidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="hunanensis" status="sp. nov.">
<emphasis id="BFD6EAD38D462468E5A21C2CFAEA7372" box="[1165,1322,1369,1395]" italics="true" pageId="6" pageNumber="407">T. hunanensis</emphasis>
</taxonomicName>
<emphasis id="BFD6EAD38D462468E41B1C2FFAB77372" bold="true" box="[1332,1399,1369,1395]" pageId="6" pageNumber="407">
<taxonomicNameLabel id="A4E557A88D462468E41B1C2FFAB77372" box="[1332,1399,1369,1395]" pageId="6" pageNumber="407" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="4AA24D428D462468E4A71C2CFF257396" authorityName="W. Horn" authorityYear="1912" class="Insecta" family="Cicindelidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis id="BFD6EAD38D462468E4A71C2CFF257396" italics="true" pageId="6" pageNumber="407">T. sauteri</emphasis>
</taxonomicName>
and 
<taxonomicName id="4AA24D428D462468E0341C08FE667396" authorityName="Tan" authorityYear="1981" box="[283,422,1406,1431]" class="Insecta" family="Cicindelidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="motoensis">
<emphasis id="BFD6EAD38D462468E0341C08FE667396" box="[283,422,1406,1431]" italics="true" pageId="6" pageNumber="407">T. motoensis</emphasis>
</taxonomicName>
by lacking lateral carinae on the apex of aedeagus (
<figureCitation id="15992A448D462468E2C61C0BFB437396" box="[1001,1155,1405,1431]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="6" pageNumber="407">Figs 132–134</figureCitation>
<emphasis id="BFD6EAD38D462468E5A51C08FB697396" box="[1162,1193,1406,1431]" italics="true" pageId="6" pageNumber="407">vs.</emphasis>
<figureCitation id="15992A448D462468E5801C0BFA8D7399" box="[1199,1357,1405,1432]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="6" pageNumber="407">Figs 152–174</figureCitation>
). From 
<taxonomicName id="4AA24D428D462468E1B81CD4FEF373BA" authorityName="W. Horn" authorityYear="1902" box="[151,307,1441,1467]" class="Insecta" family="Cicindelidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="fruhstorferi">
<emphasis id="BFD6EAD38D462468E1B81CD4FEF373BA" box="[151,307,1441,1467]" italics="true" pageId="6" pageNumber="407">T. fruhstorferi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4AA24D428D462468E0471CD4FDE373BA" authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[360,547,1441,1468]" class="Insecta" family="Cicindelidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="430" phylum="Arthropoda" rank="species" species="ziyardamensis" status="sp. nov.">
T. 
<emphasis id="BFD6EAD38D462468E0AB1CD7FDE373BA" box="[388,547,1441,1467]" italics="true" pageId="6" pageNumber="407">ziyardamensis</emphasis>
</taxonomicName>
<emphasis id="BFD6EAD38D462468E3061CD7FDA873BA" bold="true" box="[553,616,1441,1467]" pageId="6" pageNumber="407">
<taxonomicNameLabel id="A4E557A88D462468E3061CD7FDA873BA" box="[553,616,1441,1467]" pageId="6" pageNumber="407" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
a new species is easily distinguished by the shape of aedeagus (
<figureCitation id="15992A448D462468E4031CD7FF0173DE" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="6" pageNumber="407">Figs 132– 134</figureCitation>
<emphasis id="BFD6EAD38D462468E1E71CB0FF2673DE" box="[200,230,1478,1503]" italics="true" pageId="6" pageNumber="407">vs.</emphasis>
<figureCitation id="15992A448D462468E1C21CB3FE4873E1" box="[237,392,1477,1504]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="6" pageNumber="407">Figs 115–131</figureCitation>
), from 
<taxonomicName id="4AA24D428D462468E0F61CB0FDB573DE" authorityName="W. Horn" authorityYear="1902" box="[473,629,1477,1503]" class="Insecta" family="Cicindelidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="species" species="fruhstorferi">
<emphasis id="BFD6EAD38D462468E0F61CB0FDB573DE" box="[473,629,1477,1503]" italics="true" pageId="6" pageNumber="407">T. fruhstorferi</emphasis>
</taxonomicName>
additionally by wider apices of elytra (
<figureCitation id="15992A448D462468E5051CB3FB5C73DE" box="[1066,1180,1477,1503]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="6" pageNumber="407">Figs 9–11</figureCitation>
, 
<figureCitation id="15992A448D462468E5881CB3FB2D73E1" box="[1191,1261,1477,1504]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="6" pageNumber="407">90–95</figureCitation>
<emphasis id="BFD6EAD38D462468E5DB1CB0FAD273DE" box="[1268,1298,1478,1503]" italics="true" pageId="6" pageNumber="407">vs.</emphasis>
<figureCitation id="15992A448D462468E4361CB3FA5773DE" box="[1305,1431,1477,1503]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="6" pageNumber="407">Figs 1, 3–8</figureCitation>
, 
<figureCitation id="15992A448D462468E1B81C9FFF1F7002" box="[151,223,1513,1539]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="6" pageNumber="407">81–89</figureCitation>
) and by shorter hind tarsus (
<figureCitation id="15992A448D462468E33C1C9FFD707005" box="[531,688,1513,1540]" captionStart="FIGURES 175–178" captionStartId="25.[152,270,1739,1763]" captionTargetBox="[151,1435,181,1716]" captionTargetId="figure-17@25.[151,1435,181,1716]" captionTargetPageId="25" captionText="FIGURES 175–178. Morphometric features of Therates spp: 175, 177—relation of the proportions EL/PL and HTbL/HTaL (green rhombus—T. pseudovitalisi sp. n.; blue circles—T. fruhstorferi; open symbols—males; solid symbols—females); 176, 178—relation of the proportions HTbL/HTaL (small quadrates—medians; rectangles—25–75 percentiles; whiskers—min–max; significantly different samples are marked with different lower letters, according to Mann–Whitney U-test, p &lt;0.05 (176—U = 112.0, z = 5.08, p = 0.000; 178—U = 23.5, z = 3.37, p = 0.0007)); 175, 176—all specimens; 177, 178—males." figureDoi="http://doi.org/10.5281/zenodo.7758910" httpUri="https://zenodo.org/record/7758910/files/figure.png" pageId="6" pageNumber="407">Figs 175–178</figureCitation>
). From 
<taxonomicName id="4AA24D428D462468E22C1C9CFBDA7002" authorityName="Matalin et Wiesner" authorityYear="2023" box="[771,1050,1513,1539]" class="Insecta" family="Cicindelidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="407" phylum="Arthropoda" rank="subSpecies" species="pseudovitalisi" subSpecies="tibetana">
<emphasis id="BFD6EAD38D462468E22C1C9CFBDA7002" box="[771,1050,1513,1539]" italics="true" pageId="6" pageNumber="407">T. pseudovitalisi tibetana</emphasis>
</taxonomicName>
<emphasis id="BFD6EAD38D462468E5301C9FFBA77002" bold="true" box="[1055,1127,1513,1539]" pageId="6" pageNumber="407">ssp. n.</emphasis>
a nominotypical subspecies is clearly differentiated by the shape of antennomeres 9–11 (
<figureCitation id="15992A448D462468E2151F7BFC787026" box="[826,952,1549,1575]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="6" pageNumber="407">Figs 36, 37</figureCitation>
<emphasis id="BFD6EAD38D462468E2901F78FC1D7026" box="[959,989,1550,1575]" italics="true" pageId="6" pageNumber="407">vs.</emphasis>
<figureCitation id="15992A448D462468E2CB1F7BFBA47026" box="[996,1124,1549,1575]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="6" pageNumber="407">Figs 38, 39</figureCitation>
), by the proportions of hind tibiae and tarsi—mean HTbL/HTaL = 1.32 
<emphasis id="BFD6EAD38D462468E3511F44FD5C704A" box="[638,668,1586,1611]" italics="true" pageId="6" pageNumber="407">vs.</emphasis>
HTbL/HTaL = 1.25, by narrow, elongate, clearly oblique central dot (
<figureCitation id="15992A448D462468E1B01F23FED2706E" box="[159,274,1621,1647]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="6" pageNumber="407">Figs 9, 10</figureCitation>
, 
<figureCitation id="15992A448D462468E0311F23FEA4706E" box="[286,356,1621,1647]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="6" pageNumber="407">90–93</figureCitation>
<emphasis id="BFD6EAD38D462468E0441F20FE49706E" box="[363,393,1622,1647]" italics="true" pageId="6" pageNumber="407">vs.</emphasis>
<figureCitation id="15992A448D462468E0BF1F23FE24706E" box="[400,484,1621,1647]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="6" pageNumber="407">Figs 11</figureCitation>
, 
<figureCitation id="15992A448D462468E0DF1F23FDF97071" box="[496,569,1621,1648]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="6" pageNumber="407">94, 95</figureCitation>
), and by the longer elytra—mean EL/AL = 2.6 
<emphasis id="BFD6EAD38D462468E5621F20FBAC706E" box="[1101,1132,1622,1647]" italics="true" pageId="6" pageNumber="407">vs.</emphasis>
mean EL/AL = 2.48.
</paragraph>
</subSubSection>
<subSubSection id="C5B8654A8D462469E1E81F0FFAD77538" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407" type="description">
<paragraph id="8D1D36C18D462468E1E81F0FFEF170B6" blockId="6.[151,1437,1045,2008]" pageId="6" pageNumber="407">
<emphasis id="BFD6EAD38D462468E1E81F0FFE997092" bold="true" box="[199,345,1657,1683]" pageId="6" pageNumber="407">Description.</emphasis>
TL = 
<quantity id="4A5A9B248D462468E0B11F0FFD8E7095" box="[414,590,1657,1684]" metricMagnitude="-2" metricUnit="m" metricValue="1.0815" metricValueMax="1.1099999999999999" metricValueMin="1.053" pageId="6" pageNumber="407" unit="mm" value="10.815" valueMax="11.1" valueMin="10.53">10.53–11.1 mm</quantity>
in males (mean = 
<quantity id="4A5A9B248D462468E2331F0FFC507095" box="[796,912,1657,1684]" metricMagnitude="-2" metricUnit="m" metricValue="1.074" pageId="6" pageNumber="407" unit="mm" value="10.74">10.74 mm</quantity>
, n = 5), 
<quantity id="4A5A9B248D462468E2C31F0FFB4F7095" box="[1004,1167,1657,1684]" metricMagnitude="-2" metricUnit="m" metricValue="1.13" metricValueMax="1.21" metricValueMin="1.05" pageId="6" pageNumber="407" unit="mm" value="11.3" valueMax="12.1" valueMin="10.5">10.5–12.1 mm</quantity>
(mean = 
<quantity id="4A5A9B248D462468E5D61F0FFAAC7095" box="[1273,1388,1657,1684]" metricMagnitude="-2" metricUnit="m" metricValue="1.117" pageId="6" pageNumber="407" unit="mm" value="11.17">11.17 mm</quantity>
, n = 7) in females.
</paragraph>
<paragraph id="8D1D36C18D462469E1E81FB7FE84771C" blockId="6.[151,1437,1045,2008]" lastBlockId="7.[151,1437,151,933]" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407">
Head shining black-blue with purple-violet reflection; orbital plates finely striated in posterior third; frons convex, semi-circular (see above), slightly downward anteriorly (see at the side), smooth or slightly rugose, frontal sulci deep and slightly convergent in anterior 2/3, shallow and distinctly divergent in posterior third. Mandibles in males brown underside, yellow-brown topside with brown teeth and apical molar; in females dark-brown except pale basal quarter topside. Labial palpi yellow-brown or brown except dark-brown apical palpomeres; maxillary palpi dark brown with narrow light-brown apices in all palpomeres. Antennae barely extend shoulders, black, scape dark brown on anterior margin and black on posterior margin, in some females brown-black, antennomeres 2–5 with light violet tinge and narrow dark-brown apices, antennomeres 
<quantity id="4A5A9B248D462468E2791ECBFC6971D6" box="[854,937,1981,2007]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" metricValueMax="2.794" metricValueMin="2.286" pageId="6" pageNumber="407" unit="in" value="10.0" valueMax="11.0" valueMin="9.0">9–11 in</quantity>
males distinctly dilated, antennomeres 9 and 10 with wide protruding downward anterior lower margins (
<figureCitation id="15992A448D472469E21919E1FC4976B0" box="[822,905,151,177]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="7" pageNumber="408">Fig. 37</figureCitation>
), in female only slightly delated and protruding (
<figureCitation id="15992A448D472469E1B019CDFF3676D4" box="[159,246,187,213]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="7" pageNumber="408">Fig. 36</figureCitation>
). Labrum indistinctly transverse, LW/LL = 1.0–1.08 (mean = 1.04, n = 12), in males black-brown with medium-sized yellow-brown apical spot indistinctly separated from central apical teeth (
<figureCitation id="15992A448D472469E54619A9FB7B76F8" box="[1129,1211,222,249]" captionStart="FIGURES 58–64" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="7" pageNumber="408">Fig. 58</figureCitation>
), in females entirely black (
<figureCitation id="15992A448D472469E1CC1875FEF8771C" box="[227,312,258,285]" captionStart="FIGURES 58–64" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="7" pageNumber="408">Fig. 59</figureCitation>
).
</paragraph>
<paragraph id="8D1D36C18D472469E1E81851FA887764" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="408">Pronotum shining blue with violet reflection; indistinctly transverse, PW/PL = 1.0–1.11 (mean = 1.06, n = 12), apical lobe practically equal wide throughout; thorax shining black-blue with light golden-violet reflection.</paragraph>
<paragraph id="8D1D36C18D472469E1E81819FECA7418" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="408">
Femora bicolored, fore femora pale-yellow with brown violet-tinged anterior side, black with violet tinge on posterior side, middle femora brown-black with light violet tinge except pale-yellow posterior basal third, hind femora pale-yellow in basal third or quarter and black with violet tinge in apical 2/3 or 3/4; tibiae brown-black with blue-violet lustre; tarsi black with blue-violet tinge; HTbL/HTaL = 1.25–1.42 (mean = 1.32, n = 12) (
<figureCitation id="15992A448D472469E44418ADFF3D7418" captionStart="FIGURES 175–178" captionStartId="25.[152,270,1739,1763]" captionTargetBox="[151,1435,181,1716]" captionTargetId="figure-17@25.[151,1435,181,1716]" captionTargetPageId="25" captionText="FIGURES 175–178. Morphometric features of Therates spp: 175, 177—relation of the proportions EL/PL and HTbL/HTaL (green rhombus—T. pseudovitalisi sp. n.; blue circles—T. fruhstorferi; open symbols—males; solid symbols—females); 176, 178—relation of the proportions HTbL/HTaL (small quadrates—medians; rectangles—25–75 percentiles; whiskers—min–max; significantly different samples are marked with different lower letters, according to Mann–Whitney U-test, p &lt;0.05 (176—U = 112.0, z = 5.08, p = 0.000; 178—U = 23.5, z = 3.37, p = 0.0007)); 175, 176—all specimens; 177, 178—males." figureDoi="http://doi.org/10.5281/zenodo.7758910" httpUri="https://zenodo.org/record/7758910/files/figure.png" pageId="7" pageNumber="408">Figs 175–178</figureCitation>
).
</paragraph>
<paragraph id="8D1D36C18D472469E1E81B55FCE974F0" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="408">
Elytra shining black-blue with purple-violet or blue-violet reflection; indistinctly divergent toward apex, EL/ EW = 1.92–2.06 (mean = 1.98, n = 12); punctuation deep and regular between basal humps and central dot, in apical third shallow and sparse; scutellum black with blue or violet tinge; apical margin shortly cut. Elytral pattern presented by well visible pale-yellow small basal portion of humeral lunule (see in front) and elongate slightly or clearly oblique narrow wide-uniformed or slightly dilated in lateral portion central dot; basal dot very small, in some specimens indistinct or entirely absent (
<figureCitation id="15992A448D472469E37C1BA1FD0674F0" box="[595,710,727,753]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="7" pageNumber="408">Figs 9, 10</figureCitation>
, 
<figureCitation id="15992A448D472469E3FD1BA1FCDD74F0" box="[722,797,727,753]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="7" pageNumber="408">90–93</figureCitation>
).
</paragraph>
<paragraph id="8D1D36C18D472469E1E81B8DFAD77538" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="408">
Aedeagus gradually sinuate with elongate rounded apex, lacking lateral carinae (
<figureCitation id="15992A448D472469E57C1B8DFB327514" box="[1107,1266,763,789]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="7" pageNumber="408">Figs 132–134</figureCitation>
); AL = 
<quantity id="4A5A9B248D472469E4671B8CFF037538" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="7" pageNumber="408" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.5–2.7 mm</quantity>
(mean = 2.62, n = 5), EL/AL = 2.56–2.72 (mean = 2.6, n = 5), in one aberrant male EL/AL = 2.45.
</paragraph>
</subSubSection>
<subSubSection id="C5B8654A8D472469E1E81A35FEB17580" pageId="7" pageNumber="408" type="etymology">
<paragraph id="8D1D36C18D472469E1E81A35FEB17580" blockId="7.[151,1437,151,933]" pageId="7" pageNumber="408">
<emphasis id="BFD6EAD38D472469E1E81A35FE8E755C" bold="true" box="[199,334,835,861]" pageId="7" pageNumber="408">Etymology.</emphasis>
The new species is named “ 
<taxonomicName id="4AA24D428D472469E38B1A35FC80755C" authorityName="Matalin et Wiesner" authorityYear="2023" box="[676,832,835,861]" class="Insecta" family="Cicindelidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="pseudovitalisi">
<emphasis id="BFD6EAD38D472469E38B1A35FC80755C" box="[676,832,835,861]" italics="true" pageId="7" pageNumber="408">pseudovitalisi</emphasis>
</taxonomicName>
” due to it very similar outward appearance with 
<taxonomicName id="4AA24D428D472469E4A71A35FEAD7580" authority="Horn, 1913" authorityName="W. Horn" authorityYear="1913" class="Insecta" family="Cicindelidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="408" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis id="BFD6EAD38D472469E4A71A35FF237580" italics="true" pageId="7" pageNumber="408">T. vitalisi</emphasis>
<bibRefCitation id="E9334B308D472469E1C51A11FEAD7580" author="Horn, W." box="[234,365,871,897]" pageId="7" pageNumber="408" pagination="362 - 366" refId="ref25772" refString="Horn, W. (1913) Materiaux pour servir a l'etude de la faune entomologique de l'Indochine. Cicindelinae. Annales de la Societe Entomologique de Belgique, 57, 362 - 366. https: // doi. org / 10.5962 / bhl. part. 4597" type="journal article" year="1913">Horn, 1913</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C5B8654A8D472469E1E81AFCFB2875A4" box="[199,1256,906,933]" pageId="7" pageNumber="408" type="distribution">
<paragraph id="8D1D36C18D472469E1E81AFCFB2875A4" blockId="7.[151,1437,151,933]" box="[199,1256,906,933]" pageId="7" pageNumber="408">
<emphasis id="BFD6EAD38D472469E1E81AFCFEA175A5" bold="true" box="[199,353,906,932]" pageId="7" pageNumber="408">Distribution.</emphasis>
<collectingCountry id="F5B576518D472469E0481AFDFE6675A4" box="[359,422,907,933]" name="Laos" pageId="7" pageNumber="408">LAOS</collectingCountry>
: 
<collectingRegion id="4F66F8238D472469E09F1AFDFDEE75A4" box="[432,558,907,933]" country="Laos" name="Oudomxai" pageId="7" pageNumber="408">Oudomxay</collectingRegion>
, 
<collectingRegion id="4F66F8238D472469E3171AFDFD2675A4" box="[568,742,907,933]" country="Laos" name="Louangphabang" pageId="7" pageNumber="408">Luang Prabang</collectingRegion>
, 
<collectingRegion id="4F66F8238D472469E3DE1AFDFCB575A4" box="[753,885,907,933]" country="Laos" name="Houaphan" pageId="7" pageNumber="408">Houaphanh</collectingRegion>
, 
<collectingRegion id="4F66F8238D472469E2AF1AFDFC3575A4" box="[896,1013,907,933]" country="Laos" name="Phongsali" pageId="7" pageNumber="408">Phongsaly</collectingRegion>
Provinces (
<figureCitation id="15992A448D472469E5561AFDFB1C75A4" box="[1145,1244,907,933]" captionStart="FIGURES 184" captionStartId="28.[152,270,1579,1603]" captionTargetBox="[151,1435,597,1556]" captionTargetId="figure-226@28.[151,1435,597,1556]" captionTargetPageId="28" captionText="FIGURES 184. Distribution of Therates spp.: dark blue solid circle—T. vitalisi vitalisi; dark blue half-solid circle—T. vitalisi ida; lilac solid circle—T. hunnanensis sp. n.; orange solid circle—T. biserratus; pink solid circle—T. motoensis; red open circle—T. sauteri; red solid quadrat—T. fruhstorferi fruhstorferi; red half-solid quadrat—T. fruhstorferi australis ssp. n.; blue solid quadrat—T. ziyardamensis sp. n.; green solid quadrat—T. pseudovitalisi sp. n. (nominotypical subspecies); green halfsolid quadrat—T. pseudovitalisi tibetana ssp. n.; L in the symbols—according only to the literature sources. Map source— https://commons.wikimedia.org/wiki/File:East_Asia_topographic_map.png." figureDoi="http://doi.org/10.5281/zenodo.7758916" httpUri="https://zenodo.org/record/7758916/files/figure.png" pageId="7" pageNumber="408">Fig. 184</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>