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<document ID-CLB-Dataset="222069" ID-DOI="http://dx.doi.org/10.3897/fr.26.e93456" ID-GBIF-Dataset="6483d9f1-27f8-4501-9c34-75dbe4d39a05" ID-Pensoft-Pub="2193-0074-1-1" ID-Pensoft-UUID="31E88D0E24BE5C97B7D3DF99135E1514" ID-ZooBank="F8C273F5D7C54A5CBF0A56C7C3085D55" ModsDocID="2193-0074-26-1-1" checkinTime="1673408644191" checkinUser="pensoft" docAuthor="Manitkoon, Sita, Deesri, Uthumporn, Warapeang, Prapasiri, Nonsrirach, Thanit &amp; Chanthasit, Phornphen" docDate="2023" docId="04FB58BC51415E6292C05A8155902DA4" docLanguage="en" docName="FossilRecord 26(1): 1-25" docOrigin="Fossil Record 26 (1)" docPubDate="2023-01-10" docSource="http://dx.doi.org/10.3897/fr.26.e93456" docTitle="Ratchasimasaurus suranareae Basal neornithischian indet. ( Khok Sanam neornithischian)" docType="treatment" docVersion="5" id="31E88D0E24BE5C97B7D3DF99135E1514" lastPageNumber="1" masterDocId="31E88D0E24BE5C97B7D3DF99135E1514" masterDocTitle="Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications" masterLastPageNumber="25" masterPageNumber="1" pageNumber="1" updateTime="1732690605966" updateUser="ExternalLinkService">
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<mods:title id="02F1192A3B787F9B77FE197059A4CB17">Ornithischian dinosaurs in Southeast Asia: a review with palaeobiogeographic implications</mods:title>
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<mods:namePart id="EA3158C56DB457FD6686F4AD048DD5C2">Manitkoon, Sita</mods:namePart>
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<mods:namePart id="2098EB0F07CEF4B1C03A57E811CD318A">Deesri, Uthumporn</mods:namePart>
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<mods:affiliation id="144763C0D546E4D8BABF1E58283C71B0">Faculty of Science, Mahasarakham University, Khamriang, Mahasarakham, 44150, Thailand</mods:affiliation>
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<mods:namePart id="1066056355734CC80BCAA4B0E0D2E057">Warapeang, Prapasiri</mods:namePart>
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<mods:affiliation id="24D44BD5F90E13D400A5048966F1CCA9">Faculty of Science, Mahasarakham University, Khamriang, Mahasarakham, 44150, Thailand</mods:affiliation>
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<mods:namePart id="D0F738ACA1B1095AF7556E0B53FD0E4B">Nonsrirach, Thanit</mods:namePart>
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<mods:namePart id="812E734A3277D8AB450D8AAFA0761E34">Chanthasit, Phornphen</mods:namePart>
<mods:affiliation id="E193ECAA832E0EE0431C005F097D93BD">Palaeontological Research and Education Centre, Mahasarakham University, Khamriang, Mahasarakham, 44150, Thailand</mods:affiliation>
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<treatment id="04FB58BC51415E6292C05A8155902DA4" ID-GBIF-Taxon="208787392" LSID="urn:lsid:plazi:treatment:04FB58BC51415E6292C05A8155902DA4" httpUri="http://treatment.plazi.org/id/04FB58BC51415E6292C05A8155902DA4" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Stegosauridae">
<subSubSection id="605123F49C70D17D9F19515300C303E4" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="5BBC08D52B7057C88AFA69226E22191B" pageId="0" pageNumber="1">
<taxonomicName id="F9AA01654335A8F747B570086CEB7498" LSID="04FB58BC-5141-5E62-92C0-5A8155902DA4" authority="Basal neornithischian indet. (' Khok Sanam neornithischian')" authorityName="Basal neornithischian indet. ( Khok Sanam neornithischian)" family="Hadrosauridae" genus="Ratchasimasaurus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Ratchasimasaurus suranareae" order="Ornithischia" pageId="0" pageNumber="1" rank="species" species="suranareae">
Ratchasimasaurus suranareae Basal neornithischian indet. ('Khok Sanam
<normalizedToken id="20D24088C58D9871711644B7AD004421" originalValue="neornithischian">neornithischian'</normalizedToken>
)
</taxonomicName>
</paragraph>
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<subSubSection id="CD12F4310A891DE38A27DE6E5F28E978" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="3A42C1BE0BEDE3436EDF34ABD92A1772" pageId="0" pageNumber="1">Material.</paragraph>
<paragraph id="8768E32B442F5403591A0F8F4177F4E9" pageId="0" pageNumber="1">
WNM-Ks-001, an isolated tooth (Fig.
<figureCitation id="A0E63B4329C03DB3B145DB5D04A2DECB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Basal neornithischian specimens from the lower Phu Kradung Formation. (left) Left dentary (PRC 149) from Phu Noi locality, in lateral (A) and medial (B) views; (right) isolated tooth (WNM-Ks- 001) from Khok Sanam locality, in labial (if dentary) (C), lingual (D) and mesial / distal (E) views. Scale bars: 5 cm (A, B); 1 cm (C-E)." figureDoi="10.3897/fr.26.e93456.figure4" httpUri="https://binary.pensoft.net/fig/793762" pageId="0" pageNumber="1">4C-E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="10E889AABD43D0A6E411832F03A71885" pageId="0" pageNumber="1" type="locality and age">
<paragraph id="9D8280874498C9AAB5B54618F6147879" pageId="0" pageNumber="1">Locality and age.</paragraph>
<paragraph id="EB7560E6944ABC44C068C4CDD5FA7D6D" pageId="0" pageNumber="1">Khok Sanam locality, Kham Muang District, Kalasin Province; the lower Phu Kradung Formation, Late Jurassic.</paragraph>
<caption id="871158825FB3C294283418B9CF226050" doi="10.3897/fr.26.e93456.figure4" httpUri="https://binary.pensoft.net/fig/793762" pageId="0" pageNumber="1" start="Figure 4" startId="F4">
<paragraph id="AADBC03D7CDAF1B9846B84BB8162ED01" pageId="0" pageNumber="1">
<emphasis id="0FB0ABB2E5C3A96E87833ABC6F2AD14B" bold="true" pageId="0" pageNumber="1">Figure 4.</emphasis>
Basal neornithischian specimens from the lower Phu Kradung Formation. (left) Left dentary (PRC 149) from Phu Noi locality, in lateral (
<emphasis id="146FD2C14FFC8F217881653B10BB507A" bold="true" pageId="0" pageNumber="1">A</emphasis>
) and medial (
<emphasis id="7A2D0746E041F9B53E34E6277318881F" bold="true" pageId="0" pageNumber="1">B</emphasis>
) views; (right) isolated tooth (WNM-Ks-001) from Khok Sanam locality, in labial (if dentary) (
<emphasis id="638F5670677DD8AFF7D3F8AD37A596FD" bold="true" pageId="0" pageNumber="1">C</emphasis>
), lingual (
<emphasis id="A3E673071BEDA2887CD92E19868B1A34" bold="true" pageId="0" pageNumber="1">D</emphasis>
) and mesial/distal (
<emphasis id="D754CB8178D5587B7DE55194D942F2EC" bold="true" pageId="0" pageNumber="1">E</emphasis>
) views. Scale bars: 5 cm (
<emphasis id="2753E0880F2DCEF7F2B1FDA688AB71D4" bold="true" pageId="0" pageNumber="1">A, B</emphasis>
); 1 cm (
<emphasis id="B727575BDCFE4A4ED5BA4D9B0E9917A2" bold="true" pageId="0" pageNumber="1">C-E</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="EFD36AF2F7781886EE6E703F44DF0715" pageId="0" pageNumber="1" type="comments">
<paragraph id="679B2DDE1200525CF2E364964E5F9A2E" pageId="0" pageNumber="1">Comments.</paragraph>
<paragraph id="ED318637B2BB56D79A81D48040A049A3" pageId="0" pageNumber="1">
A dentary tooth shows the fan-shaped crown, and has the characters of the posterior teeth present in the dentary teeth of PRC 149 (
<bibRefCitation id="5311EC1CB34EE6015313D46A16E13ABE" author="Buffetaut, E" journalOrPublisher="Journal of Science and Technology Mahasarakham University" pageId="0" pageNumber="1" pagination="344 - 347" refId="B32" refString="Buffetaut, E, Suteethorn, S, Suteethorn, V, Deesri, U, Tong, H, 2014. Preliminary note on a small ornithopod dinosaur from the Phu Kradung Formation (terminal Jurassic - basal Cretaceous) of Phu Noi, north-eastern Thailand. Journal of Science and Technology Mahasarakham University 33: 344 - 347" title="Preliminary note on a small ornithopod dinosaur from the Phu Kradung Formation (terminal Jurassic - basal Cretaceous) of Phu Noi, north-eastern Thailand." volume="33" year="2014">Buffetaut et al. 2014</bibRefCitation>
) and of
<taxonomicName id="8B6751C77CC2944D24F0E2EE28D3B732" authorityName="Marsh" authorityYear="1877" class="Reptilia" family="Fabrosauridae" genus="Nanosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nanosaurus agilis" order="Ornithischia" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="agilis">
<emphasis id="F3ADBA9002E8F8AD2A2F4A7BF1BE5521" italics="true" pageId="0" pageNumber="1">Nanosaurus agilis</emphasis>
</taxonomicName>
(
<bibRefCitation id="E32116BA2227150D995260FF650E7D95" DOI="https://doi.org/10.31711/giw.v5.pp167-207" author="Carpenter, K" journalOrPublisher="Geology of the Intermountain West" pageId="0" pageNumber="1" pagination="167 - 207" refId="B40" refString="Carpenter, K, Galton, PM, 2018. A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA. Geology of the Intermountain West 5: 167 - 207, DOI: https://doi.org/10.31711/giw.v5.pp167-207" title="A photo documentation of bipedal ornithischian dinosaurs from the Upper Jurassic Morrison Formation, USA." url="https://doi.org/10.31711/giw.v5.pp167-207" volume="5" year="2018">Carpenter and Galton 2018</bibRefCitation>
). Although the enamelled surface is thin, a well-developed ornamentation of ridges is still noticeable on the lingual side as the enamel is asymmetrically distributed. The margin of the laterally compressed crowns bears distinct denticles. The labial side of the tooth bears no enamel and shows worn facets. The ornamented lingual side does not show a prominent median ridge, contrasting with the European Early Cretaceous
<taxonomicName id="2E75DA1D2B178880FFCD8DCAB6195D18" baseAuthorityName="Huxley" baseAuthorityYear="1870" class="Reptilia" family="Hypsilophodontidae" genus="Hypsilophodon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Hypsilophodon foxii" order="Ornithischia" pageId="0" pageNumber="1" phylum="Chordata" rank="species" species="foxii">
<emphasis id="D0BB96B13DFDF7B4A2F5A81E16ED866A" italics="true" pageId="0" pageNumber="1">Hypsilophodon foxii</emphasis>
</taxonomicName>
(
<bibRefCitation id="43A6E2E1ACB4312DE4A5106D912E24C3" author="Galton, PM" journalOrPublisher="Geneve" pageId="0" pageNumber="1" pagination="211 - 273" refId="B56" refString="Galton, PM, 2009. Notes on Neocomian (Lower Cretaceous) ornithopod dinosaurs from England - Hypsilophodon, Valdosaurus, &quot; Camptosaurus &quot;, &quot; Iguanodon &quot; - and referred specimens from Romania and elsewhere. Geneve 28: 211 - 273" title="Notes on Neocomian (Lower Cretaceous) ornithopod dinosaurs from England - Hypsilophodon, Valdosaurus, &quot; Camptosaurus &quot;, &quot; Iguanodon &quot; - and referred specimens from Romania and elsewhere." volume="28" year="2009">Galton 2009</bibRefCitation>
) and other derived neornithischians. There is moderately developed labiolingual expansion (
<normalizedToken id="F11A80CDC0481007DB6FEE8000AD742F" originalValue="cingulum">'cingulum'</normalizedToken>
) at the base of the crown, as seen in basal neornithischians (
<bibRefCitation id="FCCDB8CC255647802472E8A677A497EE" DOI="https://doi.org/10.1671/0272-4634(2005)025[0823:SODFTM]2.0.CO;2" author="Barrett, PM" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="1" refId="B10" refString="Barrett, PM, Butler, RJ, Knoll, F, 2005. Small-Bodied Ornithischian Dinosaurs from the Middle Jurassic of Sichuan, China. Source: Journal of Vertebrate Paleontology Journal of Vertebrate Paleontology 25: 823-834. https://doi.org/10.1671/0272-4634(2005)025[0823:SODFTM]2.0.CO;2" title="Small-Bodied Ornithischian Dinosaurs from the Middle Jurassic of Sichuan, China. Source: Journal of Vertebrate Paleontology Journal of Vertebrate Paleontology 25: 823 - 834." url="https://doi.org/10.1671/0272-4634(2005)025[0823:SODFTM]2.0.CO;2" year="2005">Barrett et al. 2005</bibRefCitation>
). The cylindrical root has been preserved and curves to the labial side.
</paragraph>
</subSubSection>
</treatment>
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