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<document id="F8A7268030341AE8D51F44D3D540D649" ID-CLB-Dataset="35486" ID-DOI="http://dx.doi.org/10.3897/zookeys.205.3491" ID-GBIF-Dataset="16f08073-854b-40a2-aef9-72a97c85c8bd" ID-PMC="PMC3391731" ID-Pensoft-Pub="1313-2970-205-59" ID-PubMed="22792035" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-205-59" ModsDocOrigin="ZooKeys 205" ModsDocTitle="A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae, Gnaphosidae)" checkinTime="1451248915768" checkinUser="pensoft" docAuthor="Kovblyuk, Mykola M., Kastrygina, Zoya A. &amp; Omelko, Mikhail M." docDate="2012" docId="04B5F2495FB25E5EFEECAC5A70785D74" docLanguage="en" docName="ZooKeys 205: 59-89" docOrigin="ZooKeys 205" docSource="http://dx.doi.org/10.3897/zookeys.205.3491" docTitle="Haplodrassus bohemicus Miller &amp; Buchar 1977" docType="treatment" docVersion="5" lastPageNumber="63" masterDocId="FFC2BD6FFFD62C69FFC3EC55350AFFA3" masterDocTitle="A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae, Gnaphosidae)" masterLastPageNumber="89" masterPageNumber="59" pageNumber="61" updateTime="1732937868572" updateUser="ExternalLinkService">
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<mods:title id="A5BFFBA82A8127C726FAA7A9CFC0C7E9">A review of the spider genus Haplodrassus Chamberlin, 1922 in Crimea (Ukraine) and adjacent areas (Araneae, Gnaphosidae)</mods:title>
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<mods:namePart id="209F9B2B878317D6951F25D83C41A531">Kovblyuk, Mykola M.</mods:namePart>
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<mods:namePart id="5CF0EBFB3D63D82B82F08A4484F5051C">Kastrygina, Zoya A.</mods:namePart>
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<mods:namePart id="8C874B7B8A12AA15CD94A9AD542B403A">Omelko, Mikhail M.</mods:namePart>
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<treatment id="04B5F2495FB25E5EFEECAC5A70785D74" ID-GBIF-Taxon="152036254" LSID="urn:lsid:plazi:treatment:04B5F2495FB25E5EFEECAC5A70785D74" httpUri="http://treatment.plazi.org/id/04B5F2495FB25E5EFEECAC5A70785D74" lastPageId="4" lastPageNumber="63" pageId="2" pageNumber="61" scope_family="Gnaphosidae" scope_order="Araneae">
<subSubSection id="A2FC6DDBCFDC355792052032ACF0125E" pageId="2" pageNumber="61" type="nomenclature">
<paragraph id="31EC24291B939F6C9AE1C2FA1EC67984" pageId="2" pageNumber="61">
<taxonomicName id="87A2B41B81166D9A6C2D3C810C14C696" ID-CoL="3JMHG" LSID="http://species-id.net/wiki/Haplodrassus_bohemicus" authority="Miller &amp; Buchar, 1977" authorityName="Miller &amp; Buchar" authorityYear="1977" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus bohemicus" order="Araneae" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="bohemicus">Haplodrassus bohemicus Miller &amp; Buchar, 1977</taxonomicName>
Figs 1
<normalizedToken id="50C357D7248E3F7F2D6D21F0521DFADE" originalValue="–610–">-610-</normalizedToken>
12
</paragraph>
</subSubSection>
<subSubSection id="ED8F4F177500DFED3551479E73BAF691" pageId="2" pageNumber="61" type="reference_group">
<paragraph id="F9CD733E829EDCA4FED8CF56C458645A" pageId="2" pageNumber="61">
<taxonomicName id="F7EAF93CBB51BE0192FC65262BF614F7" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus bohemicus" order="Araneae" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="bohemicus">Haplodrassus bohemicus</taxonomicName>
<bibRefCitation id="000B058B11EF44B319AD767F69543705" author="Miller, F" journalOrPublisher="Acta Universitatis Carolinae - Biologica" pageId="13" pageNumber="72" pagination="157 - 171" title="Neue spinnenarten aus der gattung Zelotes Distel und Haplodrassus Chamberlin (Araneae, Gnaphosidae)." volume="6" year="1977">Miller and Buchar 1977</bibRefCitation>
: 163, pl. II, f. 1-6 (♂♀).
</paragraph>
<paragraph id="B64CF5C1E41CC5D035150D72A3602BF0" pageId="2" pageNumber="61">
<taxonomicName id="0CA1DCBAB3332A14FCD856C5571905B3" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus bohemicus" order="Araneae" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="bohemicus">Haplodrassus bohemicus</taxonomicName>
:
<bibRefCitation id="88B4A5DE2C6A716E87C61DA0C7283F76" author="Stefanovska, D" journalOrPublisher="Historia naturalis bulgarica" pageId="14" pageNumber="73" pagination="35 - 49" title="Spiders from the Scopje Region: a faunistic and zoogeographical analysis." volume="19" year="2008">Stefanovska et al. 2008</bibRefCitation>
: 37, f. 10-15 (♂♀).
</paragraph>
</subSubSection>
<subSubSection id="F2B1CC73B51B77E38902C9D0C22B0DF6" pageId="2" pageNumber="61" type="material">
<paragraph id="9207D40AF054F8C5C59FAD95D4B6E4C9" pageId="2" pageNumber="61">Material.</paragraph>
<paragraph id="5D1CEEFBF2890D4067AEB00E3C070D72" pageId="2" pageNumber="61">UKRAINE, CRIMEA: Saky Distr.: 12 ♂♂, 2 ♀♀ (TNU), near Pribrezhnaya railway station, 30.04.-24.06.2000, M.M. Kovblyuk.</paragraph>
</subSubSection>
<subSubSection id="C0F87FF61889A33473C2A1332E716D49" pageId="3" pageNumber="62" type="additional material">
<paragraph id="E5E463ACB69CA96DE5E7C4674E73D739" pageId="3" pageNumber="62">
<pageBreakToken id="95F62F8614DAEE253B0E5E985D8E8650" pageId="3" pageNumber="62" start="start">Additional</pageBreakToken>
material.
</paragraph>
<paragraph id="2A9CD2716DB8ECE36BEB54FB83C59E25" pageId="3" pageNumber="62">
UKRAINE. Nikolaev Area: 6 ♂♂ (TNU), Pervomaysky Distr., Kuripchane Vil., 5.05.-8.06.2006, N.Yu. Polchaninova. Kherson Area: 2 ♂♂, 2 ♀♀ (TNU), Henichesk Distr., Arabatskaya strelka, 4 km S Henichesk Town, 23.05.-10.06.2011, N.A. Stasyuk. Donetsk Area: 4 ♂♂ (TNU), Pershotravnevy Distr., Belosaraiskaya Kosa, 11-22.06.2001, E.V. Prokopenko; 2 ♂♂ (TNU), Novoazovsk Distr., Khomutovo Vil., &quot;Khomutovskaya Steppe&quot;,
<geoCoordinate id="5D138FFFD22CEF8A86C978F9AC94701E" direction="north" orientation="latitude" precision="925" value="47.266666">N47°16'</geoCoordinate>
,
<geoCoordinate id="59DB50C37197CB30EFC7B463FBA88B65" direction="east" orientation="longitude" precision="925" value="38.166668">E38°10'</geoCoordinate>
, 15-20.06.2004, N.Yu. Polchaninova; 1 ♂ (TNU), Slavyansky Distr., Svyatogorsk Town,
<geoCoordinate id="F19941505F9FA1A7CCF3BA43A5EC35FC" direction="north" orientation="latitude" precision="925" value="49.033333">N49°02'</geoCoordinate>
,
<geoCoordinate id="0D15D947D7D55CE20BD23DF1A945D5E2" direction="east" orientation="longitude" precision="925" value="37.65">E37°39'</geoCoordinate>
, 8-30.06.2004, N.Yu. Polchaninova. RUSSIA. Rostov Area: 7 ♂♂, 1 ♀ (TNU),
<normalizedToken id="51D96C7CF89B917367A03DCDFBDABACC" originalValue="Ust’">Ust'</normalizedToken>
-Donetsk Distr., near Razdorskaya Vil., &quot;Pukhlyakovskye sklony&quot;, 10.05.-28.06.2004, A.V. Ponomarev.
</paragraph>
</subSubSection>
<subSubSection id="DAEAD8F3A6F1240A0D46A2A56873DD62" pageId="3" pageNumber="62" type="comparative material">
<paragraph id="D56B608ACE37EB49D1351ABCAFFB7ED7" pageId="3" pageNumber="62">Comparative material.</paragraph>
<paragraph id="6F979C5F44D51978D516EEAF9EB646B2" pageId="3" pageNumber="62">
<taxonomicName id="162BDFCB661A7D34D3E5D0C2FEF76060" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pugnans" order="Araneae" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="pugnans">Haplodrassus pugnans</taxonomicName>
(Simon, 1880): RUSSIA, Magadan Area: 1 ♂, 3 ♀♀ (ISEA,
<normalizedToken id="2642F7DE3AFC756F5B46E220E46853C5" originalValue="БИ-">BI-</normalizedToken>
930),
<normalizedToken id="485D0E9CF97B6E948C1D670AA0776BF3" originalValue="Ten’kynskyi">Ten'kynskyi</normalizedToken>
Distr., env. Sibit-Tyellakh, Aborigen Field Station, Betula, 12-22.06.1983, A.V. Avershin.
</paragraph>
</subSubSection>
<subSubSection id="AA903B7F9727E4906D2DFEB89DE509C6" pageId="3" pageNumber="62" type="diagnosis">
<paragraph id="DBFF82F0CB1CF9915F8639262C9B5649" pageId="3" pageNumber="62">Diagnosis.</paragraph>
<paragraph id="549AC8DAA67372FDA18B8062F304FCA5" pageId="3" pageNumber="62">
<taxonomicName id="9F793FD67694567CDE4023FE29658C99" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus bohemicus" order="Araneae" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="bohemicus">Haplodrassus bohemicus</taxonomicName>
is most similar to
<taxonomicName id="ECAFAC05729E491319ECF1E30D9DA96A" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pugnans" order="Araneae" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="pugnans">Haplodrassus pugnans</taxonomicName>
(Simon, 1880),
<taxonomicName id="1837D2CAFA5D2AE9846A91F2091E2638" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus signifer" order="Araneae" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="signifer">Haplodrassus signifer</taxonomicName>
(C.L. Koch, 1839) and
<taxonomicName id="98E8959FC87A7B69BDEB5B437C8D2123" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pseudosignifer" order="Araneae" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="pseudosignifer">Haplodrassus pseudosignifer</taxonomicName>
Marusiket al. 1996, but differs: 1) by the shape of RTA having a
<normalizedToken id="3E24478745D843FB81C7C29BF24F4343" originalValue="“step”">&quot;step&quot;</normalizedToken>
-like dorsal margin (RTA dorsal margin without
<normalizedToken id="120B62E9C675479567FB790C54BFDE65" originalValue="“step”">&quot;step&quot;</normalizedToken>
in
<taxonomicName id="3C49A4FC4CDD42B7132F2FF406277A4D" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pugnans" order="Araneae" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="pugnans">Haplodrassus pugnans</taxonomicName>
,
<taxonomicName id="D469F84BA4457D7276302C8F57CB43A8" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus signifer" order="Araneae" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="signifer">Haplodrassus signifer</taxonomicName>
and
<taxonomicName id="3CA077FDE21047D2383F37FFF1C10698" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pseudosignifer" order="Araneae" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="pseudosignifer">Haplodrassus pseudosignifer</taxonomicName>
); 2) by the apically directed embolus lacking a tooth (retrolaterally directed embolus with a tooth in
<taxonomicName id="5A4986A7A13A4515CA9BC11C5734FC73" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pugnans" order="Araneae" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="pugnans">Haplodrassus pugnans</taxonomicName>
,
<taxonomicName id="FE735639119100460A7436CD915996C8" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus signifer" order="Araneae" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="signifer">Haplodrassus signifer</taxonomicName>
and
<taxonomicName id="CD7E3648F61A434CDD6A07B91654F481" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pseudosignifer" order="Araneae" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="pseudosignifer">Haplodrassus pseudosignifer</taxonomicName>
); 3) by the outlines of epigynal sclerites and relative proportions of the epigynal fovea (cf. Figs 10, 12 and 13, 75, 78).
</paragraph>
</subSubSection>
<subSubSection id="91B0486C49A8A15D237A369268CD20D8" lastPageId="4" lastPageNumber="63" pageId="3" pageNumber="62" type="description">
<paragraph id="9DC247B47EB1B1B94365CFEE05333923" pageId="3" pageNumber="62">Description.</paragraph>
<paragraph id="245D50258F4DB7BAD682CEB771BDD66B" pageId="3" pageNumber="62">Male measurements (n = 5). Total length 5.8-7.5 (6.7); carapace 2.7-3.2 (3.0) long, 2.0-2.6 (2.3) wide. Diameters of eyes and distances between them: AM 0.10-0.15 (0.12), AL 0.10-0.16 (0.13), PM 0.15-0.22 (0.19), PL 0.10-0.14 (0.12), AM-AM 0.09-0.12 (0.10), AM-AL 0.03-0.04 (0.04), PM-PM 0.03-0.04 (0.03), PM-PL 0.12-0.16 (0.14), AM-PM 0.12-0.18 (0.15), AL-PL 0.10-0.16 (0.14). Distances between anterior eyes and margin of clypeus: AM-clypeus 0.18-0.22 (0.20), AL-clypeus 0.10-0.18 (0.15).</paragraph>
<paragraph id="A986D35C526AB542390B3C33861957E0" pageId="3" pageNumber="62">Length of leg segments (male):</paragraph>
<paragraph id="F3092737CB39246E03E198943FCFEB20" pageId="4" pageNumber="63">
<table id="1CD25B9667CDD5ACB4B7D92CB85C6F4A" pageId="4" pageNumber="63">
<tr id="0D36E398C5124A34D5CCD9E3702CB4DB" pageId="4" pageNumber="63">
<th id="0CB9A3F8A4B5ACF851CBAFD450DCA38F" colspan="1" pageId="4" pageNumber="63" rowspan="1">
<pageBreakToken id="6336D80271C63D13FBCDDA17F9E997B4" pageId="4" pageNumber="63" start="start">Leg</pageBreakToken>
</th>
<th id="1D9DB0F7BCC0188878E57F08531C68EE" colspan="1" pageId="4" pageNumber="63" rowspan="1">Femur</th>
<th id="C99795E31CD1A58E0554ED8DDAE8CF8E" colspan="1" pageId="4" pageNumber="63" rowspan="1">Patella</th>
<th id="6D688AA48C2D595FB554CF73CC2D9341" colspan="1" pageId="4" pageNumber="63" rowspan="1">Tibia</th>
<th id="A7F39E707E827496AFE92E242F2332C8" colspan="1" pageId="4" pageNumber="63" rowspan="1">Metatarsus</th>
<th id="2A1CD4362B662BE281192BCC4098FFE5" colspan="1" pageId="4" pageNumber="63" rowspan="1">Tarsus</th>
<th id="7DB2DC9B99E4C79F14194290A4B7D836" colspan="1" pageId="4" pageNumber="63" rowspan="1">Total</th>
</tr>
</table>
</paragraph>
<paragraph id="895A54F2390398750A77A8CEC6D15311" pageId="4" pageNumber="63">
Length of palp segments: femur 1.0-1.2 (1.1), patella 0.4-0.5 (0.5), tibia 0.3-0.5 (0.4), tarsus 1.0-1.2 (1.0). Cheliceral teeth: anterior - 2 (little; proximal tooth connected with keel of cheliceral groove), posterior - 2. Abdomen 2.9-4.0 (3.6) long, 1.8-2.2 (2.0) wide. Scutum is absent. Basal segment of anterior (inferior) spinnerets 0.5-0.6 (0.6) long. Coloration light brown, as in most
<taxonomicName id="FC95660A7E591A4EDFBFCCC093A4FA83" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus" order="Araneae" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">Haplodrassus</taxonomicName>
species.
</paragraph>
<paragraph id="517B0B36855140EAF0425A50E62AE45D" pageId="4" pageNumber="63">
Palp as in Figs 1-6. RTA with a
<normalizedToken id="5BEF13EB6D720BAB330CD5C2FD07D4B8" originalValue="“step”">&quot;step&quot;</normalizedToken>
(St) on the promargin, terminal apophysis almost straight, without distinct ridge, embolus slightly twisted and lacks a tooth.
</paragraph>
<paragraph id="140689EAB8059B872C169C8585037D9D" pageId="4" pageNumber="63">
Female measurements (n = 2). Total length 7.8-8.0; carapace 2.7-3.5 long, 2.0-2.7 wide. Abdomen 4.3-5.1 long, 2.7-3.1 wide. Coloration light brown, as in most
<taxonomicName id="7621C7083B6ED401CD35468C4FC1746A" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus" order="Araneae" pageId="4" pageNumber="63" phylum="Arthropoda" rank="genus">Haplodrassus</taxonomicName>
species.
</paragraph>
<paragraph id="82EB0370CFAE9462901EE62DE023883B" pageId="4" pageNumber="63">Epigyne as in Figs 10-12. Lateral pockets long, sub-parallel, spermathecae globular. Fovea of epigyne without narrow longitudinal groove.</paragraph>
<caption id="EDF017A2979348D592E323D84519ECCD" pageId="4" pageNumber="63">
<paragraph id="63581B73FC417AEC5FB65D47AC679CF5" pageId="4" pageNumber="63">
Figures 1-9. Males of
<taxonomicName id="7E07A6D7221FC330E7DE67AC10083A3B" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus bohemicus" order="Araneae" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="bohemicus">Haplodrassus bohemicus</taxonomicName>
(1-3 from Rostov Area, 4-6 from Crimea) and
<taxonomicName id="C8B418C0A3F18E11742E867E3C455652" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pugnans" order="Araneae" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="pugnans">Haplodrassus pugnans</taxonomicName>
(7-9 from Magadan Area): 1, 4, 7 RTA, retrolateral view 2, 5, 8 palp, ventral view 3, 6, 9 bulbus, apical view. Abbreviations: Coconductor; Emembolus; Ettooth of embolus; Mamedian apophysis; Ri ridgeof terminal apophysis; RTA retrolateral tibial apophysis; St
<normalizedToken id="0359D8041021F64A0B1EC4424CD3A4F3" originalValue="“step”">&quot;step&quot;</normalizedToken>
-like keel of RTA; Taterminal apophysis.
</paragraph>
</caption>
<caption id="6EEF910C89213125CFE1E3D3C37A435F" pageId="4" pageNumber="63">
<paragraph id="83228A8FAC470DB532F81750B494C81A" pageId="4" pageNumber="63">
Figures 10-14. Females of
<taxonomicName id="FE80422D06DFBDACB0FE88AB294054B9" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus bohemicus" order="Araneae" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="bohemicus">Haplodrassus bohemicus</taxonomicName>
(10-11 from Rostov Area, 12 from Crimea) and
<taxonomicName id="DFED5803972C87CA314EBAC800D195B9" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus pugnans" order="Araneae" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="pugnans">Haplodrassus pugnans</taxonomicName>
(13-14 from Magadan Area): 10, 12-13 epigyne, ventral view 11, 14 epigyne, dorsal view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="0ED157AED562983B35C2CC5F5CD88174" pageId="4" pageNumber="63" type="distribution">
<paragraph id="E6EE149EECEFBA70F4DEBB3DB46E3D92" pageId="4" pageNumber="63">Distribution.</paragraph>
<paragraph id="921C07812F0391608C66371B93377943" pageId="4" pageNumber="63">
Czech Republic, Macedonia, Greece, Ukraine (Nikolaev, Kherson, Donetsk Areas, Crimea), Russia (Rostov and Stavropol Areas, Kalmykya, Dagestan) (
<bibRefCitation id="94FD19C0734F8B0B517211045917195C" author="Miller, F" journalOrPublisher="Acta Universitatis Carolinae - Biologica" pageId="13" pageNumber="72" pagination="157 - 171" title="Neue spinnenarten aus der gattung Zelotes Distel und Haplodrassus Chamberlin (Araneae, Gnaphosidae)." volume="6" year="1977">Miller and Buchar 1977</bibRefCitation>
;
<bibRefCitation id="33AB8C125BEE0C5C82F1944F8F955338" author="Ponomarev, AV" journalOrPublisher="Caucasian Entomological Bulletin" pageId="14" pageNumber="73" pagination="5 - 13" title="New and rare spiders of family Gnaphosidae (Aranei) from a southeast of Europe." volume="2" year="2006">Ponomarev and Tsvetkov 2006</bibRefCitation>
;
<bibRefCitation id="9352E51EB18A925DE39D5632A9FF937C" author="Stefanovska, D" journalOrPublisher="Historia naturalis bulgarica" pageId="14" pageNumber="73" pagination="35 - 49" title="Spiders from the Scopje Region: a faunistic and zoogeographical analysis." volume="19" year="2008">Stefanovska et al. 2008</bibRefCitation>
;
<bibRefCitation id="BDC2153C4CB2C5BB006948E24534CDA1" author="Ponomarev, AV" journalOrPublisher="Ekologia zhyvotnih" pageId="14" pageNumber="73" pagination="126 - 143" title="Spiders (Arachnida, Aranei) coastal and islands territories of Northern Dagestan. Yug Rossii: ekologia, razvitie." volume="4" year="2011">Ponomarev et al. 2011</bibRefCitation>
; present data).
</paragraph>
</subSubSection>
<subSubSection id="EBF63C3C4164BDA228611F503C2C115E" pageId="4" pageNumber="63" type="comments">
<paragraph id="BC8BBE4B41CAEEEE5AFA4E6E262D82DF" pageId="4" pageNumber="63">Comments.</paragraph>
<paragraph id="3B2C94E86D2E84AAB1DD391813BA12D1" pageId="4" pageNumber="63">
<taxonomicName id="DDFA08F000231EE90B20537C9F359343" class="Arachnida" family="Gnaphosidae" genus="Haplodrassus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Haplodrassus bohemicus" order="Araneae" pageId="4" pageNumber="63" phylum="Arthropoda" rank="species" species="bohemicus">Haplodrassus bohemicus</taxonomicName>
is a new species record for the Crimean fauna.
</paragraph>
</subSubSection>
<subSubSection id="DDD842B69AD734F1F8F54B2EA62E065D" pageId="4" pageNumber="63" type="habitats">
<paragraph id="8EBC5D3D12AA97B83BCF80BB406F43E6" pageId="4" pageNumber="63">Habitats.</paragraph>
<paragraph id="9CEAE97603EED1BA4BFA5FDCC9B852A2" pageId="4" pageNumber="63">Meadows, steppes and sand dunes.</paragraph>
</subSubSection>
<subSubSection id="54220E9E98D8611BB7A3633B132F1284" pageId="4" pageNumber="63" type="phenology">
<paragraph id="55CAB4AE3AB209295A59D0C8118314AF" pageId="4" pageNumber="63">Phenology.</paragraph>
<paragraph id="630665D72234B822B3037BA89CC6BE54" pageId="4" pageNumber="63">♂♀ - V-VI. In Crimea the peak activity of adults occurs in May.</paragraph>
</subSubSection>
</treatment>
</document>
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