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<document id="FAD818E07E2DB839BF5C669F5D76A3ED" ID-ISSN="0024-4082" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630454141863" checkinUser="felipe" docAuthor="Sak, Serdar, Huys, Rony &amp; Karaytuğ, Süphan" docDate="2008" docId="047B2A1AC378967A56ABBFB4C3DBFDC2" docLanguage="en" docName="j.1096-3642.2007.00373.x.pdf" docOrigin="Zoological Journal of the Linnean Society 152" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Psammoleptastacus barani Sak &amp; Huys &amp; Karaytuğ 2008, SP. NOV." docType="treatment" docVersion="2" lastPageNumber="433" masterDocId="F8425262C3699662552DBB3BC733FF90" masterDocTitle="Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae)" masterLastPageNumber="458" masterPageNumber="409" pageNumber="426" updateTime="1699276690232" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="16601D191E3B69E9421B0EC6F8024627" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F6837888F7CC2B0CF27C77EB84007A0E">
<mods:title id="CB8401274E1437234CBBF0C18BCFB52A">Disentangling the subgeneric division of Arenopontia Kunz, 1937: resurrection of Psammoleptastacus Pennak, 1942, re-examination of Neoleptastacus spinicaudatus Nicholls, 1945, and proposal of two new genera and a new generic classification (Copepoda, Harpacticoida, Arenopontiidae)</mods:title>
</mods:titleInfo>
<mods:name id="CF8D6686CE6E5B770DB9DE50C08C396E" type="personal">
<mods:role id="2F766FD91CFA60467FC98EE7B49E9B5A">
<mods:roleTerm id="DAD671E6C04C00D7D83BA377B207CD96">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0CC9FD246E8E1B5CCD397B4241DBBF8B">Sak, Serdar</mods:namePart>
</mods:name>
<mods:name id="4D46B5ACF69F34CC1DEFD327769903B9" type="personal">
<mods:role id="20F51FDF000408266103F6FCCF5701EB">
<mods:roleTerm id="9BCFEDC2A4BC9BBC0FDE78B295F5F07E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="910C02867848427D6F4C5E3DEEBC2F4C">Huys, Rony</mods:namePart>
</mods:name>
<mods:name id="2917F2C2E77A0255A8681C7C6C35A60A" type="personal">
<mods:role id="3C984147FCDD142A1EF28299DE09771D">
<mods:roleTerm id="F1C9F9977DCE94FCD464EE973E39AF30">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BF9D926E04850B347181C1151410991B">Karaytuğ, Süphan</mods:namePart>
</mods:name>
<mods:typeOfResource id="288233E833BB163362220B6E608504DD">text</mods:typeOfResource>
<mods:relatedItem id="34B6961E60AA799C4983E0BF2BFE1FB9" type="host">
<mods:titleInfo id="F26FAE62187CB8F4F751878719AA72B9">
<mods:title id="08664A5EB68DA391F03C06D9F8DE3C32">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="B57D89CCD8F383F98ECE2F5D292D9FF5">
<mods:date id="6E7EE07EE13C2119BBC74D9ACA27195B">2008</mods:date>
<mods:detail id="A941EBF7D75F63063F4D28431BF63155" type="volume">
<mods:number id="4AAFF0CDF542382BB696CA7E2A4BBA21">152</mods:number>
</mods:detail>
<mods:extent id="9B724580D9564B144BF3DF66D742B966" unit="page">
<mods:start id="9D3E49A3DF28343D69581135A049B8BD">409</mods:start>
<mods:end id="1913C96164FD1685BFC8F3E103176BF3">458</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C4DCD6C4D5B00049284180183D497B9C">journal article</mods:classification>
<mods:identifier id="6D3F335B268EFE36151308DD3D3EC9A1" type="ISSN">0024-4082</mods:identifier>
</mods:mods>
<treatment id="047B2A1AC378967A56ABBFB4C3DBFDC2" LSID="urn:lsid:plazi:treatment:047B2A1AC378967A56ABBFB4C3DBFDC2" httpUri="http://treatment.plazi.org/id/047B2A1AC378967A56ABBFB4C3DBFDC2" lastPageId="24" lastPageNumber="433" pageId="17" pageNumber="426">
<subSubSection id="C4C8C887C378967356ABBFB4C265FB34" box="[902,1366,1167,1188]" pageId="17" pageNumber="426" type="nomenclature">
<paragraph id="8C6D9B0CC378967356ABBFB4C265FB34" blockId="17.[902,1366,1167,1188]" box="[902,1366,1167,1188]" pageId="17" pageNumber="426">
<heading id="D7252C60C378967356ABBFB4C265FB34" box="[902,1366,1167,1188]" centered="true" fontSize="9" level="2" pageId="17" pageNumber="426" reason="2">
<taxonomicName id="4BD2E08FC378967356ABBFB4C3DFFB33" authority="Sak &amp; Huys &amp; Karaytuğ, 2008" authorityName="Sak &amp; Huys &amp; Karaytuğ" authorityYear="2008" box="[902,1260,1167,1188]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="426" phylum="Arthropoda" rank="species" species="barani" status="sp. nov.">PSAMMOLEPTASTACUS BARANI</taxonomicName>
<emphasis id="BEA6471EC378967351D9BFA9C265FB34" bold="true" box="[1268,1366,1167,1188]" pageId="17" pageNumber="426">
<taxonomicNameLabel id="A595FA65C378967351D9BFA9C265FB34" box="[1268,1366,1167,1188]" pageId="17" pageNumber="426" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C4C8C887C37896735617BF8EC4B2FB7A" pageId="17" pageNumber="426" type="reference_group">
<paragraph id="8C6D9B0CC37896735617BF8EC4B2FB7A" blockId="17.[826,1441,1205,1258]" pageId="17" pageNumber="426">
<taxonomicName id="4BD2E08FC37896735617BF8EC4B2FB7A" authority="Noodt, 1955 b sensu Marinov (1971)" authorityName="Noodt, 1955 b sensu Marinov" authorityYear="1971" class="Hexanauplia" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="17" pageNumber="426" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="BEA6471EC37896735617BF8EC32EFB5B" box="[826,1053,1205,1227]" italics="true" pageId="17" pageNumber="426">Arenopontia stygia</emphasis>
<bibRefCitation id="E843E6FDC3789673511EBF8EC3EBFB5B" author="Noodt W" box="[1075,1240,1205,1227]" pageId="17" pageNumber="426" pagination="86 - 109" refId="ref25760" refString="Noodt W. 1955 b. Harpacticiden (Crust. Cop.) aus dem Sandstrand der franzosischen Biscaya-Kuste. Kieler Meeresforschungen 11: 86 - 109." type="journal article" year="1955">Noodt, 1955b</bibRefCitation>
<emphasis id="BEA6471EC378967351C0BF8DC21FFB5B" box="[1261,1324,1206,1227]" italics="true" pageId="17" pageNumber="426">sensu</emphasis>
<bibRefCitation id="E843E6FDC3789673506CBF8EC4B2FB7A" author="Marinov TM" pageId="17" pageNumber="426" pagination="43 - 87" refId="ref25342" refString="Marinov TM. 1971. Kharpatikoidi ot b'lgarskoto kraibrezhnie na Cherno more [Harpacticoids of the Bulgarian Black Sea coast]. Izvestiya na Instituta Okeanografiya i Ribno Stopanstvo, Varna 11: 43 - 87. [In Bulgarian with English and Russian summaries]." type="journal article" year="1971">Marinov (1971)</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C4C8C887C378967A5617BE29C3DBFDC2" lastPageId="24" lastPageNumber="433" pageId="17" pageNumber="426" type="materials_examined">
<paragraph id="8C6D9B0CC37896735617BE29C213FAD6" blockId="17.[826,1441,1298,1350]" pageId="17" pageNumber="426">
<materialsCitation id="3CBA9151C37896735617BE29C22FFAD6" country="Turkey" county="Black Sea" location="Bosporus" municipality="Sahilkoy" pageId="17" pageNumber="426" specimenCount="1" stateProvince="Istanbul" typeStatus="holotype">
<emphasis id="BEA6471EC37896735617BE29C4EFFAB7" box="[826,988,1298,1319]" italics="true" pageId="17" pageNumber="426">
<typeStatus id="536925AEC37896735617BE29C45DFAB7" box="[826,878,1298,1319]" pageId="17" pageNumber="426">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F4C5DB9CC378967356C4BE29C309FAB8" box="[1001,1082,1298,1320]" name="Turkey" pageId="17" pageNumber="426">Turkey</collectingCountry>
,
<collectingCounty id="650CE380C3789673517FBE29C3E3FAB7" box="[1106,1232,1298,1320]" pageId="17" pageNumber="426">Black Sea</collectingCounty>
coast,
<collectingRegion id="4E1655EEC37896735017BE29C2AEFAB8" box="[1338,1437,1298,1320]" country="Turkey" name="Istanbul" pageId="17" pageNumber="426">Istanbul</collectingRegion>
,
<collectingMunicipality id="6C090176C37896735617BE0BC4ADFAD6" box="[826,926,1328,1350]" pageId="17" pageNumber="426">Sahilköy</collectingMunicipality>
(east of
<location id="890DCDD7C3789673512ABE0AC347FAD6" LSID="urn:lsid:plazi:treatment:047B2A1AC378967A56ABBFB4C3DBFDC2:890DCDD7C3789673512ABE0AC347FAD6" box="[1031,1140,1329,1350]" country="Turkey" county="Black Sea" municipality="Sahilkoy" name="Bosporus" pageId="17" pageNumber="426" stateProvince="Istanbul">Bosporus</location>
); sandy beach
</materialsCitation>
.
</paragraph>
<paragraph id="8C6D9B0CC37896735617BE55C2A4F98C" blockId="17.[826,1441,1390,1565]" pageId="17" pageNumber="426">
<emphasis id="BEA6471EC37896735617BE55C32DFA13" box="[826,1054,1390,1411]" italics="true" pageId="17" pageNumber="426">Material examined:</emphasis>
<materialsCitation id="3CBA9151C37896735107BE55C4DDFA33" collectionCode="BUZM" pageId="17" pageNumber="426" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus id="536925AEC37896735107BE55C3A3FA14" box="[1066,1168,1390,1412]" pageId="17" pageNumber="426" type="holotype">Holotype</typeStatus>
<specimenCount id="9AD45085C378967351B3BE55C380FA14" box="[1182,1203,1390,1412]" pageId="17" pageNumber="426" type="female">♀</specimenCount>
(dissected on eight slides) (
<collectionCode id="EAC303C9C378967356BBBEB6C4D4FA33" box="[918,999,1421,1443]" pageId="17" pageNumber="426">BUZM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3CBA9151C378967356D0BEB6C2A0F98C" collectingDate="2001-05-01" collectionCode="NHM" collectorName="S. Karaytug &amp; S. Sak" pageId="17" pageNumber="426" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<typeStatus id="536925AEC378967356D0BEB6C343FA32" box="[1021,1136,1421,1442]" pageId="17" pageNumber="426" type="paratype">Paratypes</typeStatus>
are
<specimenCount id="9AD45085C37896735186BEB6C3C0FA32" box="[1195,1267,1420,1442]" pageId="17" pageNumber="426" type="female">one ♀</specimenCount>
and
<specimenCount id="9AD45085C37896735019BEB6C24CFA32" box="[1332,1407,1420,1442]" pageId="17" pageNumber="426" type="male">one ♂</specimenCount>
in alcohol (
<collectionCode id="EAC303C9C378967356B3BE97C4ECFA51" box="[926,991,1452,1473]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="17" pageNumber="426" type="Herbarium">NHM</collectionCode>
reg. nos. 2006. 1966–1967), and
<specimenCount id="9AD45085C37896735055BE97C458FA70" pageId="17" pageNumber="426" type="male">two ♂♂</specimenCount>
dissected on two and seven slides, respectively (
<collectionCode id="EAC303C9C3789673566CBED2C4B1FA6E" box="[833,898,1513,1534]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="17" pageNumber="426" type="Herbarium">NHM</collectionCode>
reg. nos. 2006. 1968–1969); all collected at type locality; leg.
<collectorName id="2127FEDAC378967356FDBD3CC36FF98D" box="[976,1116,1543,1565]" pageId="17" pageNumber="426">S. Karaytuğ</collectorName>
and
<collectorName id="2127FEDAC378967351B6BD3CC3DAF98D" box="[1179,1257,1543,1565]" pageId="17" pageNumber="426">S. Sak</collectorName>
,
<date id="F86CBDCCC378967351D5BD3CC2A0F98C" box="[1272,1427,1543,1565]" pageId="17" pageNumber="426" value="2001-05-01">
<collectingDate id="E8284424C378967351D5BD3CC2A0F98C" box="[1272,1427,1543,1565]" pageId="17" pageNumber="426" value="2001-05-01">01 May 2001</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8C6D9B0CC37896735617BD7DC488F9CB" blockId="17.[826,1441,1606,1903]" box="[826,955,1606,1627]" pageId="17" pageNumber="426">
<emphasis id="BEA6471EC37896735617BD7DC488F9CB" box="[826,955,1606,1627]" italics="true" pageId="17" pageNumber="426">Description</emphasis>
</paragraph>
<paragraph id="8C6D9B0CC37896715617BD5FC6D1FE22" blockId="17.[826,1441,1606,1903]" lastBlockId="19.[164,779,198,1903]" lastPageId="19" lastPageNumber="428" pageId="17" pageNumber="426">
<emphasis id="BEA6471EC37896735617BD5FC4A7F9E9" box="[826,916,1636,1657]" italics="true" pageId="17" pageNumber="426">Female:</emphasis>
Total body length from tip of rostrum to posterior margin of caudal rami: 330–380
<emphasis id="BEA6471EC37896735050BDBAC2B8F908" bold="true" box="[1405,1419,1665,1688]" pageId="17" pageNumber="426">M</emphasis>
m (mean
<emphasis id="BEA6471EC378967356A5BD9BC4A5F927" bold="true" box="[904,918,1696,1719]" pageId="17" pageNumber="426">=</emphasis>
361
<emphasis id="BEA6471EC378967356E1BD9BC4E9F927" bold="true" box="[972,986,1696,1719]" pageId="17" pageNumber="426">M</emphasis>
m;
<emphasis id="BEA6471EC37896735128BD99C32AF927" box="[1029,1049,1698,1719]" italics="true" pageId="17" pageNumber="426">N</emphasis>
<emphasis id="BEA6471EC3789673510DBD9BC31DF927" bold="true" box="[1056,1070,1696,1719]" pageId="17" pageNumber="426">=</emphasis>
7). Maximum width measured at P5-bearing somite. Body: slender and cylindrical, without clear distinction between prosome and urosome (
<figureCitation id="14E98789C37896735684BDC6C32EF882" box="[937,1053,1789,1811]" captionStart="Figure 8" captionStartId="18.[144,223,1840,1859]" captionTargetBox="[232,1335,194,1808]" captionTargetId="figure-0@18.[231,1336,194,1810]" captionTargetPageId="18" captionText="Figure 8. Psammoleptastacus barani sp. nov. (♀). A, habitus, dorsal view. B, habitus, lateral view. C, anal somite and caudal rami, dorsal view. D, posterior part of penultimate somite, anal somite, and left caudal ramus, lateral view." pageId="17" pageNumber="426">Fig. 8A, B</figureCitation>
). Hyaline frills of thoracic somites weakly developed and crenulated; those of genital double-somite and free abdominal somites strongly developed, and consisting of rectangular digitate lappets (
<figureCitation id="14E98789C37A96715426BBFDC6B9FF4B" box="[267,394,198,220]" captionStart="Figure 8" captionStartId="18.[144,223,1840,1859]" captionTargetBox="[232,1335,194,1808]" captionTargetId="figure-0@18.[231,1336,194,1810]" captionTargetPageId="18" captionText="Figure 8. Psammoleptastacus barani sp. nov. (♀). A, habitus, dorsal view. B, habitus, lateral view. C, anal somite and caudal rami, dorsal view. D, posterior part of penultimate somite, anal somite, and left caudal ramus, lateral view." pageId="19" pageNumber="428">Figs 8A, B</figureCitation>
,
<figureCitation id="14E98789C37A967154B6BBFDC68EFF4B" box="[411,445,198,219]" captionStart="Figure 9" captionStartId="20.[144,223,1840,1859]" captionTargetBox="[204,1364,197,1805]" captionTargetId="figure-0@20.[204,1365,194,1810]" captionTargetPageId="20" captionText="Figure 9. Psammoleptastacus barani sp. nov. A, urosome ♀, ventral view. B, urosome ♂, ventral view." pageId="19" pageNumber="428">9A</figureCitation>
,
<figureCitation id="14E98789C37A967154E3BBFDC6C2FF4C" box="[462,497,198,220]" captionStart="Figure 8" captionStartId="18.[144,223,1840,1859]" captionTargetBox="[232,1335,194,1808]" captionTargetId="figure-0@18.[231,1336,194,1810]" captionTargetPageId="18" captionText="Figure 8. Psammoleptastacus barani sp. nov. (♀). A, habitus, dorsal view. B, habitus, lateral view. C, anal somite and caudal rami, dorsal view. D, posterior part of penultimate somite, anal somite, and left caudal ramus, lateral view." pageId="19" pageNumber="428">8D</figureCitation>
). Genital double-somite (
<figureCitation id="14E98789C37A96715581BBDFC631FF6B" box="[172,258,228,251]" captionStart="Figure 9" captionStartId="20.[144,223,1840,1859]" captionTargetBox="[204,1364,197,1805]" captionTargetId="figure-0@20.[204,1365,194,1810]" captionTargetPageId="20" captionText="Figure 9. Psammoleptastacus barani sp. nov. A, urosome ♀, ventral view. B, urosome ♂, ventral view." pageId="19" pageNumber="428">Fig. 9A</figureCitation>
): slightly longer than wide; without chitinous ribs marking original segmentation; with one middorsal, two lateral, and two ventral pores. Anal somite (
<figureCitation id="14E98789C37A96715581BA7BC614FEC6" box="[172,295,320,342]" captionStart="Figure 8" captionStartId="18.[144,223,1840,1859]" captionTargetBox="[232,1335,194,1808]" captionTargetId="figure-0@18.[231,1336,194,1810]" captionTargetPageId="18" captionText="Figure 8. Psammoleptastacus barani sp. nov. (♀). A, habitus, dorsal view. B, habitus, lateral view. C, anal somite and caudal rami, dorsal view. D, posterior part of penultimate somite, anal somite, and left caudal ramus, lateral view." pageId="19" pageNumber="428">Fig. 8C, D</figureCitation>
): with two dorsal and two lateral pores. Anal operculum: with minute spinules along free distal margin (
<figureCitation id="14E98789C37A96715476BA46C683FE04" box="[347,432,381,404]" captionStart="Figure 8" captionStartId="18.[144,223,1840,1859]" captionTargetBox="[232,1335,194,1808]" captionTargetId="figure-0@18.[231,1336,194,1810]" captionTargetPageId="18" captionText="Figure 8. Psammoleptastacus barani sp. nov. (♀). A, habitus, dorsal view. B, habitus, lateral view. C, anal somite and caudal rami, dorsal view. D, posterior part of penultimate somite, anal somite, and left caudal ramus, lateral view." pageId="19" pageNumber="428">Fig. 8C</figureCitation>
). Anus: positioned subterminally between caudal rami.
</paragraph>
<caption id="D8ADCB84C37B967055BDBC0BC25BF8F0" pageId="18" pageNumber="427" startId="18.[144,223,1840,1859]" targetBox="[232,1335,194,1808]" targetPageId="18">
<paragraph id="8C6D9B0CC37B967055BDBC0BC25BF8F0" blockId="18.[144,1421,1839,1889]" pageId="18" pageNumber="427">
<emphasis id="BEA6471EC37B967055BDBC0BC5ACF8D3" bold="true" box="[144,671,1840,1859]" pageId="18" pageNumber="427">
Figure 8.
<taxonomicName id="4BD2E08FC37B96705429BC0BC571F8D3" authority="Sak &amp; Huys &amp; Karaytuğ, 2008" authorityName="Sak &amp; Huys &amp; Karaytuğ" authorityYear="2008" box="[260,578,1840,1859]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="18" pageNumber="427" phylum="Arthropoda" rank="species" species="barani" status="sp. nov.">
<emphasis id="BEA6471EC37B96705429BC0BC571F8D3" bold="true" box="[260,578,1840,1859]" italics="true" pageId="18" pageNumber="427">Psammoleptastacus barani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A595FA65C37B96705765BC0BC5ACF8D3" box="[584,671,1840,1859]" pageId="18" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(♀). A, habitus, dorsal view. B, habitus, lateral view. C, anal somite and caudal rami, dorsal view. D, posterior part of penultimate somite, anal somite, and left caudal ramus, lateral view.
</paragraph>
</caption>
<paragraph id="8C6D9B0CC37A96715591BA80C6F7FD73" blockId="19.[164,779,198,1903]" pageId="19" pageNumber="428">
Caudal rami (
<figureCitation id="14E98789C37A96715454BA80C6CDFE40" box="[377,510,443,465]" captionStart="Figure 8" captionStartId="18.[144,223,1840,1859]" captionTargetBox="[232,1335,194,1808]" captionTargetId="figure-0@18.[231,1336,194,1810]" captionTargetPageId="18" captionText="Figure 8. Psammoleptastacus barani sp. nov. (♀). A, habitus, dorsal view. B, habitus, lateral view. C, anal somite and caudal rami, dorsal view. D, posterior part of penultimate somite, anal somite, and left caudal ramus, lateral view." pageId="19" pageNumber="428">Fig. 8C, D</figureCitation>
): approximately 2.75 longer than basal width, tapering posteriorly; with one pore dorsally, one near ventral proximal margin (
<figureCitation id="14E98789C37A96715581B92DC632FDBD" box="[172,257,534,557]" captionStart="Figure 9" captionStartId="20.[144,223,1840,1859]" captionTargetBox="[204,1364,197,1805]" captionTargetId="figure-0@20.[204,1365,194,1810]" captionTargetPageId="20" captionText="Figure 9. Psammoleptastacus barani sp. nov. A, urosome ♀, ventral view. B, urosome ♂, ventral view." pageId="19" pageNumber="428">Fig. 9A</figureCitation>
), and one laterally near outer spinules; outer distal corner produced into posteriorly directed recurved spinous process, accompanied by ventral spinular row at base; dorsomedial surface with posteriorly directed spinous process arising from base of seta VII. Armature: as in
<taxonomicName id="4BD2E08FC37A967154C6B98BC5B0FD54" authorityName="Pennak" authorityYear="1942" box="[491,643,687,709]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="19" pageNumber="428" phylum="Arthropoda" rank="species" species="arenaridus">
<emphasis id="BEA6471EC37A967154C6B98BC5B0FD54" box="[491,643,687,709]" italics="true" pageId="19" pageNumber="428">P. arenaridus</emphasis>
</taxonomicName>
, but with a more foliaceous seta VII.
</paragraph>
<paragraph id="8C6D9B0CC37A96715591B9D6C69CFCAF" blockId="19.[164,779,198,1903]" pageId="19" pageNumber="428">
Rostrum (
<figureCitation id="14E98789C37A96715416B9D6C6A6FC92" box="[315,405,749,771]" captionStart="Figure 8" captionStartId="18.[144,223,1840,1859]" captionTargetBox="[232,1335,194,1808]" captionTargetId="figure-0@18.[231,1336,194,1810]" captionTargetPageId="18" captionText="Figure 8. Psammoleptastacus barani sp. nov. (♀). A, habitus, dorsal view. B, habitus, lateral view. C, anal somite and caudal rami, dorsal view. D, posterior part of penultimate somite, anal somite, and left caudal ramus, lateral view." pageId="19" pageNumber="428">Figs 8A</figureCitation>
,
<figureCitation id="14E98789C37A96715483B9D6C6ECFC93" box="[430,479,749,771]" captionStart="Figure 10" captionStartId="21.[164,243,1840,1859]" captionTargetBox="[223,1383,196,1808]" captionTargetId="figure-0@21.[222,1384,194,1810]" captionTargetPageId="21" captionText="Figure 10. Psammoleptastacus barani sp. nov. (♀). A, rostrum and antennule, dorsal view. B, antennule, ventral view. C, antenna. D, free antennary endopod. E, genital field." pageId="19" pageNumber="428">10A</figureCitation>
): broadly subtriangular, tapering apically, with two delicate sensillae and one subapical ventral pore.
</paragraph>
<paragraph id="8C6D9B0CC37A96715591B873C51EFBC2" blockId="19.[164,779,198,1903]" pageId="19" pageNumber="428">
Antennule (
<figureCitation id="14E98789C37A96715469B873C6F5FCCE" box="[324,454,840,862]" captionStart="Figure 10" captionStartId="21.[164,243,1840,1859]" captionTargetBox="[223,1383,196,1808]" captionTargetId="figure-0@21.[222,1384,194,1810]" captionTargetPageId="21" captionText="Figure 10. Psammoleptastacus barani sp. nov. (♀). A, rostrum and antennule, dorsal view. B, antennule, ventral view. C, antenna. D, free antennary endopod. E, genital field." pageId="19" pageNumber="428">Fig. 10A, B</figureCitation>
): six-segmented. Segment 1: with one seta near anterodistal margin. Segment 2: longest, about 3.5 times longer than wide. Segment 4: with long aesthetasc (30-
<emphasis id="BEA6471EC37A967154EBB898C6E7FC2A" bold="true" box="[454,468,931,954]" pageId="19" pageNumber="428">M</emphasis>
m long) fused at base with seta. Distal segment: with seven naked setae (one of which spatulate), and apical acrothek consisting of short aesthetasc (13-
<emphasis id="BEA6471EC37A967154B8B8C4C690FB86" bold="true" box="[405,419,1023,1046]" pageId="19" pageNumber="428">M</emphasis>
m long) and two slender setae. Armature formula: 1-[1], 2-[7
<emphasis id="BEA6471EC37A96715737BF26C51BFBA4" bold="true" box="[538,552,1053,1076]" pageId="19" pageNumber="428">+</emphasis>
1 plumose], 3-[4], 4-[(1
<emphasis id="BEA6471EC37A967155F2BF07C7DEFBC3" bold="true" box="[223,237,1084,1107]" pageId="19" pageNumber="428">+</emphasis>
ae)], 5-[1], 6-[7
<emphasis id="BEA6471EC37A96715488BF07C680FBC3" bold="true" box="[421,435,1084,1107]" pageId="19" pageNumber="428">+</emphasis>
acrothek].
</paragraph>
<paragraph id="8C6D9B0CC37A96715591BF67C438FA33" blockId="19.[164,779,198,1903]" pageId="19" pageNumber="428">
Antenna (
<figureCitation id="14E98789C37A96715419BF67C68DFBE1" box="[308,446,1116,1138]" captionStart="Figure 10" captionStartId="21.[164,243,1840,1859]" captionTargetBox="[223,1383,196,1808]" captionTargetId="figure-0@21.[222,1384,194,1810]" captionTargetPageId="21" captionText="Figure 10. Psammoleptastacus barani sp. nov. (♀). A, rostrum and antennule, dorsal view. B, antennule, ventral view. C, antenna. D, free antennary endopod. E, genital field." pageId="19" pageNumber="428">Fig. 10C, D</figureCitation>
): coxa small, without ornamentation. Original segmentation of allobasis marked by partial transverse surface suture; with one spinular row along exopodal margin. Exopod: onesegmented, elongate, with long naked seta apically. Free endopod: with two spinular rows on anterior surface and finer spinules at outer distal corner; lateral armature consisting of two short spines; apical armature consisting of two spines and three geniculate setae, the longest of which with spinules around geniculation and fused basally to tiny accessory seta.
</paragraph>
<paragraph id="8C6D9B0CC37A96715591BE97C623FA70" blockId="19.[164,779,198,1903]" pageId="19" pageNumber="428">
Mandible, maxillule, maxilla, and maxilliped: as in
<taxonomicName id="4BD2E08FC37A96715589BEF0C63AFA71" authorityName="Cottarelli (1975) sensu Mitwally &amp; Montagna" authorityYear="2001" box="[164,265,1483,1505]" class="Hexanauplia" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="19" pageNumber="428" phylum="Arthropoda" rank="species" species="nesaie">
<emphasis id="BEA6471EC37A96715589BEF0C63AFA71" box="[164,265,1483,1505]" italics="true" pageId="19" pageNumber="428">A. nesaie</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8C6D9B0CC37A96715591BED1C268FE88" blockId="19.[164,779,198,1903]" lastBlockId="19.[826,1442,198,1168]" pageId="19" pageNumber="428">
P1 (
<figureCitation id="14E98789C37A967155D9BED1C665F990" box="[244,342,1514,1536]" captionStart="Figure 11" captionStartId="22.[144,223,1840,1859]" captionTargetBox="[189,1379,195,1808]" captionTargetId="figure-0@22.[188,1380,194,1810]" captionTargetPageId="22" captionText="Figure 11. Psammoleptastacus barani sp. nov. (♀). A, P1, anterior view. B, P2, anterior view. C, P3, anterior view. D, P4, anterior view." pageId="19" pageNumber="428">Fig. 11A</figureCitation>
): intercoxal sclerite long and rectangular. Praecoxa: subtriangular and naked. Coxa: without ornamentation. Basis: with spinular row near bases of endopod and exopod; anterior surface with a proximal pore and a small inner basal seta. Exopod: three-segmented; about 1.2 times longer than endopod; all segments with spinules along outer margin; exp-1 longest, with long unipinnate outer spine; exp-2 without outer element; exp-3 with two unipinnate outer spines and two geniculate apical setae. Endopod: two-segmented, not prehensile; enp-1 distinctly longer than exp-1, with a serrate seta at about two-thirds of the length of the inner margin, and several spinules along outer margin; enp-2 less than half the length of enp-1, with two geniculate setae (inner one about twice as long as outer).
</paragraph>
<paragraph id="8C6D9B0CC37A9671567EBA19C238FCB1" blockId="19.[826,1442,198,1168]" pageId="19" pageNumber="428">
P2–P4 (
<figureCitation id="14E98789C37A96715694BA19C30FFEA8" box="[953,1084,290,312]" captionStart="Figure 11" captionStartId="22.[144,223,1840,1859]" captionTargetBox="[189,1379,195,1808]" captionTargetId="figure-0@22.[188,1380,194,1810]" captionTargetPageId="22" captionText="Figure 11. Psammoleptastacus barani sp. nov. (♀). A, P1, anterior view. B, P2, anterior view. C, P3, anterior view. D, P4, anterior view." pageId="19" pageNumber="428">Fig. 11B–D</figureCitation>
): intercoxal sclerites naked. Praecoxae: small and naked. Coxae: rectangular and without ornamentation. Bases: smaller than coxae, with a spinular row near base of endopod (P3–P4); anterior surface with a proximal pore; outer basal seta absent (P2), plumose (P3), or naked (P4). Exopods: three-segmented; segments with spinular ornamentation, as illustrated; inner distal seta of exp-3 sparsely bipinnate, all other elements unipinnate; P3–P4 exp-3 with anterior pore. Endopods: two-segmented, with few spinules, as illustrated. P2 enp-2: with a long, apically serrate, backwardly directed seta. Distal margin of P3 enp-2: with naked outer spine and long bipinnate inner seta. P4 enp-1: slightly shorter than exp-1; distal margin of enp-2 with long, basally fused, serrate seta, and long, unipinnate outer seta. P2–P4: spine and seta formula as for the genus.
</paragraph>
<paragraph id="8C6D9B0CC37A9671567EB811C33AFC49" blockId="19.[826,1442,198,1168]" pageId="19" pageNumber="428">
Fifth legs (
<figureCitation id="14E98789C37A967156F3B811C306FCD0" box="[990,1077,810,832]" captionStart="Figure 9" captionStartId="20.[144,223,1840,1859]" captionTargetBox="[204,1364,197,1805]" captionTargetId="figure-0@20.[204,1365,194,1810]" captionTargetPageId="20" captionText="Figure 9. Psammoleptastacus barani sp. nov. A, urosome ♀, ventral view. B, urosome ♂, ventral view." pageId="19" pageNumber="428">Fig. 9A</figureCitation>
): closely set together but not touching in ventral midline. Baseoendopod and exopod fused, forming a semicircular plate; distal margin with two short bipinnate spines flanked by two long bipinnate setae; outer basal seta long and sparsely plumose.
</paragraph>
<paragraph id="8C6D9B0CC37A9671567EB8DAC326FB00" blockId="19.[826,1442,198,1168]" pageId="19" pageNumber="428">
Genital field (
<figureCitation id="14E98789C37A9671512AB8DAC359FC67" box="[1031,1130,993,1015]" captionStart="Figure 10" captionStartId="21.[164,243,1840,1859]" captionTargetBox="[223,1383,196,1808]" captionTargetId="figure-0@21.[222,1384,194,1810]" captionTargetPageId="21" captionText="Figure 10. Psammoleptastacus barani sp. nov. (♀). A, rostrum and antennule, dorsal view. B, antennule, ventral view. C, antenna. D, free antennary endopod. E, genital field." pageId="19" pageNumber="428">Fig. 10E</figureCitation>
): with genital apertures fused forming median common slit; closed off by fused P6 forming operculum, with one minute spinous process on either side; copulatory pore located midventrally, close to genital slit; seminal receptacles difficult to discern.
</paragraph>
<paragraph id="8C6D9B0CC37A96715617BF8CC4D3FA15" blockId="19.[826,1441,1207,1903]" pageId="19" pageNumber="428">
<emphasis id="BEA6471EC37A96715617BF8CC449FB5C" box="[826,890,1207,1228]" italics="true" pageId="19" pageNumber="428">Male:</emphasis>
Total body length from tip of rostrum to posterior margin of caudal rami: 290–335
<emphasis id="BEA6471EC37A96715050BFEEC2B8FB7C" bold="true" box="[1405,1419,1237,1260]" pageId="19" pageNumber="428">M</emphasis>
m (mean
<emphasis id="BEA6471EC37A967156A5BFCFC4A5FA9B" bold="true" box="[904,918,1268,1291]" pageId="19" pageNumber="428">=</emphasis>
315
<emphasis id="BEA6471EC37A967156E1BFCFC4E9FA9B" bold="true" box="[972,986,1268,1291]" pageId="19" pageNumber="428">M</emphasis>
m;
<emphasis id="BEA6471EC37A9671510DBFCEC307FA9A" box="[1056,1076,1269,1290]" italics="true" pageId="19" pageNumber="428">N</emphasis>
<emphasis id="BEA6471EC37A96715116BFCFC37AFA9B" bold="true" box="[1083,1097,1268,1291]" pageId="19" pageNumber="428">=</emphasis>
6). Body ornamentation (
<figureCitation id="14E98789C37A9671566FBE28C4A8FAB8" box="[834,923,1299,1321]" captionStart="Figure 9" captionStartId="20.[144,223,1840,1859]" captionTargetBox="[204,1364,197,1805]" captionTargetId="figure-0@20.[204,1365,194,1810]" captionTargetPageId="20" captionText="Figure 9. Psammoleptastacus barani sp. nov. A, urosome ♀, ventral view. B, urosome ♂, ventral view." pageId="19" pageNumber="428">Figs 9B</figureCitation>
,
<figureCitation id="14E98789C37A9671568ABE28C4EBFAB9" box="[935,984,1299,1321]" captionStart="Figure 12" captionStartId="23.[164,243,1834,1853]" captionTargetBox="[243,1366,195,1802]" captionTargetId="figure-0@23.[241,1367,195,1805]" captionTargetPageId="23" captionText="Figure 12. Psammoleptastacus barani sp. nov. (♂). A, habitus, dorsal view. B, P3, anterior view. C, antennule, ventral view. D, antennule, dorsal view." pageId="19" pageNumber="428">12A</figureCitation>
): essentially as in female. Sexual dimorphism: in antennule, genital segmentation, P3 endopod, P5, and P6. Spermatophore length: approximately 45
<emphasis id="BEA6471EC37A96715698BE55C4F0FA15" bold="true" box="[949,963,1390,1413]" pageId="19" pageNumber="428">M</emphasis>
m.
</paragraph>
<paragraph id="8C6D9B0CC37A9671567EBEB5C44EF8A1" blockId="19.[826,1441,1207,1903]" pageId="19" pageNumber="428">
Antennule (
<figureCitation id="14E98789C37A967156F6BEB5C352FA33" box="[987,1121,1422,1444]" captionStart="Figure 12" captionStartId="23.[164,243,1834,1853]" captionTargetBox="[243,1366,195,1802]" captionTargetId="figure-0@23.[241,1367,195,1805]" captionTargetPageId="23" captionText="Figure 12. Psammoleptastacus barani sp. nov. (♂). A, habitus, dorsal view. B, P3, anterior view. C, antennule, ventral view. D, antennule, dorsal view." pageId="19" pageNumber="428">Fig. 12C, D</figureCitation>
): nine-segmented, haplocer; geniculation between segments 7 and 8. Segment 2 longest, about 2.2 times as long as wide; segment 4 an incomplete sclerite with one modified (fused at base) and one tiny element; segment 5 with long aesthetasc (45-
<emphasis id="BEA6471EC37A9671564BBD1DC447F9AD" bold="true" box="[870,884,1574,1597]" pageId="19" pageNumber="428">M</emphasis>
m long) fused basally to very small seta; segments 6–8 with one seta and one basally fused spiniform element. Segment 9: with one spatulate seta. Setal formula: 1-[1], 2-[7
<emphasis id="BEA6471EC37A9671518BBDBAC387F908" bold="true" box="[1190,1204,1665,1688]" pageId="19" pageNumber="428">+</emphasis>
1 plumose], 3-[4
<emphasis id="BEA6471EC37A96715052BDBAC2BEF908" bold="true" box="[1407,1421,1665,1688]" pageId="19" pageNumber="428">+</emphasis>
1 pinnate spine], 4-[1
<emphasis id="BEA6471EC37A96715116BD9BC37AF927" bold="true" box="[1083,1097,1696,1719]" pageId="19" pageNumber="428">+</emphasis>
1 modified], 5-[2
<emphasis id="BEA6471EC37A96715007BD9BC20BF927" bold="true" box="[1322,1336,1696,1719]" pageId="19" pageNumber="428">+</emphasis>
(1
<emphasis id="BEA6471EC37A96715076BD9BC25AF927" bold="true" box="[1371,1385,1696,1719]" pageId="19" pageNumber="428">+</emphasis>
ae)], 6-[1
<emphasis id="BEA6471EC37A96715640BD84C448F946" bold="true" box="[877,891,1727,1750]" pageId="19" pageNumber="428">+</emphasis>
1 modified], 7-[1
<emphasis id="BEA6471EC37A96715162BD84C36EF946" bold="true" box="[1103,1117,1727,1750]" pageId="19" pageNumber="428">+</emphasis>
1 modified], 8-[1
<emphasis id="BEA6471EC37A9671501DBD84C20DF946" bold="true" box="[1328,1342,1727,1750]" pageId="19" pageNumber="428">+</emphasis>
1 modified], 9-[7
<emphasis id="BEA6471EC37A96715698BDE6C4F0F964" bold="true" box="[949,963,1757,1780]" pageId="19" pageNumber="428">+</emphasis>
acrothek]. Acrothek consisting of short aesthetasc (13-
<emphasis id="BEA6471EC37A967156C5BDC7C4C5F883" bold="true" box="[1000,1014,1788,1811]" pageId="19" pageNumber="428">M</emphasis>
m long) fused basally to two slender setae.
</paragraph>
<paragraph id="8C6D9B0CC37A967A567EBC01C5D3FE71" blockId="19.[826,1441,1207,1903]" lastBlockId="24.[144,736,462,481]" lastPageId="24" lastPageNumber="433" pageId="19" pageNumber="428">
P3 endopod (
<figureCitation id="14E98789C37A967156C0BC01C363F8C0" box="[1005,1104,1850,1872]" captionStart="Figure 12" captionStartId="23.[164,243,1834,1853]" captionTargetBox="[243,1366,195,1802]" captionTargetId="figure-0@23.[241,1367,195,1805]" captionTargetPageId="23" captionText="Figure 12. Psammoleptastacus barani sp. nov. (♂). A, habitus, dorsal view. B, P3, anterior view. C, antennule, ventral view. D, antennule, dorsal view." pageId="19" pageNumber="428">Fig. 12B</figureCitation>
): two-segmented; enp-1 with few strong spinules along outer margin; enp-2 *Length of rami calculated as sum of segment lengths.
</paragraph>
<caption id="D8ADCB84C37D967655BDBC0BC223F8D3" box="[144,1296,1839,1859]" pageId="20" pageNumber="429" startId="20.[144,223,1840,1859]" targetBox="[204,1364,197,1805]" targetPageId="20">
<paragraph id="8C6D9B0CC37D967655BDBC0BC223F8D3" blockId="20.[144,1296,1839,1859]" box="[144,1296,1839,1859]" pageId="20" pageNumber="429">
<emphasis id="BEA6471EC37D967655BDBC0BC591F8D3" bold="true" box="[144,674,1840,1859]" pageId="20" pageNumber="429">
Figure 9.
<taxonomicName id="4BD2E08FC37D96765429BC0BC576F8D3" authority="Sak &amp; Huys &amp; Karaytuğ, 2008" authorityName="Sak &amp; Huys &amp; Karaytuğ" authorityYear="2008" box="[260,581,1840,1859]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="20" pageNumber="429" phylum="Arthropoda" rank="species" species="barani" status="sp. nov.">
<emphasis id="BEA6471EC37D96765429BC0BC576F8D3" bold="true" box="[260,581,1840,1859]" italics="true" pageId="20" pageNumber="429">Psammoleptastacus barani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A595FA65C37D96765766BC0BC591F8D3" box="[587,674,1840,1859]" pageId="20" pageNumber="429" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A, urosome ♀, ventral view. B, urosome ♂, ventral view.
</paragraph>
</caption>
<caption id="D8ADCB84C37C96775589BC0BC408F8F0" pageId="21" pageNumber="430" startId="21.[164,243,1840,1859]" targetBox="[223,1383,196,1808]" targetPageId="21">
<paragraph id="8C6D9B0CC37C96775589BC0BC408F8F0" blockId="21.[164,1441,1839,1889]" pageId="21" pageNumber="430">
<emphasis id="BEA6471EC37C96775589BC0BC5F7F8D3" bold="true" box="[164,708,1840,1859]" pageId="21" pageNumber="430">
Figure 10.
<taxonomicName id="4BD2E08FC37C9677540BBC0BC55BF8D3" authority="Sak &amp; Huys &amp; Karaytuğ, 2008" authorityName="Sak &amp; Huys &amp; Karaytuğ" authorityYear="2008" box="[294,616,1840,1859]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="21" pageNumber="430" phylum="Arthropoda" rank="species" species="barani" status="sp. nov.">
<emphasis id="BEA6471EC37C9677540BBC0BC55BF8D3" bold="true" box="[294,616,1840,1859]" italics="true" pageId="21" pageNumber="430">Psammoleptastacus barani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A595FA65C37C96775743BC0BC5F7F8D3" box="[622,708,1840,1859]" pageId="21" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(♀). A, rostrum and antennule, dorsal view. B, antennule, ventral view. C, antenna. D, free antennary endopod. E, genital field.
</paragraph>
</caption>
<caption id="D8ADCB84C37F967455BDBC0BC65CF8F0" pageId="22" pageNumber="431" startId="22.[144,223,1840,1859]" targetBox="[189,1379,195,1808]" targetPageId="22">
<paragraph id="8C6D9B0CC37F967455BDBC0BC65CF8F0" blockId="22.[144,1422,1839,1889]" pageId="22" pageNumber="431">
<emphasis id="BEA6471EC37F967455BDBC0BC59EF8D3" bold="true" box="[144,685,1840,1859]" pageId="22" pageNumber="431">
Figure 11.
<taxonomicName id="4BD2E08FC37F9674543DBC0BC562F8D3" authority="Sak &amp; Huys &amp; Karaytuğ, 2008" authorityName="Sak &amp; Huys &amp; Karaytuğ" authorityYear="2008" box="[272,593,1840,1859]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="22" pageNumber="431" phylum="Arthropoda" rank="species" species="barani" status="sp. nov.">
<emphasis id="BEA6471EC37F9674543DBC0BC562F8D3" bold="true" box="[272,593,1840,1859]" italics="true" pageId="22" pageNumber="431">Psammoleptastacus barani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A595FA65C37F9674577ABC0BC59EF8D3" box="[599,685,1840,1859]" pageId="22" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(♀). A, P1, anterior view. B, P2, anterior view. C, P3, anterior view. D, P4, anterior view.
</paragraph>
</caption>
<caption id="D8ADCB84C37E96755589BC11C563F8CB" pageId="23" pageNumber="432" startId="23.[164,243,1834,1853]" targetBox="[243,1366,195,1802]" targetPageId="23">
<paragraph id="8C6D9B0CC37E96755589BC11C563F8CB" blockId="23.[164,1441,1834,1883]" pageId="23" pageNumber="432">
<emphasis id="BEA6471EC37E96755589BC11C5F5F8AE" bold="true" box="[164,710,1834,1854]" pageId="23" pageNumber="432">
Figure 12.
<taxonomicName id="4BD2E08FC37E9675540BBC11C55AF8AD" authority="Sak &amp; Huys &amp; Karaytuğ, 2008" authorityName="Sak &amp; Huys &amp; Karaytuğ" authorityYear="2008" box="[294,617,1834,1853]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="23" pageNumber="432" phylum="Arthropoda" rank="species" species="barani" status="sp. nov.">
<emphasis id="BEA6471EC37E9675540BBC11C55AF8AD" bold="true" box="[294,617,1834,1853]" italics="true" pageId="23" pageNumber="432">Psammoleptastacus barani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A595FA65C37E9675575DBC10C5F5F8AE" box="[624,710,1835,1854]" pageId="23" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(♂). A, habitus, dorsal view. B, P3, anterior view. C, antennule, ventral view. D, antennule, dorsal view.
</paragraph>
</caption>
<caption id="D8ADCB84C371967A55BDBBF3C439FF4B" box="[144,778,200,219]" pageId="24" pageNumber="433" startId="24.[144,208,200,219]">
<paragraph id="8C6D9B0CC371967A55BDBBF3C439FF4B" blockId="24.[144,778,200,219]" box="[144,778,200,219]" pageId="24" pageNumber="433">
<emphasis id="BEA6471EC371967A55BDBBF3C7DAFF4B" bold="true" box="[144,233,200,219]" pageId="24" pageNumber="433">Table 4.</emphasis>
Diagnostic features of
<taxonomicName id="4BD2E08FC371967A54C4BBF3C439FF4B" box="[489,778,200,219]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="24" pageNumber="433" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BEA6471EC371967A54C4BBF3C584FF4B" box="[489,695,200,219]" italics="true" pageId="24" pageNumber="433">Psammoleptastacus</emphasis>
species
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8C6D9B0CC371967A57A2BA39C3DCFE36" pageId="24" pageNumber="433">
<table id="FED269ACC371699D55BDBA3AC2BEFE37" box="[144,1421,257,423]" gridcols="4" gridrows="5" pageId="24" pageNumber="433">
<tr id="32E2994EC371699D55BDBA3AC2BEFE85" box="[144,1421,257,277]" gridrow="0" pageId="24" pageNumber="433" rowspan-0="1">
<th id="7133F032C371699D57A2BA3AC42DFE85" box="[655,798,257,277]" gridcol="1" gridrow="0" pageId="24" pageNumber="433">
<taxonomicName id="4BD2E08FC371967A57A2BA39C42DFE84" authorityName="Pennak" authorityYear="1942" box="[655,798,257,276]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="24" pageNumber="433" phylum="Arthropoda" rank="species" species="arenaridus">
<emphasis id="BEA6471EC371967A57A2BA39C42DFE84" box="[655,798,257,276]" italics="true" pageId="24" pageNumber="433">P. arenaridus</emphasis>
</taxonomicName>
</th>
<th id="7133F032C371699D5686BA3AC305FE85" box="[939,1078,257,277]" gridcol="2" gridrow="0" pageId="24" pageNumber="433">
<taxonomicName id="4BD2E08FC371967A5686BA39C33EFE84" authorityName="Sak &amp; Huys &amp; Karaytuğ" authorityYear="2008" baseAuthorityName="Noodt" baseAuthorityYear="1955" box="[939,1037,258,276]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="24" pageNumber="433" phylum="Arthropoda" rank="species" species="stygius">
<emphasis id="BEA6471EC371967A5686BA39C33EFE84" box="[939,1037,258,276]" italics="true" pageId="24" pageNumber="433">P. stygius</emphasis>
</taxonomicName>
</th>
<th id="7133F032C371699D51EEBA3AC2BEFE85" box="[1219,1421,257,277]" gridcol="3" gridrow="0" pageId="24" pageNumber="433">
<emphasis id="BEA6471EC371967A51EEBA39C2BEFE85" bold="true" box="[1219,1421,257,277]" pageId="24" pageNumber="433">
<taxonomicName id="4BD2E08FC371967A51EEBA39C202FE84" authority="Sak &amp; Huys &amp; Karaytuğ, 2008" authorityName="Sak &amp; Huys &amp; Karaytuğ" authorityYear="2008" box="[1219,1329,257,277]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="24" pageNumber="433" phylum="Arthropoda" rank="species" species="barani" status="sp. nov.">
<emphasis id="BEA6471EC371967A51EEBA39C202FE84" box="[1219,1329,257,277]" italics="true" pageId="24" pageNumber="433">P. barani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A595FA65C371967A501ABA39C2BEFE85" box="[1335,1421,258,277]" pageId="24" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="32E2994EC371699D55BDBA00C2BEFEDF" box="[144,1421,315,335]" gridrow="1" pageId="24" pageNumber="433">
<th id="7133F032C371699D55BDBA00C531FEDF" box="[144,514,315,335]" gridcol="0" gridrow="1" pageId="24" pageNumber="433">P1 exopod:endopod*</th>
<td id="7133F032C371699D57A2BA00C42DFEDF" box="[655,798,315,335]" gridcol="1" gridrow="1" pageId="24" pageNumber="433">1.33</td>
<td id="7133F032C371699D5686BA00C305FEDF" box="[939,1078,315,335]" gridcol="2" gridrow="1" pageId="24" pageNumber="433">1.28</td>
<td id="7133F032C371699D51EEBA00C2BEFEDF" box="[1219,1421,315,335]" gridcol="3" gridrow="1" pageId="24" pageNumber="433">1.15</td>
</tr>
<tr id="32E2994EC371699D55BDBA63C2BEFEFC" box="[144,1421,344,364]" gridrow="2" pageId="24" pageNumber="433">
<th id="7133F032C371699D55BDBA63C531FEFC" box="[144,514,344,364]" gridcol="0" gridrow="2" pageId="24" pageNumber="433">P3 enp-2 ♂ distal spine</th>
<td id="7133F032C371699D57A2BA63C42DFEFC" box="[655,798,344,364]" gridcol="1" gridrow="2" pageId="24" pageNumber="433">fused</td>
<td id="7133F032C371699D5686BA63C305FEFC" box="[939,1078,344,364]" gridcol="2" gridrow="2" pageId="24" pageNumber="433">?</td>
<td id="7133F032C371699D51EEBA63C2BEFEFC" box="[1219,1421,344,364]" gridcol="3" gridrow="2" pageId="24" pageNumber="433">discrete</td>
</tr>
<tr id="32E2994EC371699D55BDBA4DC2BEFE19" box="[144,1421,374,393]" gridrow="3" pageId="24" pageNumber="433">
<th id="7133F032C371699D55BDBA4DC531FE19" box="[144,514,374,393]" gridcol="0" gridrow="3" pageId="24" pageNumber="433">P4</th>
<td id="7133F032C371699D57A2BA4DC42DFE19" box="[655,798,374,393]" gridcol="1" gridrow="3" pageId="24" pageNumber="433">
exp-1
<emphasis id="BEA6471EC371967A57E3BA4CC5E9FE19" box="[718,730,375,393]" italics="true" pageId="24" pageNumber="433">&lt;</emphasis>
enp-1
</td>
<td id="7133F032C371699D5686BA4DC305FE19" box="[939,1078,374,393]" gridcol="2" gridrow="3" pageId="24" pageNumber="433">
exp-1
<emphasis id="BEA6471EC371967A56C7BA4CC4C5FE19" box="[1002,1014,375,393]" italics="true" pageId="24" pageNumber="433">&lt;</emphasis>
enp-1
</td>
<td id="7133F032C371699D51EEBA4DC2BEFE19" box="[1219,1421,374,393]" gridcol="3" gridrow="3" pageId="24" pageNumber="433">
exp-1
<emphasis id="BEA6471EC371967A502FBA4CC23DFE19" box="[1282,1294,375,393]" italics="true" pageId="24" pageNumber="433">&gt;</emphasis>
enp-1
</td>
</tr>
<tr id="32E2994EC371699D55BDBAA8C2BEFE37" box="[144,1421,403,423]" gridrow="4" pageId="24" pageNumber="433">
<th id="7133F032C371699D55BDBAA8C531FE37" box="[144,514,403,423]" gridcol="0" gridrow="4" pageId="24" pageNumber="433">Caudal ramus length: basal width</th>
<td id="7133F032C371699D57A2BAA8C42DFE37" box="[655,798,403,423]" gridcol="1" gridrow="4" pageId="24" pageNumber="433">2.75</td>
<td id="7133F032C371699D5686BAA8C305FE37" box="[939,1078,403,423]" gridcol="2" gridrow="4" pageId="24" pageNumber="433">3.85</td>
<td id="7133F032C371699D51EEBAA8C2BEFE37" box="[1219,1421,403,423]" gridcol="3" gridrow="4" pageId="24" pageNumber="433">2.75</td>
</tr>
</table>
</paragraph>
<paragraph id="8C6D9B0CC371967A55BDB925C586FD3E" blockId="24.[144,759,542,869]" pageId="24" pageNumber="433">minute, with strong spinule at outer distal corner, short fine seta arising from inner distal corner (homologous with long inner distal seta of female), and naked curved apical spine, discrete at base (homologous with outer distal spine of female).</paragraph>
<paragraph id="8C6D9B0CC371967A5585B98DC7F9FC9A" blockId="24.[144,759,542,869]" pageId="24" pageNumber="433">
P5 (
<figureCitation id="14E98789C371967A55F5B98CC61EFD5D" box="[216,301,695,717]" captionStart="Figure 9" captionStartId="20.[144,223,1840,1859]" captionTargetBox="[204,1364,197,1805]" captionTargetId="figure-0@20.[204,1365,194,1810]" captionTargetPageId="20" captionText="Figure 9. Psammoleptastacus barani sp. nov. A, urosome ♀, ventral view. B, urosome ♂, ventral view." pageId="24" pageNumber="433">Fig. 9B</figureCitation>
): with armature as in female, but with innermost element of distal margin much shorter and bare.
</paragraph>
<paragraph id="8C6D9B0CC371967A5585B829C7E1FCF5" blockId="24.[144,759,542,869]" pageId="24" pageNumber="433">
Sixth legs (
<figureCitation id="14E98789C371967A540AB829C64EFCB8" box="[295,381,786,808]" captionStart="Figure 9" captionStartId="20.[144,223,1840,1859]" captionTargetBox="[204,1364,197,1805]" captionTargetId="figure-0@20.[204,1365,194,1810]" captionTargetPageId="20" captionText="Figure 9. Psammoleptastacus barani sp. nov. A, urosome ♀, ventral view. B, urosome ♂, ventral view." pageId="24" pageNumber="433">Fig. 9B</figureCitation>
): asymmetrical, with smallest P6 closing off functional gonopore; each with two pinnate setae.
</paragraph>
<paragraph id="8C6D9B0CC371967A55BDB8A3C565FC7C" blockId="24.[144,758,917,1004]" pageId="24" pageNumber="433">
<emphasis id="BEA6471EC371967A55BDB8A3C63EFC3D" box="[144,269,920,941]" italics="true" pageId="24" pageNumber="433">Etymology:</emphasis>
The species is named after Prof. İbrahim Baran, Dokuz Eylül University, in recognition of his contributions to herpetology in
<collectingCountry id="F4C5DB9CC371967A572DB8EDC561FC7C" box="[512,594,982,1004]" name="Turkey" pageId="24" pageNumber="433">Turkey</collectingCountry>
.
</paragraph>
<paragraph id="8C6D9B0CC371967A55BDBF24C3DBFDC2" blockId="24.[144,759,1055,1903]" lastBlockId="24.[806,1421,541,594]" pageId="24" pageNumber="433">
<emphasis id="BEA6471EC371967A55BDBF24C577FBA4" box="[144,580,1055,1076]" italics="true" pageId="24" pageNumber="433">
Remarks:
<taxonomicName id="4BD2E08FC371967A542ABF24C577FBA4" authority="Sak &amp; Huys &amp; Karaytuğ, 2008" authorityName="Sak &amp; Huys &amp; Karaytuğ" authorityYear="2008" box="[263,580,1055,1076]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="24" pageNumber="433" phylum="Arthropoda" rank="species" species="barani" status="sp. nov.">Psammoleptastacus barani</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A595FA65C371967A5766BF24C5ACFBA4" box="[587,671,1055,1076]" pageId="24" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
differs from its two known congeners in the longer P1 endopod, and the length of P4 enp-2, which is shorter than, or at most as long as, the proximal exopod segment, instead of being distinctly longer. It is similar to
<taxonomicName id="4BD2E08FC371967A5423BF83C695FB5D" authorityName="Pennak" authorityYear="1942" box="[270,422,1208,1229]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="24" pageNumber="433" phylum="Arthropoda" rank="species" species="arenaridus">
<emphasis id="BEA6471EC371967A5423BF83C695FB5D" box="[270,422,1208,1229]" italics="true" pageId="24" pageNumber="433">P. arenaridus</emphasis>
</taxonomicName>
in the length of the caudal ramus, but deviates from it in the detailed morphology of the male P3 endopod.
<bibRefCitation id="E843E6FDC371967A54F0BFCEC5BFFA9B" author="Marinov TM" box="[477,652,1269,1291]" pageId="24" pageNumber="433" pagination="43 - 87" refId="ref25342" refString="Marinov TM. 1971. Kharpatikoidi ot b'lgarskoto kraibrezhnie na Cherno more [Harpacticoids of the Bulgarian Black Sea coast]. Izvestiya na Instituta Okeanografiya i Ribno Stopanstvo, Varna 11: 43 - 87. [In Bulgarian with English and Russian summaries]." type="journal article" year="1971">Marinov (1971)</bibRefCitation>
recorded a few specimens from the beaches of the Arkutino region (south of Sozopol) along the Bulgarian coast, which he attributed to
<taxonomicName id="4BD2E08FC371967A54B3BE6AC533FAF6" authorityName="Noodt, 1955 b sensu Marinov" authorityYear="1971" box="[414,512,1361,1382]" class="Hexanauplia" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="24" pageNumber="433" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="BEA6471EC371967A54B3BE6AC533FAF6" box="[414,512,1361,1382]" italics="true" pageId="24" pageNumber="433">A. stygia</emphasis>
</taxonomicName>
. He noted the difference in the relative length of the P1 endopod between his specimens and
<bibRefCitation id="E843E6FDC371967A5446BEB5C52DFA34" author="Noodt W" box="[363,542,1422,1444]" pageId="24" pageNumber="433" pagination="86 - 109" refId="ref25760" refString="Noodt W. 1955 b. Harpacticiden (Crust. Cop.) aus dem Sandstrand der franzosischen Biscaya-Kuste. Kieler Meeresforschungen 11: 86 - 109." type="journal article" year="1955">Noodt’s (1955b)</bibRefCitation>
<typeStatus id="536925AEC371967A570BBEB5C564FA33" box="[550,599,1422,1443]" pageId="24" pageNumber="433">type</typeStatus>
population of
<taxonomicName id="4BD2E08FC371967A55BDBE96C7C9FA52" authorityName="Sak &amp; Huys &amp; Karaytuğ" authorityYear="2008" baseAuthorityName="Noodt" baseAuthorityYear="1955" box="[144,250,1453,1474]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="24" pageNumber="433" phylum="Arthropoda" rank="species" species="stygius">
<emphasis id="BEA6471EC371967A55BDBE96C7C9FA52" box="[144,250,1453,1474]" italics="true" pageId="24" pageNumber="433">P. stygius</emphasis>
</taxonomicName>
. The similarity in the relative length of the P1 endopod, P4 endopod, and caudal ramus, the P3 endopodal sexual dimorphism, and the female P5, leave little doubt that
<bibRefCitation id="E843E6FDC371967A548ABD33C56FF98E" author="Marinov TM" box="[423,604,1544,1566]" pageId="24" pageNumber="433" pagination="43 - 87" refId="ref25342" refString="Marinov TM. 1971. Kharpatikoidi ot b'lgarskoto kraibrezhnie na Cherno more [Harpacticoids of the Bulgarian Black Sea coast]. Izvestiya na Instituta Okeanografiya i Ribno Stopanstvo, Varna 11: 43 - 87. [In Bulgarian with English and Russian summaries]." type="journal article" year="1971">Marinov (1971)</bibRefCitation>
was dealing with
<taxonomicName id="4BD2E08FC371967A55E0BD1CC607F9AC" authority="Sak &amp; Huys &amp; Karaytuğ, 2008" authorityName="Sak &amp; Huys &amp; Karaytuğ" authorityYear="2008" box="[205,308,1575,1596]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="24" pageNumber="433" phylum="Arthropoda" rank="species" species="barani" status="sp. nov.">
<emphasis id="BEA6471EC371967A55E0BD1CC607F9AC" box="[205,308,1575,1596]" italics="true" pageId="24" pageNumber="433">P. barani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A595FA65C371967A5416BD1CC6BCF9AC" box="[315,399,1575,1596]" pageId="24" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
The only discrepancy between both descriptions is found in the male P5, which has a much longer innermost seta in the Bulgarian material.
<bibRefCitation id="E843E6FDC371967A55E5BDB8C6B4F909" author="Apostolov A" box="[200,391,1667,1689]" pageId="24" pageNumber="433" pagination="88 - 110" refId="ref23964" refString="Apostolov A. 1973. Sur divers Harpacticoides (Copepodes) de la mer Noire. Zoologischer Anzeiger 190: 88 - 110." type="journal article" year="1973">Apostolov (1973)</bibRefCitation>
claimed that considerable variability was found in the P5 of his Black Sea
<taxonomicName id="4BD2E08FC371967A55BDBDFBC605F945" authorityName="Kunz, 1937 sensu Rao &amp; Ganapati" authorityYear="1969" box="[144,310,1728,1749]" class="Hexanauplia" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="24" pageNumber="433" phylum="Arthropoda" rank="species" species="subterranea">
<emphasis id="BEA6471EC371967A55BDBDFBC605F945" box="[144,310,1728,1749]" italics="true" pageId="24" pageNumber="433">A. subterranea</emphasis>
</taxonomicName>
(no localities specified), but it is conceivable that his drawings of the female P5 were based on two different species: his
<figureCitation id="14E98789C371967A5708BDC6C59BF883" box="[549,680,1789,1811]" captionStart="Figure 18" captionStartId="36.[144,223,1842,1861]" captionTargetBox="[228,1336,194,1806]" captionTargetId="figure-0@36.[227,1339,194,1810]" captionTargetPageId="36" captionText="Figure 18. Mesopontia dillonbeachia (Lang, 1965) comb. nov. (♀). A, habitus, dorsal view. B, habitus ovigerous specimen, lateral view. C, posterior part of penultimate somite, anal somite, and caudal rami, dorsal view. D, anal somite and right caudal ramus, lateral view." pageId="24" pageNumber="433">Figure 18-5</figureCitation>
shows a P5 of the
<taxonomicName id="4BD2E08FC371967A543FBC27C6AEF8A1" authorityName="Kunz" authorityYear="1937" box="[274,413,1820,1841]" class="Hexanauplia" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="24" pageNumber="433" phylum="Arthropoda" rank="genus">
<emphasis id="BEA6471EC371967A543FBC27C6AEF8A1" box="[274,413,1820,1841]" italics="true" pageId="24" pageNumber="433">Arenopontia</emphasis>
</taxonomicName>
<typeStatus id="536925AEC371967A5489BC27C6EBF8A1" box="[420,472,1820,1841]" pageId="24" pageNumber="433">type</typeStatus>
, whereas his
<figureCitation id="14E98789C371967A575EBC27C5C5F8A1" box="[627,758,1819,1842]" captionStart="Figure 18" captionStartId="36.[144,223,1842,1861]" captionTargetBox="[228,1336,194,1806]" captionTargetId="figure-0@36.[227,1339,194,1810]" captionTargetPageId="36" captionText="Figure 18. Mesopontia dillonbeachia (Lang, 1965) comb. nov. (♀). A, habitus, dorsal view. B, habitus ovigerous specimen, lateral view. C, posterior part of penultimate somite, anal somite, and caudal rami, dorsal view. D, anal somite and right caudal ramus, lateral view." pageId="24" pageNumber="433">Figure 18-6</figureCitation>
was almost certainly based on the species previously identified by
<bibRefCitation id="E843E6FDC371967A5400BC62C6D3F8FF" author="Marinov TM" box="[301,480,1881,1903]" pageId="24" pageNumber="433" pagination="43 - 87" refId="ref25342" refString="Marinov TM. 1971. Kharpatikoidi ot b'lgarskoto kraibrezhnie na Cherno more [Harpacticoids of the Bulgarian Black Sea coast]. Izvestiya na Instituta Okeanografiya i Ribno Stopanstvo, Varna 11: 43 - 87. [In Bulgarian with English and Russian summaries]." type="journal article" year="1971">Marinov (1971)</bibRefCitation>
as
<taxonomicName id="4BD2E08FC371967A573DBC62C541F8FE" authorityName="Noodt, 1955 b sensu Marinov" authorityYear="1971" box="[528,626,1881,1902]" class="Hexanauplia" family="Arenopontiidae" genus="Arenopontia" kingdom="Animalia" order="Harpacticoida" pageId="24" pageNumber="433" phylum="Arthropoda" rank="species" species="stygia">
<emphasis id="BEA6471EC371967A573DBC62C541F8FE" box="[528,626,1881,1902]" italics="true" pageId="24" pageNumber="433">A. stygia</emphasis>
</taxonomicName>
. Note that the
<typeStatus id="536925AEC371967A5678B925C4B5FDA3" box="[853,902,542,563]" pageId="24" pageNumber="433">type</typeStatus>
locality of
<taxonomicName id="4BD2E08FC371967A5127B925C342FDA2" authority="Sak &amp; Huys &amp; Karaytuğ, 2008" authorityName="Sak &amp; Huys &amp; Karaytuğ" authorityYear="2008" box="[1034,1137,541,563]" class="Hexanauplia" family="Arenopontiidae" genus="Psammoleptastacus" kingdom="Animalia" order="Harpacticoida" pageId="24" pageNumber="433" phylum="Arthropoda" rank="species" species="barani" status="sp. nov.">
<emphasis id="BEA6471EC371967A5127B925C342FDA2" box="[1034,1137,541,563]" italics="true" pageId="24" pageNumber="433">P. barani</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A595FA65C371967A5155B925C3FFFDA3" box="[1144,1228,542,563]" pageId="24" pageNumber="433" rank="species">sp. nov.</taxonomicNameLabel>
is in close proximity to the Bulgarian collecting sites.
</paragraph>
</subSubSection>
</treatment>
</document>
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