474 lines
47 KiB
XML
474 lines
47 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/BDJ.8.e57572" ID-PMC="PMC7511464" ID-Pensoft-Pub="1314-2828-8-e57572" ID-Pensoft-UUID="9627C5A06E285DBFB389A57DFEE8433F" ID-PubMed="33013175" ID-ZooBank="CE3CFA74ABAA48A49EB22776C731837F" ModsDocID="1314-2828-8-e57572" checkinTime="1600294130755" checkinUser="pensoft" docAuthor="Yanez-Rivera, Beatriz, Tovar-Hernandez, Maria Ana, Galvan-Villa, Cristian Moises & Rios-Jara, Eduardo" docDate="2020" docId="043A950F89BA54CB8CE21C179A8A8673" docLanguage="en" docName="BiodivDatJour 8: e57572" docOrigin="Biodiversity Data Journal 8" docSource="http://dx.doi.org/10.3897/BDJ.8.e57572" docTitle="Clymenura scutata Tovar-Hernandez and Yanez-Rivera 2020, sp. n." docType="treatment" docVersion="2" id="9627C5A06E285DBFB389A57DFEE8433F" lastPageNumber="57572" masterDocId="9627C5A06E285DBFB389A57DFEE8433F" masterDocTitle="Tubicolous polychaete worms (Annelida) from Bahia de Chamela Islands Sanctuary, Mexico, with the description of a new bamboo worm" masterLastPageNumber="57572" masterPageNumber="57572" pageNumber="57572" updateTime="1668128532144" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Tubicolous polychaete worms (Annelida) from Bahia de Chamela Islands Sanctuary, Mexico, with the description of a new bamboo worm</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yanez-Rivera, Beatriz</mods:namePart>
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<mods:affiliation>CONACYT- Centro de Investigacion en Alimentacion y Desarrollo A. C. (CIAD), Unidad Mazatlan en Acuicultura y Manejo Ambiental, Mazatlan, Mexico</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-3192-2142</mods:nameIdentifier>
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<mods:nameIdentifier type="email">beyariv@ciad.mx</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tovar-Hernandez, Maria Ana</mods:namePart>
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<mods:affiliation>Universidad Autonoma de Nuevo Leon, Facultad de Ciencias Biologicas, Laboratorio de Biosistematica, San Nicolas de los Garza, Nuevo Leon, Mexico</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-5263-2830</mods:nameIdentifier>
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<mods:nameIdentifier type="email">maria_ana_tovar@yahoo.com</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Galvan-Villa, Cristian Moises</mods:namePart>
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<mods:affiliation>Universidad de Guadalajara, Centro Universitario de Ciencias Biologicas y Agropecuarias, Departamento de Ecologia, Zapopan, Mexico</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0003-1927-2500</mods:nameIdentifier>
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rios-Jara, Eduardo</mods:namePart>
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<mods:affiliation>Universidad de Guadalajara, Centro Universitario de Ciencias Biologicas y Agropecuarias, Departamento de Ecologia, Zapopan, Mexico</mods:affiliation>
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</mods:name>
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<mods:title>Biodiversity Data Journal</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>8</mods:number>
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<mods:start>57572</mods:start>
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<mods:end>57572</mods:end>
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<mods:url>http://dx.doi.org/10.3897/BDJ.8.e57572</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/BDJ.8.e57572</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1314-2828-8-e57572</mods:identifier>
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<subSubSection pageId="0" pageNumber="57572" type="nomenclature">
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<paragraph pageId="0" pageNumber="57572">
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<taxonomicName LSID="043A950F-89BA-54CB-8CE2-1C179A8A8673" authority="Tovar-Hernández and Yáñez-Rivera, 2020" authorityName="Tovar-Hernandez and Yanez-Rivera" authorityYear="2020" class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymenura scutata" pageId="0" pageNumber="57572" phylum="Annelida" rank="species" species="scutata" status="sp. n.">
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Clymenura scutata
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<normalizedToken originalValue="Tovar-Hernández">Tovar-Hernandez</normalizedToken>
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and
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<normalizedToken originalValue="Yáñez-Rivera">Yanez-Rivera</normalizedToken>
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, 2020
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</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="57572">sp. n.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="57572" type="materials_examined">
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<paragraph pageId="0" pageNumber="57572">Materials</paragraph>
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<paragraph pageId="0" pageNumber="57572">
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<materialsCitation collectingDate="2013-6-27" collectingMethod="Scuba dive" collectionCode="UDG, LEMA" collectorName="Beatriz Yanez-Rivera" country="Mexico" latitude="19.549166" location="Mexico" longLatPrecision="21" longitude="-105.10555" pageId="0" pageNumber="57572" specimenCode="LEMA-PO 153" specimenCount="1" typeStatus="Holotype">
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<emphasis bold="true" pageId="0" pageNumber="57572">Type status:</emphasis>
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<typeStatus pageId="0" pageNumber="57572">Holotype</typeStatus>
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.
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<emphasis bold="true" pageId="0" pageNumber="57572">Occurrence:</emphasis>
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catalogNumber:
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<specimenCode pageId="0" pageNumber="57572">LEMA-PO153</specimenCode>
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; recordedBy:
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<collectorName pageId="0" pageNumber="57572">
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Beatriz
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<normalizedToken originalValue="Yáñez-Rivera">Yanez-Rivera</normalizedToken>
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</collectorName>
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; lifeStage:
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<specimenType pageId="0" pageNumber="57572">Adult</specimenType>
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;
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<emphasis bold="true" pageId="0" pageNumber="57572">Taxon:</emphasis>
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phylum: Annelida; class: Polychaeta; family: Maldanidae; genus: Clymenura;
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<emphasis bold="true" pageId="0" pageNumber="57572">Location:</emphasis>
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higherGeographyID: Pacific Ocean; higherGeography: Tropical Eastern Pacific; continent: America; islandGroup: Islas de Chamela; island: Isla Cocinas; country:
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<collectingCountry pageId="0" pageNumber="57572">
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<normalizedToken originalValue="México">Mexico</normalizedToken>
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</collectingCountry>
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; countryCode: MX; stateProvince: Jalisco; municipality: La Huerta; maximumDepthInMeters: 7; verbatimLatitude:
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<geoCoordinate degrees="19" direction="north" minutes="32" orientation="latitude" precision="15" seconds="57" value="19.549166">
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19°32
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<normalizedToken originalValue="’57’’">'57''</normalizedToken>
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N
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</geoCoordinate>
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; verbatimLongitude:
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<geoCoordinate degrees="105" direction="west" minutes="06" orientation="longitude" precision="15" seconds="20" value="-105.10555">
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105°06
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<normalizedToken originalValue="’20’’">'20''</normalizedToken>
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W
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</geoCoordinate>
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;
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<emphasis bold="true" pageId="0" pageNumber="57572">Identification:</emphasis>
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identifiedBy:
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<determinerName pageId="0" pageNumber="57572">
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<normalizedToken originalValue="María">Maria</normalizedToken>
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Ana
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||
<normalizedToken originalValue="Tovar-Hernández">Tovar-Hernandez</normalizedToken>
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||
</determinerName>
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||
;
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||
<emphasis bold="true" pageId="0" pageNumber="57572">Event:</emphasis>
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samplingProtocol:
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<collectingMethod pageId="0" pageNumber="57572">Scuba dive</collectingMethod>
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; eventDate:
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<collectingDate pageId="0" pageNumber="57572" value="2013-6-27">June 27, 2013</collectingDate>
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; year: 2013; month: 6; day: 27; habitat: On rock; fieldNumber: Site 16;
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||
<emphasis bold="true" pageId="0" pageNumber="57572">Record Level:</emphasis>
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language: Spanish; institutionID: Universidad de Guadalajara; collectionID:
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<normalizedToken originalValue="Colección">Coleccion</normalizedToken>
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<normalizedToken originalValue="Biológica">Biologica</normalizedToken>
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del Laboratorio de Ecosistemas Marinos y Acuicultura, Universidad de Guadalajara,
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<normalizedToken originalValue="México">Mexico</normalizedToken>
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; institutionCode:
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<collectionCode pageId="0" pageNumber="57572">UDG</collectionCode>
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; collectionCode:
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<collectionCode pageId="0" pageNumber="57572">LEMA</collectionCode>
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</materialsCitation>
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<materialsCitation collectingDate="2013-6-27" collectingMethod="Scuba dive" collectionCode="UANL" collectorName="Beatriz Yanez-Rivera" country="Mexico" latitude="19.549166" location="Mexico" longLatPrecision="21" longitude="-105.10555" pageId="0" pageNumber="57572" specimenCode="UANL 8144" specimenCount="3" specimenCount-adult="3" typeStatus="Paratype">
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<emphasis bold="true" pageId="0" pageNumber="57572">Type status:</emphasis>
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||
<typeStatus pageId="0" pageNumber="57572">Paratype</typeStatus>
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||
.
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||
<emphasis bold="true" pageId="0" pageNumber="57572">Occurrence:</emphasis>
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catalogNumber:
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<specimenCode pageId="0" pageNumber="57572">UANL 8144</specimenCode>
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; recordedBy:
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<collectorName pageId="0" pageNumber="57572">
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||
Beatriz
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<normalizedToken originalValue="Yáñez-Rivera">Yanez-Rivera</normalizedToken>
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</collectorName>
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; individualCount:
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<specimenCount pageId="0" pageNumber="57572">3</specimenCount>
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; lifeStage:
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<specimenType pageId="0" pageNumber="57572">Adult</specimenType>
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;
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<emphasis bold="true" pageId="0" pageNumber="57572">Taxon:</emphasis>
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||
phylum: Annelida; class: Polychaeta; family: Maldanidae; genus: Clymenura;
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||
<emphasis bold="true" pageId="0" pageNumber="57572">Location:</emphasis>
|
||
higherGeographyID: Pacific Ocean; higherGeography: Tropical Eastern Pacific; continent: America; islandGroup: Islas de Chamela; island: Isla Cocinas; country:
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<collectingCountry pageId="0" pageNumber="57572">
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<normalizedToken originalValue="México">Mexico</normalizedToken>
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</collectingCountry>
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; countryCode: MX; stateProvince: Jalisco; municipality: La Huerta; maximumDepthInMeters: 7; verbatimLatitude:
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<geoCoordinate degrees="19" direction="north" minutes="32" orientation="latitude" precision="15" seconds="57" value="19.549166">
|
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19°32
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<normalizedToken originalValue="’57’’">'57''</normalizedToken>
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N
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</geoCoordinate>
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; verbatimLongitude:
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<geoCoordinate degrees="105" direction="west" minutes="06" orientation="longitude" precision="15" seconds="20" value="-105.10555">
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105°06
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<normalizedToken originalValue="’20’’">'20''</normalizedToken>
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W
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;
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<emphasis bold="true" pageId="0" pageNumber="57572">Identification:</emphasis>
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identifiedBy:
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<determinerName pageId="0" pageNumber="57572">
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<normalizedToken originalValue="María">Maria</normalizedToken>
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Ana
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<normalizedToken originalValue="Tovar-Hernández">Tovar-Hernandez</normalizedToken>
|
||
</determinerName>
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||
;
|
||
<emphasis bold="true" pageId="0" pageNumber="57572">Event:</emphasis>
|
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samplingProtocol:
|
||
<collectingMethod pageId="0" pageNumber="57572">Scuba dive</collectingMethod>
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||
; eventDate:
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||
<collectingDate pageId="0" pageNumber="57572" value="2013-6-27">June 27, 2013</collectingDate>
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||
; year: 2013; month: 6; day: 27; habitat: On rock; fieldNumber: Site 16;
|
||
<emphasis bold="true" pageId="0" pageNumber="57572">Record Level:</emphasis>
|
||
language: Spanish; institutionID: Universidad
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||
<normalizedToken originalValue="Autónoma">Autonoma</normalizedToken>
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||
de Nuevo
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||
<normalizedToken originalValue="León">Leon</normalizedToken>
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||
; collectionID: UANL, NL INV 0002-05-09; institutionCode:
|
||
<collectionCode pageId="0" pageNumber="57572">UANL</collectionCode>
|
||
; collectionCode:
|
||
<collectionCode pageId="0" pageNumber="57572">UANL</collectionCode>
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||
</materialsCitation>
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||
<materialsCitation collectingDate="2013-6-27" collectingMethod="Dredge" collectionCode="UNAM-ICML, ICML-EMU" collectorName="Beatriz Yanez-Rivera" country="Mexico" latitude="19.551943" location="Mexico" longLatPrecision="21" longitude="-105.10861" pageId="0" pageNumber="57572" specimenCode="ICML-EMU- 12758" specimenCount="1" specimenCount-adult="1" typeStatus="Paratype">
|
||
<emphasis bold="true" pageId="0" pageNumber="57572">Type status:</emphasis>
|
||
<typeStatus pageId="0" pageNumber="57572">Paratype</typeStatus>
|
||
.
|
||
<emphasis bold="true" pageId="0" pageNumber="57572">Occurrence:</emphasis>
|
||
catalogNumber:
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||
<specimenCode pageId="0" pageNumber="57572">ICML-EMU-12758</specimenCode>
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; recordedBy:
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<collectorName pageId="0" pageNumber="57572">
|
||
Beatriz
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<normalizedToken originalValue="Yáñez-Rivera">Yanez-Rivera</normalizedToken>
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</collectorName>
|
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; individualCount:
|
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<specimenCount pageId="0" pageNumber="57572">1</specimenCount>
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; lifeStage:
|
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<specimenType pageId="0" pageNumber="57572">Adult</specimenType>
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;
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||
<emphasis bold="true" pageId="0" pageNumber="57572">Taxon:</emphasis>
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phylum: Annelida; class: Polychaeta; family: Maldanidae; genus: Clymenura;
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<emphasis bold="true" pageId="0" pageNumber="57572">Location:</emphasis>
|
||
higherGeographyID: Pacific Ocean; higherGeography: Tropical Eastern Pacific; continent: America; islandGroup: Islas de Chamela; island: Isla Cocinas; country:
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<collectingCountry pageId="0" pageNumber="57572">
|
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<normalizedToken originalValue="México">Mexico</normalizedToken>
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</collectingCountry>
|
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; countryCode: MX; stateProvince: Jalisco; municipality: La Huerta; maximumDepthInMeters: 8; verbatimLatitude:
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<geoCoordinate degrees="19" direction="north" minutes="33" orientation="latitude" precision="15" seconds="07" value="19.551943">
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||
19°33
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<normalizedToken originalValue="’07’’">'07''</normalizedToken>
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N
|
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</geoCoordinate>
|
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; verbatimLongitude:
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<geoCoordinate degrees="105" direction="west" minutes="06" orientation="longitude" precision="15" seconds="31" value="-105.10861">
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105°06
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<normalizedToken originalValue="’31’’">'31''</normalizedToken>
|
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W
|
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</geoCoordinate>
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;
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<emphasis bold="true" pageId="0" pageNumber="57572">Identification:</emphasis>
|
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identifiedBy:
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<determinerName pageId="0" pageNumber="57572">
|
||
<normalizedToken originalValue="María">Maria</normalizedToken>
|
||
Ana
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||
<normalizedToken originalValue="Tovar-Hernández">Tovar-Hernandez</normalizedToken>
|
||
</determinerName>
|
||
;
|
||
<emphasis bold="true" pageId="0" pageNumber="57572">Event:</emphasis>
|
||
samplingProtocol:
|
||
<collectingMethod pageId="0" pageNumber="57572">Dredge</collectingMethod>
|
||
; eventDate:
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||
<collectingDate pageId="0" pageNumber="57572" value="2013-6-27">June 27, 2013</collectingDate>
|
||
; year: 2013; month: 6; day: 27; habitat: On rock; fieldNumber: Site 14;
|
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<emphasis bold="true" pageId="0" pageNumber="57572">Record Level:</emphasis>
|
||
language: Spanish; institutionID: Universidad Nacional
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||
<normalizedToken originalValue="Autónoma">Autonoma</normalizedToken>
|
||
de
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||
<normalizedToken originalValue="México">Mexico</normalizedToken>
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||
, Instituto de Ciencias del Mar y
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<normalizedToken originalValue="Limnología">Limnologia</normalizedToken>
|
||
; collectionID:
|
||
<normalizedToken originalValue="Colección">Coleccion</normalizedToken>
|
||
Regional de Invertebrados Marinos; institutionCode:
|
||
<collectionCode pageId="0" pageNumber="57572">UNAM-ICML</collectionCode>
|
||
; collectionCode:
|
||
<collectionCode pageId="0" pageNumber="57572">ICML-EMU</collectionCode>
|
||
</materialsCitation>
|
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</paragraph>
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</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="57572" type="description">
|
||
<paragraph pageId="0" pageNumber="57572">Description</paragraph>
|
||
<paragraph pageId="0" pageNumber="57572">
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Preserved holotype complete, with body pale without any distinctive pigmentation pattern (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F5999123" captionText="Figure 2. Clymenura scutata sp. n. A) Entire worm, lateral view, tube removed, B-C) cephalic plaque, latero-dorsal views, D-E) posterior end and anal plaque, lateral and ventral view, respectively, F) anal plaque, posterior view, G) worm regenerating cephalic plaque and first anterior segments. A-D, F) Holotype LEMA-PO 153, E, G) Paratype UANL 8144. Scale bars: A) 2 mm, B-E, G) 1 mm, F) 0.6 mm." figureDoi="10.3897/BDJ.8.e57572.figure2" httpUri="https://binary.pensoft.net/fig/437506" pageId="0" pageNumber="57572">2</figureCitation>
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A). Body 32 mm length (complete paratypes: 28 mm in paratype 2 (UANL-8144), 20 mm in paratype 4 (ICML-EMU-12758); incomplete paratypes: 28 mm in paratype 1 (UANL-8144), 10 mm in paratype 3 (UANL-8144); 1 mm width (1 mm in paratypes 2 (UANL-8144) and 4 (ICML-EMU-12758); 0.5 mm in paratype 2 (UANL-8144), 0.8 mm in paratype 3 (UANL-8144). Body with 14 chaetigers (complete paratypes: 20 chaetigers in paratype 2 (UANL-8144); 15 chaetigers in paratype 4 (ICML-EMU-12758), two preanal achaetigerous segments with tori (two preanal achaetigers segments in all paratypes, except in paratype 1 (UANL-8144) whose posterior end is missing). Well-developed cephalic plaque, oval, with lateral margins smooth, entire (no incisions or notches) (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F5999123" captionText="Figure 2. Clymenura scutata sp. n. A) Entire worm, lateral view, tube removed, B-C) cephalic plaque, latero-dorsal views, D-E) posterior end and anal plaque, lateral and ventral view, respectively, F) anal plaque, posterior view, G) worm regenerating cephalic plaque and first anterior segments. A-D, F) Holotype LEMA-PO 153, E, G) Paratype UANL 8144. Scale bars: A) 2 mm, B-E, G) 1 mm, F) 0.6 mm." figureDoi="10.3897/BDJ.8.e57572.figure2" httpUri="https://binary.pensoft.net/fig/437506" pageId="0" pageNumber="57572">2</figureCitation>
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B-C) and posterior margin V-shaped (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F5999123" captionText="Figure 2. Clymenura scutata sp. n. A) Entire worm, lateral view, tube removed, B-C) cephalic plaque, latero-dorsal views, D-E) posterior end and anal plaque, lateral and ventral view, respectively, F) anal plaque, posterior view, G) worm regenerating cephalic plaque and first anterior segments. A-D, F) Holotype LEMA-PO 153, E, G) Paratype UANL 8144. Scale bars: A) 2 mm, B-E, G) 1 mm, F) 0.6 mm." figureDoi="10.3897/BDJ.8.e57572.figure2" httpUri="https://binary.pensoft.net/fig/437506" pageId="0" pageNumber="57572">2</figureCitation>
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C). Nuchal organs straight, parallel, almost full length of plaque (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F5999123" captionText="Figure 2. Clymenura scutata sp. n. A) Entire worm, lateral view, tube removed, B-C) cephalic plaque, latero-dorsal views, D-E) posterior end and anal plaque, lateral and ventral view, respectively, F) anal plaque, posterior view, G) worm regenerating cephalic plaque and first anterior segments. A-D, F) Holotype LEMA-PO 153, E, G) Paratype UANL 8144. Scale bars: A) 2 mm, B-E, G) 1 mm, F) 0.6 mm." figureDoi="10.3897/BDJ.8.e57572.figure2" httpUri="https://binary.pensoft.net/fig/437506" pageId="0" pageNumber="57572">2</figureCitation>
|
||
C). Palpode well developed with rounded tip (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F5999123" captionText="Figure 2. Clymenura scutata sp. n. A) Entire worm, lateral view, tube removed, B-C) cephalic plaque, latero-dorsal views, D-E) posterior end and anal plaque, lateral and ventral view, respectively, F) anal plaque, posterior view, G) worm regenerating cephalic plaque and first anterior segments. A-D, F) Holotype LEMA-PO 153, E, G) Paratype UANL 8144. Scale bars: A) 2 mm, B-E, G) 1 mm, F) 0.6 mm." figureDoi="10.3897/BDJ.8.e57572.figure2" httpUri="https://binary.pensoft.net/fig/437506" pageId="0" pageNumber="57572">2</figureCitation>
|
||
C). No ocelli. No segmental collars (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F5999123" captionText="Figure 2. Clymenura scutata sp. n. A) Entire worm, lateral view, tube removed, B-C) cephalic plaque, latero-dorsal views, D-E) posterior end and anal plaque, lateral and ventral view, respectively, F) anal plaque, posterior view, G) worm regenerating cephalic plaque and first anterior segments. A-D, F) Holotype LEMA-PO 153, E, G) Paratype UANL 8144. Scale bars: A) 2 mm, B-E, G) 1 mm, F) 0.6 mm." figureDoi="10.3897/BDJ.8.e57572.figure2" httpUri="https://binary.pensoft.net/fig/437506" pageId="0" pageNumber="57572">2</figureCitation>
|
||
A, Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F5999127" captionText="Figure 3. Clymenura scutata sp. n. A) Worm with cephalic plaque and first three chaetigers under regeneration, B) glandular shield in chaetiger 8, C) manubriavicular uncini from 1 st chaetiger, D) manubriavicular uncinus from 11 chaetiger, E) notochaetae from 11 chaetiger, F) third chaetiger, G) last chaetiger. A, G) Paratype UANL 8144, B-F) Holotype LEMA-PO 153. Scale bars: A-B) 1 mm, C-E) 1000 X magnification, F-G) 0.5 mm." figureDoi="10.3897/BDJ.8.e57572.figure3" httpUri="https://binary.pensoft.net/fig/448090" pageId="0" pageNumber="57572">3</figureCitation>
|
||
A). Ventral shield of chaetiger 8 half-oval shaped, revealed with methyl green (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F5999127" captionText="Figure 3. Clymenura scutata sp. n. A) Worm with cephalic plaque and first three chaetigers under regeneration, B) glandular shield in chaetiger 8, C) manubriavicular uncini from 1 st chaetiger, D) manubriavicular uncinus from 11 chaetiger, E) notochaetae from 11 chaetiger, F) third chaetiger, G) last chaetiger. A, G) Paratype UANL 8144, B-F) Holotype LEMA-PO 153. Scale bars: A-B) 1 mm, C-E) 1000 X magnification, F-G) 0.5 mm." figureDoi="10.3897/BDJ.8.e57572.figure3" httpUri="https://binary.pensoft.net/fig/448090" pageId="0" pageNumber="57572">3</figureCitation>
|
||
A-B). Chaetigers 11 and 12 are the longest (paratype 1, UANL-8144: chaetigers 10 and 11; paratype 2, UANL-8144: chaetigers 13-15). Notochaetae capillaries of one type, smooth by only slightly winged (Fig. 3E); those from anterior segments short (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F5999127" captionText="Figure 3. Clymenura scutata sp. n. A) Worm with cephalic plaque and first three chaetigers under regeneration, B) glandular shield in chaetiger 8, C) manubriavicular uncini from 1 st chaetiger, D) manubriavicular uncinus from 11 chaetiger, E) notochaetae from 11 chaetiger, F) third chaetiger, G) last chaetiger. A, G) Paratype UANL 8144, B-F) Holotype LEMA-PO 153. Scale bars: A-B) 1 mm, C-E) 1000 X magnification, F-G) 0.5 mm." figureDoi="10.3897/BDJ.8.e57572.figure3" httpUri="https://binary.pensoft.net/fig/448090" pageId="0" pageNumber="57572">3</figureCitation>
|
||
F), whereas those from posterior chaetigers as long was two times the length of capillaries from anterior segments (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F5999127" captionText="Figure 3. Clymenura scutata sp. n. A) Worm with cephalic plaque and first three chaetigers under regeneration, B) glandular shield in chaetiger 8, C) manubriavicular uncini from 1 st chaetiger, D) manubriavicular uncinus from 11 chaetiger, E) notochaetae from 11 chaetiger, F) third chaetiger, G) last chaetiger. A, G) Paratype UANL 8144, B-F) Holotype LEMA-PO 153. Scale bars: A-B) 1 mm, C-E) 1000 X magnification, F-G) 0.5 mm." figureDoi="10.3897/BDJ.8.e57572.figure3" httpUri="https://binary.pensoft.net/fig/448090" pageId="0" pageNumber="57572">3</figureCitation>
|
||
G). Single row of manubriavicular uncini present, starting in chaetiger 1 with 3-4 small teeth above the main fang, subrostral bristles or hairs or barbules not seen (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F5999127" captionText="Figure 3. Clymenura scutata sp. n. A) Worm with cephalic plaque and first three chaetigers under regeneration, B) glandular shield in chaetiger 8, C) manubriavicular uncini from 1 st chaetiger, D) manubriavicular uncinus from 11 chaetiger, E) notochaetae from 11 chaetiger, F) third chaetiger, G) last chaetiger. A, G) Paratype UANL 8144, B-F) Holotype LEMA-PO 153. Scale bars: A-B) 1 mm, C-E) 1000 X magnification, F-G) 0.5 mm." figureDoi="10.3897/BDJ.8.e57572.figure3" httpUri="https://binary.pensoft.net/fig/448090" pageId="0" pageNumber="57572">3</figureCitation>
|
||
C) and long and nearly straight handles. Manubriavicular uncini from median and posterior chaetigers similar shape to those present in chaetiger 1, but with developed subrostral bristles or hairs or barbules (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F5999127" captionText="Figure 3. Clymenura scutata sp. n. A) Worm with cephalic plaque and first three chaetigers under regeneration, B) glandular shield in chaetiger 8, C) manubriavicular uncini from 1 st chaetiger, D) manubriavicular uncinus from 11 chaetiger, E) notochaetae from 11 chaetiger, F) third chaetiger, G) last chaetiger. A, G) Paratype UANL 8144, B-F) Holotype LEMA-PO 153. Scale bars: A-B) 1 mm, C-E) 1000 X magnification, F-G) 0.5 mm." figureDoi="10.3897/BDJ.8.e57572.figure3" httpUri="https://binary.pensoft.net/fig/448090" pageId="0" pageNumber="57572">3</figureCitation>
|
||
D) and handles curved. First and second preanal segments about 1/2 and 2/3 length of last chaetigerous segment, respectively, with achaetous reduced torus (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F5999123" captionText="Figure 2. Clymenura scutata sp. n. A) Entire worm, lateral view, tube removed, B-C) cephalic plaque, latero-dorsal views, D-E) posterior end and anal plaque, lateral and ventral view, respectively, F) anal plaque, posterior view, G) worm regenerating cephalic plaque and first anterior segments. A-D, F) Holotype LEMA-PO 153, E, G) Paratype UANL 8144. Scale bars: A) 2 mm, B-E, G) 1 mm, F) 0.6 mm." figureDoi="10.3897/BDJ.8.e57572.figure2" httpUri="https://binary.pensoft.net/fig/437506" pageId="0" pageNumber="57572">2</figureCitation>
|
||
A-B). Pygidial funnel well developed (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F5999123" captionText="Figure 2. Clymenura scutata sp. n. A) Entire worm, lateral view, tube removed, B-C) cephalic plaque, latero-dorsal views, D-E) posterior end and anal plaque, lateral and ventral view, respectively, F) anal plaque, posterior view, G) worm regenerating cephalic plaque and first anterior segments. A-D, F) Holotype LEMA-PO 153, E, G) Paratype UANL 8144. Scale bars: A) 2 mm, B-E, G) 1 mm, F) 0.6 mm." figureDoi="10.3897/BDJ.8.e57572.figure2" httpUri="https://binary.pensoft.net/fig/437506" pageId="0" pageNumber="57572">2</figureCitation>
|
||
D-F), ending with 28 distinct triangular cirri (26-29 in paratypes), well separated and alternated short with long (twice longer than short ones) and with the ventral cirrus long (about twice the length of the long alternates) (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F5999123" captionText="Figure 2. Clymenura scutata sp. n. A) Entire worm, lateral view, tube removed, B-C) cephalic plaque, latero-dorsal views, D-E) posterior end and anal plaque, lateral and ventral view, respectively, F) anal plaque, posterior view, G) worm regenerating cephalic plaque and first anterior segments. A-D, F) Holotype LEMA-PO 153, E, G) Paratype UANL 8144. Scale bars: A) 2 mm, B-E, G) 1 mm, F) 0.6 mm." figureDoi="10.3897/BDJ.8.e57572.figure2" httpUri="https://binary.pensoft.net/fig/437506" pageId="0" pageNumber="57572">2</figureCitation>
|
||
A-B). Pygidial funnel with a distinctive callus ring (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F5999123" captionText="Figure 2. Clymenura scutata sp. n. A) Entire worm, lateral view, tube removed, B-C) cephalic plaque, latero-dorsal views, D-E) posterior end and anal plaque, lateral and ventral view, respectively, F) anal plaque, posterior view, G) worm regenerating cephalic plaque and first anterior segments. A-D, F) Holotype LEMA-PO 153, E, G) Paratype UANL 8144. Scale bars: A) 2 mm, B-E, G) 1 mm, F) 0.6 mm." figureDoi="10.3897/BDJ.8.e57572.figure2" httpUri="https://binary.pensoft.net/fig/437506" pageId="0" pageNumber="57572">2</figureCitation>
|
||
A). Anus terminal, located just in the centre of the funnel (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F5999123" captionText="Figure 2. Clymenura scutata sp. n. A) Entire worm, lateral view, tube removed, B-C) cephalic plaque, latero-dorsal views, D-E) posterior end and anal plaque, lateral and ventral view, respectively, F) anal plaque, posterior view, G) worm regenerating cephalic plaque and first anterior segments. A-D, F) Holotype LEMA-PO 153, E, G) Paratype UANL 8144. Scale bars: A) 2 mm, B-E, G) 1 mm, F) 0.6 mm." figureDoi="10.3897/BDJ.8.e57572.figure2" httpUri="https://binary.pensoft.net/fig/437506" pageId="0" pageNumber="57572">2</figureCitation>
|
||
F). Glandular pattern revealed with methyl green shows blue bands in anterior segments (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F5999127" captionText="Figure 3. Clymenura scutata sp. n. A) Worm with cephalic plaque and first three chaetigers under regeneration, B) glandular shield in chaetiger 8, C) manubriavicular uncini from 1 st chaetiger, D) manubriavicular uncinus from 11 chaetiger, E) notochaetae from 11 chaetiger, F) third chaetiger, G) last chaetiger. A, G) Paratype UANL 8144, B-F) Holotype LEMA-PO 153. Scale bars: A-B) 1 mm, C-E) 1000 X magnification, F-G) 0.5 mm." figureDoi="10.3897/BDJ.8.e57572.figure3" httpUri="https://binary.pensoft.net/fig/448090" pageId="0" pageNumber="57572">3</figureCitation>
|
||
A) and glandular shield on chaetiger 8 stains deep blue (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F5999127" captionText="Figure 3. Clymenura scutata sp. n. A) Worm with cephalic plaque and first three chaetigers under regeneration, B) glandular shield in chaetiger 8, C) manubriavicular uncini from 1 st chaetiger, D) manubriavicular uncinus from 11 chaetiger, E) notochaetae from 11 chaetiger, F) third chaetiger, G) last chaetiger. A, G) Paratype UANL 8144, B-F) Holotype LEMA-PO 153. Scale bars: A-B) 1 mm, C-E) 1000 X magnification, F-G) 0.5 mm." figureDoi="10.3897/BDJ.8.e57572.figure3" httpUri="https://binary.pensoft.net/fig/448090" pageId="0" pageNumber="57572">3</figureCitation>
|
||
B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="57572" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="57572">Diagnosis</paragraph>
|
||
<paragraph pageId="0" pageNumber="57572">A worm of 32 mm length, 1 mm width, with 14 chaetigers, two pre-anal achaetous segments with tori, manubriavicular uncini present from chaetiger 1 with 3-4 teeth above the main fang without barbules; manubriavicular uncini similar to that of chaetiger 1, but the number of barbules increases towards the most posterior segments. Cephalic plaque oval with entire, smooth margins; rounded palpode; nuchal organs straight, parallel, almost full length of plaque. No ocelli, no segmental collars. Anal funnel present with 28 distinct alternating triangular cirri: short and long (twice longer than short ones) and with the mid-ventral cirrus longest (about twice the length of the adjacent long). Chaetiger 8 glandular shield extending anteriorly and ventrally from notochaetal fascicle forming a half-oval.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="57572" type="etymology">
|
||
<paragraph pageId="0" pageNumber="57572">Etymology</paragraph>
|
||
<paragraph pageId="0" pageNumber="57572">
|
||
From the Latin
|
||
<taxonomicName authorityName="Tovar-Hernandez and Yanez-Rivera" authorityYear="2020" class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="scutata" pageId="0" pageNumber="57572" phylum="Annelida" rank="species" species="scutata">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">scutata</emphasis>
|
||
</taxonomicName>
|
||
, meaning armed with a shield and referring to the glandular shield on chaetiger 8.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="57572" type="taxon discussion">
|
||
<paragraph pageId="0" pageNumber="57572">Taxon discussion</paragraph>
|
||
<paragraph pageId="0" pageNumber="57572">
|
||
The specimens, here reviewed, match with the emendation of
|
||
<taxonomicName class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymenura" pageId="0" pageNumber="57572" phylum="Annelida" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Clymenura</emphasis>
|
||
</taxonomicName>
|
||
provided by
|
||
<bibRefCitation author="Read, G. B." journalOrPublisher="Zootaxa" pageId="0" pageNumber="57572" pagination="39 - 52" refId="B6001948" refString="Read, G. B., 2011. A new Clymenura (Polychaeta: Maldanidae) from the intertidal of Banks Peninsula, New Zealand, with a reassessment of Leiochone Grube, 1868 and Clymenura Verrill, 1900. Zootaxa 2934: 39 - 52" title="A new Clymenura (Polychaeta: Maldanidae) from the intertidal of Banks Peninsula, New Zealand, with a reassessment of Leiochone Grube, 1868 and Clymenura Verrill, 1900" volume="2934" year="2011">Read (2011)</bibRefCitation>
|
||
. The genus
|
||
<taxonomicName class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymenura" pageId="0" pageNumber="57572" phylum="Annelida" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Clymenura</emphasis>
|
||
</taxonomicName>
|
||
has never been reported in Mexico. The closest record to the Mexican Pacific of a
|
||
<taxonomicName class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymenura" pageId="0" pageNumber="57572" phylum="Annelida" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Clymenura</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<taxonomicName lsidName="C. gracilis" pageId="0" pageNumber="57572" rank="species" species="gracilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">C. gracilis</emphasis>
|
||
</taxonomicName>
|
||
Hartman, 1969 (
|
||
<bibRefCitation author="Hartman, O." journalOrPublisher="Allan Hancock Foundation, University of Southern California, Los Angeles" pageId="0" pageNumber="57572" refId="B5999756" refString="Hartman, O., 1969. Atlas of the sedentariate polychaetous annelids from California. Allan Hancock Foundation, University of Southern California, Los Angeles" title="Atlas of the sedentariate polychaetous annelids from California" year="1969">Hartman 1969</bibRefCitation>
|
||
), a species originally described from Santa Monica, California. However,
|
||
<bibRefCitation author="Read, G. B." journalOrPublisher="Zootaxa" pageId="0" pageNumber="57572" pagination="39 - 52" refId="B6001948" refString="Read, G. B., 2011. A new Clymenura (Polychaeta: Maldanidae) from the intertidal of Banks Peninsula, New Zealand, with a reassessment of Leiochone Grube, 1868 and Clymenura Verrill, 1900. Zootaxa 2934: 39 - 52" title="A new Clymenura (Polychaeta: Maldanidae) from the intertidal of Banks Peninsula, New Zealand, with a reassessment of Leiochone Grube, 1868 and Clymenura Verrill, 1900" volume="2934" year="2011">Read (2011)</bibRefCitation>
|
||
considers
|
||
<taxonomicName lsidName="C. gracilis" pageId="0" pageNumber="57572" rank="species" species="gracilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">C. gracilis</emphasis>
|
||
</taxonomicName>
|
||
as
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">incertae sedis</emphasis>
|
||
because the holotype lacks of ventral shield on chaetiger 8; consequently, it cannot be a member of
|
||
<taxonomicName class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymenura" pageId="0" pageNumber="57572" phylum="Annelida" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Clymenura</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="57572">
|
||
Amongst the nine valid taxa of
|
||
<taxonomicName class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymenura" pageId="0" pageNumber="57572" phylum="Annelida" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Clymenura</emphasis>
|
||
</taxonomicName>
|
||
worldwide, two species have been described from America:
|
||
<taxonomicName lsidName="C. cirrata" pageId="0" pageNumber="57572" rank="species" species="cirrata">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">C. cirrata</emphasis>
|
||
</taxonomicName>
|
||
(Ehlers, 1887 as
|
||
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1864" class="Polychaeta" family="Maldanidae" genus="Clymene" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymene" pageId="0" pageNumber="57572" phylum="Annelida" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Clymene</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation author="Ehlers, E" journalOrPublisher="Encyclopedique de Roret" pageId="0" pageNumber="57572" pagination="1 - 335" refId="B5999550" refString="Ehlers, E, 1887. Reports on the results of dredging, under the direction of L. F. Pourtales, during the years 1868-1870, and of Alexander Agassiz, in the Gulf of Mexico (1877-78), and in the Caribbean Sea (1878-79), in the U.S. Coast Survey steamer "Blake", Lieut-Com. C. D. Sigsbee, U.S.N. and Commander J. R. Bartlett, U.S.N., commanding. XXXI. Report on the Annelids. In: Memoirs of the Museum of Comparative Zoology, Harvard.: 1 - 335" title="Reports on the results of dredging, under the direction of L. F. Pourtales, during the years 1868 - 1870, and of Alexander Agassiz, in the Gulf of Mexico (1877 - 78), and in the Caribbean Sea (1878 - 79), in the U. S. Coast Survey steamer " Blake ", Lieut-Com. C. D. Sigsbee, U. S. N. and Commander J. R. Bartlett, U. S. N., commanding. XXXI. Report on the Annelids." volumeTitle="Memoirs of the Museum of Comparative Zoology, Harvard." year="1887">Ehlers 1887</bibRefCitation>
|
||
) from Florida and
|
||
<taxonomicName lsidName="C. columbiana" pageId="0" pageNumber="57572" rank="species" species="columbiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">C. columbiana</emphasis>
|
||
</taxonomicName>
|
||
(Berkeley, 1929 as
|
||
<taxonomicName genus="Leichone" lsidName="Leichone" pageId="0" pageNumber="57572" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Leichone</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation author="Berkeley, E" journalOrPublisher="Contributions to Canadian Biology and Fisheries" pageId="0" pageNumber="57572" pagination="307 - 316" refId="B5999314" refString="Berkeley, E, 1929. Polychaetous annelids from the Nanaimo District part 4. Chaetopteridae to Maldanidae. Contributions to Canadian Biology and Fisheries 4 (1): 307 - 316" title="Polychaetous annelids from the Nanaimo District part 4. Chaetopteridae to Maldanidae" volume="4" year="1929">Berkeley 1929</bibRefCitation>
|
||
) to western Canada (Table
|
||
<tableCitation captionStart="Table 2" captionStartId="T6002427" captionText="Table 2. Main features of some Clymenura species." httpUri="http://table.plazi.org/id/B66383E64A9FB7BA36937C6B6BD8FC02" pageId="0" pageNumber="57572" tableUuid="B66383E64A9FB7BA36937C6B6BD8FC02">2</tableCitation>
|
||
).
|
||
<taxonomicName authorityName="Tovar-Hernandez and Yanez-Rivera" authorityYear="2020" class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymenura scutata" pageId="0" pageNumber="57572" phylum="Annelida" rank="species" species="scutata">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Clymenura scutata</emphasis>
|
||
</taxonomicName>
|
||
sp. n., and
|
||
<taxonomicName lsidName="C. columbiana" pageId="0" pageNumber="57572" rank="species" species="columbiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">C. columbiana</emphasis>
|
||
</taxonomicName>
|
||
have an anal plaque with a long mid-ventral cirrus; however,
|
||
<taxonomicName authorityName="Tovar-Hernandez and Yanez-Rivera" authorityYear="2020" class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymenura scutata" pageId="0" pageNumber="57572" phylum="Annelida" rank="species" species="scutata">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Clymenura scutata</emphasis>
|
||
</taxonomicName>
|
||
sp. n. differs from
|
||
<taxonomicName lsidName="C. columbiana" pageId="0" pageNumber="57572" rank="species" species="columbiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">C. columbiana</emphasis>
|
||
</taxonomicName>
|
||
by:
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="57572">lateral margins of the cephalic plaque are entire,</paragraph>
|
||
<paragraph pageId="0" pageNumber="57572">does not present prostomial ocelli,</paragraph>
|
||
<paragraph pageId="0" pageNumber="57572">it has a marked dentition of uncini in first thoracic chaetiger and</paragraph>
|
||
<paragraph pageId="0" pageNumber="57572">
|
||
it has two preanal achaetous segments (cephalic plaque notched laterally, prostomial eyes present, reduced or reminiscent dentition of uncini in first thoracic chaetiger and three pre-anal achaetous segments in
|
||
<taxonomicName lsidName="C. columbiana" pageId="0" pageNumber="57572" rank="species" species="columbiana">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">C. columbiana</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="57572">
|
||
<taxonomicName baseAuthorityName="Ehlers" baseAuthorityYear="1887" class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymenura cirrata" pageId="0" pageNumber="57572" phylum="Annelida" rank="species" species="cirrata">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Clymenura cirrata</emphasis>
|
||
</taxonomicName>
|
||
is characterised by the presence of four long anal cirri (only the mid-ventral is long in
|
||
<taxonomicName authorityName="Tovar-Hernandez and Yanez-Rivera" authorityYear="2020" class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymenura scutata" pageId="0" pageNumber="57572" phylum="Annelida" rank="species" species="scutata">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Clymenura scutata</emphasis>
|
||
</taxonomicName>
|
||
sp. n.); the lateral margins or the cephalic plaque are notched (entire in
|
||
<taxonomicName authorityName="Tovar-Hernandez and Yanez-Rivera" authorityYear="2020" class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymenura scutata" pageId="0" pageNumber="57572" phylum="Annelida" rank="species" species="scutata">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Clymenura scutata</emphasis>
|
||
</taxonomicName>
|
||
sp. n.); and collars are present in chaetigers 2-4 (absent in
|
||
<taxonomicName authorityName="Tovar-Hernandez and Yanez-Rivera" authorityYear="2020" class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymenura scutata" pageId="0" pageNumber="57572" phylum="Annelida" rank="species" species="scutata">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Clymenura scutata</emphasis>
|
||
</taxonomicName>
|
||
sp. n.).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="57572">
|
||
The rest of the species in the genus were described from high north European latitudes and the Northern Pacific region: Novaya Zemlya (one species), Japan (three species), Norway (one species), the Laptev Sea (one species), except for
|
||
<taxonomicName authorityName="Read" authorityYear="2011" class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymenura snaiko" pageId="0" pageNumber="57572" phylum="Annelida" rank="species" species="snaiko">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Clymenura snaiko</emphasis>
|
||
</taxonomicName>
|
||
Read, 2011 (
|
||
<bibRefCitation author="Read, G. B." journalOrPublisher="Zootaxa" pageId="0" pageNumber="57572" pagination="39 - 52" refId="B6001948" refString="Read, G. B., 2011. A new Clymenura (Polychaeta: Maldanidae) from the intertidal of Banks Peninsula, New Zealand, with a reassessment of Leiochone Grube, 1868 and Clymenura Verrill, 1900. Zootaxa 2934: 39 - 52" title="A new Clymenura (Polychaeta: Maldanidae) from the intertidal of Banks Peninsula, New Zealand, with a reassessment of Leiochone Grube, 1868 and Clymenura Verrill, 1900" volume="2934" year="2011">Read 2011</bibRefCitation>
|
||
), that was described from New Zealand (see
|
||
<bibRefCitation author="Read, G. B." journalOrPublisher="Zootaxa" pageId="0" pageNumber="57572" pagination="39 - 52" refId="B6001948" refString="Read, G. B., 2011. A new Clymenura (Polychaeta: Maldanidae) from the intertidal of Banks Peninsula, New Zealand, with a reassessment of Leiochone Grube, 1868 and Clymenura Verrill, 1900. Zootaxa 2934: 39 - 52" title="A new Clymenura (Polychaeta: Maldanidae) from the intertidal of Banks Peninsula, New Zealand, with a reassessment of Leiochone Grube, 1868 and Clymenura Verrill, 1900" volume="2934" year="2011">Read 2011</bibRefCitation>
|
||
to further details).
|
||
<taxonomicName authorityName="Read" authorityYear="2011" class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymenura snaiko" pageId="0" pageNumber="57572" phylum="Annelida" rank="species" species="snaiko">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Clymenura snaiko</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Tovar-Hernandez and Yanez-Rivera" authorityYear="2020" class="Polychaeta" family="Maldanidae" genus="Clymenura" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Clymenura scutata" pageId="0" pageNumber="57572" phylum="Annelida" rank="species" species="scutata">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">Clymenura scutata</emphasis>
|
||
</taxonomicName>
|
||
sp. n., have the mid-ventral anal cirrus long, but the Mexican species have alternated cirri, whereas all other cirri are similar in length in
|
||
<taxonomicName lsidName="C. snaiko" pageId="0" pageNumber="57572" rank="species" species="snaiko">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">C. snaiko</emphasis>
|
||
</taxonomicName>
|
||
. In addition, the lateral margins of the cephalic plaque are notched in
|
||
<taxonomicName lsidName="C. snaiko" pageId="0" pageNumber="57572" rank="species" species="snaiko">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">C. snaiko</emphasis>
|
||
</taxonomicName>
|
||
versus entire in
|
||
<taxonomicName lsidName="C. scutata" pageId="0" pageNumber="57572" rank="species" species="scutata">
|
||
<emphasis italics="true" pageId="0" pageNumber="57572">C. scutata</emphasis>
|
||
</taxonomicName>
|
||
sp. n.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="57572">
|
||
Referring to intraspecific variation, a paratype was found under regeneration of the anterior end of the body (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F5999123" captionText="Figure 2. Clymenura scutata sp. n. A) Entire worm, lateral view, tube removed, B-C) cephalic plaque, latero-dorsal views, D-E) posterior end and anal plaque, lateral and ventral view, respectively, F) anal plaque, posterior view, G) worm regenerating cephalic plaque and first anterior segments. A-D, F) Holotype LEMA-PO 153, E, G) Paratype UANL 8144. Scale bars: A) 2 mm, B-E, G) 1 mm, F) 0.6 mm." figureDoi="10.3897/BDJ.8.e57572.figure2" httpUri="https://binary.pensoft.net/fig/437506" pageId="0" pageNumber="57572">2</figureCitation>
|
||
G). The cephalic plaque and the first three segments are a half narrower than segment 4 and those subsequent. However, the glandular shield on chaetiger 8 is present, it maintain the half-oval shape. As the number of chaetigers (and size) varies in all paratypes, their use, therefore, cannot be considered diagnostic.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |