170 lines
17 KiB
XML
170 lines
17 KiB
XML
<document id="9D8757530DB8B074AA3CBF9ED09B850A" ID-DOI="10.1649/1066.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5369967" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630268265367" checkinUser="valdenar" docAuthor="Ratcliffe, Brett C." docDate="1821" docId="041BEE3DB05DA77CFEB1FBAEFC5DFA2E" docLanguage="en" docName="1066.1.pdf" docOrigin="The Coleopterists Bulletin 62 (2)" docSource="http://dx.doi.org/10.1649/1066.1" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Cyclocephala perplexa Ratcliffe 2008, new species" docType="treatment" docVersion="3" lastPageNumber="241" masterDocId="F8229645B04CA76EFFDDFFBAFF8DFFAC" masterDocTitle="More New Species ofCyclocephalaDejean, 1821 from South America (Scarabaeidae: Dynastinae: Cyclocephalini)" masterLastPageNumber="241" masterPageNumber="221" pageNumber="221" updateTime="1699205293535" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="B6017BCF679E4CC0EC350077307B860E">
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<mods:title id="7480B481B792274723AB37F1FA14FD1A">More New Species ofCyclocephalaDejean, 1821 from South America (Scarabaeidae: Dynastinae: Cyclocephalini)</mods:title>
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<mods:name id="12D1987BB8B048E0A90C5C6E6CA77472" type="personal">
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<mods:roleTerm id="28FBDD21C44C83D0105CBF9D714290C1">Author</mods:roleTerm>
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<mods:namePart id="AEF6BBFEE6A6138370F3FCCDF9FECF40">Ratcliffe, Brett C.</mods:namePart>
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<mods:typeOfResource id="6F3B263E31F629904442205211D3763F">text</mods:typeOfResource>
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<mods:title id="F584FF478DD54705AB552DE04E062F6C">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="F1A0D4CC333FFE92FED8EC1C0E87875D">
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<mods:date id="9F8DFCD53C6D8B27696E237517A75DF8">1821</mods:date>
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<mods:detail id="2D37FED2FCBA7266270EBDD7B9ACC462" type="pubDate">
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<mods:number id="040EB6ECBDA88B93C63521D62D201ED7">2008-06-30</mods:number>
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<mods:detail id="83818D656E8D5982AA77DD234DC419D5" type="volume">
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<mods:number id="EB4A24FB670DF5C2C21DC180CB1108FE">62</mods:number>
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<mods:detail id="5A59FF6E50C0AC74A531749B9D3B0538" type="issue">
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<mods:number id="E4458BAECB82169705836B58C0E07207">2</mods:number>
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<mods:extent id="447D4D508737B6CBF090095296CCEC2A" unit="page">
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<mods:start id="A3C5514E5D14DAD0E55725BDC87505F8">221</mods:start>
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<mods:end id="39A8F03369AFCA3B298886E23D219FFF">241</mods:end>
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</mods:extent>
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<mods:location id="50DAF22F486B19271A92FDD67EE21EED">
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<mods:url id="5ACF85CA57F44BEC7077AC3DE9CC39BC">http://dx.doi.org/10.1649/1066.1</mods:url>
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</mods:location>
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<mods:classification id="4BFA1E165EF9D70C8983A624230B9DA9">journal article</mods:classification>
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<mods:identifier id="6F6739A4FF43ECEBDF349248A0D02F7A" type="DOI">10.1649/1066.1</mods:identifier>
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<mods:identifier id="3DBC47A924DB7907B5F3235F7F354408" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="88C55CFA05E71EBAF27F45317986C429" type="Zenodo-Dep">5369967</mods:identifier>
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</mods:mods>
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<treatment id="041BEE3DB05DA77CFEB1FBAEFC5DFA2E" LSID="urn:lsid:plazi:treatment:041BEE3DB05DA77CFEB1FBAEFC5DFA2E" httpUri="http://treatment.plazi.org/id/041BEE3DB05DA77CFEB1FBAEFC5DFA2E" lastPageId="18" pageId="17">
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<subSubSection id="C4A80CA0B05DA77FFEB1FBAEFCA2FB86" box="[364,815,1044,1066]" pageId="17" type="nomenclature">
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<paragraph id="8C0D5F2BB05DA77FFEB1FBAEFCA2FB86" blockId="17.[364,815,1044,1093]" box="[364,815,1044,1066]" pageId="17">
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<heading id="D745E847B05DA77FFEB1FBAEFCA2FB86" box="[364,815,1044,1066]" centered="true" fontSize="9" level="2" pageId="17" reason="2">
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<emphasis id="BEC68339B05DA77FFEB1FBAEFCA2FB86" bold="true" box="[364,815,1044,1066]" pageId="17">
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<taxonomicName id="4BB224A8B05DA77FFEB1FBAEFD3DFB86" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2008" box="[364,688,1044,1066]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="species" species="perplexa" status="sp. nov.">
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<emphasis id="BEC68339B05DA77FFEB1FBAEFDC2FB86" bold="true" box="[364,591,1044,1066]" italics="true" pageId="17">Cyclocephala perplexa</emphasis>
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Ratcliffe
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</taxonomicName>
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,
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<taxonomicNameLabel id="A5F53E42B05DA77FFD60FBAEFCA2FB86" box="[701,815,1044,1066]" pageId="17" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C4A80CA0B05DA77FFDD6FB95FD1CFBE9" box="[523,657,1071,1093]" pageId="17" type="description">
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<paragraph id="8C0D5F2BB05DA77FFDD6FB95FD1CFBE9" blockId="17.[364,815,1044,1093]" box="[523,657,1071,1093]" pageId="17">
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(
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<figureCitation id="148943AEB05DA77FFDCFFB95FD07FBE9" box="[530,650,1071,1093]" captionStart="Figs" captionStartId="18.[167,206,858,877]" captionTargetBox="[156,976,195,825]" captionTargetId="figure-260@18.[146,976,194,836]" captionTargetPageId="18" captionText="Figs. 34–36. Cyclocephala perplexa: 34) dorsal view of holotype; 35–36) parameres" figureDoi="http://doi.org/10.5281/zenodo.5369983" httpUri="https://zenodo.org/record/5369983/files/figure.png" pageId="17">Figs. 34–36</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C4A80CA0B05DA77CFF1AFBDAFCFDFBA1" lastPageId="18" pageId="17" type="materials_examined">
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<paragraph id="8C0D5F2BB05DA77FFF1AFBDAFDCBFB00" blockId="17.[175,1005,1120,1570]" pageId="17">
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<emphasis id="BEC68339B05DA77FFF1AFBDAFEE8FBDA" bold="true" box="[199,357,1120,1142]" pageId="17">Type Material.</emphasis>
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<materialsCitation id="3CDA5576B05DA77FFEAFFBDAFD96FB3D" collectingDate="1925-09" country="Bolivia" location="San Jose" pageId="17" specimenCount="1" stateProvince="La Paz" typeStatus="holotype">
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<typeStatus id="5309E189B05DA77FFEAFFBDAFE5FFBDA" box="[370,466,1120,1142]" pageId="17" type="holotype">Holotype</typeStatus>
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male, labeled ‘‘
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<location id="896D09F0B05DA77FFD5CFBDAFD68FBDA" LSID="urn:lsid:plazi:treatment:041BEE3DB05DA77CFEB1FBAEFC5DFA2E:896D09F0B05DA77FFD5CFBDAFD68FBDA" box="[641,741,1120,1142]" country="Bolivia" name="San Jose" pageId="17" stateProvince="La Paz">San Jose</location>
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, Dept.
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<collectingRegion id="4E7691C9B05DA77FFCE1FBDAFC01FBDA" box="[828,908,1120,1142]" country="Bolivia" name="La Paz" pageId="17">La Paz</collectingRegion>
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,
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<collectingCountry id="F4A51FBBB05DA77FFC40FBDAFC65FBDA" box="[925,1000,1120,1142]" name="Bolivia" pageId="17">Bolivia</collectingCountry>
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,
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<date id="F80C79EBB05DA77FFF72FBC1FED0FB3D" box="[175,349,1147,1169]" pageId="17" value="1925-09">
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<collectingDate id="E8488003B05DA77FFF72FBC1FED0FB3D" box="[175,349,1147,1169]" pageId="17" value="1925-09">September 1925</collectingDate>
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</date>
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, GLHarrington.’’
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</materialsCitation>
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<materialsCitation id="3CDA5576B05DA77FFDF6FBC1FDCFFB00" collectorName="Lincoln, NE" country="Bolivia" location="University of Nebraska State Museum" pageId="17" specimenCount="1" stateProvince="La Paz" typeStatus="holotype">
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<typeStatus id="5309E189B05DA77FFDF6FBC1FD06FB3D" box="[555,651,1147,1169]" pageId="17" type="holotype">Holotype</typeStatus>
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deposited at the
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<location id="896D09F0B05DA77FFC80FBC1FE3FFB00" LSID="urn:lsid:plazi:treatment:041BEE3DB05DA77CFEB1FBAEFC5DFA2E:896D09F0B05DA77FFC80FBC1FE3FFB00" country="Bolivia" name="University of Nebraska State Museum" pageId="17" stateProvince="La Paz">University of Nebraska State Museum</location>
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,
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<collectorName id="21473AFDB05DA77FFE60FB2CFDCFFB00" box="[445,578,1174,1196]" pageId="17">Lincoln, NE</collectorName>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8C0D5F2BB05DA77CFF1AFB0BFE09FC29" blockId="17.[175,1005,1120,1570]" lastBlockId="18.[146,975,858,901]" lastPageId="18" pageId="17">
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<emphasis id="BEC68339B05DA77FFF1AFB0BFEA7FB6B" bold="true" box="[199,298,1201,1223]" pageId="17">
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<typeStatus id="5309E189B05DA77FFF1AFB0BFEABFB6B" box="[199,294,1201,1223]" pageId="17" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Male. Length
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<quantity id="4B4AF2CEB05DA77FFE12FB0BFDA7FB6B" box="[463,554,1201,1223]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="17" unit="mm" value="12.5">12.5 mm</quantity>
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; width across humeri
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<quantity id="4B4AF2CEB05DA77FFCC4FB0BFCE4FB6B" box="[793,873,1201,1223]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="17" unit="mm" value="7.2">7.2 mm</quantity>
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. Color dark testaceous, shining, except for black, longitudinal, externally undulating band on pronotum either side of middle and broad, piceous margins (all) of each elytron.
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<emphasis id="BEC68339B05DA77FFF72FAB8FF69FABB" box="[175,228,1282,1303]" italics="true" pageId="17">Head</emphasis>
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: Surface completely, densely rugopunctate, setigerous; setae moderately dense, moderately long, tawny. Frontoclypeal suture arcuate. Clypeus with apex broadly rounded on angles, subtruncate, with distinct marginal bead. Eyes small, interocular width equals 3.8 transverse eye diameters. Antenna with 10 segments, club 3- segmented, long, subequal in length to segments 1–7.
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<emphasis id="BEC68339B05DA77FFD31FAD6FCC3FA2D" box="[748,846,1388,1409]" italics="true" pageId="17">Pronotum</emphasis>
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: Surface finely shagreened, moderately densely punctate; punctures moderate in size, setigerous; setae moderate in density and length, tawny. Base with complete marginal bead. Posterior angles broadly rounded.
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<emphasis id="BEC68339B05DA77FFDD2FA07FDC0FA7E" box="[527,589,1469,1490]" italics="true" pageId="17">Elytra</emphasis>
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: Surface coarsely, irregularly roughened, rows of punctures indistinct, setigerous; setae moderate in density and length, tawny.
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<taxonomicName id="4BB224A8B05DA77FFF25FA48FE26F9A4" authority=": Surface" authorityName="Surface" box="[248,427,1522,1544]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="17" phylum="Chordata" rank="genus">
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<emphasis id="BEC68339B05DA77FFF25FA48FED8F9AB" box="[248,341,1522,1543]" italics="true" pageId="17">Pygidium</emphasis>
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: Surface
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</taxonomicName>
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opaque, finely shagreened, with small punctures moderate in density, setigerous; setae moderately dense, long, tawny. In lateral view, surface 238 (caudal and lateral views).
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</paragraph>
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<caption id="D8CD0FA3B05EA77CFF7AFCE0FC42FCC1" ID-DOI="http://doi.org/10.5281/zenodo.5369983" ID-Zenodo-Dep="5369983" box="[167,975,858,877]" httpUri="https://zenodo.org/record/5369983/files/figure.png" pageId="18" startId="18.[167,206,858,877]" targetBox="[156,976,195,825]" targetPageId="18">
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<paragraph id="8C0D5F2BB05EA77CFF7AFCE0FC42FCC1" blockId="18.[146,975,858,901]" box="[167,975,858,877]" pageId="18">
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<emphasis id="BEC68339B05EA77CFF7AFCE0FE95FCC1" bold="true" box="[167,280,858,877]" pageId="18">Figs. 34–36.</emphasis>
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<taxonomicName id="4BB224A8B05EA77CFEF0FCE0FE77FCC1" authorityName="Ratcliffe" authorityYear="2008" box="[301,506,858,877]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="18" phylum="Arthropoda" rank="species" species="perplexa">
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<emphasis id="BEC68339B05EA77CFEF0FCE0FE77FCC1" box="[301,506,858,877]" italics="true" pageId="18">Cyclocephala perplexa</emphasis>
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</taxonomicName>
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:
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<emphasis id="BEC68339B05EA77CFDD6FCE0FDA5FCC1" bold="true" box="[523,552,858,877]" pageId="18">34)</emphasis>
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dorsal view of holotype;
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<emphasis id="BEC68339B05EA77CFCF7FCE0FCEBFCC1" bold="true" box="[810,870,858,877]" pageId="18">35–36)</emphasis>
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parameres
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</paragraph>
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</caption>
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<paragraph id="8C0D5F2BB05EA77CFF4FFC1DFCFDFBA1" blockId="18.[146,976,935,1410]" pageId="18">
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nearly flat.
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<emphasis id="BEC68339B05EA77CFED0FC1DFEB1FC10" box="[269,316,935,956]" italics="true" pageId="18">Legs</emphasis>
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: Protibia tridentate, basal tooth slightly removed from others. Protarsi missing. Metatarsi missing.
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<emphasis id="BEC68339B05EA77CFE25FC78FDB7FC7B" box="[504,570,962,983]" italics="true" pageId="18">Venter</emphasis>
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: Prosternal process long, columnar, apex expanded, transversely oval, with transverse, raised ‘‘button’’ on anterior half. Thoracic sternites with long, dense, tawny setae.
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<emphasis id="BEC68339B05EA77CFDA2FC4DFD65FBA0" box="[639,744,1015,1036]" italics="true" pageId="18">Parameres</emphasis>
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:
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<figureCitation id="148943AEB05EA77CFD28FC4DFCE1FBA1" box="[757,876,1015,1037]" captionStart="Figs" captionStartId="18.[167,206,858,877]" captionTargetBox="[156,976,195,825]" captionTargetId="figure-260@18.[146,976,194,836]" captionTargetPageId="18" captionText="Figs. 34–36. Cyclocephala perplexa: 34) dorsal view of holotype; 35–36) parameres" figureDoi="http://doi.org/10.5281/zenodo.5369983" httpUri="https://zenodo.org/record/5369983/files/figure.png" pageId="18">Figs. 35–36</figureCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C4A80CA0B05EA77CFF77FBABFCE8FBF1" pageId="18" type="etymology">
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<paragraph id="8C0D5F2BB05EA77CFF77FBABFCE8FBF1" blockId="18.[146,976,935,1410]" pageId="18">
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<emphasis id="BEC68339B05EA77CFF77FBABFE90FB8B" bold="true" box="[170,285,1041,1063]" pageId="18">Etymology.</emphasis>
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The combination of character states for this species is unique (especially the short, robust body form) and the absence of all the tarsomeres presents some degree of uncertainty. Hence, this species is perplexing.
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</paragraph>
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</subSubSection>
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<subSubSection id="C4A80CA0B05EA77CFF77FBD8FE62FB6B" pageId="18" type="distribution">
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<paragraph id="8C0D5F2BB05EA77CFF77FBD8FE62FB6B" blockId="18.[146,976,935,1410]" pageId="18">
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<emphasis id="BEC68339B05EA77CFF77FBD8FEA5FBD4" bold="true" box="[170,296,1122,1144]" pageId="18">Distribution.</emphasis>
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<taxonomicName id="4BB224A8B05EA77CFEEDFBD8FD9FFBDB" authorityName="Ratcliffe" authorityYear="2008" box="[304,530,1122,1143]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="18" phylum="Arthropoda" rank="species" species="perplexa">
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<emphasis id="BEC68339B05EA77CFEEDFBD8FD9FFBDB" box="[304,530,1122,1143]" italics="true" pageId="18">Cyclocephala perplexa</emphasis>
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</taxonomicName>
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is known only from, presumably, San José near the Rio
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<collectingRegion id="4E7691C9B05EA77CFEC5FBC6FECBFB3E" box="[280,326,1148,1170]" country="Bolivia" name="Beni" pageId="18">Beni</collectingRegion>
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to the northeast of
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<collectingRegion id="4E7691C9B05EA77CFDCCFBC6FDD4FB3E" box="[529,601,1148,1170]" country="Bolivia" name="La Paz" pageId="18">La Paz</collectingRegion>
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at an elevation of 500–1,000 meters. The habitat has, no doubt, been drastically changed since the specimen was collected more than 80 years ago.
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</paragraph>
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</subSubSection>
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<subSubSection id="C4A80CA0B05EA77CFF77FB76FC5DFA2E" pageId="18" type="diagnosis">
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<paragraph id="8C0D5F2BB05EA77CFF77FB76FC5DFA2E" blockId="18.[146,976,935,1410]" pageId="18">
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<emphasis id="BEC68339B05EA77CFF77FB76FE9FFB4E" bold="true" box="[170,274,1228,1250]" pageId="18">Diagnosis.</emphasis>
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The combination of relatively short and broad body form, coarse elytral sculpturing, color pattern, long antennal club, small eyes, dense setae both dorsally and ventrally, and the simple parameres is unique for this species. This specimen of
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<taxonomicName id="4BB224A8B05EA77CFECFFAA6FE08FA9D" authorityName="Ratcliffe" authorityYear="2008" box="[274,389,1308,1329]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="18" phylum="Arthropoda" rank="species" species="perplexa">
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<emphasis id="BEC68339B05EA77CFECFFAA6FE08FA9D" box="[274,389,1308,1329]" italics="true" pageId="18">C. perplexa</emphasis>
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</taxonomicName>
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will not key past couplet
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<quantity id="4B4AF2CEB05EA77CFD4DFAA6FD4EFA9E" box="[656,707,1308,1330]" metricMagnitude="0" metricUnit="m" metricValue="1.3462" pageId="18" unit="in" value="53.0">53 in</quantity>
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<bibRefCitation id="E82322DAB05EA77CFD16FAA6FCE9FA9E" author="Endrodi" box="[715,868,1308,1330]" pageId="18" refId="ref10514" refString="Endrodi, S. 1985. The Dynastinae of the World. W. Junk Publishers, Dordrecht. 800 pp." type="book" year="1985">Endrödi (1985)</bibRefCitation>
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because it lacks protarsi. However, it is possible to go past this point because the parameres are different than those of species that occur in this section of the key. At couplet 100, the absence of protarsal claws makes it impossible to proceed any further in the key.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |