treatments-xml/data/04/1B/EE/041BEE3DB04AA769FE82FB90FCEEFB27.xml
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<document id="8E65555F9EC6BEA351CD9205B7C4CE86" ID-DOI="10.1649/1066.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5369967" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630268265367" checkinUser="valdenar" docAuthor="Ratcliffe, Brett C." docDate="1821" docId="041BEE3DB04AA769FE82FB90FCEEFB27" docLanguage="en" docName="1066.1.pdf" docOrigin="The Coleopterists Bulletin 62 (2)" docSource="http://dx.doi.org/10.1649/1066.1" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Cyclocephala chera Ratcliffe 2008, new species" docType="treatment" docVersion="4" lastPageNumber="241" masterDocId="F8229645B04CA76EFFDDFFBAFF8DFFAC" masterDocTitle="More New Species ofCyclocephalaDejean, 1821 from South America (Scarabaeidae: Dynastinae: Cyclocephalini)" masterLastPageNumber="241" masterPageNumber="221" pageNumber="221" updateTime="1699205293535" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="529DC4EE4E6AFACF02B33993477E4186">More New Species ofCyclocephalaDejean, 1821 from South America (Scarabaeidae: Dynastinae: Cyclocephalini)</mods:title>
</mods:titleInfo>
<mods:name id="1481E1666C72A2430706FD4E74E95380" type="personal">
<mods:role id="83D78C059C8A54D1BF48DDE4698301FE">
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<mods:namePart id="BDB1B89FCDD43F9F7FBB7DC307E4DB3A">Ratcliffe, Brett C.</mods:namePart>
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<mods:title id="AB302CBB745F959DF1672CAAA1B1BD25">The Coleopterists Bulletin</mods:title>
</mods:titleInfo>
<mods:part id="7E8D0C8965958662EFA8BD109618344B">
<mods:date id="7B47D2BE7C37BDAEC5A3F587EFF15D29">1821</mods:date>
<mods:detail id="5D1CA3D7BB790F383AA8A695F0229CD0" type="pubDate">
<mods:number id="F96FC47985E9D588696B62DDA962D138">2008-06-30</mods:number>
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<mods:classification id="0833421DCEF765AC54A7A98415267EC7">journal article</mods:classification>
<mods:identifier id="47D9FF357354DD8E412CB66C273BD168" type="DOI">10.1649/1066.1</mods:identifier>
<mods:identifier id="EFC4C31F1287EF871AB3B759F10E3137" type="ISSN">1938-4394</mods:identifier>
<mods:identifier id="2AFB86C7441B748FC4933CAD112B1349" type="Zenodo-Dep">5369967</mods:identifier>
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<treatment id="041BEE3DB04AA769FE82FB90FCEEFB27" ID-DOI="http://doi.org/10.5281/zenodo.5461600" ID-Zenodo-Dep="5461600" LSID="urn:lsid:plazi:treatment:041BEE3DB04AA769FE82FB90FCEEFB27" httpUri="http://treatment.plazi.org/id/041BEE3DB04AA769FE82FB90FCEEFB27" lastPageId="7" pageId="6">
<subSubSection id="C4A80CA0B04AA768FE82FB90FC8FFBEC" box="[351,770,1066,1088]" pageId="6" type="nomenclature">
<paragraph id="8C0D5F2BB04AA768FE82FB90FC8FFBEC" blockId="6.[351,770,1066,1116]" box="[351,770,1066,1088]" pageId="6">
<heading id="D745E847B04AA768FE82FB90FC8FFBEC" box="[351,770,1066,1088]" centered="true" fontSize="9" level="2" pageId="6" reason="2">
<emphasis id="BEC68339B04AA768FE82FB90FC8FFBEC" bold="true" box="[351,770,1066,1088]" pageId="6">
<taxonomicName id="4BB224A8B04AA768FE82FB90FD0EFBEC" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2008" box="[351,643,1066,1088]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="chera" status="sp. nov.">
<emphasis id="BEC68339B04AA768FE82FB90FDAFFBEC" bold="true" box="[351,546,1066,1088]" italics="true" pageId="6">Cyclocephala chera</emphasis>
Ratcliffe
</taxonomicName>
,
<taxonomicNameLabel id="A5F53E42B04AA768FD52FB90FC8FFBEC" box="[655,770,1066,1088]" pageId="6" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C4A80CA0B04AA768FDD7FBFCFDDAFBF0" box="[522,599,1094,1116]" pageId="6" type="description">
<paragraph id="8C0D5F2BB04AA768FDD7FBFCFDDAFBF0" blockId="6.[351,770,1066,1116]" box="[522,599,1094,1116]" pageId="6">
(
<figureCitation id="148943AEB04AA768FDCFFBFCFDC2FBF0" box="[530,591,1094,1116]" captionStart="Fig" captionStartId="6.[167,198,785,804]" captionText="Fig. 8. Cyclocephala chera, dorsal view of paratype." pageId="6">Fig. 8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C4A80CA0B04AA769FF77FBCDFDBDFE4A" lastPageId="7" pageId="6" type="materials_examined">
<paragraph id="8C0D5F2BB04AA768FF77FBCDFE40FB54" blockId="6.[146,976,1143,1570]" pageId="6">
<emphasis id="BEC68339B04AA768FF77FBCDFECAFB21" bold="true" box="[170,327,1143,1165]" pageId="6">Type Material.</emphasis>
<materialsCitation id="3CDA5576B04AA768FE8EFBCDFE40FB54" collectionCode="R" collectorName="Davis &amp; Pogue &amp; Solis. &amp; Five" country="Guyana" elevation="275" latitude="9.516666" location="Burra Burra" longLatPrecision="21" longitude="-9.983334" pageId="6" specimenCount="2" typeStatus="holotype">
<typeStatus id="5309E189B04AA768FE8EFBCDFE3EFB21" box="[339,435,1143,1165]" pageId="6" type="holotype">Holotype</typeStatus>
female, labeled
<collectingCountry id="F4A51FBBB04AA768FDAEFBCDFD61FB21" box="[627,748,1143,1165]" name="Guyana" pageId="6">GUYANA</collectingCountry>
: White Water Cmp.,
<location id="896D09F0B04AA768FF4FFB28FE93FB04" LSID="urn:lsid:plazi:treatment:041BEE3DB04AA769FE82FB90FCEEFB27:896D09F0B04AA768FF4FFB28FE93FB04" box="[146,286,1170,1192]" country="Guyana" latitude="9.516666" longLatPrecision="21" longitude="-9.983334" name="Burra Burra" pageId="6">Burra Burra</location>
(sic)
<collectionCode id="EAA3C7EEB04AA768FEA9FB28FE04FB04" box="[372,393,1170,1192]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="6">R</collectionCode>
. Iwokrama Res.,
<quantity id="4B4AF2CEB04AA768FDB4FB28FD56FB04" box="[617,731,1170,1192]" metricMagnitude="2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="6" unit="m" value="275.0" valueMax="300.0" valueMin="250.0">
<elevation id="079FB818B04AA768FDB4FB28FD56FB04" box="[617,731,1170,1192]" metricMagnitude="2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="6" unit="m" value="275.0" valueMax="300.0" valueMin="250.0">250300 m</elevation>
</quantity>
, 16.II.02, 4
<emphasis id="BEC68339B04AA768FCB1FB28FCF8FB0B" bold="true" box="[876,885,1170,1191]" italics="true" pageId="6">°</emphasis>
40
<geoCoordinate id="E98639ECB04AA768FC50FB2BFC45FB04" box="[909,968,1169,1192]" degrees="9" direction="north" minutes="31" orientation="latitude" pageId="6" precision="15" seconds="0" value="9.516666">
<emphasis id="BEC68339B04AA768FC50FB2BFC18FB0B" bold="true" box="[909,917,1169,1191]" pageId="6">9</emphasis>
31
<emphasis id="BEC68339B04AA768FC70FB2BFC38FB0B" bold="true" box="[941,949,1169,1191]" pageId="6">0</emphasis>
N
</geoCoordinate>
/ 58
<emphasis id="BEC68339B04AA768FF77FB16FF3EFB6D" bold="true" box="[170,179,1196,1217]" italics="true" pageId="6">°</emphasis>
40
<geoCoordinate id="E98639ECB04AA768FF16FB16FE80FB6E" box="[203,269,1196,1218]" degrees="9" direction="west" minutes="59" orientation="longitude" pageId="6" precision="15" seconds="0" value="-9.983334">
<emphasis id="BEC68339B04AA768FF16FB16FF5EFB6E" bold="true" box="[203,211,1196,1218]" pageId="6">9</emphasis>
59
<emphasis id="BEC68339B04AA768FF36FB16FF7EFB6E" bold="true" box="[235,243,1196,1218]" pageId="6">0</emphasis>
W
</geoCoordinate>
,
<collectorName id="21473AFDB04AA768FEC2FB16FED0FB6E" box="[287,349,1196,1218]" pageId="6">Davis</collectorName>
,
<collectorName id="21473AFDB04AA768FEB2FB16FE23FB6E" box="[367,430,1196,1218]" pageId="6">Pogue</collectorName>
&amp;
<collectorName id="21473AFDB04AA768FE03FB16FD94FB6E" box="[478,537,1196,1218]" pageId="6">Solis.</collectorName>
<collectorName id="21473AFDB04AA768FDEDFB16FDD0FB6E" box="[560,605,1196,1218]" pageId="6">Five</collectorName>
<typeStatus id="5309E189B04AA768FDB6FB16FD43FB6E" box="[619,718,1196,1218]" pageId="6" type="paratype">paratypes</typeStatus>
with same data;
<specimenCount id="9AB494A2B04AA768FC41FB16FF78FB72" pageId="6" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
with same data but with date of 15.II.02, and
<specimenCount id="9AB494A2B04AA768FD07FB72FCEDFB72" box="[730,864,1224,1246]" pageId="6" type="generic" typeStatus="paratype">one paratype</specimenCount>
with same data but with date of 14.II.02.
</materialsCitation>
</paragraph>
<paragraph id="8C0D5F2BB04AA768FF77FB44FEA7FAC9" blockId="6.[146,976,1143,1570]" pageId="6">
<materialsCitation id="3CDA5576B04AA768FF77FB44FEA7FAC9" collectorName="Lincoln, NE &amp; Brett C. Ratcliffe Collection" country="United States of America" location="Washington" pageId="6" specimenCount="3" stateProvince="Washington" typeStatus="holotype">
<typeStatus id="5309E189B04AA768FF77FB44FE87FAB8" box="[170,266,1278,1300]" pageId="6" type="holotype">Holotype</typeStatus>
and
<specimenCount id="9AB494A2B04AA768FE91FB44FE7DFAB8" box="[332,496,1278,1300]" pageId="6" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
deposited at the
<collectingCountry id="F4A51FBBB04AA768FD60FB44FD75FAB8" box="[701,760,1278,1300]" name="United States of America" pageId="6">U. S.</collectingCountry>
National Museum,
<collectingRegion id="4E7691C9B04AA768FF4FFAA3FE9CFA83" box="[146,273,1305,1327]" country="United States of America" name="Washington" pageId="6">Washington</collectingRegion>
, D. C.;
<specimenCount id="9AB494A2B04AA768FEBFFAA3FE7FFA83" box="[354,498,1305,1327]" pageId="6" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
deposited at the University of
<collectingRegion id="4E7691C9B04AA768FCEFFAA3FC42FA83" box="[818,975,1305,1327]" country="United States of America" name="Nebraska" pageId="6">Nebraska State</collectingRegion>
Museum,
<collectorName id="21473AFDB04AA768FF24FA8EFEF6FAE6" box="[249,379,1332,1354]" pageId="6">Lincoln, NE</collectorName>
;
<specimenCount id="9AB494A2B04AA768FE5AFA8EFD91FAE6" box="[391,540,1332,1354]" pageId="6" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
placed in the
<collectorName id="21473AFDB04AA768FD70FA8EFC42FAE6" box="[685,975,1332,1354]" pageId="6">Brett C. Ratcliffe Collection</collectorName>
(
<collectorName id="21473AFDB04AA768FF44FAF5FE93FAC9" box="[153,286,1359,1381]" pageId="6">Lincoln, NE</collectorName>
).
</materialsCitation>
</paragraph>
<paragraph id="8C0D5F2BB04AA769FF77FAD1FDBDFE4A" blockId="6.[146,976,1143,1570]" lastBlockId="7.[175,1005,194,1163]" lastPageId="7" pageId="6">
<emphasis id="BEC68339B04AA768FF77FAD1FE80FA2D" bold="true" box="[170,269,1387,1409]" pageId="6">
<typeStatus id="5309E189B04AA768FF77FAD1FE84FA2D" box="[170,265,1387,1409]" pageId="6" type="holotype">Holotype</typeStatus>
.
</emphasis>
Female. Length
<quantity id="4B4AF2CEB04AA768FE15FAD1FDA9FA2D" box="[456,548,1387,1409]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="6" unit="mm" value="15.2">15.2 mm</quantity>
; width across humeri
<quantity id="4B4AF2CEB04AA768FCC8FAD1FCEBFA2D" box="[789,870,1387,1409]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="6" unit="mm" value="9.1">9.1 mm</quantity>
. Color of clypeus, tibiae, pygidium, and venter dark brown; pronotum, scutellum, and femora light orange; frons black with small, orangish subtriangular spot at center apex; elytra black and weakly suffused with dark orange on disc behind humeri.
<emphasis id="BEC68339B04AA768FF4FFA6DFF45FA40" box="[146,200,1495,1516]" italics="true" pageId="6">Head</emphasis>
: Frons with moderately dense, moderately large punctures. Frontoclypeal suture biarcuate, distinct. Clypeus with surface roughened, vaguely rugopunctate; apex semicircular with thin, marginal bead. Interocular width equals 2.7 transverse eye diameters. Antenna 10-segmented, club slightly longer than segments 27.
<emphasis id="BEC68339B04BA769FE87FF67FE30FF5E" box="[346,445,221,242]" italics="true" pageId="7">Pronotum</emphasis>
: Surface weakly shagreened, punctate; punctures moderate in density and size. Base without marginal bead. Posterior angles broadly rounded.
<emphasis id="BEC68339B04BA769FEB1FEA9FE26FE84" box="[364,427,275,296]" italics="true" pageId="7">Elytra</emphasis>
: Surface weakly shagreened, punctate; punctures large, dense, double rows of punctate striae distinct. Apical angles with sparse, minute setae.
<taxonomicName id="4BB224A8B04BA769FF30FEF3FE28FEF3" authority=": Surface" authorityName="Surface" box="[237,421,329,351]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="7" phylum="Chordata" rank="genus">
<emphasis id="BEC68339B04BA769FF30FEF3FEC1FEF2" box="[237,332,329,350]" italics="true" pageId="7">Pygidium</emphasis>
: Surface
</taxonomicName>
vaguely shagreened, punctate; punctures small, moderate in density, setigerous; setae minute, tawny. In lateral view, surface weakly convex.
<emphasis id="BEC68339B04BA769FF72FEC5FF53FE38" box="[175,222,383,404]" italics="true" pageId="7">Legs</emphasis>
: Protibia tridentate, basal tooth small and strongly removed from apical teeth. Protarsus simple. Metatarsus subequal in length to metatibia.
<emphasis id="BEC68339B04BA769FC7FFE20FC68FE03" box="[930,997,410,431]" italics="true" pageId="7">Venter</emphasis>
: Prosternal process long, columnar, apex transversely oval and flat with a raised, transverse button on anterior half.
</paragraph>
</subSubSection>
<subSubSection id="C4A80CA0B04BA769FF1AFE51FC31FD9B" pageId="7" type="description">
<paragraph id="8C0D5F2BB04BA769FF1AFE51FC31FD9B" blockId="7.[175,1005,194,1163]" pageId="7">
<emphasis id="BEC68339B04BA769FF1AFE51FEA0FDAD" bold="true" box="[199,301,491,513]" pageId="7">Variation.</emphasis>
Females (
<specimenCount id="9AB494A2B04BA769FE47FE51FD9BFDAD" box="[410,534,491,513]" pageId="7" type="generic" typeStatus="paratypes">9 paratypes</specimenCount>
). Length
<quantity id="4B4AF2CEB04BA769FDA7FE51FC80FDAD" box="[634,781,491,513]" metricMagnitude="-2" metricUnit="m" metricValue="1.435" metricValueMax="1.48" metricValueMin="1.39" pageId="7" unit="mm" value="14.35" valueMax="14.8" valueMin="13.9">13.914.8 mm</quantity>
; width across humeri
<quantity id="4B4AF2CEB04BA769FF72FDBCFEA6FDB0" box="[175,299,518,540]" metricMagnitude="-3" metricUnit="m" metricValue="8.05" metricValueMax="8.5" metricValueMin="7.6" pageId="7" unit="mm" value="8.05" valueMax="8.5" valueMin="7.6">7.68.5 mm</quantity>
. The
<typeStatus id="5309E189B04BA769FEBAFDBCFE47FDB0" box="[359,458,518,540]" pageId="7" type="paratype">paratypes</typeStatus>
do not differ significantly from the
<typeStatus id="5309E189B04BA769FC94FDBCFC2EFDB0" box="[841,931,518,540]" pageId="7" type="holotype">holotype</typeStatus>
except that the pronotal punctures become denser on the sides in some specimens.
</paragraph>
</subSubSection>
<subSubSection id="C4A80CA0B04BA769FF1AFD87FC60FDC2" pageId="7" type="etymology">
<paragraph id="8C0D5F2BB04BA769FF1AFD87FC60FDC2" blockId="7.[175,1005,194,1163]" pageId="7">
<emphasis id="BEC68339B04BA769FF1AFD87FEB7FDFF" bold="true" box="[199,314,573,595]" pageId="7">Etymology.</emphasis>
The specific epithet is the Greek word
<taxonomicName id="4BB224A8B04BA769FD14FD87FD72FDFE" authorityName="Ratcliffe" authorityYear="2008" box="[713,767,573,594]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="chera">
<emphasis id="BEC68339B04BA769FD14FD87FD72FDFE" box="[713,767,573,594]" italics="true" pageId="7">chera</emphasis>
</taxonomicName>
, meaning a widow and is used here as a noun in apposition in reference to the
<typeStatus id="5309E189B04BA769FD01FDE2FC85FDC2" box="[732,776,600,622]" pageId="7">type</typeStatus>
series of females only.
</paragraph>
</subSubSection>
<subSubSection id="C4A80CA0B04BA769FF1AFDC9FC01FD13" pageId="7" type="distribution">
<paragraph id="8C0D5F2BB04BA769FF1AFDC9FC01FD13" blockId="7.[175,1005,194,1163]" pageId="7">
<emphasis id="BEC68339B04BA769FF1AFDC9FECBFD25" bold="true" box="[199,326,627,649]" pageId="7">Distribution.</emphasis>
<taxonomicName id="4BB224A8B04BA769FE92FDCEFD9EFD25" authorityName="Ratcliffe" authorityYear="2008" box="[335,531,628,649]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="chera">
<emphasis id="BEC68339B04BA769FE92FDCEFD9EFD25" box="[335,531,628,649]" italics="true" pageId="7">Cyclocephala chera</emphasis>
</taxonomicName>
is known only from
<collectingCountry id="F4A51FBBB04BA769FD29FDC9FCC4FD25" box="[756,841,627,649]" name="Guyana" pageId="7">Guyana</collectingCountry>
. The specimens were taken in lowland rainforest at the Iwokrama Field Station along the Burroburro River nearly in the middle of the country at its narrowest point.
</paragraph>
</subSubSection>
<subSubSection id="C4A80CA0B04BA769FF1AFD7FFCEEFB27" pageId="7" type="diagnosis">
<paragraph id="8C0D5F2BB04BA769FF1AFD7FFC27FCD1" blockId="7.[175,1005,194,1163]" pageId="7">
<emphasis id="BEC68339B04BA769FF1AFD7FFEA2FD77" bold="true" box="[199,303,709,731]" pageId="7">Diagnosis.</emphasis>
Normally, I am hesitant to describe a new species of
<taxonomicName id="4BB224A8B04BA769FCBAFD7FFC61FD76" authorityName="Dejean" authorityYear="1821" box="[871,1004,709,730]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="genus">
<emphasis id="BEC68339B04BA769FCBAFD7FFC61FD76" box="[871,1004,709,730]" italics="true" pageId="7">Cyclocephala</emphasis>
</taxonomicName>
based upon females only, because it is usually the male characters of enlarged protarsi, protibial teeth, pygidium, and parameres that provide the most reliable features for distinguishing species. I make an exception in this case because (1) the body form and coloration are so distinctive, (2) it will not key out in
<bibRefCitation id="E82322DAB04BA769FC45FC8BFF62FCCE" author="Endrodi" pageId="7" refId="ref10514" refString="Endrodi, S. 1985. The Dynastinae of the World. W. Junk Publishers, Dordrecht. 800 pp." type="book" year="1985">Endrödi (1985)</bibRefCitation>
nor is it described in any new species descriptions that have appeared since
<bibRefCitation id="E82322DAB04BA769FF72FCDDFEC3FCD1" author="Endrodi" box="[175,334,871,893]" pageId="7" refId="ref10514" refString="Endrodi, S. 1985. The Dynastinae of the World. W. Junk Publishers, Dordrecht. 800 pp." type="book" year="1985">Endrödi (1985)</bibRefCitation>
, and (3) I have not seen anything like it in any collection.
</paragraph>
<paragraph id="8C0D5F2BB04BA769FF1AFC38FCEEFB27" blockId="7.[175,1005,194,1163]" pageId="7">
The broadly oval and vaulted body form most closely resembles
<taxonomicName id="4BB224A8B04BA769FCB1FC39FF61FC18" authority="Sharp" authorityName="Sharp" authorityYear="1877" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="conspicua">
<emphasis id="BEC68339B04BA769FCB1FC39FC61FC34" box="[876,1004,899,920]" italics="true" pageId="7">C. conspicua</emphasis>
Sharp
</taxonomicName>
and
<taxonomicName id="4BB224A8B04BA769FEFBFC24FD4AFC18" authority="Heyne and Taschenberg. In" authorityName="Heyne and Taschenberg. In" box="[294,711,926,948]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="gregaria">
<emphasis id="BEC68339B04BA769FEFBFC24FE17FC1F" box="[294,410,926,947]" italics="true" pageId="7">C. gregaria</emphasis>
Heyne and Taschenberg. In
</taxonomicName>
those species, however, the elytral margin is thickened into a distinct swelling or flange, whereas the margin in
<taxonomicName id="4BB224A8B04BA769FF14FC6EFE91FC45" authorityName="Ratcliffe" authorityYear="2008" box="[201,284,980,1001]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="chera">
<emphasis id="BEC68339B04BA769FF14FC6EFE91FC45" box="[201,284,980,1001]" italics="true" pageId="7">C. chera</emphasis>
</taxonomicName>
is imperceptibly thickened. Moreover, those species have characteristic black patterns on the pronotum and/or elytra, whereas
<taxonomicName id="4BB224A8B04BA769FD26FC55FCDFFBA8" authorityName="Ratcliffe" authorityYear="2008" box="[763,850,1007,1028]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="chera">
<emphasis id="BEC68339B04BA769FD26FC55FCDFFBA8" box="[763,850,1007,1028]" italics="true" pageId="7">C. chera</emphasis>
</taxonomicName>
does not. The darkest and relatively uncommon forms of
<taxonomicName id="4BB224A8B04BA769FDA1FBB0FC07FBB3" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" box="[636,906,1033,1055]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="complanata">
<emphasis id="BEC68339B04BA769FDA1FBB0FC9DFBB3" box="[636,784,1034,1055]" italics="true" pageId="7">C. complanata</emphasis>
Burmeister
</taxonomicName>
resemble the color in
<taxonomicName id="4BB224A8B04BA769FEE9FB9FFE06FB96" authorityName="Ratcliffe" authorityYear="2008" box="[308,395,1061,1082]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="chera">
<emphasis id="BEC68339B04BA769FEE9FB9FFE06FB96" box="[308,395,1061,1082]" italics="true" pageId="7">C. chera</emphasis>
</taxonomicName>
, but
<taxonomicName id="4BB224A8B04BA769FE1AFB9FFDD1FB96" authorityName="Burmeister" authorityYear="1847" box="[455,604,1061,1082]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="complanata">
<emphasis id="BEC68339B04BA769FE1AFB9FFDD1FB96" box="[455,604,1061,1082]" italics="true" pageId="7">C. complanata</emphasis>
</taxonomicName>
has the base of the pronotum with a marginal bead, has a more slender body form, and occurs only in Mesoamerica. In short, I know of nothing else like
<taxonomicName id="4BB224A8B04BA769FDF8FBE1FDF4FBDC" authorityName="Ratcliffe" authorityYear="2008" box="[549,633,1115,1136]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="7" phylum="Arthropoda" rank="species" species="chera">
<emphasis id="BEC68339B04BA769FDF8FBE1FDF4FBDC" box="[549,633,1115,1136]" italics="true" pageId="7">C. chera</emphasis>
</taxonomicName>
. In
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, it will key on so far as couplet
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the female key, and then no other choices fit.
</paragraph>
</subSubSection>
</treatment>
</document>