267 lines
25 KiB
XML
267 lines
25 KiB
XML
<document id="095E958A8759634DFB4900D2C35DC555" ID-DOI="10.1649/1066.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5369967" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630268265367" checkinUser="valdenar" docAuthor="Ratcliffe, Brett C." docDate="1821" docId="041BEE3DB048A768FE8EFA1FFDFDFBA9" docLanguage="en" docName="1066.1.pdf" docOrigin="The Coleopterists Bulletin 62 (2)" docSource="http://dx.doi.org/10.1649/1066.1" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Cyclocephala carlsoni Ratcliffe 2008, new species" docType="treatment" docVersion="3" lastPageNumber="241" masterDocId="F8229645B04CA76EFFDDFFBAFF8DFFAC" masterDocTitle="More New Species ofCyclocephalaDejean, 1821 from South America (Scarabaeidae: Dynastinae: Cyclocephalini)" masterLastPageNumber="241" masterPageNumber="221" pageNumber="221" updateTime="1699205293535" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="F25E04F5F9488781CE52966717318050" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1927565DB49B39882BF62249697B0E27">
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<mods:title id="B8239DDF9DD2581D54815693117B7055">More New Species ofCyclocephalaDejean, 1821 from South America (Scarabaeidae: Dynastinae: Cyclocephalini)</mods:title>
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<mods:name id="95F10707C0876347AC54F44B631DF1D1" type="personal">
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<mods:role id="FD379AC4BF3CF7275DD9A4B7B266F448">
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<mods:roleTerm id="4C0BF5B210710D1659ACFC04B669959E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="3E9F40E3F459AD911A6EA8B361BCBE49">Ratcliffe, Brett C.</mods:namePart>
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<mods:typeOfResource id="163913A26DB7CBA422CD2B3E598CDB7E">text</mods:typeOfResource>
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<mods:title id="1CF5C69AEFD295133FFD5D2FD8E77D40">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="8D4A93BA269388DFA2D0E185B4F7BE0D">
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<mods:date id="8CCDBBA2AB00C5721085B21BD33FBB83">1821</mods:date>
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<mods:detail id="1245B0CACBF13B0B5A1EFCCDE61BA9E1" type="pubDate">
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<mods:number id="023DA5068890102EEA320F5369FCBD79">2008-06-30</mods:number>
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</mods:detail>
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<mods:detail id="A14568F953B6C2C4E8695FAB717A88C4" type="volume">
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<mods:number id="977C45CD7F9B1ADDD2DAC030ACB1DD7B">62</mods:number>
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</mods:detail>
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<mods:detail id="E2E93A0D6CC06E4C086685692DB42200" type="issue">
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<mods:number id="8B526EDE2E749AA8486B53E836E616C8">2</mods:number>
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</mods:detail>
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<mods:extent id="C78F6F05966E89769D6804C5536D199D" unit="page">
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<mods:start id="8996A570805C087481708F8B152D0EE9">221</mods:start>
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<mods:end id="13B9ED2CC54F3E4F4C7930AB16EF3B55">241</mods:end>
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</mods:extent>
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</mods:part>
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<mods:location id="4EC3890476760A4425F513BF4CA0827B">
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<mods:url id="160BAD32DD8E2B6BC43EA13A3ECEB16F">http://dx.doi.org/10.1649/1066.1</mods:url>
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</mods:location>
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<mods:classification id="9398E1DC48E822B13DB72F4889D9B65B">journal article</mods:classification>
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<mods:identifier id="13DDE947D80C60CA10BA3E05DC73E87E" type="DOI">10.1649/1066.1</mods:identifier>
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<mods:identifier id="6DD580F85531778850AC9747BC61CE39" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="39594937A732B47497D7A29C8BA72107" type="Zenodo-Dep">5369967</mods:identifier>
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</mods:mods>
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<treatment id="041BEE3DB048A768FE8EFA1FFDFDFBA9" LSID="urn:lsid:plazi:treatment:041BEE3DB048A768FE8EFA1FFDFDFBA9" httpUri="http://treatment.plazi.org/id/041BEE3DB048A768FE8EFA1FFDFDFBA9" lastPageId="6" pageId="4">
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<subSubSection id="C4A80CA0B048A76AFE8EFA1FFC83FA17" box="[339,782,1445,1467]" pageId="4" type="nomenclature">
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<paragraph id="8C0D5F2BB048A76AFE8EFA1FFC83FA17" blockId="4.[339,782,1445,1495]" box="[339,782,1445,1467]" pageId="4">
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<heading id="D745E847B048A76AFE8EFA1FFC83FA17" box="[339,782,1445,1467]" centered="true" fontSize="9" level="2" pageId="4" reason="2">
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<emphasis id="BEC68339B048A76AFE8EFA1FFC83FA17" bold="true" box="[339,782,1445,1467]" pageId="4">
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<taxonomicName id="4BB224A8B048A76AFE8EFA1FFD02FA17" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2008" box="[339,655,1445,1467]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="4" phylum="Arthropoda" rank="species" species="carlsoni" status="sp. nov.">
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<emphasis id="BEC68339B048A76AFE8EFA1FFDA3FA17" bold="true" box="[339,558,1445,1467]" italics="true" pageId="4">Cyclocephala carlsoni</emphasis>
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Ratcliffe
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</taxonomicName>
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,
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<taxonomicNameLabel id="A5F53E42B048A76AFD46FA1FFC83FA17" box="[667,782,1445,1467]" pageId="4" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C4A80CA0B048A76AFE27FA7BFDEAFA7B" box="[506,615,1473,1495]" pageId="4" type="description">
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<paragraph id="8C0D5F2BB048A76AFE27FA7BFDEAFA7B" blockId="4.[339,782,1445,1495]" box="[506,615,1473,1495]" pageId="4">
|
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(
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<figureCitation id="148943AEB048A76AFDDCFA7BFDECFA7B" box="[513,609,1473,1495]" captionStart="Figs" captionStartId="4.[167,206,898,917]" captionTargetBox="[146,976,194,876]" captionTargetId="figure-217@4.[146,976,194,876]" captionTargetPageId="4" captionText="Figs. 5–7. Cyclocephala carlsoni: 5) dorsal view of holotype; 6–7) parameres (caudal and lateral views)." figureDoi="http://doi.org/10.5281/zenodo.5369971" httpUri="https://zenodo.org/record/5369971/files/figure.png" pageId="4">Figs. 5–7</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C4A80CA0B048A76BFF77FA4BFE05FB09" lastPageId="5" pageId="4" type="materials_examined">
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<paragraph id="8C0D5F2BB048A76AFF77FA4BFC42F9AB" blockId="4.[146,975,1521,1570]" box="[170,975,1521,1543]" pageId="4">
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<emphasis id="BEC68339B048A76AFF77FA4BFEC8F9AB" bold="true" box="[170,325,1521,1543]" pageId="4">Type Material.</emphasis>
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<materialsCitation id="3CDA5576B048A76AFE8DFA4BFC42F9AB" box="[336,975,1521,1543]" collectorName="de Regina" country="French Guiana" location="Rue de Regina" pageId="4" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5309E189B048A76AFE8DFA4BFE23F9AB" box="[336,430,1521,1543]" pageId="4" type="holotype">Holotype</typeStatus>
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male, labeled ‘‘
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<collectingCountry id="F4A51FBBB048A76AFD92FA4BFD4FF9AB" box="[591,706,1521,1543]" name="French Guiana" pageId="4">GUYANE</collectingCountry>
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,
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<location id="896D09F0B048A76AFD12FA4BFCE1F9AB" LSID="urn:lsid:plazi:treatment:041BEE3DB048A768FE8EFA1FFDFDFBA9:896D09F0B048A76AFD12FA4BFCE1F9AB" box="[719,876,1521,1543]" country="French Guiana" name="Rue de Regina" pageId="4">
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Rue
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<collectorName id="21473AFDB048A76AFCDFFA4BFCE1F9AB" box="[770,876,1521,1543]" pageId="4">de Regina</collectorName>
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||
</location>
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||
,
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||
<quantity id="4B4AF2CEB048A76AFCA4FA4BFC34F9AB" box="[889,953,1521,1543]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="4" unit="km" value="40.0">40 km</quantity>
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S
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</materialsCitation>
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</paragraph>
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||
<paragraph id="8C0D5F2BB048A76BFF4FF9B6FC60FEA2" blockId="4.[146,975,1521,1570]" lastBlockId="5.[175,1005,194,1570]" lastPageId="5" pageId="4">
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<materialsCitation id="3CDA5576B048A76BFF4FF9B6FC60FEA2" collectingDate="1992-09-23" collectingDateMax="2004-09-18" collectingDateMin="1992-09-23" collectionCode="BL, MN, R" collectorName="de Belizon & Rojkoff." country="French Guiana" lastPageId="5" location="Piste de Coralie" municipality="Piste de Belizon" pageId="4" specimenCount="2" specimenCount-female="1" typeStatus="allotype">
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Cayenne,
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<date id="F80C79EBB048A76AFF25F9B6FE08F98E" box="[248,389,1548,1570]" pageId="4" value="1992-09-23">
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<collectingDate id="E8488003B048A76AFF25F9B6FE08F98E" box="[248,389,1548,1570]" pageId="4" value="1992-09-23">23 Sept. 1992</collectingDate>
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||
</date>
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, bl&mv, DCCarlson, FTHovore, PHSullivan collectors.’’
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<typeStatus id="5309E189B049A76BFF72FF78FE88FF74" box="[175,261,194,216]" pageId="5" type="allotype">Allotype</typeStatus>
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female labeled ‘‘GUYANE,
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||
<location id="896D09F0B049A76BFDFCFF78FD31FF74" LSID="urn:lsid:plazi:treatment:041BEE3DB048A768FE8EFA1FFDFDFBA9:896D09F0B049A76BFDFCFF78FD31FF74" box="[545,700,194,216]" country="French Guiana" municipality="Piste de Belizon" name="Piste de Coralie" pageId="5">Piste de Coralie</location>
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, Pk 2,
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<date id="F80C79EBB049A76BFD21FF78FC0CFF74" box="[764,897,194,216]" pageId="5" value="1992-09-27">
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<collectingDate id="E8488003B049A76BFD21FF78FC0CFF74" box="[764,897,194,216]" pageId="5" value="1992-09-27">27 Sept. 1992</collectingDate>
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</date>
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||
,
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<collectionCode id="EAA3C7EEB049A76BFC57FF78FC24FF74" box="[906,937,194,216]" pageId="5">BL</collectionCode>
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&
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<collectionCode id="EAA3C7EEB049A76BFC60FF78FC64FF74" box="[957,1001,194,216]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="5">MN</collectionCode>
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||
, DCCarlson, FTHovore, PHSullivan collectors.’’ One male and
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||
<specimenCount id="9AB494A2B049A76BFCFEFF67FC02FF5F" box="[803,911,221,243]" pageId="5" type="female">one female</specimenCount>
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||
<typeStatus id="5309E189B049A76BFC48FF67FC61FF5F" box="[917,1004,221,243]" pageId="5" type="paratype">paratype</typeStatus>
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labeled ‘‘
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<collectingCountry id="F4A51FBBB049A76BFED9FF42FE6DFEA2" box="[260,480,248,270]" name="French Guiana" pageId="5">FRENCH GUIANA</collectingCountry>
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, PK 15/15
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<collectingMunicipality id="6C69C551B049A76BFD96FF42FD64FEA2" box="[587,745,248,270]" pageId="5">
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Piste
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<collectorName id="21473AFDB049A76BFDA2FF42FD64FEA2" box="[639,745,248,270]" pageId="5">de Belizon</collectorName>
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||
</collectingMunicipality>
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||
,
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||
<date id="F80C79EBB049A76BFD2FFF42FCE9FEA2" box="[754,868,248,270]" pageId="5" value="2004-09-18">
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||
<collectingDate id="E8488003B049A76BFD2FFF42FCE9FEA2" box="[754,868,248,270]" pageId="5" value="2004-09-18">IX-18-2004</collectingDate>
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||
</date>
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||
,
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||
<collectionCode id="EAA3C7EEB049A76BFCB3FF42FC0EFEA2" box="[878,899,248,270]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5">R</collectionCode>
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.
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<collectorName id="21473AFDB049A76BFC56FF42FC68FEA2" box="[907,997,248,270]" pageId="5">Rojkoff.</collectorName>
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||
’’
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8C0D5F2BB049A76BFF1AFEAEFD7BFECC" blockId="5.[175,1005,194,1570]" pageId="5">
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||
<materialsCitation id="3CDA5576B049A76BFF1AFEAEFD7FFECC" collectionCode="CA" collectorName="Lincoln, NE & B. C. Ratcliffe & Lincoln NE" location="Nebraska State" pageId="5" specimenCount="3" stateProvince="Nebraska" typeStatus="holotype">
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||
<typeStatus id="5309E189B049A76BFF1AFEAEFEAAFE86" box="[199,295,276,298]" pageId="5" type="holotype">Holotype</typeStatus>
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and
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<typeStatus id="5309E189B049A76BFE80FEAEFE22FE86" box="[349,431,276,298]" pageId="5" type="allotype">allotype</typeStatus>
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(property of David Carlson, Fair Oaks,
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<collectionCode id="EAA3C7EEB049A76BFC8AFEAEFCF1FE86" box="[855,892,276,298]" country="USA" name="Chicago Academy of Sciences" pageId="5" type="Museum">CA</collectionCode>
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||
) deposited at the University of
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<collectingRegion id="4E7691C9B049A76BFE4AFE95FDB6FEE9" box="[407,571,303,325]" country="United States of America" name="Nebraska" pageId="5">Nebraska State</collectingRegion>
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Museum,
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<collectorName id="21473AFDB049A76BFD69FE95FCB3FEE9" box="[692,830,303,325]" pageId="5">Lincoln, NE</collectorName>
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. Two
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<typeStatus id="5309E189B049A76BFC54FE95FC61FEE9" box="[905,1004,303,325]" pageId="5" type="paratype">paratypes</typeStatus>
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deposited in the
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<collectorName id="21473AFDB049A76BFE81FEF0FE7FFECC" box="[348,498,330,352]" pageId="5">B. C. Ratcliffe</collectorName>
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collection (
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<collectorName id="21473AFDB049A76BFDB6FEF0FD67FECC" box="[619,746,330,352]" pageId="5">Lincoln NE</collectorName>
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)
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</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8C0D5F2BB049A76BFF1AFEDFFDBAFC1E" blockId="5.[175,1005,194,1570]" pageId="5">
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||
<emphasis id="BEC68339B049A76BFF1AFEDFFEA7FED7" bold="true" box="[199,298,357,379]" pageId="5">
|
||
<typeStatus id="5309E189B049A76BFF1AFEDFFEABFED7" box="[199,294,357,379]" pageId="5" type="holotype">Holotype</typeStatus>
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||
.
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||
</emphasis>
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||
Male. Length 10.0 mm; width across humeri
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<quantity id="4B4AF2CEB049A76BFC96FEDFFC11FED7" box="[843,924,357,379]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="5" unit="mm" value="5.2">5.2 mm</quantity>
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||
. Color testaceous except for black frons, 2 broad, longitudinal bands on pronotum, posterior borders of scutellum and elytral suture, 3 black spots on each elytron (post-humeral, post-scutellar, center of disc behind middle; and apices of femora, tibiae, and protibial teeth). Tarsi, pygidium, abdominal sternites light yellowish brown.
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||
<emphasis id="BEC68339B049A76BFEDAFE56FEB0FDAD" box="[263,317,492,513]" italics="true" pageId="5">Head</emphasis>
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||
: Surface of frons densely punctate, punctures small. Clypeus vaguely roughened or rugulopunctate; apex broadly truncate, feebly emarginate at middle, weakly reflexed. Frontoclypeal suture biarcuate, distinct. Interocular width equals 3.4 transverse eye diameters. Antenna 10-segmented, club subequal in length to segments 2–7.
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||
<emphasis id="BEC68339B049A76BFE3EFDE2FDCBFDC1" box="[483,582,600,621]" italics="true" pageId="5">Pronotum</emphasis>
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||
: Surface moderately densely punctate, punctures small. Posterior angles broadly rounded. Base lacking marginal bead.
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||
<emphasis id="BEC68339B049A76BFF72FD34FF63FD0F" box="[175,238,654,675]" italics="true" pageId="5">Elytra</emphasis>
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||
: Surface weakly shagreened, punctate; punctures minute and moderately large mixed, paired punctate striae indistinct.
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||
<taxonomicName id="4BB224A8B049A76BFDA2FD13FCBBFD13" authority=": Surface" authorityName="Surface" box="[639,822,681,703]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="genus">
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||
<emphasis id="BEC68339B049A76BFDA2FD13FD53FD12" box="[639,734,681,702]" italics="true" pageId="5">Pygidium</emphasis>
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||
: Surface
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||
</taxonomicName>
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||
densely punctate; punctures minute, surface becoming rugulose in angles. In lateral view, surface evenly convex.
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<emphasis id="BEC68339B049A76BFE97FD65FEF4FD58" box="[330,377,735,756]" italics="true" pageId="5">Legs</emphasis>
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||
: Protibia tridentate, basal tooth small, slightly removed from other teeth. Protarsus strongly enlarged; tarsomeres 2–4 each slightly larger than preceding; 5th subequal in length to tarsomeres 2–4; median claw enlarged, strongly curved, apex widely split into broad and small rami. Posterior tibia short, stout, expanded towards apex; apex with fringe of short, stout spinules. Metatarsus subequal in length to metatibia.
|
||
<emphasis id="BEC68339B049A76BFD4BFCDCFD54FCD7" box="[662,729,870,891]" italics="true" pageId="5">Venter</emphasis>
|
||
: Prosternal process long, slender, columnar, apex a small, oval disc with raised, transversely oval ‘‘button’’ on anterior 2/3.
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||
<emphasis id="BEC68339B049A76BFE85FC26FE4EFC1D" box="[344,451,924,945]" italics="true" pageId="5">Parameres</emphasis>
|
||
:
|
||
<figureCitation id="148943AEB049A76BFE0FFC26FDBFFC1E" box="[466,562,924,946]" captionStart="Figs" captionStartId="4.[167,206,898,917]" captionTargetBox="[146,976,194,876]" captionTargetId="figure-217@4.[146,976,194,876]" captionTargetPageId="4" captionText="Figs. 5–7. Cyclocephala carlsoni: 5) dorsal view of holotype; 6–7) parameres (caudal and lateral views)." figureDoi="http://doi.org/10.5281/zenodo.5369971" httpUri="https://zenodo.org/record/5369971/files/figure.png" pageId="5">Figs. 6–7</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C0D5F2BB049A76BFF1AFC0DFE05FB09" blockId="5.[175,1005,194,1570]" pageId="5">
|
||
<emphasis id="BEC68339B049A76BFF1AFC0DFEAFFC61" bold="true" box="[199,290,951,973]" pageId="5">
|
||
<typeStatus id="5309E189B049A76BFF1AFC0DFE93FC61" box="[199,286,951,973]" pageId="5" type="allotype">Allotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Female. Length 10.0 mm; width across humeri
|
||
<quantity id="4B4AF2CEB049A76BFCCEFC0DFCE8FC61" box="[787,869,951,973]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="5" unit="mm" value="5.4">5.4 mm</quantity>
|
||
. As
|
||
<typeStatus id="5309E189B049A76BFC4EFC0DFC60FC61" box="[915,1005,951,973]" pageId="5" type="holotype">holotype</typeStatus>
|
||
except in the following respects: Color and pattern similar except pronotum with small, black spot near lateral margin on each side and subapical swelling of elytral margin piceous.
|
||
<emphasis id="BEC68339B049A76BFEB7FBB3FE2DFBB2" box="[362,416,1033,1054]" italics="true" pageId="5">Head</emphasis>
|
||
: Entire surface punctate to rugopunctate, punctures moderate in size. Interocular width equals 2.1 transverse eye diameters.
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<emphasis id="BEC68339B049A76BFC7BFB9EFC68FB95" box="[934,997,1060,1081]" italics="true" pageId="5">Elytra</emphasis>
|
||
: Epipleuron (in ventral view) attenuated at level of sternite 4. Elytral margin laterad of apical umbone swollen into weak knob.
|
||
<taxonomicName id="4BB224A8B049A76BFD07FBE0FCEDFBC3" authority=": In" authorityName="In" box="[730,864,1113,1135]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="5" phylum="Chordata" rank="genus">
|
||
<emphasis id="BEC68339B049A76BFD07FBE0FCB4FBC3" box="[730,825,1114,1135]" italics="true" pageId="5">Pygidium</emphasis>
|
||
: In
|
||
</taxonomicName>
|
||
lateral view, surface weakly convex.
|
||
<emphasis id="BEC68339B049A76BFE71FBCFFE51FB26" box="[428,476,1141,1162]" italics="true" pageId="5">Legs</emphasis>
|
||
: Protibia with teeth subequally spaced. Protarsus simple, not enlarged.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B049A76BFF1AFB11FD8EFA81" pageId="5" type="description">
|
||
<paragraph id="8C0D5F2BB049A76BFF1AFB11FC26FB5B" blockId="5.[175,1005,194,1570]" pageId="5">
|
||
<emphasis id="BEC68339B049A76BFF1AFB11FEA0FB6D" bold="true" box="[199,301,1195,1217]" pageId="5">Variation.</emphasis>
|
||
Males (
|
||
<specimenCount id="9AB494A2B049A76BFE5BFB11FE75FB6D" box="[390,504,1195,1217]" pageId="5" type="generic" typeStatus="paratype">1 paratype</specimenCount>
|
||
). Length
|
||
<quantity id="4B4AF2CEB049A76BFD83FB11FD23FB6D" box="[606,686,1195,1217]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="5" unit="mm" value="9.3">9.3 mm</quantity>
|
||
; width across humeri
|
||
<quantity id="4B4AF2CEB049A76BFC45FB11FC64FB6D" box="[920,1001,1195,1217]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="5" unit="mm" value="5.2">5.2 mm</quantity>
|
||
. As
|
||
<typeStatus id="5309E189B049A76BFF0CFB7CFEA6FB70" box="[209,299,1222,1244]" pageId="5" type="holotype">holotype</typeStatus>
|
||
except in the following respects: Head and pronotum with punctures dense, moderately large. Clypeus rugulose. Elytra with discal spot absent.
|
||
</paragraph>
|
||
<paragraph id="8C0D5F2BB049A76BFF1AFB47FD8EFA81" blockId="5.[175,1005,194,1570]" pageId="5">
|
||
Females (
|
||
<specimenCount id="9AB494A2B049A76BFEF2FB47FE2FFABF" box="[303,418,1277,1299]" pageId="5" type="generic" typeStatus="paratype">1 paratype</specimenCount>
|
||
). Length
|
||
<quantity id="4B4AF2CEB049A76BFDD7FB47FDEAFABF" box="[522,615,1277,1299]" metricMagnitude="-2" metricUnit="m" metricValue="1.03" pageId="5" unit="mm" value="10.3">10.3 mm</quantity>
|
||
; width across humeri
|
||
<quantity id="4B4AF2CEB049A76BFC8BFB47FC25FABF" box="[854,936,1277,1299]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="5" unit="mm" value="5.6">5.6 mm</quantity>
|
||
. Head with clypeus transversely rugose.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B049A76BFF1AFA89FC2FFA19" pageId="5" type="etymology">
|
||
<paragraph id="8C0D5F2BB049A76BFF1AFA89FC2FFA19" blockId="5.[175,1005,194,1570]" pageId="5">
|
||
<emphasis id="BEC68339B049A76BFF1AFA89FEB7FAE5" bold="true" box="[199,314,1331,1353]" pageId="5">Etymology.</emphasis>
|
||
The species is named in honor of David Carlson (Fair Oaks, CA), one of the collectors of this new species, who has loaned me numerous specimens for study over the years. This patronym is also crafted in grateful appreciation of his permission for allowing
|
||
<typeStatus id="5309E189B049A76BFE06FA3EFD8AFA36" box="[475,519,1412,1434]" pageId="5">type</typeStatus>
|
||
specimens from his personal collection to be deposited in the collection of the University of
|
||
<collectingRegion id="4E7691C9B049A76BFD46FA25FCB4FA19" box="[667,825,1439,1461]" country="United States of America" name="Nebraska" pageId="5">Nebraska State</collectingRegion>
|
||
Museum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B049A76BFF1AFA01FD19FA40" pageId="5" type="distribution">
|
||
<paragraph id="8C0D5F2BB049A76BFF1AFA01FD19FA40" blockId="5.[175,1005,194,1570]" pageId="5">
|
||
<emphasis id="BEC68339B049A76BFF1AFA01FECBFA7D" bold="true" box="[199,326,1467,1489]" pageId="5">Distribution.</emphasis>
|
||
<taxonomicName id="4BB224A8B049A76BFE92FA01FDA1FA7C" authorityName="Ratcliffe" authorityYear="2008" box="[335,556,1467,1488]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="carlsoni">
|
||
<emphasis id="BEC68339B049A76BFE92FA01FDA1FA7C" box="[335,556,1467,1488]" italics="true" pageId="5">Cyclocephala carlsoni</emphasis>
|
||
</taxonomicName>
|
||
is endemic to
|
||
<collectingCountry id="F4A51FBBB049A76BFD16FA01FCE1FA7D" box="[715,876,1467,1489]" name="French Guiana" pageId="5">French Guiana</collectingCountry>
|
||
, and all the specimens were taken from lowland rainforest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B049A768FF1AFA4BFDFDFBA9" lastPageId="6" pageId="5" type="diagnosis">
|
||
<paragraph id="8C0D5F2BB049A768FF1AFA4BFDFDFBA9" blockId="5.[175,1005,194,1570]" lastBlockId="6.[146,975,845,1029]" lastPageId="6" pageId="5">
|
||
<emphasis id="BEC68339B049A76BFF1AFA4BFEA2F9AB" bold="true" box="[199,303,1521,1543]" pageId="5">Diagnosis.</emphasis>
|
||
<taxonomicName id="4BB224A8B049A76BFEE8FA48FD82F9AB" authorityName="Ratcliffe" authorityYear="2008" box="[309,527,1522,1543]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="5" phylum="Arthropoda" rank="species" species="carlsoni">
|
||
<emphasis id="BEC68339B049A76BFEE8FA48FD82F9AB" box="[309,527,1522,1543]" italics="true" pageId="5">Cyclocephala carlsoni</emphasis>
|
||
</taxonomicName>
|
||
males will key as far as couplet
|
||
<quantity id="4B4AF2CEB049A76BFC89FA4BFC1FF9AB" box="[852,914,1521,1543]" metricMagnitude="0" metricUnit="m" metricValue="8.9662" pageId="5" unit="in" value="353.0">353 in</quantity>
|
||
<bibRefCitation id="E82322DAB049A76BFC45FA4BFF7CF98E" author="Endrodi" pageId="5" refId="ref10514" refString="Endrodi, S. 1985. The Dynastinae of the World. W. Junk Publishers, Dordrecht. 800 pp." type="book" year="1985">Endrödi (1985)</bibRefCitation>
|
||
, at which point no other choices apply. The female
|
||
<typeStatus id="5309E189B049A76BFCD9F9B6FCDBF98E" box="[772,854,1548,1570]" pageId="5" type="allotype">allotype</typeStatus>
|
||
will key as far 226 as couplet 261, where choices no longer fit. While this species has similar color and pattern to other species of
|
||
<taxonomicName id="4BB224A8B04AA768FE2FFCD2FDFAFCD1" authorityName="Dejean" authorityYear="1821" box="[498,631,872,893]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BEC68339B04AA768FE2FFCD2FDFAFCD1" box="[498,631,872,893]" italics="true" pageId="6">Cyclocephala</emphasis>
|
||
</taxonomicName>
|
||
, it is the combination of these characters (and others mentioned in the description) with its small size that makes this species distinctive. I have examined the literature for all of the new species of
|
||
<taxonomicName id="4BB224A8B04AA768FF4FFC03FE9AFC62" authorityName="Dejean" authorityYear="1821" box="[146,279,953,974]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BEC68339B04AA768FF4FFC03FE9AFC62" box="[146,279,953,974]" italics="true" pageId="6">Cyclocephala</emphasis>
|
||
</taxonomicName>
|
||
described since
|
||
<bibRefCitation id="E82322DAB04AA768FE12FC03FDFCFC63" author="Endrodi" box="[463,625,953,975]" pageId="6" refId="ref10514" refString="Endrodi, S. 1985. The Dynastinae of the World. W. Junk Publishers, Dordrecht. 800 pp." type="book" year="1985">Endrödi (1985)</bibRefCitation>
|
||
, particularly those from
|
||
<collectingCountry id="F4A51FBBB04AA768FC5BFC03FF53FC46" name="French Guiana" pageId="6">French Guiana</collectingCountry>
|
||
by Roger-Paul Dechambre, and none of them correspond with the combination of character states of
|
||
<taxonomicName id="4BB224A8B04AA768FE20FC55FDE6FBA8" authorityName="Ratcliffe" authorityYear="2008" box="[509,619,1007,1028]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="6" phylum="Arthropoda" rank="species" species="carlsoni">
|
||
<emphasis id="BEC68339B04AA768FE20FC55FDE6FBA8" box="[509,619,1007,1028]" italics="true" pageId="6">C. carlsoni</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |