322 lines
32 KiB
XML
322 lines
32 KiB
XML
<document id="617F6020AD23EB4FDC0D7839F76CA1DF" ID-DOI="10.1649/1066.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5369967" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630268265367" checkinUser="valdenar" docAuthor="Ratcliffe, Brett C." docDate="1821" docId="041BEE3DB047A760FE8CFA1FFF3FFF3D" docLanguage="en" docName="1066.1.pdf" docOrigin="The Coleopterists Bulletin 62 (2)" docSource="http://dx.doi.org/10.1649/1066.1" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Cyclocephala hielkemaorum Ratcliffe 2008, new species" docType="treatment" docVersion="3" lastPageNumber="241" masterDocId="F8229645B04CA76EFFDDFFBAFF8DFFAC" masterDocTitle="More New Species ofCyclocephalaDejean, 1821 from South America (Scarabaeidae: Dynastinae: Cyclocephalini)" masterLastPageNumber="241" masterPageNumber="221" pageNumber="221" updateTime="1699205293535" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="2E0CD07C55091D525C2540979F655C9F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CB61FD68BE8AABFE7C0BEABDDC4FEDE3">
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<mods:title id="2CABA1C02A96043748BD3F10944407D3">More New Species ofCyclocephalaDejean, 1821 from South America (Scarabaeidae: Dynastinae: Cyclocephalini)</mods:title>
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</mods:titleInfo>
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<mods:name id="89DC74246933ABA624054D2F70BED6CF" type="personal">
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<mods:role id="0B33EB9EC3BAF3CD434BED97894294F3">
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<mods:roleTerm id="7861F119D09B2D8B25FF2D789912AE3D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FD8E5C62EE72A80EBF7C7C8350422E79">Ratcliffe, Brett C.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="65B97FA553726474F16EDCBE33EC7459">text</mods:typeOfResource>
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<mods:titleInfo id="7BECB29F0E3171FC43885CB11379003F">
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<mods:title id="258F22292678F34C44017F7370926E9F">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="A8B2843F625B1E904D6B701612AB4747">
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<mods:date id="D1F96B6F3FB5E4C03B5C05D1A4C062B5">1821</mods:date>
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<mods:detail id="538635FF1775BAFEED1427962B6A442E" type="pubDate">
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<mods:number id="4F13462D79953F405B985129D6447CB1">2008-06-30</mods:number>
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</mods:detail>
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<mods:detail id="EE40398BEAB4DCBE563083D9B2AF3F23" type="volume">
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<mods:number id="67BD9088219ED1A635A1B522B8E9DD51">62</mods:number>
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</mods:detail>
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<mods:detail id="2D4F6BB1A25CE5CCC7B405D41F9F77BD" type="issue">
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<mods:number id="C3EF804EFB84986E8DE50C887D5CC439">2</mods:number>
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</mods:detail>
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<mods:extent id="F4792A52F9D689312B5C4671029271D3" unit="page">
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<mods:start id="98A48DCCDD8066F5878951F1020B5A9A">221</mods:start>
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<mods:end id="453E0E58FB77BD2EBBEEEE67110F6632">241</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="77E8FF283CCE8ED05D06DD1D4380BA44">
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<mods:url id="BBC552B89CFCCC090F57A7616BC09841">http://dx.doi.org/10.1649/1066.1</mods:url>
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</mods:location>
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<mods:classification id="6DE83D3F1F66564DA308FE1AD1291211">journal article</mods:classification>
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<mods:identifier id="675E316BB3BA747D219DDF6A83569183" type="DOI">10.1649/1066.1</mods:identifier>
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<mods:identifier id="D82A5B0527E04F1A52ABEF344B9B08C0" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="0B9CBF7695854AD73DA47794416283FC" type="Zenodo-Dep">5369967</mods:identifier>
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</mods:mods>
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<treatment id="041BEE3DB047A760FE8CFA1FFF3FFF3D" LSID="urn:lsid:plazi:treatment:041BEE3DB047A760FE8CFA1FFF3FFF3D" httpUri="http://treatment.plazi.org/id/041BEE3DB047A760FE8CFA1FFF3FFF3D" lastPageId="14" pageId="11">
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<subSubSection id="C4A80CA0B047A765FE8CFA1FFCC7FA17" box="[337,842,1445,1467]" pageId="11" type="nomenclature">
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<paragraph id="8C0D5F2BB047A765FE8CFA1FFCC7FA17" blockId="11.[337,842,1445,1494]" box="[337,842,1445,1467]" pageId="11">
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<heading id="D745E847B047A765FE8CFA1FFCC7FA17" box="[337,842,1445,1467]" centered="true" fontSize="9" level="2" pageId="11" reason="2">
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<emphasis id="BEC68339B047A765FE8CFA1FFCC7FA17" bold="true" box="[337,842,1445,1467]" pageId="11">
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||
<taxonomicName id="4BB224A8B047A765FE8CFA1FFD41FA17" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2008" box="[337,716,1445,1467]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="hielkemaorum" status="sp. nov.">
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||
<emphasis id="BEC68339B047A765FE8CFA1FFDE6FA17" bold="true" box="[337,619,1445,1467]" italics="true" pageId="11">Cyclocephala hielkemaorum</emphasis>
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Ratcliffe
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</taxonomicName>
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,
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<taxonomicNameLabel id="A5F53E42B047A765FD05FA1FFCC7FA17" box="[728,842,1445,1467]" pageId="11" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C4A80CA0B047A765FDD6FA7AFD1CFA7A" box="[523,657,1472,1494]" pageId="11" type="description">
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<paragraph id="8C0D5F2BB047A765FDD6FA7AFD1CFA7A" blockId="11.[337,842,1445,1494]" box="[523,657,1472,1494]" pageId="11">
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(
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<figureCitation id="148943AEB047A765FDCFFA7AFD07FA7A" box="[530,650,1472,1494]" captionStart="Figs" captionStartId="12.[167,206,1259,1278]" captionTargetBox="[146,976,194,1231]" captionTargetId="figure-65@12.[146,976,194,1237]" captionTargetPageId="12" captionText="Figs. 21–25. Cyclocephala hielkemaorum: 21) dorsal view of holotype; 22–23) parameres (caudal and lateral views); 24–25) right protarsus, median view, of 24) C. hielkemaorum and 25) C. longitarsis." figureDoi="http://doi.org/10.5281/zenodo.5369977" httpUri="https://zenodo.org/record/5369977/files/figure.png" pageId="11">Figs. 21–24</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C4A80CA0B047A763FF1AFA4BFE9AFC46" lastPageId="13" pageId="11" type="materials_examined">
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<paragraph id="8C0D5F2BB047A765FF1AFA4BFC61F9AB" blockId="11.[175,1004,1521,1570]" box="[199,1004,1521,1543]" pageId="11">
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<emphasis id="BEC68339B047A765FF1AFA4BFED3F9AB" bold="true" box="[199,350,1521,1543]" pageId="11">Type Material.</emphasis>
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<materialsCitation id="3CDA5576B047A765FEB8FA4BFC61F9AB" box="[357,1004,1521,1543]" collectingDate="2005-07-03" country="Guyana" location="Roura" municipality="Cacao" pageId="11" specimenCount="1" typeStatus="holotype">
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<typeStatus id="5309E189B047A765FEB8FA4BFE48F9AB" box="[357,453,1521,1543]" pageId="11" type="holotype">Holotype</typeStatus>
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male, labeled ‘‘FR.
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<collectingCountry id="F4A51FBBB047A765FD4EFA4BFD6AF9AB" box="[659,743,1521,1543]" name="Guyana" pageId="11">Guyana</collectingCountry>
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,
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<location id="896D09F0B047A765FD2FFA4BFCBAF9AB" LSID="urn:lsid:plazi:treatment:041BEE3DB047A760FE8CFA1FFF3FFF3D:896D09F0B047A765FD2FFA4BFCBAF9AB" box="[754,823,1521,1543]" country="Guyana" municipality="Cacao" name="Roura" pageId="11">Roura</location>
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:
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<collectingMunicipality id="6C69C551B047A765FC9DFA4BFC0EF9AB" box="[832,899,1521,1543]" pageId="11">Cacao</collectingMunicipality>
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,
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<date id="F80C79EBB047A765FC53FA4BFC65F9AB" box="[910,1000,1521,1543]" pageId="11" value="2005-07-03">
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<collectingDate id="E8488003B047A765FC53FA4BFC65F9AB" box="[910,1000,1521,1543]" pageId="11" value="2005-07-03">3-7-2005</collectingDate>
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</date>
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,
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</materialsCitation>
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||
</paragraph>
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<paragraph id="8C0D5F2BB047A765FF72F9B6FC61F98E" blockId="11.[175,1004,1521,1570]" box="[175,1004,1548,1570]" pageId="11">
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A. J. Hielkema//On mercury-light//
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<taxonomicName id="4BB224A8B047A765FDC4F9B6FD7AF98E" box="[537,759,1548,1570]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="undefined-g">Cyclocephala spec. G</taxonomicName>
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. det. M. A. Hielkema.’’
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</paragraph>
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<paragraph id="8C0D5F2BB040A762FF4FFFC4FD4CFA7D" blockId="12.[146,178,126,145]" lastBlockId="12.[146,975,1387,1570]" pageId="12">
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<materialsCitation id="3CDA5576B040A762FF4FFFC4FD6AFA30" collectingDate="2005-06-27" collectorName="A. J. Hielkema & Single" country="Guyana" elevation="300" location="Roura" municipality="Chateau Cacao" pageId="12" specimenCount="232" specimenCount-female="232" typeStatus="allotype">
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<specimenCount id="9AB494A2B040A762FF4FFFC4FEB4FA2D" box="[146,313,126,1409]" pageId="12" type="female" typeStatus="Allotype">232 Allotype female</specimenCount>
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, labeled ‘‘FR
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<collectingCountry id="F4A51FBBB040A762FE12FAD1FDC9FA2D" box="[463,580,1387,1409]" name="Guyana" pageId="12">GUYANA</collectingCountry>
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,
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<location id="896D09F0B040A762FD8CFAD1FD1BFA2D" LSID="urn:lsid:plazi:treatment:041BEE3DB047A760FE8CFA1FFF3FFF3D:896D09F0B040A762FD8CFAD1FD1BFA2D" box="[593,662,1387,1409]" country="Guyana" municipality="Chateau Cacao" name="Roura" pageId="12">Roura</location>
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:
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<quantity id="4B4AF2CEB040A762FD7FFAD1FD6FFA2D" box="[674,738,1387,1409]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="12" unit="m" value="300.0">
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||
<elevation id="079FB818B040A762FD7FFAD1FD6FFA2D" box="[674,738,1387,1409]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="12" unit="m" value="300.0">300 m</elevation>
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</quantity>
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W of
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<collectingMunicipality id="6C69C551B040A762FCF4FAD1FC46FA2D" box="[809,971,1387,1409]" pageId="12">Chateau Cacao</collectingMunicipality>
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, 25/
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<date id="F80C79EBB040A762FF6FFA3CFE9AFA30" box="[178,279,1414,1436]" pageId="12" value="2005-06-27">
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||
<collectingDate id="E8488003B040A762FF6FFA3CFE9AFA30" box="[178,279,1414,1436]" pageId="12" value="2005-06-27">27-6-2005</collectingDate>
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||
</date>
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||
,
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<collectorName id="21473AFDB040A762FEFFFA3CFE31FA30" box="[290,444,1414,1436]" pageId="12">A. J. Hielkema</collectorName>
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//flying at light.’’
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<collectorName id="21473AFDB040A762FDACFA3CFD22FA30" box="[625,687,1414,1436]" pageId="12">Single</collectorName>
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||
male
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||
</materialsCitation>
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||
<materialsCitation id="3CDA5576B040A762FD33FA3CFD34FA7D" collectingDate="2005-07-23" collectorName="A. J. Hielkema" country="Guyana" determinerName="M. A. Hielkema." location="Roura" municipality="Cacao" pageId="12" specimenCount="1" typeStatus="paratype">
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||
<typeStatus id="5309E189B040A762FD33FA3CFCC5FA30" box="[750,840,1414,1436]" pageId="12" type="paratype">paratype</typeStatus>
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labeled ‘‘FR
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<collectingCountry id="F4A51FBBB040A762FF4FFA1AFE8AFA1A" box="[146,263,1440,1462]" name="Guyana" pageId="12">GUYANA</collectingCountry>
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||
,
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||
<location id="896D09F0B040A762FEC7FA1AFED2FA1A" LSID="urn:lsid:plazi:treatment:041BEE3DB047A760FE8CFA1FFF3FFF3D:896D09F0B040A762FEC7FA1AFED2FA1A" box="[282,351,1440,1462]" country="Guyana" municipality="Cacao" name="Roura" pageId="12">Roura</location>
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||
:
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<collectingMunicipality id="6C69C551B040A762FEACFA1AFE39FA1A" box="[369,436,1440,1462]" pageId="12">Cacao</collectingMunicipality>
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||
,
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<date id="F80C79EBB040A762FE15FA1AFDA3FA1A" box="[456,558,1440,1462]" pageId="12" value="2005-07-23">
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<collectingDate id="E8488003B040A762FE15FA1AFDA3FA1A" box="[456,558,1440,1462]" pageId="12" value="2005-07-23">23-7-2005</collectingDate>
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</date>
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,
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<collectorName id="21473AFDB040A762FD9CFA1AFD61FA1A" box="[577,748,1440,1462]" pageId="12">A. J. Hielkema</collectorName>
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//on mercury-light in forest//
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<taxonomicName id="4BB224A8B040A762FF06FA01FE22FA7D" box="[219,431,1467,1489]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="undetermined">Cyclocephala species</taxonomicName>
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L, det.
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<determinerName id="79D1B3EFB040A762FDD9FA01FD34FA7D" box="[516,697,1467,1489]" pageId="12">M. A. Hielkema.</determinerName>
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</materialsCitation>
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||
’’
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</paragraph>
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<caption id="D8CD0FA3B040A762FF7AFB51FEBEFA82" ID-DOI="http://doi.org/10.5281/zenodo.5369977" ID-Zenodo-Dep="5369977" httpUri="https://zenodo.org/record/5369977/files/figure.png" pageId="12" startId="12.[167,206,1259,1278]" targetBox="[146,976,194,1231]" targetPageId="12">
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||
<paragraph id="8C0D5F2BB040A762FF7AFB51FEBEFA82" blockId="12.[146,975,1259,1327]" pageId="12">
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||
<emphasis id="BEC68339B040A762FF7AFB51FE95FB52" bold="true" box="[167,280,1259,1278]" pageId="12">Figs. 21–25.</emphasis>
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||
<taxonomicName id="4BB224A8B040A762FEF0FB56FDA8FB53" authorityName="Ratcliffe" authorityYear="2008" box="[301,549,1260,1279]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="hielkemaorum">
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||
<emphasis id="BEC68339B040A762FEF0FB56FDA8FB53" box="[301,549,1260,1279]" italics="true" pageId="12">Cyclocephala hielkemaorum</emphasis>
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||
</taxonomicName>
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||
:
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<emphasis id="BEC68339B040A762FDEDFB51FDC1FB52" bold="true" box="[560,588,1259,1278]" pageId="12">21)</emphasis>
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||
dorsal view of holotype;
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||
<emphasis id="BEC68339B040A762FCEDFB51FCE0FB52" bold="true" box="[816,877,1259,1278]" pageId="12">22–23)</emphasis>
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||
parameres (caudal and lateral views);
|
||
<emphasis id="BEC68339B040A762FE58FAB9FE4CFABA" bold="true" box="[389,449,1283,1302]" pageId="12">24–25)</emphasis>
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||
right protarsus, median view, of
|
||
<emphasis id="BEC68339B040A762FD33FAB9FC87FABA" bold="true" box="[750,778,1283,1302]" pageId="12">24)</emphasis>
|
||
<taxonomicName id="4BB224A8B040A762FCCDFAB9FC25FABA" authorityName="Ratcliffe" authorityYear="2008" box="[784,936,1283,1302]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="hielkemaorum">
|
||
<emphasis id="BEC68339B040A762FCCDFAB9FC25FABA" box="[784,936,1283,1302]" italics="true" pageId="12">C. hielkemaorum</emphasis>
|
||
</taxonomicName>
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||
and
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||
<emphasis id="BEC68339B040A762FF4FFAA1FF23FA82" bold="true" box="[146,174,1307,1326]" pageId="12">25)</emphasis>
|
||
<taxonomicName id="4BB224A8B040A762FF68FAA6FEA1FA83" authorityName="Dechambre" authorityYear="1999" box="[181,300,1308,1327]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="12" phylum="Arthropoda" rank="species" species="longitarsis">
|
||
<emphasis id="BEC68339B040A762FF68FAA6FEA1FA83" box="[181,300,1308,1327]" italics="true" pageId="12">C. longitarsis</emphasis>
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||
</taxonomicName>
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||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8C0D5F2BB040A762FF77FA6DFE93F98E" blockId="12.[146,975,1387,1570]" pageId="12">
|
||
<materialsCitation id="3CDA5576B040A762FF77FA6DFE95F9AB" collectorName="Lincoln, NE" location="University of Nebraska State Museum" pageId="12" specimenCount="2" typeStatus="holotype">
|
||
<typeStatus id="5309E189B040A762FF77FA6DFE87FA41" box="[170,266,1495,1517]" pageId="12" type="holotype">Holotype</typeStatus>
|
||
and
|
||
<typeStatus id="5309E189B040A762FEE3FA6DFE1DFA41" box="[318,400,1495,1517]" pageId="12" type="allotype">allotype</typeStatus>
|
||
deposited at the
|
||
<location id="896D09F0B040A762FDE2FA6DFC46FA41" LSID="urn:lsid:plazi:treatment:041BEE3DB047A760FE8CFA1FFF3FFF3D:896D09F0B040A762FDE2FA6DFC46FA41" box="[575,971,1495,1517]" name="University of Nebraska State Museum" pageId="12">University of Nebraska State Museum</location>
|
||
,
|
||
<collectorName id="21473AFDB040A762FF4FFA4BFE95F9AB" box="[146,280,1521,1543]" pageId="12">Lincoln, NE</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3CDA5576B040A762FEF9FA4BFE97F98E" country="Netherlands" location="Meindert Hielkema" municipality="Gouda" pageId="12" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="5309E189B040A762FEF9FA4BFEF2F9AB" box="[292,383,1521,1543]" pageId="12" type="paratype">Paratype</typeStatus>
|
||
deposited in the
|
||
<location id="896D09F0B040A762FDE7FA4BFC89F9AB" LSID="urn:lsid:plazi:treatment:041BEE3DB047A760FE8CFA1FFF3FFF3D:896D09F0B040A762FDE7FA4BFC89F9AB" box="[570,772,1521,1543]" country="Netherlands" municipality="Gouda" name="Meindert Hielkema" pageId="12">Meindert Hielkema</location>
|
||
collection (
|
||
<collectingMunicipality id="6C69C551B040A762FC5CFA4BFC46F9AB" box="[897,971,1521,1543]" pageId="12">Gouda</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F4A51FBBB040A762FF4FF9B6FE9EF98E" box="[146,275,1548,1570]" name="Netherlands" pageId="12">Netherlands</collectingCountry>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C0D5F2BB041A763FF1AFF78FEA0FD5A" blockId="13.[175,1005,194,1546]" pageId="13">
|
||
<emphasis id="BEC68339B041A763FF1AFF78FEA7FF74" bold="true" box="[199,298,194,216]" pageId="13">
|
||
<typeStatus id="5309E189B041A763FF1AFF78FEABFF74" box="[199,294,194,216]" pageId="13" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Male. Length 14.0 mm; width across humeri
|
||
<quantity id="4B4AF2CEB041A763FC96FF78FC11FF74" box="[843,924,194,216]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="13" unit="mm" value="6.7">6.7 mm</quantity>
|
||
. Color testaceous except for 3 small black spots on each elytron (post-humeral, postscutellar, center of disc behind middle), and apices of femora, tibiae, protibial teeth, and tarsomeres black.
|
||
<emphasis id="BEC68339B041A763FE09FEAEFD87FE85" box="[468,522,276,297]" italics="true" pageId="13">Head</emphasis>
|
||
: Surface finely shagreened, completely, densely punctate; punctures small, those on frons slightly larger than those on clypeus. Frontoclypeal suture biarcuate, weak. Clypeus with apex narrowly parabolic, with thin bead on margin. Interocular width equals 3.0 transverse eye diameters. Antenna 10-segmented, club subequal in length to segments 2–7.
|
||
<emphasis id="BEC68339B041A763FC5FFE3AFC68FE39" box="[898,997,384,405]" italics="true" pageId="13">Pronotum</emphasis>
|
||
: Surface minutely shagreened, completely, densely punctate, punctures similar in size to those of frons. Base with complete marginal bead. Posterior angles broadly rounded.
|
||
<emphasis id="BEC68339B041A763FECCFE6BFEDDFE4A" box="[273,336,465,486]" italics="true" pageId="13">Elytra</emphasis>
|
||
: Surface finely shagreened, punctate; punctures moderately large, dense; double rows of punctate striae distinct, punctures in those rows setigerous, setae short and tawny.
|
||
<taxonomicName id="4BB224A8B041A763FE16FDBDFD1DFDB1" authority=": Surface" authorityName="Surface" box="[459,656,519,541]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="13" phylum="Chordata" rank="genus">
|
||
<emphasis id="BEC68339B041A763FE16FDBDFDA7FDB0" box="[459,554,519,540]" italics="true" pageId="13">Pygidium</emphasis>
|
||
: Surface
|
||
</taxonomicName>
|
||
vaguely shagreened, punctate; punctures moderately large, moderately dense, setigerous; setae moderately dense, long, tawny. In lateral view, surface evenly convex.
|
||
<emphasis id="BEC68339B041A763FC89FD84FC09FDFF" box="[852,900,574,595]" italics="true" pageId="13">Legs</emphasis>
|
||
: Protibia bidentate, with slight swelling where basal, 3rd tooth would be. Protarsus strongly enlarged; tarsomeres 2–4 each slightly larger than preceding; 4th with large, ventral lobe (
|
||
<figureCitation id="148943AEB041A763FEE1FD35FE04FD09" box="[316,393,655,677]" captionStart="Figs" captionStartId="12.[167,206,1259,1278]" captionTargetBox="[146,976,194,1231]" captionTargetId="figure-65@12.[146,976,194,1237]" captionTargetPageId="12" captionText="Figs. 21–25. Cyclocephala hielkemaorum: 21) dorsal view of holotype; 22–23) parameres (caudal and lateral views); 24–25) right protarsus, median view, of 24) C. hielkemaorum and 25) C. longitarsis." figureDoi="http://doi.org/10.5281/zenodo.5369977" httpUri="https://zenodo.org/record/5369977/files/figure.png" pageId="13">Fig. 24</figureCitation>
|
||
), 5th subequal in length to tarsomeres 2–4, nearly flat on ventral surface; median claw enlarged, strongly bent, apex entire. Metatarsus twice as long as metatibia.
|
||
<emphasis id="BEC68339B041A763FE1DFD7FFD8EFD76" box="[448,515,709,730]" italics="true" pageId="13">Venter</emphasis>
|
||
: Prosternal process slender, conical.
|
||
<emphasis id="BEC68339B041A763FCA6FD7FFC6BFD76" box="[891,998,709,730]" italics="true" pageId="13">Parameres</emphasis>
|
||
:
|
||
<figureCitation id="148943AEB041A763FF72FD5AFEA5FD5A" box="[175,296,736,758]" captionStart="Figs" captionStartId="12.[167,206,1259,1278]" captionTargetBox="[146,976,194,1231]" captionTargetId="figure-65@12.[146,976,194,1237]" captionTargetPageId="12" captionText="Figs. 21–25. Cyclocephala hielkemaorum: 21) dorsal view of holotype; 22–23) parameres (caudal and lateral views); 24–25) right protarsus, median view, of 24) C. hielkemaorum and 25) C. longitarsis." figureDoi="http://doi.org/10.5281/zenodo.5369977" httpUri="https://zenodo.org/record/5369977/files/figure.png" pageId="13">Figs. 22–23</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C0D5F2BB041A763FF1AFD41FE9AFC46" blockId="13.[175,1005,194,1546]" pageId="13">
|
||
<emphasis id="BEC68339B041A763FF1AFD41FEAFFCBD" bold="true" box="[199,290,763,785]" pageId="13">
|
||
<typeStatus id="5309E189B041A763FF1AFD41FE93FCBD" box="[199,286,763,785]" pageId="13" type="allotype">Allotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Female. Length
|
||
<quantity id="4B4AF2CEB041A763FE0FFD41FDA3FCBD" box="[466,558,763,785]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="13" unit="mm" value="12.6">12.6 mm</quantity>
|
||
; width across humeri
|
||
<quantity id="4B4AF2CEB041A763FCCEFD41FCE9FCBD" box="[787,868,763,785]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="13" unit="mm" value="6.1">6.1 mm</quantity>
|
||
. As
|
||
<typeStatus id="5309E189B041A763FC4EFD41FC60FCBD" box="[915,1005,763,785]" pageId="13" type="holotype">holotype</typeStatus>
|
||
except in the following respects: Color and pattern similar except elytra with postmedian, small, black spot absent, and swelling of elytral margin darkened.
|
||
<emphasis id="BEC68339B041A763FF72FCF7FF63FCCE" box="[175,238,845,866]" italics="true" pageId="13">Elytra</emphasis>
|
||
: Surface with setae minute or abraded away. Epipleuron (in ventral view) gradually tapering towards apex. Lateral margin slightly swollen at level of sternites 1–2.
|
||
<taxonomicName id="4BB224A8B041A763FEE2FC39FE74FC35" authority=": Surface" authorityName="Surface" box="[319,505,899,921]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="13" phylum="Chordata" rank="genus">
|
||
<emphasis id="BEC68339B041A763FEE2FC39FE13FC34" box="[319,414,899,920]" italics="true" pageId="13">Pygidium</emphasis>
|
||
: Surface
|
||
</taxonomicName>
|
||
glabrous, densely punctate, punctures small. In lateral view, surface nearly flat.
|
||
<emphasis id="BEC68339B041A763FDC2FC24FDC3FC1F" box="[543,590,926,947]" italics="true" pageId="13">Legs</emphasis>
|
||
: Protibia tridentate, teeth subequally spaced. Protarsus simple, not enlarged. Metatarsus slightly longer than metatibia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B041A763FF1AFC4AFCAAFB8D" pageId="13" type="description">
|
||
<paragraph id="8C0D5F2BB041A763FF1AFC4AFCAAFB8D" blockId="13.[175,1005,194,1546]" pageId="13">
|
||
<emphasis id="BEC68339B041A763FF1AFC4AFEA0FBAA" bold="true" box="[199,301,1008,1030]" pageId="13">Variation.</emphasis>
|
||
Males (
|
||
<specimenCount id="9AB494A2B041A763FE59FC4AFE79FBAA" box="[388,500,1008,1030]" pageId="13" type="generic" typeStatus="paratype">1 paratype</specimenCount>
|
||
). Length
|
||
<quantity id="4B4AF2CEB041A763FD8AFC4AFD3EFBAA" box="[599,691,1008,1030]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="13" unit="mm" value="13.9">13.9 mm</quantity>
|
||
; width across humeri
|
||
<quantity id="4B4AF2CEB041A763FC45FC4AFC64FBAA" box="[920,1001,1008,1030]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="13" unit="mm" value="6.6">6.6 mm</quantity>
|
||
. The
|
||
<typeStatus id="5309E189B041A763FF02FBB1FEB4FB8D" box="[223,313,1035,1057]" pageId="13" type="paratype">paratype</typeStatus>
|
||
does not differ significantly from the
|
||
<typeStatus id="5309E189B041A763FD1BFBB1FCAEFB8D" box="[710,803,1035,1057]" pageId="13" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B041A763FF1AFB9DFD16FBDF" pageId="13" type="etymology">
|
||
<paragraph id="8C0D5F2BB041A763FF1AFB9DFD16FBDF" blockId="13.[175,1005,194,1546]" pageId="13">
|
||
<emphasis id="BEC68339B041A763FF1AFB9DFEB7FB91" bold="true" box="[199,314,1063,1085]" pageId="13">Etymology.</emphasis>
|
||
This species is named in honor of Meindert Hielkema and his son, Auke, who were responsible for collecting this species in
|
||
<collectingCountry id="F4A51FBBB041A763FD21FBF8FC15FBF4" box="[764,920,1090,1112]" name="French Guiana" pageId="13">French Guiana</collectingCountry>
|
||
and for generously sending me the specimens for study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B041A763FF1AFBC2FDDAFB06" pageId="13" type="distribution">
|
||
<paragraph id="8C0D5F2BB041A763FF1AFBC2FDDAFB06" blockId="13.[175,1005,194,1546]" pageId="13">
|
||
<emphasis id="BEC68339B041A763FF1AFBC2FECBFB22" bold="true" box="[199,326,1144,1166]" pageId="13">Distribution.</emphasis>
|
||
<taxonomicName id="4BB224A8B041A763FE8EFBC3FDFEFB22" authorityName="Ratcliffe" authorityYear="2008" box="[339,627,1145,1166]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="hielkemaorum">
|
||
<emphasis id="BEC68339B041A763FE8EFBC3FDFEFB22" box="[339,627,1145,1166]" italics="true" pageId="13">Cyclocephala hielkemaorum</emphasis>
|
||
</taxonomicName>
|
||
is known only from the lowland rainforest near Cacao in
|
||
<collectingCountry id="F4A51FBBB041A763FE6EFB2EFDD9FB06" box="[435,596,1172,1194]" name="French Guiana" pageId="13">French Guiana</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B041A760FF1AFB15FF3FFF3D" lastPageId="14" pageId="13" type="diagnosis">
|
||
<paragraph id="8C0D5F2BB041A763FF1AFB15FD00F9A6" blockId="13.[175,1005,194,1546]" pageId="13">
|
||
<emphasis id="BEC68339B041A763FF1AFB15FEA2FB69" bold="true" box="[199,303,1199,1221]" pageId="13">Diagnosis.</emphasis>
|
||
<taxonomicName id="4BB224A8B041A763FEEAFB15FDC2FB68" authorityName="Ratcliffe" authorityYear="2008" box="[311,591,1199,1220]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="hielkemaorum">
|
||
<emphasis id="BEC68339B041A763FEEAFB15FDC2FB68" box="[311,591,1199,1220]" italics="true" pageId="13">Cyclocephala hielkemaorum</emphasis>
|
||
</taxonomicName>
|
||
will key to couplets 240/
|
||
<quantity id="4B4AF2CEB041A763FC8EFB15FC1CFB69" box="[851,913,1199,1221]" metricMagnitude="0" metricUnit="m" metricValue="6.1213999999999995" pageId="13" unit="in" value="241.0">241 in</quantity>
|
||
<bibRefCitation id="E82322DAB041A763FC45FB15FF7CFB4C" author="Endrodi" pageId="13" refId="ref10514" refString="Endrodi, S. 1985. The Dynastinae of the World. W. Junk Publishers, Dordrecht. 800 pp." type="book" year="1985">Endrödi (1985)</bibRefCitation>
|
||
, which end with
|
||
<taxonomicName id="4BB224A8B041A763FE61FB71FD41FB4C" authority="Hohne" authorityName="Hohne" authorityYear="1923" box="[444,716,1226,1248]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="quadripunctata">
|
||
<emphasis id="BEC68339B041A763FE61FB71FDF5FB4C" box="[444,632,1227,1248]" italics="true" pageId="13">C. quadripunctata</emphasis>
|
||
Höhne
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4BB224A8B041A763FCD3FB71FC65FB4C" authority="Endrodi" authorityName="Endrodi" authorityYear="1966" box="[782,1000,1226,1248]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="peruana">
|
||
<emphasis id="BEC68339B041A763FCD3FB71FC0EFB4C" box="[782,899,1227,1248]" italics="true" pageId="13">C. peruana</emphasis>
|
||
Endrödi
|
||
</taxonomicName>
|
||
, respectively. Both of these species have a black frons, different parameres, and are smaller in size.
|
||
<taxonomicName id="4BB224A8B041A763FEB7FABBFD01FABA" authorityName="Ratcliffe" authorityYear="2008" box="[362,652,1281,1302]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="hielkemaorum">
|
||
<emphasis id="BEC68339B041A763FEB7FABBFD01FABA" box="[362,652,1281,1302]" italics="true" pageId="13">Cyclocephala hielkemaorum</emphasis>
|
||
</taxonomicName>
|
||
is most similar, including the parameres, to
|
||
<taxonomicName id="4BB224A8B041A763FE9BFAA6FDC0FA9E" authority="Dechambre" authorityName="Dechambre" authorityYear="1999" box="[326,589,1308,1330]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="longitarsis">
|
||
<emphasis id="BEC68339B041A763FE9BFAA6FE43FA9D" box="[326,462,1308,1329]" italics="true" pageId="13">C. longitarsis</emphasis>
|
||
Dechambre
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F4A51FBBB041A763FD4CFAA6FD61FA9E" box="[657,748,1308,1330]" name="Ecuador" pageId="13">Ecuador</collectingCountry>
|
||
. However, the protarsus of the males is significantly different: in
|
||
<taxonomicName id="4BB224A8B041A763FD9AFA8DFD7CFAE0" authorityName="Ratcliffe" authorityYear="2008" box="[583,753,1335,1356]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="hielkemaorum">
|
||
<emphasis id="BEC68339B041A763FD9AFA8DFD7CFAE0" box="[583,753,1335,1356]" italics="true" pageId="13">C. hielkemaorum</emphasis>
|
||
</taxonomicName>
|
||
, the 4th tarsomere has a large, ventral lobe, and the 5th tarsomere is nearly flat on its venter (
|
||
<figureCitation id="148943AEB041A763FC49FAE8FC6CFAC4" box="[916,993,1362,1384]" captionStart="Figs" captionStartId="12.[167,206,1259,1278]" captionTargetBox="[146,976,194,1231]" captionTargetId="figure-65@12.[146,976,194,1237]" captionTargetPageId="12" captionText="Figs. 21–25. Cyclocephala hielkemaorum: 21) dorsal view of holotype; 22–23) parameres (caudal and lateral views); 24–25) right protarsus, median view, of 24) C. hielkemaorum and 25) C. longitarsis." figureDoi="http://doi.org/10.5281/zenodo.5369977" httpUri="https://zenodo.org/record/5369977/files/figure.png" pageId="13">Fig. 24</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="4BB224A8B041A763FEFFFAD7FE25FA2E" authorityName="Dechambre" authorityYear="1999" box="[290,424,1389,1410]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="longitarsis">
|
||
<emphasis id="BEC68339B041A763FEFFFAD7FE25FA2E" box="[290,424,1389,1410]" italics="true" pageId="13">C. longitarsis</emphasis>
|
||
</taxonomicName>
|
||
the 4th tarsomere is not expanded into a large lobe, and the 5th tarsomere is concave on its venter with the ventrolateral edge strongly toothed (
|
||
<figureCitation id="148943AEB041A763FEC4FA19FEE8FA15" box="[281,357,1443,1465]" captionStart="Figs" captionStartId="12.[167,206,1259,1278]" captionTargetBox="[146,976,194,1231]" captionTargetId="figure-65@12.[146,976,194,1237]" captionTargetPageId="12" captionText="Figs. 21–25. Cyclocephala hielkemaorum: 21) dorsal view of holotype; 22–23) parameres (caudal and lateral views); 24–25) right protarsus, median view, of 24) C. hielkemaorum and 25) C. longitarsis." figureDoi="http://doi.org/10.5281/zenodo.5369977" httpUri="https://zenodo.org/record/5369977/files/figure.png" pageId="13">Fig. 25</figureCitation>
|
||
). In addition, these two species are broadly separated geographically.
|
||
<taxonomicName id="4BB224A8B041A763FE88FA05FDFCFA78" authorityName="Ratcliffe" authorityYear="2008" box="[341,625,1471,1492]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="hielkemaorum">
|
||
<emphasis id="BEC68339B041A763FE88FA05FDFCFA78" box="[341,625,1471,1492]" italics="true" pageId="13">Cyclocephala hielkemaorum</emphasis>
|
||
</taxonomicName>
|
||
is known from northeastern South America on the east side of the Andes, and
|
||
<taxonomicName id="4BB224A8B041A763FDAEFA60FD74FA43" authorityName="Dechambre" authorityYear="1999" box="[627,761,1498,1519]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="13" phylum="Arthropoda" rank="species" species="longitarsis">
|
||
<emphasis id="BEC68339B041A763FDAEFA60FD74FA43" box="[627,761,1498,1519]" italics="true" pageId="13">C. longitarsis</emphasis>
|
||
</taxonomicName>
|
||
is known from western South America on the west side of the Andes.
|
||
</paragraph>
|
||
<paragraph id="8C0D5F2BB042A760FF4FFFC4FF3FFF3D" blockId="14.[146,178,126,145]" box="[146,178,126,145]" pageId="14">234</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |