340 lines
33 KiB
XML
340 lines
33 KiB
XML
<document id="6103E4B004B2BE0B886859C9AA90A1C9" ID-DOI="10.1649/1066.1" ID-ISSN="1938-4394" ID-Zenodo-Dep="5369967" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630268265367" checkinUser="valdenar" docAuthor="Ratcliffe, Brett C." docDate="1821" docId="041BEE3DB045A765FEB5FC4EFDDBFAD3" docLanguage="en" docName="1066.1.pdf" docOrigin="The Coleopterists Bulletin 62 (2)" docSource="http://dx.doi.org/10.1649/1066.1" docStyle="DocumentStyle:76939A7658C5F80CF4BB6818E728552A.2:TheColeopBull.2003-2009.journal_article.1cover" docStyleId="76939A7658C5F80CF4BB6818E728552A" docStyleName="TheColeopBull.2003-2009.journal_article.1cover" docStyleVersion="2" docTitle="Cyclocephala compacta Ratcliffe 2008, new species" docType="treatment" docVersion="4" lastPageNumber="241" masterDocId="F8229645B04CA76EFFDDFFBAFF8DFFAC" masterDocTitle="More New Species ofCyclocephalaDejean, 1821 from South America (Scarabaeidae: Dynastinae: Cyclocephalini)" masterLastPageNumber="241" masterPageNumber="221" pageNumber="221" updateTime="1699205293535" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="8347786B26B3D57704B5DC394380A00C">
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<mods:title id="0A857A4B4BDA2ACD41D2C3C21283E880">More New Species ofCyclocephalaDejean, 1821 from South America (Scarabaeidae: Dynastinae: Cyclocephalini)</mods:title>
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<mods:name id="170A3A909EEC97C05721838AEAE5A686" type="personal">
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<mods:role id="CDFFAC49C54DF3AD32AB2D5C451BC48C">
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<mods:roleTerm id="C369F9AD3673DA3554FD580C20FAE086">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A53F8787B2EAE7306C288315F1E3C480">Ratcliffe, Brett C.</mods:namePart>
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<mods:typeOfResource id="B3E5190FF963A13E4A3BF78FE1A11699">text</mods:typeOfResource>
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<mods:titleInfo id="BF28F22AAE7D0AA438827D9A1597BB3F">
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<mods:title id="76F55DC697B3192B4E89B4A8492CF5C9">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="01DA9A86B2FA99FA378C1459F71AD80D">
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<mods:date id="7802270CBBBDFA6ADD9879A41B5EA161">1821</mods:date>
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<mods:detail id="1A893728AF8FB82FF3DCE7552F07BC2B" type="pubDate">
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<mods:number id="6D13373504F9CF3A4E55A0AF72954AAC">2008-06-30</mods:number>
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</mods:detail>
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<mods:detail id="73AA4684DBA8774388CE70EDAF648D39" type="volume">
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<mods:number id="E742E2A83E1469393D40C6D6BEB85C62">62</mods:number>
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</mods:detail>
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<mods:detail id="48AA89F6C3DAEB1E32F15B98906B1B98" type="issue">
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<mods:number id="36E2EF8FC35E6F1200EFF495FE5ABB37">2</mods:number>
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</mods:detail>
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<mods:extent id="531077B8C87FA88A97BC6FE274FA9B19" unit="page">
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<mods:start id="498B176F356098F40749B3995CCC6117">221</mods:start>
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<mods:end id="4CBB5E62C9933B579F95483A3B83210C">241</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="6BC366CE4B33249E3B945ECDB724864F">
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<mods:url id="B431A2E9B4B8A49E1F2AC375006908B7">http://dx.doi.org/10.1649/1066.1</mods:url>
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</mods:location>
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<mods:classification id="FB3203D01981B216525EC5A3AD339990">journal article</mods:classification>
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<mods:identifier id="7FFCC226EA783226CD216F22B6D49F2C" type="DOI">10.1649/1066.1</mods:identifier>
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<mods:identifier id="566E24E1633EFA0418347BA6237687DA" type="ISSN">1938-4394</mods:identifier>
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<mods:identifier id="AE21FEE2AFB93FF3411D629ADCF8B788" type="Zenodo-Dep">5369967</mods:identifier>
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</mods:mods>
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<treatment id="041BEE3DB045A765FEB5FC4EFDDBFAD3" ID-DOI="http://doi.org/10.5281/zenodo.5461604" ID-Zenodo-Dep="5461604" LSID="urn:lsid:plazi:treatment:041BEE3DB045A765FEB5FC4EFDDBFAD3" httpUri="http://treatment.plazi.org/id/041BEE3DB045A765FEB5FC4EFDDBFAD3" lastPageId="11" pageId="9">
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<subSubSection id="C4A80CA0B045A767FEB5FC4EFCB9FBA6" box="[360,820,1012,1034]" pageId="9" type="nomenclature">
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<paragraph id="8C0D5F2BB045A767FEB5FC4EFCB9FBA6" blockId="9.[360,820,1012,1062]" box="[360,820,1012,1034]" pageId="9">
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<heading id="D745E847B045A767FEB5FC4EFCB9FBA6" box="[360,820,1012,1034]" centered="true" fontSize="9" level="2" pageId="9" reason="2">
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<emphasis id="BEC68339B045A767FEB5FC4EFCB9FBA6" bold="true" box="[360,820,1012,1034]" pageId="9">
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<taxonomicName id="4BB224A8B045A767FEB5FC4EFD38FBA6" authority="Ratcliffe" authorityName="Ratcliffe" authorityYear="2008" box="[360,693,1012,1034]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="9" phylum="Arthropoda" rank="species" species="compacta" status="sp. nov.">
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<emphasis id="BEC68339B045A767FEB5FC4EFDD9FBA6" bold="true" box="[360,596,1012,1034]" italics="true" pageId="9">Cyclocephala compacta</emphasis>
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Ratcliffe
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</taxonomicName>
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||
,
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<taxonomicNameLabel id="A5F53E42B045A767FD1CFC4EFCB9FBA6" box="[705,820,1012,1034]" pageId="9" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C4A80CA0B045A767FDD6FBAAFD1CFB8A" box="[523,657,1040,1062]" pageId="9" type="description">
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<paragraph id="8C0D5F2BB045A767FDD6FBAAFD1CFB8A" blockId="9.[360,820,1012,1062]" box="[523,657,1040,1062]" pageId="9">
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(
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<figureCitation id="148943AEB045A767FDCFFBAAFD04FB8A" box="[530,649,1040,1062]" captionStart="Figs" captionStartId="10.[167,206,1170,1189]" captionTargetBox="[146,975,194,1148]" captionTargetId="figure-136@10.[146,976,194,1148]" captionTargetPageId="10" captionText="Figs. 17–20. Cyclocephala compacta: 17) dorsal view of holotype; 18–19) parameres" figureDoi="http://doi.org/10.5281/zenodo.5369975" httpUri="https://zenodo.org/record/5369975/files/figure.png" pageId="9">Figs. 17–20</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C4A80CA0B045A765FF1AFBFBFDF9FD9B" lastPageId="11" pageId="9" type="materials_examined">
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<paragraph id="8C0D5F2BB045A767FF1AFBFBFCEAFB6F" blockId="9.[175,1005,1089,1570]" pageId="9">
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<emphasis id="BEC68339B045A767FF1AFBFBFEE5FBFB" bold="true" box="[199,360,1089,1111]" pageId="9">Type Material.</emphasis>
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<materialsCitation id="3CDA5576B045A767FEA4FBFBFEA3FB21" collectingDate="1993-10-05" collectingDateMax="1993-10-15" collectingDateMin="1993-10-05" collectorName="S. Ariquemes & C. & K. Messenger." country="Brazil" latitude="-10.535833" location="Fazenda Rancho Grande" municipality="Ariquemes" pageId="9" specimenCount="1" stateProvince="Rondonia" typeStatus="holotype">
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<typeStatus id="5309E189B045A767FEA4FBFBFE54FBFB" box="[377,473,1089,1111]" pageId="9" type="holotype">Holotype</typeStatus>
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||
male, labeled ‘‘
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<collectingCountry id="F4A51FBBB045A767FD49FBFBFD7BFBFB" box="[660,758,1089,1111]" name="Brazil" pageId="9">BRASIL</collectingCountry>
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:
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<collectingRegion id="4E7691C9B045A767FCD4FBFBFCFFFBFB" box="[777,882,1089,1111]" country="Brazil" name="Rondonia" pageId="9">Rondonia</collectingRegion>
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||
,
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||
<quantity id="4B4AF2CEB045A767FC5AFBFBFC45FBFB" box="[903,968,1089,1111]" metricMagnitude="4" metricUnit="m" metricValue="6.2" pageId="9" unit="km" value="62.0">62 km</quantity>
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||
<collectorName id="21473AFDB045A767FC05FBFBFEADFBDE" pageId="9">
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S.
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<collectingMunicipality id="6C69C551B045A767FF72FBE6FEADFBDE" box="[175,288,1116,1138]" pageId="9">Ariquemes</collectingMunicipality>
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||
</collectorName>
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||
,
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<location id="896D09F0B045A767FEEDFBE6FDB3FBDE" LSID="urn:lsid:plazi:treatment:041BEE3DB045A765FEB5FC4EFDDBFAD3:896D09F0B045A767FEEDFBE6FDB3FBDE" box="[304,574,1116,1138]" country="Brazil" latitude="-10.535833" municipality="Ariquemes" name="Fazenda Rancho Grande" pageId="9" stateProvince="Rondonia">Fazenda Rancho Grande</location>
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,
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||
<geoCoordinate id="E98639ECB045A767FD93FBE6FD13FBDE" box="[590,670,1116,1138]" degrees="10" direction="south" minutes="32" orientation="latitude" pageId="9" precision="15" seconds="9" value="-10.535833">
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||
10
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<emphasis id="BEC68339B045A767FDBBFBE6FDE2FBDD" bold="true" box="[614,623,1116,1137]" italics="true" pageId="9">°</emphasis>
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||
32
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<emphasis id="BEC68339B045A767FD5AFBE6FD02FBDE" bold="true" box="[647,655,1116,1138]" pageId="9">9</emphasis>
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||
S
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||
</geoCoordinate>
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||
, 62
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||
<emphasis id="BEC68339B045A767FD1BFBE6FD42FBDD" bold="true" box="[710,719,1116,1137]" italics="true" pageId="9">°</emphasis>
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||
49
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||
<emphasis id="BEC68339B045A767FD3AFBE6FD62FBDE" bold="true" box="[743,751,1116,1138]" pageId="9">9</emphasis>
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||
,
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||
<date id="F80C79EBB045A767FCDCFBE6FC0FFBDE" box="[769,898,1116,1138]" pageId="9" value="1993-10-05" valueMax="1993-10-15" valueMin="1993-10-05">
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||
<collectingDate id="E8488003B045A767FCDCFBE6FC0FFBDE" box="[769,898,1116,1138]" pageId="9" value="1993-10-05" valueMax="1993-10-15" valueMin="1993-10-05">X-5-15-1993</collectingDate>
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</date>
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,
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<collectorName id="21473AFDB045A767FC4CFBE6FEABFB21" pageId="9">C. & K. Messenger.</collectorName>
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||
’’
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||
</materialsCitation>
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||
<materialsCitation id="3CDA5576B045A767FEE8FBCDFCEAFB6F" collectingDate="1994-09-19" collectingDateMax="1994-09-31" collectingDateMin="1994-09-19" collectorName="A. Reifschneider." country="Brazil" elevation="165" latitude="-10.535833" location="Fazenda Rancho Grande" municipality="Five" pageId="9" specimenCount="3" stateProvince="Rondonia" typeStatus="allotype">
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||
<typeStatus id="5309E189B045A767FEE8FBCDFE00FB21" box="[309,397,1143,1165]" pageId="9" type="allotype">Allotype</typeStatus>
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||
and
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||
<specimenCount id="9AB494A2B045A767FE1FFBCDFDBAFB21" box="[450,567,1143,1165]" pageId="9" type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
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with same data.
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<collectingMunicipality id="6C69C551B045A767FD3AFBCDFC99FB21" box="[743,788,1143,1165]" pageId="9">Five</collectingMunicipality>
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||
additional
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||
<typeStatus id="5309E189B045A767FC54FBCDFC61FB21" box="[905,1004,1143,1165]" pageId="9" type="paratype">paratypes</typeStatus>
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||
labeled ‘‘
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||
<collectingCountry id="F4A51FBBB045A767FED3FB28FEFDFB04" box="[270,368,1170,1192]" name="Brazil" pageId="9">BRASIL</collectingCountry>
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||
:
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||
<collectingRegion id="4E7691C9B045A767FE5DFB28FE64FB04" box="[384,489,1170,1192]" country="Brazil" name="Rondonia" pageId="9">Rondonia</collectingRegion>
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||
,
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||
<locationDeviation id="9DBF4609B045A767FE26FB28FD59FB04" box="[507,724,1170,1192]" location="Ariquemes" pageId="9">
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||
<quantity id="4B4AF2CEB045A767FE26FB28FDB6FB04" box="[507,571,1170,1192]" metricMagnitude="4" metricUnit="m" metricValue="6.2" pageId="9" unit="km" value="62.0">62 km</quantity>
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||
S Ariquemes
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</locationDeviation>
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||
,
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||
<quantity id="4B4AF2CEB045A767FD39FB28FCABFB04" box="[740,806,1170,1192]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="9" unit="m" value="165.0">
|
||
<elevation id="079FB818B045A767FD39FB28FCABFB04" box="[740,806,1170,1192]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="9" unit="m" value="165.0">165 m</elevation>
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||
</quantity>
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||
,
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||
<location id="896D09F0B045A767FCE5FB28FF72FB6F" LSID="urn:lsid:plazi:treatment:041BEE3DB045A765FEB5FC4EFDDBFAD3:896D09F0B045A767FCE5FB28FF72FB6F" country="Brazil" latitude="-10.535833" municipality="Five" name="Fazenda Rancho Grande" pageId="9" stateProvince="Rondonia">Fazenda Rancho Grande</location>
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||
,
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||
<geoCoordinate id="E98639ECB045A767FED6FB17FED6FB6F" box="[267,347,1196,1219]" degrees="10" direction="south" minutes="32" orientation="latitude" pageId="9" precision="15" seconds="9" value="-10.535833">
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||
10
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<emphasis id="BEC68339B045A767FEFEFB17FEA1FB6E" bold="true" box="[291,300,1197,1218]" italics="true" pageId="9">°</emphasis>
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||
32
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<emphasis id="BEC68339B045A767FE99FB16FEC1FB6E" bold="true" box="[324,332,1196,1218]" pageId="9">9</emphasis>
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||
S
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</geoCoordinate>
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, 62
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||
<emphasis id="BEC68339B045A767FE5DFB17FE04FB6E" bold="true" box="[384,393,1197,1218]" italics="true" pageId="9">°</emphasis>
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||
49
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||
<emphasis id="BEC68339B045A767FE7CFB16FE24FB6E" bold="true" box="[417,425,1196,1218]" pageId="9">9</emphasis>
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||
,
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||
<date id="F80C79EBB045A767FE6AFB17FDE7FB6F" box="[439,618,1197,1219]" pageId="9" value="1994-09-19" valueMax="1994-09-31" valueMin="1994-09-19">
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||
<collectingDate id="E8488003B045A767FE6AFB17FDE7FB6F" box="[439,618,1197,1219]" pageId="9" value="1994-09-19" valueMax="1994-09-31" valueMin="1994-09-19">19–31 Sept. 1994</collectingDate>
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||
</date>
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, coll.
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<collectorName id="21473AFDB045A767FD75FB17FCD2FB6F" box="[680,863,1197,1219]" pageId="9">A. Reifschneider.</collectorName>
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||
’’
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8C0D5F2BB045A767FF1AFB72FC5BFAB8" blockId="9.[175,1005,1089,1570]" pageId="9">
|
||
<materialsCitation id="3CDA5576B045A767FF1AFB72FC5CFAB8" collectorName="Lincoln, NE & Alex Reifschneider Collection & B. C. Ratcliffe Collection" location="Nebraska State" pageId="9" specimenCount="2" stateProvince="Nebraska" typeStatus="holotype">
|
||
<typeStatus id="5309E189B045A767FF1AFB72FEA8FB72" box="[199,293,1224,1246]" pageId="9" type="holotype">Holotype</typeStatus>
|
||
,
|
||
<typeStatus id="5309E189B045A767FEF0FB72FEF0FB72" box="[301,381,1224,1246]" pageId="9" type="allotype">allotype</typeStatus>
|
||
, and
|
||
<specimenCount id="9AB494A2B045A767FE73FB72FDBBFB72" box="[430,566,1224,1246]" pageId="9" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
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||
deposited at the University of
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<collectingRegion id="4E7691C9B045A767FC85FB72FC61FB72" box="[856,1004,1224,1246]" country="United States of America" name="Nebraska" pageId="9">Nebraska State</collectingRegion>
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||
Museum,
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||
<collectorName id="21473AFDB045A767FECCFB59FE03FB55" box="[273,398,1251,1273]" pageId="9">Lincoln, NE</collectorName>
|
||
. Two
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||
<typeStatus id="5309E189B045A767FE14FB59FDAAFB55" box="[457,551,1251,1273]" pageId="9" type="paratype">paratypes</typeStatus>
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||
deposited in the
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<collectorName id="21473AFDB045A767FD11FB59FC61FB55" box="[716,1004,1251,1273]" pageId="9">Alex Reifschneider Collection</collectorName>
|
||
and
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||
<specimenCount id="9AB494A2B045A767FF06FB44FEE7FAB8" box="[219,362,1278,1300]" pageId="9" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
|
||
deposited in the
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<collectorName id="21473AFDB045A767FDCEFB44FC86FAB8" box="[531,779,1278,1300]" pageId="9">B. C. Ratcliffe Collection</collectorName>
|
||
(both
|
||
<collectorName id="21473AFDB045A767FC90FB44FC46FAB8" box="[845,971,1278,1300]" pageId="9">Lincoln, NE</collectorName>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C0D5F2BB045A764FF1AFAA0FC84FB11" blockId="9.[175,1005,1089,1570]" lastBlockId="10.[146,975,1170,1213]" lastPageId="10" pageId="9">
|
||
<emphasis id="BEC68339B045A767FF1AFAA0FEA7FA9C" bold="true" box="[199,298,1306,1328]" pageId="9">
|
||
<typeStatus id="5309E189B045A767FF1AFAA0FEABFA9C" box="[199,294,1306,1328]" pageId="9" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Male. Length
|
||
<quantity id="4B4AF2CEB045A767FE3EFAA0FDB2FA9C" box="[483,575,1306,1328]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="9" unit="mm" value="12.2">12.2 mm</quantity>
|
||
; width across humeri
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||
<quantity id="4B4AF2CEB045A767FC96FAA0FC11FA9C" box="[843,924,1306,1328]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="9" unit="mm" value="7.2">7.2 mm</quantity>
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||
. Color testaceous except for black frons; dark testaceous clypeus with black bead on apex; 6 small, black spots on pronotum (
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||
<quantity id="4B4AF2CEB045A767FD80FAEAFD0BFACA" box="[605,646,1360,1382]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="9" unit="in" value="4.0">4 in</quantity>
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||
transverse row and 1 each behind median spots in transverse row); elytra with curved, black macula behind humerus, short black line near middle, and small black spot mesad of black line; pygidium black either side of middle; protibial teeth black; femora and tarsi dark testaceous; meso- and metathorax and abdominal sternites piceous.
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||
<emphasis id="BEC68339B045A767FCB1FA06FC2FFA7D" box="[876,930,1468,1489]" italics="true" pageId="9">Head</emphasis>
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||
: Frons with dense, moderately large punctures. Frontoclypeal suture biarcuate, distinct. Clypeus with surface vaguely, transversely rugopunctate; apex semicircular with thin, marginal bead. Interocular width equals 3.0 transverse eye diameters. 230 (caudal and lateral views);
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||
<emphasis id="BEC68339B046A764FE57FB10FE2BFB11" bold="true" box="[394,422,1194,1213]" pageId="10">20)</emphasis>
|
||
pronotum showing variation in spots.
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||
</paragraph>
|
||
<caption id="D8CD0FA3B046A764FF7AFB28FC42FB09" ID-DOI="http://doi.org/10.5281/zenodo.5369975" ID-Zenodo-Dep="5369975" box="[167,975,1170,1190]" httpUri="https://zenodo.org/record/5369975/files/figure.png" pageId="10" startId="10.[167,206,1170,1189]" targetBox="[146,975,194,1148]" targetPageId="10">
|
||
<paragraph id="8C0D5F2BB046A764FF7AFB28FC42FB09" blockId="10.[146,975,1170,1213]" box="[167,975,1170,1190]" pageId="10">
|
||
<emphasis id="BEC68339B046A764FF7AFB28FE95FB09" bold="true" box="[167,280,1170,1189]" pageId="10">Figs. 17–20.</emphasis>
|
||
<taxonomicName id="4BB224A8B046A764FEF0FB29FD8CFB0A" authorityName="Ratcliffe" authorityYear="2008" box="[301,513,1171,1190]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="10" phylum="Arthropoda" rank="species" species="compacta">
|
||
<emphasis id="BEC68339B046A764FEF0FB29FD8CFB0A" box="[301,513,1171,1190]" italics="true" pageId="10">Cyclocephala compacta</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<emphasis id="BEC68339B046A764FDCFFB28FDA3FB09" bold="true" box="[530,558,1170,1189]" pageId="10">17)</emphasis>
|
||
dorsal view of holotype;
|
||
<emphasis id="BEC68339B046A764FCF6FB28FCEAFB09" bold="true" box="[811,871,1170,1189]" pageId="10">18–19)</emphasis>
|
||
parameres
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8C0D5F2BB046A765FF4FFB5CFCF5FEA2" blockId="10.[146,975,1254,1570]" lastBlockId="11.[175,1005,194,1407]" lastPageId="11" pageId="10">
|
||
Antenna 10-segmented, club slightly longer than segments 2–7.
|
||
<emphasis id="BEC68339B046A764FCCEFB5CFCFBFB57" box="[787,886,1254,1275]" italics="true" pageId="10">Pronotum</emphasis>
|
||
: Surface weakly shagreened, moderately punctate at center; punctures moderately large, becoming denser on sides. Base with complete marginal bead. Posterior angles broadly rounded.
|
||
<emphasis id="BEC68339B046A764FE92FA8CFE03FAE7" box="[335,398,1334,1355]" italics="true" pageId="10">Elytra</emphasis>
|
||
: Surface weakly shagreened, punctate; punctures large, dense, double rows of punctate striae indistinct. Apical angles with sparse, minute setae.
|
||
<taxonomicName id="4BB224A8B046A764FF02FAD6FE28FA2E" authority=": Surface" authorityName="Surface" box="[223,421,1388,1410]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="10" phylum="Chordata" rank="genus">
|
||
<emphasis id="BEC68339B046A764FF02FAD6FEB3FA2D" box="[223,318,1388,1409]" italics="true" pageId="10">Pygidium</emphasis>
|
||
: Surface
|
||
</taxonomicName>
|
||
opaque, vaguely shagreened, vaguely punctate; punctures dense, moderately large, shallow, setigerous; setae short, tawny. In lateral view, surface regularly convex.
|
||
<emphasis id="BEC68339B046A764FDC4FA1BFDC5FA1A" box="[537,584,1441,1462]" italics="true" pageId="10">Legs</emphasis>
|
||
: Protibia tridentate, basal tooth small and strongly removed from apical teeth; median tooth with anterior edge perpendicular to shaft of tibia (
|
||
<figureCitation id="148943AEB046A764FE05FA6DFDA9FA41" box="[472,548,1495,1517]" captionStart="Figs" captionStartId="10.[167,206,1170,1189]" captionTargetBox="[146,975,194,1148]" captionTargetId="figure-136@10.[146,976,194,1148]" captionTargetPageId="10" captionText="Figs. 17–20. Cyclocephala compacta: 17) dorsal view of holotype; 18–19) parameres" figureDoi="http://doi.org/10.5281/zenodo.5369975" httpUri="https://zenodo.org/record/5369975/files/figure.png" pageId="10">Fig. 17</figureCitation>
|
||
). Protarsus strongly enlarged; tarsomeres 2–4 each slightly larger than preceding; 4th with subtriangular, ventral lobe, 5th subequal in length to tarsomeres 1–4; median claw enlarged, strongly bent, apex narrowly split into large and small rami. Metatarsus a little longer than metatibia.
|
||
<emphasis id="BEC68339B047A765FF72FF67FF7FFF5E" box="[175,242,221,242]" italics="true" pageId="11">Venter</emphasis>
|
||
: Prosternal process long, columnar, apex transversely oval and flat with a raised, transverse ‘‘button’’ on anterior half.
|
||
<emphasis id="BEC68339B047A765FD5FFF42FD60FEA1" box="[642,749,248,269]" italics="true" pageId="11">Parameres</emphasis>
|
||
:
|
||
<figureCitation id="148943AEB047A765FD26FF42FCF9FEA2" box="[763,884,248,270]" captionStart="Figs" captionStartId="10.[167,206,1170,1189]" captionTargetBox="[146,975,194,1148]" captionTargetId="figure-136@10.[146,976,194,1148]" captionTargetPageId="10" captionText="Figs. 17–20. Cyclocephala compacta: 17) dorsal view of holotype; 18–19) parameres" figureDoi="http://doi.org/10.5281/zenodo.5369975" httpUri="https://zenodo.org/record/5369975/files/figure.png" pageId="11">Figs. 18–19</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C0D5F2BB047A765FF1AFEAEFDF9FD9B" blockId="11.[175,1005,194,1407]" pageId="11">
|
||
<emphasis id="BEC68339B047A765FF1AFEAEFEAFFE86" bold="true" box="[199,290,276,298]" pageId="11">
|
||
<typeStatus id="5309E189B047A765FF1AFEAEFE93FE86" box="[199,286,276,298]" pageId="11" type="allotype">Allotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Female. Length
|
||
<quantity id="4B4AF2CEB047A765FE0EFEAEFDA2FE86" box="[467,559,276,298]" metricMagnitude="-2" metricUnit="m" metricValue="1.16" pageId="11" unit="mm" value="11.6">11.6 mm</quantity>
|
||
; width across humeri
|
||
<quantity id="4B4AF2CEB047A765FCCEFEAEFCE8FE86" box="[787,869,276,298]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="11" unit="mm" value="6.8">6.8 mm</quantity>
|
||
. As
|
||
<typeStatus id="5309E189B047A765FC4EFEAEFC60FE86" box="[915,1005,276,298]" pageId="11" type="holotype">holotype</typeStatus>
|
||
except in the following respects: Color and pattern similar except pronotum with only 1 small, black spot behind each anterior angle, elytra with transverse band on disc reduced a spot, and swelling of elytral margin darkened.
|
||
<emphasis id="BEC68339B047A765FC8FFEDFFC1CFED6" box="[850,913,357,378]" italics="true" pageId="11">Elytra</emphasis>
|
||
: Surface with minute setae absent from near apex. Epipleuron (in ventral view) gradually tapering towards apex, abruptly narrowed at level of sternite 1. Lateral margin slightly swollen at level of sternites 1–2.
|
||
<taxonomicName id="4BB224A8B047A765FDAAFE0CFCBBFE60" authority=": Surface" authorityName="Surface" box="[631,822,438,460]" class="Actinopterygii" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="11" phylum="Chordata" rank="genus">
|
||
<emphasis id="BEC68339B047A765FDAAFE0CFD5BFE67" box="[631,726,438,459]" italics="true" pageId="11">Pygidium</emphasis>
|
||
: Surface
|
||
</taxonomicName>
|
||
shiny, punctate; punctures minute and moderately large mixed, moderate in density on disc, dense on sides, setigerous; setae sparse on sides, minute, tawny, absent elsewhere. In lateral view, surface weakly convex.
|
||
<emphasis id="BEC68339B047A765FD92FDBDFDF2FDB0" box="[591,639,519,540]" italics="true" pageId="11">Legs</emphasis>
|
||
: Protarsus simple, not enlarged. Metatarsus subequal in length to metatibia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B047A765FF1AFD87FD5CFCCE" pageId="11" type="description">
|
||
<paragraph id="8C0D5F2BB047A765FF1AFD87FC1CFD13" blockId="11.[175,1005,194,1407]" pageId="11">
|
||
<emphasis id="BEC68339B047A765FF1AFD87FEA0FDFF" bold="true" box="[199,301,573,595]" pageId="11">Variation.</emphasis>
|
||
Males (
|
||
<specimenCount id="9AB494A2B047A765FE54FD87FD85FDFF" box="[393,520,573,595]" pageId="11" type="generic" typeStatus="paratypes">4 paratypes</specimenCount>
|
||
). Length 12.3–14.0 mm; width across humeri
|
||
<quantity id="4B4AF2CEB047A765FF72FDE2FEA6FDC2" box="[175,299,600,622]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.4" metricValueMin="7.1" pageId="11" unit="mm" value="7.25" valueMax="7.4" valueMin="7.1">7.1–7.4 mm</quantity>
|
||
. The
|
||
<typeStatus id="5309E189B047A765FEBAFDE2FE47FDC2" box="[359,458,600,622]" pageId="11" type="paratype">paratypes</typeStatus>
|
||
do not differ significantly from the
|
||
<typeStatus id="5309E189B047A765FC94FDE2FC2EFDC2" box="[841,931,600,622]" pageId="11" type="holotype">holotype</typeStatus>
|
||
except for variation in the number of spots on the pronotum: 1 as
|
||
<typeStatus id="5309E189B047A765FCF8FDC9FC0CFD25" box="[805,897,627,649]" pageId="11" type="holotype">holotype</typeStatus>
|
||
, 1 lacking posterior 2 spots, and 2 with 6 spots in anterior row (
|
||
<figureCitation id="148943AEB047A765FD0FFD34FCCFFD08" box="[722,834,654,676]" captionStart="Figs" captionStartId="10.[167,206,1170,1189]" captionTargetBox="[146,975,194,1148]" captionTargetId="figure-136@10.[146,976,194,1148]" captionTargetPageId="10" captionText="Figs. 17–20. Cyclocephala compacta: 17) dorsal view of holotype; 18–19) parameres" figureDoi="http://doi.org/10.5281/zenodo.5369975" httpUri="https://zenodo.org/record/5369975/files/figure.png" pageId="11">Fig. 20a–c</figureCitation>
|
||
). All of the male
|
||
<typeStatus id="5309E189B047A765FF72FD13FE9FFD13" box="[175,274,681,703]" pageId="11" type="paratype">paratypes</typeStatus>
|
||
have a black, subtriangular mark on the base of the clypeus.
|
||
</paragraph>
|
||
<paragraph id="8C0D5F2BB047A765FF1AFD7FFD5CFCCE" blockId="11.[175,1005,194,1407]" pageId="11">
|
||
Females (
|
||
<specimenCount id="9AB494A2B047A765FEEEFD7FFE3BFD77" box="[307,438,709,731]" pageId="11" type="generic" typeStatus="paratypes">4 paratypes</specimenCount>
|
||
). Length 11.0–13.0 mm; width across humeri
|
||
<quantity id="4B4AF2CEB047A765FC1FFD7FFE8CFD5A" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="7.4" metricValueMin="6.6" pageId="11" unit="mm" value="7.0" valueMax="7.4" valueMin="6.6">6.6– 7.4 mm</quantity>
|
||
. The female
|
||
<typeStatus id="5309E189B047A765FE4AFD5AFE77FD5A" box="[407,506,736,758]" pageId="11" type="paratype">paratypes</typeStatus>
|
||
do not differ significantly from the
|
||
<typeStatus id="5309E189B047A765FC47FD5AFC61FD5A" box="[922,1004,736,758]" pageId="11" type="allotype">allotype</typeStatus>
|
||
except for variation in the number of spots on the pronotum and elytra:
|
||
<specimenCount id="9AB494A2B047A765FC3DFD41FE98FC80" pageId="11" type="generic">2 specimens</specimenCount>
|
||
with 4 spots in transverse row on pronotum and
|
||
<specimenCount id="9AB494A2B047A765FCFBFCACFC2CFC80" box="[806,929,790,812]" pageId="11" type="generic">2 specimens</specimenCount>
|
||
with 4 spots in transverse row on pronotum followed by 2 additional spots in second row. Two specimens with 2 discal spots instead of 1.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B047A765FF1AFCDDFC60FC34" pageId="11" type="etymology">
|
||
<paragraph id="8C0D5F2BB047A765FF1AFCDDFC60FC34" blockId="11.[175,1005,194,1407]" pageId="11">
|
||
<emphasis id="BEC68339B047A765FF1AFCDDFEB7FCD1" bold="true" box="[199,314,871,893]" pageId="11">Etymology.</emphasis>
|
||
The name is derived from the Latin
|
||
<emphasis id="BEC68339B047A765FD19FCD2FCA6FCD1" box="[708,811,872,893]" italics="true" pageId="11">compactus</emphasis>
|
||
, meaning thick or compact, in reference to the relatively compact and stout body form of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B047A765FF1AFC24FDE2FB97" pageId="11" type="distribution">
|
||
<paragraph id="8C0D5F2BB047A765FF1AFC24FDE2FB97" blockId="11.[175,1005,194,1407]" pageId="11">
|
||
<emphasis id="BEC68339B047A765FF1AFC24FECBFC18" bold="true" box="[199,326,926,948]" pageId="11">Distribution.</emphasis>
|
||
<taxonomicName id="4BB224A8B047A765FE90FC24FDB5FC1F" authorityName="Ratcliffe" authorityYear="2008" box="[333,568,926,947]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="compacta">
|
||
<emphasis id="BEC68339B047A765FE90FC24FDB5FC1F" box="[333,568,926,947]" italics="true" pageId="11">Cyclocephala compacta</emphasis>
|
||
</taxonomicName>
|
||
is known from the Schmidt farm south of Ariquemes,
|
||
<collectingRegion id="4E7691C9B047A765FEF6FC02FE19FC62" box="[299,404,952,974]" country="Brazil" name="Rondonia" pageId="11">Rondônia</collectingRegion>
|
||
,
|
||
<collectingCountry id="F4A51FBBB047A765FE7DFC03FE6DFC63" box="[416,480,953,975]" name="Brazil" pageId="11">Brazil</collectingCountry>
|
||
. This area has been largely cut over for agriculture except for the Schmidt farm, which is now a relict island of biodiversity. Emmel and Austin (1990) reported this area had the highest species diversity of butterflies in the world, and a number of new species of scarabs (
|
||
<bibRefCitation id="E82322DAB047A765FD37FBB0FCF0FB8C" author="Ratcliffe" box="[746,893,1034,1056]" pageId="11" pagination="179 - 190" refId="ref10608" refString="Ratcliffe, B. C. 1992. New species and country records of Brazilian Cyclocephala (Coleoptera: Scarabaeidae: Dynastinae). Tidschrift voor Entomologie 136: 179 - 190." type="journal article" year="1992">Ratcliffe 1992</bibRefCitation>
|
||
) have also been described from this single, rich locale.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4A80CA0B047A765FF1AFBFAFDDBFAD3" pageId="11" type="diagnosis">
|
||
<paragraph id="8C0D5F2BB047A765FF1AFBFAFDDBFAD3" blockId="11.[175,1005,194,1407]" pageId="11">
|
||
<emphasis id="BEC68339B047A765FF1AFBFAFEA2FBFA" bold="true" box="[199,303,1088,1110]" pageId="11">Diagnosis.</emphasis>
|
||
Males of
|
||
<taxonomicName id="4BB224A8B047A765FE43FBFBFD93FBFA" authorityName="Ratcliffe" authorityYear="2008" box="[414,542,1089,1110]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="compacta">
|
||
<emphasis id="BEC68339B047A765FE43FBFBFD93FBFA" box="[414,542,1089,1110]" italics="true" pageId="11">C. compacta</emphasis>
|
||
</taxonomicName>
|
||
with six spots on the pronotum will key to couplet
|
||
<quantity id="4B4AF2CEB047A765FEDEFBE1FEC9FBDD" box="[259,324,1115,1137]" metricMagnitude="0" metricUnit="m" metricValue="3.3019999999999996" pageId="11" unit="in" value="130.0">130 in</quantity>
|
||
<bibRefCitation id="E82322DAB047A765FE90FBE1FE61FBDD" author="Endrodi" box="[333,492,1115,1137]" pageId="11" refId="ref10514" refString="Endrodi, S. 1985. The Dynastinae of the World. W. Junk Publishers, Dordrecht. 800 pp." type="book" year="1985">Endrödi (1985)</bibRefCitation>
|
||
, which is
|
||
<taxonomicName id="4BB224A8B047A765FD85FBE6FC60FBDD" authority="Mannerheim." authorityName="Mannerheim." box="[600,1005,1115,1137]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="quatuordecempunctata">
|
||
<emphasis id="BEC68339B047A765FD85FBE6FCD4FBDD" box="[600,857,1116,1137]" italics="true" pageId="11">C. quatuordecempunctata</emphasis>
|
||
Mannerheim.
|
||
</taxonomicName>
|
||
<taxonomicName id="4BB224A8B047A765FF72FBCDFD91FB20" box="[175,540,1143,1164]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="quatuordecempunctata">
|
||
<emphasis id="BEC68339B047A765FF72FBCDFD91FB20" box="[175,540,1143,1164]" italics="true" pageId="11">Cyclocephala quatuordecempunctata</emphasis>
|
||
</taxonomicName>
|
||
, however, has elytral setae, no pygidial spots, and parameres different from those of
|
||
<taxonomicName id="4BB224A8B047A765FD9CFB28FD32FB0B" authorityName="Ratcliffe" authorityYear="2008" box="[577,703,1170,1191]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="compacta">
|
||
<emphasis id="BEC68339B047A765FD9CFB28FD32FB0B" box="[577,703,1170,1191]" italics="true" pageId="11">C. compacta</emphasis>
|
||
</taxonomicName>
|
||
. Specimens with only two or four spots on the pronotum do not go through the key. This kind of variation illustrates the difficulty in constructing a key that relies on the numbers of spots on the pronotum or elytra; specimens will not properly key if only one character state is mentioned. On the other hand, Mannerheim probably did not have a series that showed this variation nor possibly did Endrödi when he created the key. The parameres of
|
||
<taxonomicName id="4BB224A8B047A765FE66FA8EFDCDFAE5" authorityName="Ratcliffe" authorityYear="2008" box="[443,576,1332,1353]" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="compacta">
|
||
<emphasis id="BEC68339B047A765FE66FA8EFDCDFAE5" box="[443,576,1332,1353]" italics="true" pageId="11">C. compacta</emphasis>
|
||
</taxonomicName>
|
||
are somewhat similar to those of
|
||
<taxonomicName id="4BB224A8B047A765FC08FA8EFE1FFAC8" authority="Endrodi" authorityName="Endrodi" authorityYear="1975" class="Insecta" family="Dynastidae" genus="Cyclocephala" kingdom="Animalia" order="Coleoptera" pageId="11" phylum="Arthropoda" rank="species" species="howdenannae">
|
||
<emphasis id="BEC68339B047A765FC08FA8EFEBEFAC8" italics="true" pageId="11">C. howdenannae</emphasis>
|
||
Endrödi
|
||
</taxonomicName>
|
||
, but that species has an emarginate clypeus, no basal bead on the pronotum, and is larger (
|
||
<quantity id="4B4AF2CEB047A765FE22FAD3FDC6FAD3" box="[511,587,1385,1407]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="11" unit="mm" value="18.0">18 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |