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<document id="1E973B004D1197BD522605E39F03F83F" ID-DOI="10.1080/14772019.2016.1151958" ID-ISSN="1478-0941" ID-PMC="PMC5221741" ID-PubMed="28077930" ID-Zenodo-Dep="10883098" ID-ZooBank="A30E5364-0003-4467-B902-43A41AD456CC" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1711483431746" checkinUser="felipe" docAuthor="Schwarzhans, Werner, Mors, Thomas, Engelbrecht, Andrea, Reguero, Marcelo &amp; Kriwet, Jurgen" docDate="2017" docId="041B87CAFFAAFFFED61FFD2CE0B1D961" docLanguage="en" docName="JSystPalaeontol.15.2.147-170.pdf" docOrigin="Journal of Systematic Palaeontology 15 (2)" docSource="http://dx.doi.org/10.1080/14772019.2016.1151958" docStyle="DocumentStyle:602474738E3D100F0F13788B53F61545.3:JSystPalaeontol.2016-.journal_article.open" docStyleId="602474738E3D100F0F13788B53F61545" docStyleName="JSystPalaeontol.2016-.journal_article.open" docStyleVersion="3" docTitle="Notoberyx cionei Schwarzhans &amp; Mors &amp; Engelbrecht &amp; Reguero &amp; Kriwet 2017, sp. nov." docType="treatment" docVersion="3" lastPageNumber="160" masterDocId="F822FFB2FFA4FFF0D753FFE4E35DDA07" masterDocTitle="Before the freeze: otoliths from the Eocene of Seymour Island, Antarctica, reveal dominance of gadiform fishes (Teleostei)" masterLastPageNumber="170" masterPageNumber="147" pageNumber="160" updateTime="1711973403844" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B22C5FDA4D3341DAFA47A313795DD74B">Before the freeze: otoliths from the Eocene of Seymour Island, Antarctica, reveal dominance of gadiform fishes (Teleostei)</mods:title>
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<mods:namePart id="974A0C543CF4FFE66AF80A20B3407727">Schwarzhans, Werner</mods:namePart>
<mods:affiliation id="43D52E8BCD115491A1171795793BCC05">Ahrensburger Weg 103, D- 22359 Hamburg, Germany; &amp; Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, DK- 2100 Copenhagen, Denmark;</mods:affiliation>
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<mods:namePart id="8EB2D9A825D9B83E1645184A03B73924">Mors, Thomas</mods:namePart>
<mods:affiliation id="FD8CA9F48CC654EAA8DCD780C154056A">Swedish Museum of Natural History, Department of Palaeobiology, P. O. Box 5007, SE- 10405 Stockholm, Sweden;</mods:affiliation>
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<mods:namePart id="820DB4BC09DA4758FF1C4643E4F588E5">Engelbrecht, Andrea</mods:namePart>
<mods:affiliation id="1A92F94BBFA1785E86E70087CF74A498">University of Vienna, Department of Palaeontology, Althanstrasse 14, 1090 Vienna, Austria;</mods:affiliation>
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<mods:namePart id="0FE92C9E87E00B4D96BED06DD2DB0232">Reguero, Marcelo</mods:namePart>
<mods:affiliation id="F82DA3EB7C72F7A12F8B9166D776021E">Museo de La Plata, Division ́ Paleontolog ́ ıa de Vertebrados, Paseo del Bosque s / n, B 1900 FWA La Plata, Argentina</mods:affiliation>
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<mods:namePart id="9A8D9349782187DBA8DBF6F2E4D6DF14">Kriwet, Jurgen</mods:namePart>
<mods:affiliation id="9AD4B97CC087374C83151324DE47B5EC">University of Vienna, Department of Palaeontology, Althanstrasse 14, 1090 Vienna, Austria;</mods:affiliation>
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<heading id="D74581B0FFAAFFFED61FFD2CE13CD8D8" box="[332,609,712,735]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="160" reason="8">
<taxonomicName id="4BB24D5FFFAAFFFED61FFD2CE15AD8D8" authority="Schwarzhans &amp; Mors &amp; Engelbrecht &amp; Reguero &amp; Kriwet, 2017" authorityName="Schwarzhans &amp; Mors &amp; Engelbrecht &amp; Reguero &amp; Kriwet" authorityYear="2017" box="[332,519,712,735]" family="Trachichthyidae" genus="Notoberyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Beryciformes" pageId="14" pageNumber="160" phylum="Chordata" rank="species" species="cionei" status="sp. nov.">
<emphasis id="BEC6EACEFFAAFFFED61FFD2CE15AD8D8" bold="true" box="[332,519,712,735]" italics="true" pageId="14" pageNumber="160">Notoberyx cionei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5F557B5FFAAFFFED55EFD2CE13CD8D8" box="[525,609,712,735]" pageId="14" pageNumber="160" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="C4A86557FFAAFFFED62AFD0CE169D8F8" box="[377,564,744,767]" pageId="14" pageNumber="160" type="description">
<paragraph id="8C0D36DCFFAAFFFED62AFD0CE169D8F8" blockId="14.[332,609,712,767]" box="[377,564,744,767]" pageId="14" pageNumber="160">
(
<figureCitation id="14892A59FFAAFFFED6D2FD0CE2A2D8F8" box="[385,511,744,767]" captionStart="Figure 5" captionStartId="13.[159,227,1403,1424]" captionTargetBox="[175,1452,183,1370]" captionTargetId="figure-122@13.[175,1452,183,1371]" captionTargetPageId="13" captionText="Figure 5. Drawings of Eocene otoliths from Seymour Island. A—C, Hoplobrotula? antipoda sp. nov.; A, B, holotype, NRM-PZ P.15984, mirror imaged; A, inner face; B, ventral view; C, paratype, NRM-PZ P.15985, inner face. D—G, Notoberyx cionei gen. nov., sp. nov.; D—F, holotype, NRM-PZ P.15987; D, anterior view; E, inner face; F, ventral view; G, paratype, NRM-PZ P.15988, inner face. H, Centroberyx sp., NRM-PZ P.15986, mirror imaged, inner face. I, J, Acanthopterygii indet., NRM-PZ P.15990, mirror imaged; I, inner face, J, ventral view. K, L, Percoidei indet., NRM-PZ P.15992, mirror imaged; K, inner face; L, ventral view. M, N, Haemulidae? indet., NRM-PZ P.15993, mirror imaged; M, inner face; N, ventral view. O, P, Sparidae? indet., NRM-PZ P.15994, mirror imaged; O, inner face; P, ventral view. Q, R, Cepola anderssoni sp. nov., holotype, NRM-PZ P.15996, mirror imaged; Q, inner face; R, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10883112" httpUri="https://zenodo.org/record/10883112/files/figure.png" pageId="14" pageNumber="160">Figs 5D—G</figureCitation>
,
<figureCitation id="14892A59FFAAFFFED558FD0CE171D8F8" box="[523,556,744,767]" captionStart="Figure 6" captionStartId="15.[159,227,1815,1836]" captionTargetBox="[216,1409,183,1783]" captionTargetId="figure-1@15.[216,1410,183,1783]" captionTargetPageId="15" captionText="Figure 6. Eocene otoliths from Seymour Island. A, Argentina antarctica sp. nov., holotype, NRM-PZ P.15964, mirror imaged, inner face. B, C, Diaphus? marambionis sp. nov., holotype, NRM-PZ P.15966; B, inner face; C, ventral view. D—F, Macruronus eastmani sp. nov., holotype, NRM-PZ P.15970, mirror imaged; D, inner face; E, ventral view; F, outer face. G—I, Palimphemus seymourensis sp. nov., holotype, NRM-PZ P.15973, mirror imaged; G, inner face; H, ventral view; I, outer face. J—L, Coelorinchus nordenskjoeldi sp. nov., holotype, NRM-PZ P.15978; J, inner face; K, ventral view; L, outer face. M, N, Coelorinchus balushkini sp. nov., holotype, NRM-PZ P.15976; M, inner face; N, ventral view. O, P, Hoplobrotula? antipoda sp. nov., holotype, NRM-PZ P.15984, mirror imaged; O, inner face; P, ventral view. Q, Notoberyx cionei gen. nov., sp. nov.; holotype, NRM-PZ P.15987, inner face. R, Cepola anderssoni sp. nov., holotype, NRM-PZ P.15996, mirror imaged, inner face." figureDoi="http://doi.org/10.5281/zenodo.10883114" httpUri="https://zenodo.org/record/10883114/files/figure.png" pageId="14" pageNumber="160">6O</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C4A86557FFAAFFFED7CCFCCFE194D9F2" pageId="14" pageNumber="160" type="materials_examined">
<paragraph id="8C0D36DCFFAAFFFED7CCFCCFE20FD985" blockId="14.[159,781,811,898]" pageId="14" pageNumber="160">
<emphasis id="BEC6EACEFFAAFFFED7CCFCCFE257D945" bold="true" box="[159,266,811,834]" pageId="14" pageNumber="160">Material.</emphasis>
<materialsCitation id="3CDA3C81FFAAFFFED64BFCCFE391D965" pageId="14" pageNumber="160" specimenCode="NRM-PZ P.15987" specimenCount="1" typeStatus="holotype">
<typeStatus id="5309887EFFAAFFFED64BFCCFE2DDD945" box="[280,384,811,834]" pageId="14" pageNumber="160" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DC149EA7FFAAFFFED6C5FCCFE13BD945" box="[406,614,811,834]" pageId="14" pageNumber="160">NRM-PZ P.15987</specimenCode>
(
<figureCitation id="14892A59FFAAFFFED5D1FCCFE054D945" box="[642,777,811,834]" captionStart="Figure 5" captionStartId="13.[159,227,1403,1424]" captionTargetBox="[175,1452,183,1370]" captionTargetId="figure-122@13.[175,1452,183,1371]" captionTargetPageId="13" captionText="Figure 5. Drawings of Eocene otoliths from Seymour Island. A—C, Hoplobrotula? antipoda sp. nov.; A, B, holotype, NRM-PZ P.15984, mirror imaged; A, inner face; B, ventral view; C, paratype, NRM-PZ P.15985, inner face. D—G, Notoberyx cionei gen. nov., sp. nov.; D—F, holotype, NRM-PZ P.15987; D, anterior view; E, inner face; F, ventral view; G, paratype, NRM-PZ P.15988, inner face. H, Centroberyx sp., NRM-PZ P.15986, mirror imaged, inner face. I, J, Acanthopterygii indet., NRM-PZ P.15990, mirror imaged; I, inner face, J, ventral view. K, L, Percoidei indet., NRM-PZ P.15992, mirror imaged; K, inner face; L, ventral view. M, N, Haemulidae? indet., NRM-PZ P.15993, mirror imaged; M, inner face; N, ventral view. O, P, Sparidae? indet., NRM-PZ P.15994, mirror imaged; O, inner face; P, ventral view. Q, R, Cepola anderssoni sp. nov., holotype, NRM-PZ P.15996, mirror imaged; Q, inner face; R, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10883112" httpUri="https://zenodo.org/record/10883112/files/figure.png" pageId="14" pageNumber="160">Figs 5D—F</figureCitation>
,
<figureCitation id="14892A59FFAAFFFED7CCFCAFE39ED965" box="[159,195,843,866]" captionStart="Figure 6" captionStartId="15.[159,227,1815,1836]" captionTargetBox="[216,1409,183,1783]" captionTargetId="figure-1@15.[216,1410,183,1783]" captionTargetPageId="15" captionText="Figure 6. Eocene otoliths from Seymour Island. A, Argentina antarctica sp. nov., holotype, NRM-PZ P.15964, mirror imaged, inner face. B, C, Diaphus? marambionis sp. nov., holotype, NRM-PZ P.15966; B, inner face; C, ventral view. D—F, Macruronus eastmani sp. nov., holotype, NRM-PZ P.15970, mirror imaged; D, inner face; E, ventral view; F, outer face. G—I, Palimphemus seymourensis sp. nov., holotype, NRM-PZ P.15973, mirror imaged; G, inner face; H, ventral view; I, outer face. J—L, Coelorinchus nordenskjoeldi sp. nov., holotype, NRM-PZ P.15978; J, inner face; K, ventral view; L, outer face. M, N, Coelorinchus balushkini sp. nov., holotype, NRM-PZ P.15976; M, inner face; N, ventral view. O, P, Hoplobrotula? antipoda sp. nov., holotype, NRM-PZ P.15984, mirror imaged; O, inner face; P, ventral view. Q, Notoberyx cionei gen. nov., sp. nov.; holotype, NRM-PZ P.15987, inner face. R, Cepola anderssoni sp. nov., holotype, NRM-PZ P.15996, mirror imaged, inner face." figureDoi="http://doi.org/10.5281/zenodo.10883114" httpUri="https://zenodo.org/record/10883114/files/figure.png" pageId="14" pageNumber="160">6O</figureCitation>
)
</materialsCitation>
.
<typeStatus id="5309887EFFAAFFFED7B0FCAFE212D965" box="[227,335,843,866]" pageId="14" pageNumber="160" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9AB4FD55FFAAFFFED636FCAFE14FD965" box="[357,530,843,866]" count="2" pageId="14" pageNumber="160" type="generic">two specimens</specimenCount>
, NRM-PZ P.15988— 15989 (
<figureCitation id="14892A59FFAAFFFED7A2FC8FE21BD985" box="[241,326,875,898]" captionStart="Figure 5" captionStartId="13.[159,227,1403,1424]" captionTargetBox="[175,1452,183,1370]" captionTargetId="figure-122@13.[175,1452,183,1371]" captionTargetPageId="13" captionText="Figure 5. Drawings of Eocene otoliths from Seymour Island. A—C, Hoplobrotula? antipoda sp. nov.; A, B, holotype, NRM-PZ P.15984, mirror imaged; A, inner face; B, ventral view; C, paratype, NRM-PZ P.15985, inner face. D—G, Notoberyx cionei gen. nov., sp. nov.; D—F, holotype, NRM-PZ P.15987; D, anterior view; E, inner face; F, ventral view; G, paratype, NRM-PZ P.15988, inner face. H, Centroberyx sp., NRM-PZ P.15986, mirror imaged, inner face. I, J, Acanthopterygii indet., NRM-PZ P.15990, mirror imaged; I, inner face, J, ventral view. K, L, Percoidei indet., NRM-PZ P.15992, mirror imaged; K, inner face; L, ventral view. M, N, Haemulidae? indet., NRM-PZ P.15993, mirror imaged; M, inner face; N, ventral view. O, P, Sparidae? indet., NRM-PZ P.15994, mirror imaged; O, inner face; P, ventral view. Q, R, Cepola anderssoni sp. nov., holotype, NRM-PZ P.15996, mirror imaged; Q, inner face; R, ventral view." figureDoi="http://doi.org/10.5281/zenodo.10883112" httpUri="https://zenodo.org/record/10883112/files/figure.png" pageId="14" pageNumber="160">Fig. 5G</figureCitation>
).
</paragraph>
<paragraph id="8C0D36DCFFAAFFFED7CCFC7BE194D9F2" blockId="14.[159,781,927,1013]" pageId="14" pageNumber="160">
<emphasis id="BEC6EACEFFAAFFFED7CCFC7BE271D9B1" bold="true" box="[159,300,927,950]" pageId="14" pageNumber="160">Occurrence.</emphasis>
Telm 5 unit;
<taxonomicName id="4BB24D5FFFAAFFFED688FC7BE17FD9B1" authorityName="Scopoli" authorityYear="1777" box="[475,546,927,950]" class="Gastropoda" family="Naticidae" genus="Natica" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="160" phylum="Mollusca" rank="genus">
<emphasis id="BEC6EACEFFAAFFFED688FC7BE17FD9B1" box="[475,546,927,950]" italics="true" pageId="14" pageNumber="160">Natica</emphasis>
</taxonomicName>
horizon,
<taxonomicName id="4BB24D5FFFAAFFFED5CAFC7BE1A4D9B1" box="[665,761,927,950]" class="Bivalvia" family="Cucullaeidae" genus="Cucullea" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Arcida" pageId="14" pageNumber="160" phylum="Mollusca" rank="genus">
<emphasis id="BEC6EACEFFAAFFFED5CAFC7BE1A4D9B1" box="[665,761,927,950]" italics="true" pageId="14" pageNumber="160">Cucullea</emphasis>
</taxonomicName>
I member, La Meseta Formation, late Ypresian, early Eocene. Site IAA 2/95, Seymour Island,
<collectingCountry id="F4A5764CFFAAFFFED501FC3AE198D9F2" box="[594,709,990,1013]" name="Antarctica" pageId="14" pageNumber="160">Antarctica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C4A86557FFAAFFFED7CCFBF6E1A0DE6E" pageId="14" pageNumber="160" type="etymology">
<paragraph id="8C0D36DCFFAAFFFED7CCFBF6E1A0DE6E" blockId="14.[159,781,1042,1129]" pageId="14" pageNumber="160">
<emphasis id="BEC6EACEFFAAFFFED7CCFBF6E27CDE2E" bold="true" box="[159,289,1042,1065]" pageId="14" pageNumber="160">Etymology.</emphasis>
Named in honour of Alberto Cione (La Plata,
<taxonomicName id="4BB24D5FFFAAFFFED7B1FBD6E212DE4E" authorityName="Linnaeus" authorityYear="1758" box="[226,335,1074,1097]" family="Argentinidae" genus="Argentina" kingdom="Animalia" order="Osmeriformes" pageId="14" pageNumber="160" phylum="Chordata" rank="genus">Argentina</taxonomicName>
) for his many contributions to the knowledge of fossil fishes from South America and
<collectingCountry id="F4A5764CFFAAFFFED5D5FBB6E1A4DE6E" box="[646,761,1106,1129]" name="Antarctica" pageId="14" pageNumber="160">Antarctica</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C4A86557FFAAFFFED7CCFB62E211DF1B" pageId="14" pageNumber="160" type="diagnosis">
<paragraph id="8C0D36DCFFAAFFFED7CCFB62E211DF1B" blockId="14.[159,782,1158,1308]" pageId="14" pageNumber="160">
<emphasis id="BEC6EACEFFAAFFFED7CCFB62E249DE9A" bold="true" box="[159,276,1158,1181]" pageId="14" pageNumber="160">Diagnosis.</emphasis>
OL:OH = 0.7—0.75. Dorsal rim high, broad; ventral rim deep, broad. Rostrum short, blunt, about equal length of antirostrum. CaL:OsL = 1.4—1.5. Ostium narrow, only slightly wider than cauda; cauda distinctly turned upwards.
</paragraph>
</subSubSection>
<subSubSection id="C4A86557FFAAFFFED7CCFADDE637DB13" pageId="14" pageNumber="160" type="description">
<paragraph id="8C0D36DCFFAAFFFED7CCFADDE10DDCA8" blockId="14.[159,782,1337,1966]" pageId="14" pageNumber="160">
<emphasis id="BEC6EACEFFAAFFFED7CCFADDE277DF57" bold="true" box="[159,298,1337,1360]" pageId="14" pageNumber="160">Description.</emphasis>
Large, robust, high-bodied, oval otoliths up to about
<quantity id="4B4A9B39FFAAFFFED653FABDE21DDF77" box="[256,320,1369,1392]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="160" unit="mm" value="6.0">6 mm</quantity>
in length (
<typeStatus id="5309887EFFAAFFFED6EAFABDE148DF77" box="[441,533,1369,1392]" pageId="14" pageNumber="160" type="holotype">holotype</typeStatus>
<quantity id="4B4A9B39FFAAFFFED54DFABDE12CDF77" box="[542,625,1369,1392]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="14" pageNumber="160" unit="mm" value="5.8">5.8 mm</quantity>
long). OL:OT = 3.0. Dorsal rim high, much expanded and broad, undulating; rounded pre- and postdorsal angles close to anterior and posterior limits of dorsal rim, postdorsal angle usually somewhat pronounced. Ventral rim deep, nearly as broad and expanded as dorsal rim, but more gently curved and smooth. Anterior rim blunt, nearly vertically cut, with very short and massive rostrum not reaching beyond length of antirostrum; with broad, shallow excisura in between. Posterior rim likewise blunt and nearly vertically cut, its tip shifted dorsally above level of caudal tip.
</paragraph>
<paragraph id="8C0D36DCFFAAFFFED7E9F95CE637DB13" blockId="14.[159,782,1337,1966]" lastBlockId="14.[845,1466,189,276]" pageId="14" pageNumber="160">Inner face bent along the horizontal axis, nearly straight in vertical direction. Sulcus slightly supramedian, wide, moderately deep. Ostium anteriorly opened, slightly deepened, distinctly shorter than cauda and only slightly wider. OsH:CaH = 1.15—1.3. Cauda long, bent upwards, terminating close to posterior rim of otolith, its colliculum with well-marked ventral rim. Dorsal depression wide, large, well defined towards sulcus; dorsal field with some short radial furrows near the dorsal margin. Ventral field smooth, with distinct ventral furrow very close to ventral rim of otolith. Outer face with broad central umbo.</paragraph>
</subSubSection>
<subSubSection id="C4A86557FFAAFFFED41EFED5E0B1D961" pageId="14" pageNumber="160" type="discussion">
<paragraph id="8C0D36DCFFAAFFFED41EFED5E6C0D860" blockId="14.[845,1467,305,870]" pageId="14" pageNumber="160">
<emphasis id="BEC6EACEFFAAFFFED41EFED5E0E6DB4F" bold="true" box="[845,955,305,328]" pageId="14" pageNumber="160">Remarks.</emphasis>
These highly diagnostic otoliths differs from the only congener,
<taxonomicName id="4BB24D5FFFAAFFFED371FEB5E7C6DB6F" baseAuthorityName="Schwarzhans" baseAuthorityYear="2007" box="[1058,1179,337,360]" family="Trachichthyidae" genus="Notoberyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Beryciformes" pageId="14" pageNumber="160" phylum="Chordata" rank="species" species="madseni">
<emphasis id="BEC6EACEFFAAFFFED371FEB5E7C6DB6F" box="[1058,1179,337,360]" italics="true" pageId="14" pageNumber="160">N. madseni</emphasis>
</taxonomicName>
from the Northern Hemisphere, in being even more compressed (OL:OH = 0.7—0.75 vs. 0.8—0.85), the ostium being slightly wider than the cauda (vs. not being wider at all) and the absence of the peculiar deepening of the rear portion of the cauda as observed in
<taxonomicName id="4BB24D5FFFAAFFFED4BEFE15E738D80F" baseAuthorityName="Schwarzhans" baseAuthorityYear="2007" box="[1005,1125,497,520]" family="Trachichthyidae" genus="Notoberyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Beryciformes" pageId="14" pageNumber="160" phylum="Chordata" rank="species" species="madseni">
<emphasis id="BEC6EACEFFAAFFFED4BEFE15E738D80F" box="[1005,1125,497,520]" italics="true" pageId="14" pageNumber="160">N. madseni</emphasis>
</taxonomicName>
. The latter character could possibly support placing
<taxonomicName id="4BB24D5FFFAAFFFED36DFDF5E7E7D82F" baseAuthorityName="Schwarzhans" baseAuthorityYear="2007" box="[1086,1210,529,552]" family="Trachichthyidae" genus="Notoberyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Beryciformes" pageId="14" pageNumber="160" phylum="Chordata" rank="species" species="madseni">
<emphasis id="BEC6EACEFFAAFFFED36DFDF5E7E7D82F" box="[1086,1210,529,552]" italics="true" pageId="14" pageNumber="160">N. madseni</emphasis>
</taxonomicName>
in a genus of its own, once more species with this morphological pattern have become known to support a further taxonomic division.
</paragraph>
<paragraph id="8C0D36DCFFAAFFFED434FD94E0B1D961" blockId="14.[845,1467,305,870]" pageId="14" pageNumber="160">
The two species here attributed to
<taxonomicName id="4BB24D5FFFAAFFFED3A2FD94E63DD880" authorityName="Schwarzhans &amp; Mors &amp; Engelbrecht &amp; Reguero &amp; Kriwet" authorityYear="2017" box="[1265,1376,624,647]" family="Trachichthyidae" genus="Notoberyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Beryciformes" pageId="14" pageNumber="160" phylum="Chordata" rank="genus">
<emphasis id="BEC6EACEFFAAFFFED3A2FD94E63DD880" box="[1265,1376,624,647]" italics="true" pageId="14" pageNumber="160">Notoberyx</emphasis>
</taxonomicName>
show a bipolar temperate distribution with one species each in the early Eocene of
<collectingCountry id="F4A5764CFFAAFFFED354FD54E72BD8C0" box="[1031,1142,688,711]" name="Antarctica" pageId="14" pageNumber="160">Antarctica</collectingCountry>
and
<collectingCountry id="F4A5764CFFAAFFFED3E1FD54E645D8C0" box="[1202,1304,688,711]" name="Denmark" pageId="14" pageNumber="160">Denmark</collectingCountry>
, subject, however, to further finds of these highly diagnostic otoliths. The Danish Eocene otoliths of
<taxonomicName id="4BB24D5FFFAAFFFED3E3FD14E670D900" baseAuthorityName="Schwarzhans" baseAuthorityYear="2007" box="[1200,1325,752,775]" family="Trachichthyidae" genus="Notoberyx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Beryciformes" pageId="14" pageNumber="160" phylum="Chordata" rank="species" species="madseni">
<emphasis id="BEC6EACEFFAAFFFED3E3FD14E670D900" box="[1200,1325,752,775]" italics="true" pageId="14" pageNumber="160">N. madseni</emphasis>
</taxonomicName>
were exclusively found in burrow concretions of stomatopods, apparently indicating that these fishes represented their preferred prey.
</paragraph>
</subSubSection>
</treatment>
</document>