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<document id="8E8A24484F90ADBDF5A32054881CC24A" ID="10.11646/zootaxa.4147.4.4" ID-DOI="10.11646/zootaxa.4147.4.4" ID-GBIF-Dataset="db022185-3c2d-4978-8951-d0c23ccdb47a" ID-ISSN="1175-5326" ID-Zenodo-Dep="264988" ID-ZooBank="8B25DA4D-0B7A-4568-8E7C-642EB932F2D0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1470379029729" checkinUser="plazi" docAuthor="Arango, Stephania Sandoval, Obando, Ranulfo González &amp; Aldrete, Alfonso Neri García" docDate="2016" docId="03FFFA7A6D07FFB5269A2385FC0D57A8" docLanguage="en" docName="zootaxa.4147.4.4.pdf" docOrigin="Zootaxa 4147 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Graphocaecilius turneri Arango, Obando &amp; Aldrete, 2016, n. sp." docType="treatment" docVersion="7" lastPageNumber="429" masterDocId="FFC682026D0FFFBD260D2312FFB1534F" masterDocTitle="New species of Graphocaecilius Enderlein (Psocodea: Psocoptera: Lachesillidae) from Colombia" masterLastPageNumber="432" masterPageNumber="421" pageNumber="429" updateTime="1698367001650" updateUser="plazi">
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<mods:title id="EB73DD027384F947F3811108614708AC">New species of Graphocaecilius Enderlein (Psocodea: Psocoptera: Lachesillidae) from Colombia</mods:title>
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<mods:namePart id="45E55D291ECFB8E181A773DE9A7CE435">Arango, Stephania Sandoval</mods:namePart>
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<mods:namePart id="27FB39FABB1D3B20B22436FD61BBEB7B">Obando, Ranulfo González</mods:namePart>
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<mods:namePart id="286F4D83EB31E8DD311C44CEABC77E6D">Aldrete, Alfonso Neri García</mods:namePart>
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<mods:date id="FB6430625901E8418B1E55725C023DD1">2016</mods:date>
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<treatment id="03FFFA7A6D07FFB5269A2385FC0D57A8" ID-DOI="http://doi.org/10.5281/zenodo.6085910" ID-GBIF-Taxon="123514465" ID-Zenodo-Dep="6085910" LSID="urn:lsid:plazi:treatment:03FFFA7A6D07FFB5269A2385FC0D57A8" httpUri="http://treatment.plazi.org/id/03FFFA7A6D07FFB5269A2385FC0D57A8" lastPageNumber="429" pageId="8" pageNumber="429">
<subSubSection id="C34C18E76D07FFB5269A2385FE96539C" pageId="8" pageNumber="429" type="nomenclature">
<paragraph id="8BE94B6C6D07FFB5269A2385FDF853FD" blockId="8.[151,585,151,211]" box="[151,585,151,178]" pageId="8" pageNumber="429">
<heading id="D0A1FC006D07FFB5269A2385FDF853FD" bold="true" box="[151,585,151,178]" fontSize="11" level="1" pageId="8" pageNumber="429" reason="1">
<emphasis id="B922977E6D07FFB5269A2385FDF853FD" bold="true" box="[151,585,151,178]" pageId="8" pageNumber="429">
<taxonomicName id="4C5630EF6D07FFB5269A2385FE0B53FE" ID-CoL="6L3CJ" box="[151,442,151,177]" class="Insecta" family="Lachesillidae" genus="Graphocaecilius" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="429" phylum="Arthropoda" rank="species" species="turneri" status="sp. nov.">
<emphasis id="B922977E6D07FFB5269A2385FE0B53FE" bold="true" box="[151,442,151,177]" italics="true" pageId="8" pageNumber="429">Graphocaecilius turneri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2112A056D07FFB527CC238AFDB253FD" box="[449,515,152,178]" pageId="8" pageNumber="429" rank="species">n. sp.</taxonomicNameLabel>
Male
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE94B6C6D07FFB5269A23A8FE96539C" blockId="8.[151,585,151,211]" box="[151,295,186,211]" pageId="8" pageNumber="429">
(
<figureCitation id="136D57E96D07FFB5269223A8FEAF539C" box="[159,286,186,211]" captionStart="FIGURES 21 25" captionStartId="9.[151,264,1781,1804]" captionTargetBox="[247,1374,262,1764]" captionTargetId="figure@9.[225,1376,194,1764]" captionTargetPageId="9" captionText="FIGURES 21 25. Graphocaecilius turneri (Male). 21. Fore- and hind wings. 22. Front view of head. 23. Clunium, paraprocts and epiproct. 24. Phallosome. 25. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264993/files/figure.png" pageId="8" pageNumber="429">Figs 2125</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34C18E76D07FFB5269A2210FEEC5110" pageId="8" pageNumber="429" type="diagnosis">
<paragraph id="8BE94B6C6D07FFB5269A2210FE1C52BC" blockId="8.[151,1437,258,1255]" pageId="8" pageNumber="429">
<emphasis id="B922977E6D07FFB5269A2210FEA05254" bold="true" box="[151,273,258,283]" pageId="8" pageNumber="429">Diagnosis.</emphasis>
Differing from
<taxonomicName id="4C5630EF6D07FFB527C72211FDCA5255" box="[458,635,259,282]" class="Insecta" family="Lachesillidae" genus="Graphocaecilius" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="429" phylum="Arthropoda" rank="species" species="interpretatus">
<emphasis id="B922977E6D07FFB527C72211FDCA5255" box="[458,635,259,282]" italics="true" pageId="8" pageNumber="429">G. interpretatus</emphasis>
</taxonomicName>
,
<emphasis id="B922977E6D07FFB524862211FD5A5255" box="[651,747,259,282]" italics="true" pageId="8" pageNumber="429">G. p i c t u s</emphasis>
, and
<taxonomicName id="4C5630EF6D07FFB5253D2211FC7C5254" box="[816,973,259,283]" class="Insecta" family="Lachesillidae" genus="Graphocaecilius" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="429" phylum="Arthropoda" rank="species" species="trypetoides">
<emphasis id="B922977E6D07FFB5253D2211FC7C5254" box="[816,973,259,283]" italics="true" pageId="8" pageNumber="429">G. trypetoides</emphasis>
</taxonomicName>
, in having the forewings without brown spots. Differing from
<taxonomicName id="4C5630EF6D07FFB527812235FDAC5271" box="[396,541,294,318]" class="Insecta" family="Lachesillidae" genus="Graphocaecilius" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="429" phylum="Arthropoda" rank="species" species="enderleini">
<emphasis id="B922977E6D07FFB527812235FDAC5271" box="[396,541,294,318]" italics="true" pageId="8" pageNumber="429">G. enderleini</emphasis>
</taxonomicName>
, in having the distal half of the pterostigma pigmented and in having the areola postica narrower. Differing fom
<taxonomicName id="4C5630EF6D07FFB527F22259FDDD522C" box="[511,620,331,355]" class="Insecta" family="Lachesillidae" genus="Graphocaecilius" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="429" phylum="Arthropoda" rank="species" species="lacteus">
<emphasis id="B922977E6D07FFB527F22259FDDD522C" box="[511,620,331,355]" italics="true" pageId="8" pageNumber="429">G. lacteus</emphasis>
</taxonomicName>
, in having the distal half of the pterostigma pigmented, and in having the areola postica distinctly narrower. Differing from
<taxonomicName id="4C5630EF6D07FFB524C0227DFCD852C9" box="[717,873,367,390]" class="Insecta" family="Lachesillidae" genus="Graphocaecilius" kingdom="Animalia" order="Psocodea" pageId="8" pageNumber="429" phylum="Arthropoda" rank="species" species="peruvianus">
<emphasis id="B922977E6D07FFB524C0227DFCD852C9" box="[717,873,367,390]" italics="true" pageId="8" pageNumber="429">G. peruvianus</emphasis>
</taxonomicName>
, in having the phallosome anteriorly unstemmed, in the shape of the endophallic sclerites, and in lacking clunial projections over the area of the epiproct and paraprocts. Differing from
<emphasis id="B922977E6D07FFB527CB22A5FD935281" box="[454,546,439,462]" italics="true" pageId="8" pageNumber="429">G. va r t y i</emphasis>
in the shape of the hypandrium, in lacking clunial posterior projections, and in the endophallic sclerites.
</paragraph>
<paragraph id="8BE94B6C6D07FFB526CA22ECFEEC5110" blockId="8.[151,1437,258,1255]" pageId="8" pageNumber="429">
<emphasis id="B922977E6D07FFB526CA22ECFEBB5158" bold="true" box="[199,266,510,535]" pageId="8" pageNumber="429">Color</emphasis>
. Body dark brown, compound eyes black, legs yellow, forewings yellow, veins brown; pterostigma brown yellowish, hindwings hyaline, with a yellowish hue. Tergal lobes of meso- and metathorax dark brown, abdomen creamy.
</paragraph>
</subSubSection>
<subSubSection id="C34C18E76D07FFB526CA2178FE32577C" pageId="8" pageNumber="429" type="description">
<paragraph id="8BE94B6C6D07FFB526CA2178FC535088" blockId="8.[151,1437,258,1255]" pageId="8" pageNumber="429">
<emphasis id="B922977E6D07FFB526CA2178FED051CC" bold="true" box="[199,353,618,643]" pageId="8" pageNumber="429">Morphology.</emphasis>
As in diagnosis plus the following: Head vertex almost straight, compound eyes prominent, upper borders above the level of the vertex, with interommatidial setae (
<figureCitation id="136D57E96D07FFB525DE219CFB9851E8" box="[979,1065,654,679]" captionStart="FIGURES 21 25" captionStartId="9.[151,264,1781,1804]" captionTargetBox="[247,1374,262,1764]" captionTargetId="figure@9.[225,1376,194,1764]" captionTargetPageId="9" captionText="FIGURES 21 25. Graphocaecilius turneri (Male). 21. Fore- and hind wings. 22. Front view of head. 23. Clunium, paraprocts and epiproct. 24. Phallosome. 25. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264993/files/figure.png" pageId="8" pageNumber="429">Fig. 22</figureCitation>
). Forewing L/W: 3.0. Forewing pterostigma elongate, much wider posteriorly, with setae in the membrane; lp/wp: 3.38. Areola postica triangular (
<figureCitation id="136D57E96D07FFB5269221C4FF4951A0" box="[159,248,726,751]" captionStart="FIGURES 21 25" captionStartId="9.[151,264,1781,1804]" captionTargetBox="[247,1374,262,1764]" captionTargetId="figure@9.[225,1376,194,1764]" captionTargetPageId="9" captionText="FIGURES 21 25. Graphocaecilius turneri (Male). 21. Fore- and hind wings. 22. Front view of head. 23. Clunium, paraprocts and epiproct. 24. Phallosome. 25. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264993/files/figure.png" pageId="8" pageNumber="429">Fig. 21</figureCitation>
). Hindwing L/W: 2.59 (
<figureCitation id="136D57E96D07FFB5241421C4FDC051A0" box="[537,625,726,751]" captionStart="FIGURES 21 25" captionStartId="9.[151,264,1781,1804]" captionTargetBox="[247,1374,262,1764]" captionTargetId="figure@9.[225,1376,194,1764]" captionTargetPageId="9" captionText="FIGURES 21 25. Graphocaecilius turneri (Male). 21. Fore- and hind wings. 22. Front view of head. 23. Clunium, paraprocts and epiproct. 24. Phallosome. 25. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264993/files/figure.png" pageId="8" pageNumber="429">Fig. 21</figureCitation>
). Hypandrium broad, setose, posterior projection with postero-lateral corners rounded distally (
<figureCitation id="136D57E96D07FFB527B821E8FDB9505C" box="[437,520,762,787]" captionStart="FIGURES 21 25" captionStartId="9.[151,264,1781,1804]" captionTargetBox="[247,1374,262,1764]" captionTargetId="figure@9.[225,1376,194,1764]" captionTargetPageId="9" captionText="FIGURES 21 25. Graphocaecilius turneri (Male). 21. Fore- and hind wings. 22. Front view of head. 23. Clunium, paraprocts and epiproct. 24. Phallosome. 25. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264993/files/figure.png" pageId="8" pageNumber="429">Fig. 25</figureCitation>
). Phallosome slender, unstemmed anteriorly, three times as long as wide; external parameres slender, wide proximally, narrowing distally; aedeagal arch as illustrated; endophallus with an anterior group of minute denticles and nine posterior sclerites, disposed as illustrated (
<figureCitation id="136D57E96D07FFB525F32050FBE35014" box="[1022,1106,834,859]" captionStart="FIGURES 21 25" captionStartId="9.[151,264,1781,1804]" captionTargetBox="[247,1374,262,1764]" captionTargetId="figure@9.[225,1376,194,1764]" captionTargetPageId="9" captionText="FIGURES 21 25. Graphocaecilius turneri (Male). 21. Fore- and hind wings. 22. Front view of head. 23. Clunium, paraprocts and epiproct. 24. Phallosome. 25. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264993/files/figure.png" pageId="8" pageNumber="429">Fig. 24</figureCitation>
). Posterior border of clunium not projected over the areas of paraprocts and epiproct (
<figureCitation id="136D57E96D07FFB5253F2074FC3A5030" box="[818,907,870,895]" captionStart="FIGURES 21 25" captionStartId="9.[151,264,1781,1804]" captionTargetBox="[247,1374,262,1764]" captionTargetId="figure@9.[225,1376,194,1764]" captionTargetPageId="9" captionText="FIGURES 21 25. Graphocaecilius turneri (Male). 21. Fore- and hind wings. 22. Front view of head. 23. Clunium, paraprocts and epiproct. 24. Phallosome. 25. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264993/files/figure.png" pageId="8" pageNumber="429">Fig. 23</figureCitation>
). Paraprocts (
<figureCitation id="136D57E96D07FFB522202074FB325030" box="[1069,1155,870,895]" captionStart="FIGURES 21 25" captionStartId="9.[151,264,1781,1804]" captionTargetBox="[247,1374,262,1764]" captionTargetId="figure@9.[225,1376,194,1764]" captionTargetPageId="9" captionText="FIGURES 21 25. Graphocaecilius turneri (Male). 21. Fore- and hind wings. 22. Front view of head. 23. Clunium, paraprocts and epiproct. 24. Phallosome. 25. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264993/files/figure.png" pageId="8" pageNumber="429">Fig. 23</figureCitation>
) broad, with few setae, restricted to the latero-posterior areas; sensory fields with 17 trichobothria in basal rosettes. Epiproct (
<figureCitation id="136D57E96D07FFB523352098FA3E50EC" box="[1336,1423,906,931]" captionStart="FIGURES 21 25" captionStartId="9.[151,264,1781,1804]" captionTargetBox="[247,1374,262,1764]" captionTargetId="figure@9.[225,1376,194,1764]" captionTargetPageId="9" captionText="FIGURES 21 25. Graphocaecilius turneri (Male). 21. Fore- and hind wings. 22. Front view of head. 23. Clunium, paraprocts and epiproct. 24. Phallosome. 25. Hypandrium. Scales in mm." httpUri="https://zenodo.org/record/264993/files/figure.png" pageId="8" pageNumber="429">Fig. 23</figureCitation>
), triangular, with rounded apex, and three macrosetae in the posterior border.
</paragraph>
<paragraph id="8BE94B6C6D07FFB526CA20C0FE32577C" blockId="8.[151,1437,258,1255]" pageId="8" pageNumber="429">
<emphasis id="B922977E6D07FFB526CA20C0FEC750A4" bold="true" box="[199,374,978,1003]" pageId="8" pageNumber="429">Measurements</emphasis>
(in microns). FW: 2475, HW: 1750, F: 550, T: 912, t1: 287, t2: 137, ctt1: 16,
<collectingCountry id="F3410BFC6D07FFB522F320C1FA9950A4" box="[1278,1320,979,1003]" name="Mexico" pageId="8" pageNumber="429">Mx</collectingCountry>
4: 170, f1: 490, f2: 335, f3: 290, f4: 220, f5: 160, f6: 145, f7: 130, f8: 120, f9: 110, f10: 105, f11: 110, IO: 300, D: 295, d: 210, IO/d: 1.43, PO: 0.71.
</paragraph>
</subSubSection>
<subSubSection id="C34C18E76D07FFB526CA272CFC0D57A8" pageId="8" pageNumber="429" type="etymology">
<paragraph id="8BE94B6C6D07FFB526CA272CFB885734" blockId="8.[151,1437,258,1255]" pageId="8" pageNumber="429">
<emphasis id="B922977E6D07FFB526CA272CFEFF5718" bold="true" box="[199,334,1086,1111]" pageId="8" pageNumber="429">Etymology.</emphasis>
This species is dedicated to Dr. Bryan D. Turner, formerly of Kings College, London, in recognition to his important contributions to the study of Neotropical Psocoptera.
</paragraph>
<paragraph id="8BE94B6C6D07FFB526CA2794FC0D57A8" blockId="8.[151,1437,258,1255]" pageId="8" pageNumber="429">
<materialsCitation id="3B3E41316D07FFB526CA2794FE53578C" ID-GBIF-Occurrence="1290996069" collectorName="Santiago de Cali" country="Colombia" latitude="3.4919167" location="Valle del Cauca" pageId="8" pageNumber="429" specimenCount="1" stateProvince="Valle del Cauca" typeStatus="holotype">
<emphasis id="B922977E6D07FFB526CA2794FE1557D0" bold="true" box="[199,420,1158,1183]" pageId="8" pageNumber="429">Specimens studied</emphasis>
.
<typeStatus id="54EDF5CE6D07FFB527BE2794FDAA57D0" box="[435,539,1158,1183]" pageId="8" pageNumber="429" type="holotype">Holotype</typeStatus>
male.
<collectingCountry id="F3410BFC6D07FFB524642795FCB357D0" box="[617,770,1159,1183]" name="Colombia" pageId="8" pageNumber="429">COLOMBIA</collectingCountry>
.
<collectingRegion id="4992858E6D07FFB525032794FC7357D0" box="[782,962,1158,1183]" country="Colombia" name="Valle del Cauca" pageId="8" pageNumber="429">Valle del Cauca</collectingRegion>
,
<collectorName id="26A32EBA6D07FFB525C22794FB3D57D0" box="[975,1164,1158,1183]" pageId="8" pageNumber="429">Santiago de Cali</collectorName>
, San Antonio, ca. Finca San Francisco,
<geoCoordinate id="EE622DAB6D07FFB5274427B9FE53578C" box="[329,482,1195,1219]" direction="north" orientation="latitude" pageId="8" pageNumber="429" precision="1" value="3.4919167">3°2930.9N</geoCoordinate>
</materialsCitation>
:
<materialsCitation id="3B3E41316D07FFB527E227B8FC0D57A8" ID-GBIF-Occurrence="1290996060" collectingDate="2012-11-04" collectorName="Santiago de Cali" country="Colombia" elevation="1860" latitude="3.4919167" location="Valle del Cauca" longitude="-76.61917" pageId="8" pageNumber="429" specimenCount="1" specimenCount-male="1" stateProvince="Valle del Cauca" typeStatus="paratype">
<geoCoordinate id="EE622DAB6D07FFB527E227B8FD29578C" box="[495,664,1194,1219]" direction="west" orientation="longitude" pageId="8" pageNumber="429" precision="1" value="-76.61917">76°3709.0W</geoCoordinate>
,
<quantity id="4CAEE6896D07FFB524AA27B8FCB0578D" box="[679,769,1194,1219]" metricMagnitude="3" metricUnit="m" metricValue="1.86" pageId="8" pageNumber="429" unit="m" value="1860.0">
<elevation id="007BAC5F6D07FFB524AA27B8FCB0578D" box="[679,769,1194,1219]" metricMagnitude="3" metricUnit="m" metricValue="1.86" pageId="8" pageNumber="429" unit="m" value="1860.0">1860 m</elevation>
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,
<date id="FFE86DAC6D07FFB5251D27B9FC3F578C" box="[784,910,1195,1219]" pageId="8" pageNumber="429" value="2012-11-04">
<collectingDate id="EFAC94446D07FFB5251D27B9FC3F578C" box="[784,910,1195,1219]" pageId="8" pageNumber="429" value="2012-11-04">4. XI.2012</collectingDate>
</date>
, MUSENUV slide code 26169, R. González.
<typeStatus id="54EDF5CE6D07FFB5269A27DDFF4F57A8" box="[151,254,1231,1255]" pageId="8" pageNumber="429" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D5080E56D07FFB5270527DDFEE457A8" box="[264,341,1230,1255]" pageId="8" pageNumber="429" type="male">1 male</specimenCount>
, same data as holotype, MUSENUV slide code 26170.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>