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<document id="5852833F0D93F4CC9EA54936E1274734" ID-DOI="10.23885/181433262019151-2534" ID-ISSN="2713-1785" ID-Zenodo-Dep="8145596" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.tables_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1689300242523" checkinUser="felipe" docAuthor="Kaplin, V. G." docDate="2019" docId="03FFED66C4435E39FEB03C35D060FD72" docLanguage="en" docName="CaucEntomolBull.15.1.25-34.pdf" docOrigin="Caucasian Entomological Bulletin (Caucas. entomol. bull.) 15 (1)" docSource="http://dx.doi.org/10.23885/181433262019151-2534" docStyle="DocumentStyle:6A2F040C6DCB19A13F917346BF1BCB54.1:CaucEntomolBull.2017-.journal_article" docStyleId="6A2F040C6DCB19A13F917346BF1BCB54" docStyleName="CaucEntomolBull.2017-.journal_article" docStyleVersion="1" docTitle="Trigoniophthalmus tseyi Kaplin 2019, sp. n." docType="treatment" docVersion="1" lastPageNumber="31" masterDocId="FFC6951EC4475E3EFF943826D55BFFA8" masterDocTitle="New species of bristletails of the genus Trigoniophthalmus Verhoeff, 1910 (Archaeognatha: Machilidae) from North Ossetia - Alania (Russia)" masterLastPageNumber="34" masterPageNumber="25" pageNumber="28" updateTime="1689615317153" updateUser="diego" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="6BEDDD0AA124910B188913C42ADDF144">New species of bristletails of the genus Trigoniophthalmus Verhoeff, 1910 (Archaeognatha: Machilidae) from North Ossetia - Alania (Russia)</mods:title>
</mods:titleInfo>
<mods:name id="3D34E1E41F41A56667B93527F28D60B3" type="personal">
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<mods:namePart id="2205706E3CD6BA6215604D467BA7BADA">Kaplin, V. G.</mods:namePart>
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<mods:title id="FC92099E97DD0A2BC178A340F2C3F845">Caucasian Entomological Bulletin</mods:title>
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<mods:date id="659A26D5FC68BDB337091C75BEA8836E">2019</mods:date>
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<mods:title id="2D7DEAA0214F12E8E262E6F29DC5D644">Caucas. entomol. bull.</mods:title>
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<treatment id="03FFED66C4435E39FEB03C35D060FD72" LSID="urn:lsid:plazi:treatment:03FFED66C4435E39FEB03C35D060FD72" httpUri="http://treatment.plazi.org/id/03FFED66C4435E39FEB03C35D060FD72" lastPageId="7" lastPageNumber="31" pageId="4" pageNumber="28">
<subSubSection id="C34C0FFBC4435E3AFEB03C35D7C4FB80" box="[292,671,1043,1065]" pageId="4" pageNumber="28" type="nomenclature">
<paragraph id="8BE95C70C4435E3AFEB03C35D7C4FB80" blockId="4.[292,671,1043,1065]" box="[292,671,1043,1065]" pageId="4" pageNumber="28">
<heading id="D0A1EB1CC4435E3AFEB03C35D7C4FB80" box="[292,671,1043,1065]" centered="true" fontSize="9" level="2" pageId="4" pageNumber="28" reason="3">
<taxonomicName id="4C5627F3C4435E3AFEB03C35D704FB81" authority="Kaplin" authorityName="Kaplin" authorityYear="2019" box="[292,607,1043,1065]" class="Insecta" family="Machilidae" genus="Trigoniophthalmus" kingdom="Animalia" order="Archaeognatha" pageId="4" pageNumber="28" phylum="Arthropoda" rank="species" species="tseyi" status="sp. n.">
<emphasis id="B9228062C4435E3AFEB03C35D74EFB81" box="[292,533,1043,1065]" italics="true" pageId="4" pageNumber="28">Trigoniophthalmus tseyi</emphasis>
Kaplin
</taxonomicName>
,
<emphasis id="B9228062C4435E3AFDFC3C35D7C4FB80" bold="true" box="[616,671,1043,1064]" pageId="4" pageNumber="28">
<taxonomicNameLabel id="A2113D19C4435E3AFDFC3C35D7C4FB80" box="[616,671,1043,1064]" pageId="4" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C0FFBC4435E3AFE303C16D77BFBED" box="[420,544,1071,1094]" pageId="4" pageNumber="28" type="description">
<paragraph id="8BE95C70C4435E3AFE303C16D77BFBED" blockId="4.[420,544,1071,1094]" box="[420,544,1071,1094]" pageId="4" pageNumber="28">
(
<figureCitation id="136D40F5C4435E3AFE3F3C16D742FBED" box="[427,537,1071,1094]" captionStart-0="Figs 1623" captionStart-1="Figs 2428" captionStartId-0="6.[237,268,1876,1893]" captionStartId-1="8.[237,268,1896,1913]" captionTargetBox-0="[192,1394,273,1833]" captionTargetBox-1="[198,1386,277,1875]" captionTargetId-0="figure-1@6.[189,1398,272,1848]" captionTargetId-1="figure-104@8.[189,1398,272,1877]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionText-0="Figs 1623. Trigoniophthalmus tseyi sp. n., male, holotype. 16 distal chain of antennal flagellum; 17 apex of cercus; 18 eyes and paired ocelli; 19 maxillary palp; 20 labial palp;21 distal part of mandible; 22 fore leg (part); 23 male genitalia with urocoxite IX. Scale bars 0.1 mm. Рис. 1623. Trigoniophthalmus tseyi sp. n., самец, гоΛотип. 16 цепочка ΑистаΛьной части жгутика усиков; 17 вершина церки; 18 гΛаза и парные гΛазки; 19 нижнечеΛюстной щупик; 20 нижнегубной щупик; 21 ΑистаΛьная часть верхней чеΛюсти; 22 переΑняя нога (часть); 23 поΛовой аппарат самца с кокситом IX сегмента брюшка.Масштабные Λинейки 0.1 мм." captionText-1="Figs 2428. Trigoniophthalmus tseyi sp. n., female, paratype. 24 urosternite and urocoxites IV; 25 urosternite and urocoxites VII; 26 ovipositor with urocoxite IX; 27 apex of anterior gonapophysis of ovipositor; 28 apex of posterior gonapophysis of ovipositor.Scale bars 0.1 mm. Рис. 2428. Trigoniophthalmus tseyi sp. n., самка, паратип. 24 стернит и кокситы IV сегмента брюшка; 25 стернит и кокситы VII сегмента брюшка; 26 яйцекΛаΑ с кокситом IX сегмента брюшка; 27 вершина переΑних створок яйцекΛаΑа; 28 вершина заΑних створок яйцекΛаΑа. Масштабные Λинейки 0.1 мм." figureDoi-0="http://doi.org/10.5281/zenodo.8145602" figureDoi-1="http://doi.org/10.5281/zenodo.8145604" httpUri-0="https://zenodo.org/record/8145602/files/figure.png" httpUri-1="https://zenodo.org/record/8145604/files/figure.png" pageId="4" pageNumber="28">Figs 1628</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34C0FFBC4435E3AFF793C43D499FB10" pageId="4" pageNumber="28" type="materials_examined">
<paragraph id="8BE95C70C4435E3AFF793C43D499FB10" blockId="4.[188,776,1124,1631]" pageId="4" pageNumber="28">
<emphasis id="B9228062C4435E3AFF793C43D462FBDE" bold="true" box="[237,313,1125,1142]" pageId="4" pageNumber="28">Material.</emphasis>
<materialsCitation id="3B3E562DC4435E3AFED43C43D775FB0A" collectingDate="2018-04-28" collectionCode="V" collectorName="G. Kaplin" country="Russia" county="Alagir Distr." elevation="1950" latitude="42.791943" location="Recom" longLatPrecision="19" longitude="43.90389" municipality="Verkhniy Tsey" pageId="4" pageNumber="28" specimenCount="1" specimenCount-male="1" stateProvince="North Ossetia" typeStatus="holotype">
<typeStatus id="54EDE2D2C4435E3AFED43C43D4D1FBDE" box="[320,394,1125,1142]" pageId="4" pageNumber="28">Holotype</typeStatus>
,
<specimenCount id="9D5097F9C4435E3AFE013C40D4F8FBD0" box="[405,419,1126,1144]" pageId="4" pageNumber="28" type="male"></specimenCount>
(in slides):
<collectingCountry id="F3411CE0C4435E3AFD923C43D761FBDE" box="[518,570,1125,1142]" name="Russia" pageId="4" pageNumber="28">Russia</collectingCountry>
,
<collectingRegion id="49929292C4435E3AFDD03C43D7EEFBDE" box="[580,693,1125,1142]" country="Russia" name="North Ossetia" pageId="4" pageNumber="28">North Ossetia</collectingRegion>
<emphasis id="B9228062C4435E3AFD2A3C42D79CFBDE" box="[702,711,1124,1142]" italics="true" pageId="4" pageNumber="28"></emphasis>
Alania,
<collectingCounty id="628824FCC4435E3AFF293C5DD445FB24" box="[189,286,1147,1164]" pageId="4" pageNumber="28">Alagir Distr.</collectingCounty>
,
<collectingMunicipality id="6B8DC60AC4435E3AFEB63C5DD4CBFB24" box="[290,400,1147,1164]" pageId="4" pageNumber="28">Verkhniy Tsey</collectingMunicipality>
settl.,environs of
<location id="8E890AABC4435E3AFD8D3C5AD714FB24" LSID="urn:lsid:plazi:treatment:03FFED66C4435E39FEB03C35D060FD72:8E890AABC4435E3AFD8D3C5AD714FB24" box="[537,591,1148,1164]" country="Russia" county="Alagir Distr." latitude="42.791943" longLatPrecision="19" longitude="43.90389" municipality="Verkhniy Tsey" name="Recom" pageId="4" pageNumber="28" stateProvince="North Ossetia">Recom</location>
sanctuary,
<geoCoordinate id="EE623AB7C4435E3AFD313C5DD7A5FB24" box="[677,766,1146,1164]" degrees="42" direction="north" minutes="47" orientation="latitude" pageId="4" pageNumber="28" precision="15" seconds="31" value="42.791943">
42
<emphasis id="B9228062C4435E3AFD2C3C5CD7E4FB24" box="[696,703,1146,1164]" italics="true" pageId="4" pageNumber="28">°</emphasis>
4731ʺN
</geoCoordinate>
<emphasis id="B9228062C4435E3AFC963C5CD65CFB24" box="[770,775,1146,1164]" italics="true" pageId="4" pageNumber="28">/</emphasis>
<geoCoordinate id="EE623AB7C4435E3AFF293CB7D44FFB09" box="[189,276,1168,1186]" degrees="43" direction="east" minutes="54" orientation="longitude" pageId="4" pageNumber="28" precision="15" seconds="14" value="43.90389">
43
<emphasis id="B9228062C4435E3AFF443CB6D58CFB0A" box="[208,215,1168,1186]" italics="true" pageId="4" pageNumber="28">°</emphasis>
5414ʺE
</geoCoordinate>
,
<quantity id="4CAEF195C4435E3AFE8A3CB7D406FB09" box="[286,349,1169,1186]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="4" pageNumber="28" unit="m" value="1950.0">
<elevation id="007BBB43C4435E3AFE8A3CB7D406FB09" box="[286,349,1169,1186]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="4" pageNumber="28" unit="m" value="1950.0">1950 m</elevation>
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,
<date id="FFE87AB0C4435E3AFEFC3CB7D4E0FB0A" box="[360,443,1169,1186]" pageId="4" pageNumber="28" value="2018-04-28">
<collectingDate id="EFAC8358C4435E3AFEFC3CB7D4E0FB0A" box="[360,443,1169,1186]" pageId="4" pageNumber="28" value="2018-04-28">28.04.2018</collectingDate>
</date>
(
<collectionCode id="ED47C4B5C4435E3AFE5C3CB7D48CFB0A" box="[456,471,1169,1186]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="4" pageNumber="28" type="Museum">V</collectionCode>
.
<collectorName id="26A339A6C4435E3AFE4E3CB7D772FB0A" box="[474,553,1168,1186]" pageId="4" pageNumber="28">G. Kaplin</collectorName>
)
</materialsCitation>
.
<materialsCitation id="3B3E562DC4435E3AFDAC3CB7D4E5FB10" collectingDate="2018-04-28" collectionCode="V" collectorName="G. Kaplin" country="Russia" county="Alagir Distr." elevation="1950" latitude="42.791943" location="Recom" longLatPrecision="19" longitude="43.90389" municipality="Verkhniy Tsey" pageId="4" pageNumber="28" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="North Ossetia" typeStatus="paratype">
<typeStatus id="54EDE2D2C4435E3AFDAC3CB7D7DDFB09" box="[568,646,1169,1185]" pageId="4" pageNumber="28">Paratypes</typeStatus>
:
<specimenCount id="9D5097F9C4435E3AFD043CB7D7FCFB0B" box="[656,679,1169,1187]" pageId="4" pageNumber="28" type="male">1♂</specimenCount>
,
<specimenCount id="9D5097F9C4435E3AFD273CB7D790FB0B" box="[691,715,1169,1187]" pageId="4" pageNumber="28" type="female">2♀</specimenCount>
(
<specimenCount id="9D5097F9C4435E3AFD4E3CB7D7ABFB0B" box="[730,752,1169,1187]" pageId="4" pageNumber="28" type="female">1♀</specimenCount>
in slides), same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C0FFBC4435E39FF793C98D1B3FE96" lastPageId="7" lastPageNumber="31" pageId="4" pageNumber="28" type="description">
<paragraph id="8BE95C70C4435E3AFF793C98D438F984" blockId="4.[188,776,1124,1631]" pageId="4" pageNumber="28">
<emphasis id="B9228062C4435E3AFF793C98D439FB79" bold="true" box="[237,354,1214,1233]" pageId="4" pageNumber="28">Description.</emphasis>
Body length: male
<quantity id="4CAEF195C4435E3AFD9C3C98D739FB79" box="[520,610,1214,1233]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" pageId="4" pageNumber="28" unit="mm" value="7.25" valueMax="7.5" valueMin="7.0">77.5 mm</quantity>
, female
<quantity id="4CAEF195C4435E3AFD3C3C98D65FFB79" box="[680,772,1214,1233]" metricMagnitude="-3" metricUnit="m" metricValue="7.65" metricValueMax="8.0" metricValueMin="7.3" pageId="4" pageNumber="28" unit="mm" value="7.65" valueMax="8.0" valueMin="7.3">7.38 mm</quantity>
. Body width: males and females
<quantity id="4CAEF195C4435E3AFE4B3CF1D715FB42" box="[479,590,1239,1258]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.1" metricValueMin="1.8" pageId="4" pageNumber="28" unit="mm" value="1.95" valueMax="2.1" valueMin="1.8">1.82.1 mm</quantity>
. General body color whitish. Antennal base, frons, maxillae, mandibles, hypopharynx with brown hypodermal pigment of weak and medium intensity. Color of scales on surface of body brownish. Antennae slightly shorter than body. Distal chains of flagellum divided into 912 articles in male and female (
<figureCitation id="136D40F5C4435E3AFE603D75D768FACE" box="[500,563,1363,1382]" captionStart="Figs 1623" captionStartId="6.[237,268,1876,1893]" captionTargetBox="[192,1394,273,1833]" captionTargetId="figure-1@6.[189,1398,272,1848]" captionTargetPageId="6" captionText="Figs 1623. Trigoniophthalmus tseyi sp. n., male, holotype. 16 distal chain of antennal flagellum; 17 apex of cercus; 18 eyes and paired ocelli; 19 maxillary palp; 20 labial palp;21 distal part of mandible; 22 fore leg (part); 23 male genitalia with urocoxite IX. Scale bars 0.1 mm. Рис. 1623. Trigoniophthalmus tseyi sp. n., самец, гоΛотип. 16 цепочка ΑистаΛьной части жгутика усиков; 17 вершина церки; 18 гΛаза и парные гΛазки; 19 нижнечеΛюстной щупик; 20 нижнегубной щупик; 21 ΑистаΛьная часть верхней чеΛюсти; 22 переΑняя нога (часть); 23 поΛовой аппарат самца с кокситом IX сегмента брюшка.Масштабные Λинейки 0.1 мм." figureDoi="http://doi.org/10.5281/zenodo.8145602" httpUri="https://zenodo.org/record/8145602/files/figure.png" pageId="4" pageNumber="28">Fig. 16</figureCitation>
). Clypeus of male with long thin bristles. Cercus approximately 0.410.46 body length, including about 17 articles in both sexes. Apex of cercus with one spike (
<figureCitation id="136D40F5C4435E3AFF613DBBD46AFA18" box="[245,305,1437,1456]" captionStart="Figs 1623" captionStartId="6.[237,268,1876,1893]" captionTargetBox="[192,1394,273,1833]" captionTargetId="figure-1@6.[189,1398,272,1848]" captionTargetPageId="6" captionText="Figs 1623. Trigoniophthalmus tseyi sp. n., male, holotype. 16 distal chain of antennal flagellum; 17 apex of cercus; 18 eyes and paired ocelli; 19 maxillary palp; 20 labial palp;21 distal part of mandible; 22 fore leg (part); 23 male genitalia with urocoxite IX. Scale bars 0.1 mm. Рис. 1623. Trigoniophthalmus tseyi sp. n., самец, гоΛотип. 16 цепочка ΑистаΛьной части жгутика усиков; 17 вершина церки; 18 гΛаза и парные гΛазки; 19 нижнечеΛюстной щупик; 20 нижнегубной щупик; 21 ΑистаΛьная часть верхней чеΛюсти; 22 переΑняя нога (часть); 23 поΛовой аппарат самца с кокситом IX сегмента брюшка.Масштабные Λинейки 0.1 мм." figureDoi="http://doi.org/10.5281/zenodo.8145602" httpUri="https://zenodo.org/record/8145602/files/figure.png" pageId="4" pageNumber="28">Fig.17</figureCitation>
). Articles of cerci, except apical one in female and two distal ones in male, with 1 or 2 colorless supporting macrochaetae on the inner side. Cerci of female also with 3 or 4 macrochaetae on outer side, male with only 1 macrochaeta in such position. Supporting macrochaetae also present on lateral sides of articles of caudal filament.
</paragraph>
<paragraph id="8BE95C70C4435E3AFF793E14D145FC9D" blockId="4.[188,776,1124,1631]" lastBlockId="4.[811,1401,676,1631]" pageId="4" pageNumber="28">
Compound eyes black (in ethanol). Ratio of length to width of compound eye about
<quantity id="4CAEF195C4435E3AFEEF3E6AD4B2F9F6" box="[379,489,1611,1631]" metricMagnitude="-2" metricUnit="m" metricValue="2.6162" metricValueMax="2.6416" metricValueMin="2.5908" pageId="4" pageNumber="28" unit="in" value="1.03" valueMax="1.04" valueMin="1.02">1.021.04 in</quantity>
males and
<quantity id="4CAEF195C4435E3AFDDF3E6DD7E2F9F6" box="[587,697,1611,1630]" metricMagnitude="-2" metricUnit="m" metricValue="2.667" metricValueMax="2.6924" metricValueMin="2.6416" pageId="4" pageNumber="28" unit="in" value="1.05" valueMax="1.06" valueMin="1.04">1.041.06 in</quantity>
females; ratio of length of contact line to length of eye
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both sexes. Paired ocelli submedian, pear-shaped, brown with a welldefined white border, 1.41.6 times as wide as long in both sexes. Distance between inner margins of ocelli about 0.160.18 and between their outer margins 0.600.65 total width of compound eyes in both sexes (
<figureCitation id="136D40F5C4435E3AFC423B04D148FC9D" box="[982,1043,802,821]" captionStart="Figs 1623" captionStartId="6.[237,268,1876,1893]" captionTargetBox="[192,1394,273,1833]" captionTargetId="figure-1@6.[189,1398,272,1848]" captionTargetPageId="6" captionText="Figs 1623. Trigoniophthalmus tseyi sp. n., male, holotype. 16 distal chain of antennal flagellum; 17 apex of cercus; 18 eyes and paired ocelli; 19 maxillary palp; 20 labial palp;21 distal part of mandible; 22 fore leg (part); 23 male genitalia with urocoxite IX. Scale bars 0.1 mm. Рис. 1623. Trigoniophthalmus tseyi sp. n., самец, гоΛотип. 16 цепочка ΑистаΛьной части жгутика усиков; 17 вершина церки; 18 гΛаза и парные гΛазки; 19 нижнечеΛюстной щупик; 20 нижнегубной щупик; 21 ΑистаΛьная часть верхней чеΛюсти; 22 переΑняя нога (часть); 23 поΛовой аппарат самца с кокситом IX сегмента брюшка.Масштабные Λинейки 0.1 мм." figureDoi="http://doi.org/10.5281/zenodo.8145602" httpUri="https://zenodo.org/record/8145602/files/figure.png" pageId="4" pageNumber="28">Fig. 18</figureCitation>
).
</paragraph>
<paragraph id="8BE95C70C4435E3AFCC83B1DD6F5FBE3" blockId="4.[811,1401,676,1631]" pageId="4" pageNumber="28">
Apical article of maxillary palp 0.77 (male) or 0.82 (female) times as long as preceding one. Dorsal surface of 7
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, 6
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and 5
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articles of maxillary palp in both sexes with 1113, 1113 and 4 hyaline spines, respectively. Ventral surface of 27
<superScript id="7C23F138C4435E3AFAB53BA0D077FC3A" attach="left" box="[1313,1324,902,914]" fontSize="5" pageId="4" pageNumber="28">th</superScript>
articles of male maxillary palp with relatively numerous and long thin chaetae, on sixth and especially seventh articles such chaetae distributed more sparse (
<figureCitation id="136D40F5C4435E3AFB873BF5D108FC4E" box="[1043,1107,979,998]" captionStart="Figs 1623" captionStartId="6.[237,268,1876,1893]" captionTargetBox="[192,1394,273,1833]" captionTargetId="figure-1@6.[189,1398,272,1848]" captionTargetPageId="6" captionText="Figs 1623. Trigoniophthalmus tseyi sp. n., male, holotype. 16 distal chain of antennal flagellum; 17 apex of cercus; 18 eyes and paired ocelli; 19 maxillary palp; 20 labial palp;21 distal part of mandible; 22 fore leg (part); 23 male genitalia with urocoxite IX. Scale bars 0.1 mm. Рис. 1623. Trigoniophthalmus tseyi sp. n., самец, гоΛотип. 16 цепочка ΑистаΛьной части жгутика усиков; 17 вершина церки; 18 гΛаза и парные гΛазки; 19 нижнечеΛюстной щупик; 20 нижнегубной щупик; 21 ΑистаΛьная часть верхней чеΛюсти; 22 переΑняя нога (часть); 23 поΛовой аппарат самца с кокситом IX сегмента брюшка.Масштабные Λинейки 0.1 мм." figureDoi="http://doi.org/10.5281/zenodo.8145602" httpUri="https://zenodo.org/record/8145602/files/figure.png" pageId="4" pageNumber="28">Fig. 19</figureCitation>
). Similar long thin chaetae also present on dorsal surface of 2
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and 3
<superScript id="7C23F138C4435E3AFBEC3BCDD1D8FC5F" attach="left" box="[1144,1155,1003,1015]" fontSize="5" pageId="4" pageNumber="28">rd</superScript>
articles of male labial palp. Apical article of labial palp triangularly oval, 2.2 (male) or 2.3 (female) times as long as wide (
<figureCitation id="136D40F5C4435E3AFBDB3C39D1D4FB9A" box="[1103,1167,1055,1074]" captionStart="Figs 1623" captionStartId="6.[237,268,1876,1893]" captionTargetBox="[192,1394,273,1833]" captionTargetId="figure-1@6.[189,1398,272,1848]" captionTargetPageId="6" captionText="Figs 1623. Trigoniophthalmus tseyi sp. n., male, holotype. 16 distal chain of antennal flagellum; 17 apex of cercus; 18 eyes and paired ocelli; 19 maxillary palp; 20 labial palp;21 distal part of mandible; 22 fore leg (part); 23 male genitalia with urocoxite IX. Scale bars 0.1 mm. Рис. 1623. Trigoniophthalmus tseyi sp. n., самец, гоΛотип. 16 цепочка ΑистаΛьной части жгутика усиков; 17 вершина церки; 18 гΛаза и парные гΛазки; 19 нижнечеΛюстной щупик; 20 нижнегубной щупик; 21 ΑистаΛьная часть верхней чеΛюсти; 22 переΑняя нога (часть); 23 поΛовой аппарат самца с кокситом IX сегмента брюшка.Масштабные Λинейки 0.1 мм." figureDoi="http://doi.org/10.5281/zenodo.8145602" httpUri="https://zenodo.org/record/8145602/files/figure.png" pageId="4" pageNumber="28">Fig. 20</figureCitation>
). Mandibles with 4 distal teeth (
<figureCitation id="136D40F5C4435E3AFCF13C1ED6F9FBE3" box="[869,930,1080,1099]" captionStart="Figs 1623" captionStartId="6.[237,268,1876,1893]" captionTargetBox="[192,1394,273,1833]" captionTargetId="figure-1@6.[189,1398,272,1848]" captionTargetPageId="6" captionText="Figs 1623. Trigoniophthalmus tseyi sp. n., male, holotype. 16 distal chain of antennal flagellum; 17 apex of cercus; 18 eyes and paired ocelli; 19 maxillary palp; 20 labial palp;21 distal part of mandible; 22 fore leg (part); 23 male genitalia with urocoxite IX. Scale bars 0.1 mm. Рис. 1623. Trigoniophthalmus tseyi sp. n., самец, гоΛотип. 16 цепочка ΑистаΛьной части жгутика усиков; 17 вершина церки; 18 гΛаза и парные гΛазки; 19 нижнечеΛюстной щупик; 20 нижнегубной щупик; 21 ΑистаΛьная часть верхней чеΛюсти; 22 переΑняя нога (часть); 23 поΛовой аппарат самца с кокситом IX сегмента брюшка.Масштабные Λинейки 0.1 мм." figureDoi="http://doi.org/10.5281/zenodo.8145602" httpUri="https://zenodo.org/record/8145602/files/figure.png" pageId="4" pageNumber="28">Fig. 21</figureCitation>
).
</paragraph>
<paragraph id="8BE95C70C4435E3AFCC83C74D159FAD3" blockId="4.[811,1401,676,1631]" pageId="4" pageNumber="28">
Fore femur and tibia of male and female widened, without sensory field (
<figureCitation id="136D40F5C4435E3AFC333C4DD6BDFBD6" box="[935,998,1131,1150]" captionStart="Figs 1623" captionStartId="6.[237,268,1876,1893]" captionTargetBox="[192,1394,273,1833]" captionTargetId="figure-1@6.[189,1398,272,1848]" captionTargetPageId="6" captionText="Figs 1623. Trigoniophthalmus tseyi sp. n., male, holotype. 16 distal chain of antennal flagellum; 17 apex of cercus; 18 eyes and paired ocelli; 19 maxillary palp; 20 labial palp;21 distal part of mandible; 22 fore leg (part); 23 male genitalia with urocoxite IX. Scale bars 0.1 mm. Рис. 1623. Trigoniophthalmus tseyi sp. n., самец, гоΛотип. 16 цепочка ΑистаΛьной части жгутика усиков; 17 вершина церки; 18 гΛаза и парные гΛазки; 19 нижнечеΛюстной щупик; 20 нижнегубной щупик; 21 ΑистаΛьная часть верхней чеΛюсти; 22 переΑняя нога (часть); 23 поΛовой аппарат самца с кокситом IX сегмента брюшка.Масштабные Λинейки 0.1 мм." figureDoi="http://doi.org/10.5281/zenodo.8145602" httpUri="https://zenodo.org/record/8145602/files/figure.png" pageId="4" pageNumber="28">Fig. 22</figureCitation>
). Ratios of length to width of femur, tibia and tarsus as shown in
<tableCitation id="C6D469CBC4435E3AFC443CA2D149FB3F" box="[976,1042,1156,1175]" captionStart="Table 4" captionStartId="4.[237,279,1683,1700]" captionTargetId="graphics-953@4.[189,1398,1733,2018]" captionTargetPageId="4" captionText="Table 4. Ratios of length to width of leg segments of Trigoniophthalmus tseyi sp. n. ТабΛица 4. Отношения ΑΛины к ширине сегментов ног Trigoniophthalmus tseyi sp. n." pageId="4" pageNumber="28">Table 4</tableCitation>
. Ratio of length of 3
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tarsomere of tarsus to total length of tarsus
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both sexes. Coxa of male with long, thin bristles, missing from the femur, tibia and tarsus. Ventral surface of femur, tibia and tarsus with colorless spine-like chaetae (
<tableCitation id="C6D469CBC4435E3AFCA73CCFD622FB54" box="[819,889,1257,1276]" captionStart="Table 5" captionStartId="7.[237,279,790,807]" captionTargetId="graphics-301@7.[189,1397,841,1163]" captionTargetPageId="7" captionText="Table 5. Number of spine-like setae on the legs of Trigoniophthalmus tseyi sp. n. ТабΛица 5. КоΛичество игΛовиΑных щетинок на ногах Trigoniophthalmus tseyi sp. n." pageId="4" pageNumber="28">Table 5</tableCitation>
). Middle and hind legs with coxal styli. Length of styli
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(female) or
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(male). Ratio of length of styli to width of middle and hind coxae about
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both sexes. Praetarsus with 2 claws and support cone-shaped projection between them. Ratios of length to width of projection 1.92.1, widths of projection and pretarsus about 0.3.
</paragraph>
<paragraph id="8BE95C70C4435E39FCC83DA7D779FE0D" blockId="4.[811,1401,676,1631]" lastBlockId="7.[189,776,273,729]" lastPageId="7" lastPageNumber="31" pageId="4" pageNumber="28">
Posterior margin of pronotum with a deep notch. In both sexes, urites I and VVII with 1 + 1 eversible vesicles, but urites IIIV with 2 + 2 eversible vesicles (
<figureCitation id="136D40F5C4435E3AFB263D95D041FA6E" box="[1202,1306,1459,1478]" captionStart="Figs 2428" captionStartId="8.[237,268,1896,1913]" captionTargetBox="[198,1386,277,1875]" captionTargetId="figure-104@8.[189,1398,272,1877]" captionTargetPageId="8" captionText="Figs 2428. Trigoniophthalmus tseyi sp. n., female, paratype. 24 urosternite and urocoxites IV; 25 urosternite and urocoxites VII; 26 ovipositor with urocoxite IX; 27 apex of anterior gonapophysis of ovipositor; 28 apex of posterior gonapophysis of ovipositor.Scale bars 0.1 mm. Рис. 2428. Trigoniophthalmus tseyi sp. n., самка, паратип. 24 стернит и кокситы IV сегмента брюшка; 25 стернит и кокситы VII сегмента брюшка; 26 яйцекΛаΑ с кокситом IX сегмента брюшка; 27 вершина переΑних створок яйцекΛаΑа; 28 вершина заΑних створок яйцекΛаΑа. Масштабные Λинейки 0.1 мм." figureDoi="http://doi.org/10.5281/zenodo.8145604" httpUri="https://zenodo.org/record/8145604/files/figure.png" pageId="4" pageNumber="28">Figs 24, 25</figureCitation>
). In male, posterior angle of urosternites II, IIIVI, VII approximately 78°, 7173° and 68°; but in female, 84°, 7375° and 65°, respectively. In male, posterior angle of urosternite VIII about 84°. Ratios of lengths of styli (without apical spines) and urocoxites IIIX as shown in
<tableCitation id="C6D469CBC4435E3AFCB93E14D634F9ED" box="[813,879,1586,1605]" captionStart="Table 6" captionStartId="7.[237,279,1220,1237]" captionTargetId="graphics-523@7.[189,1397,1269,1617]" captionTargetPageId="7" captionText="Table 6. Length ratios of urosternites and urocoxites of Trigoniophthalmus tseyi sp. n. ТабΛица 6. Отношения ΑΛины стернитов и кокситов брюшка Trigoniophthalmus tseyi sp. n." pageId="4" pageNumber="28">Table 6</tableCitation>
. Inner posterior lobes of urocoxites VII between eversible vesicles of female protruding (
<figureCitation id="136D40F5C4435E3AFBDF3E6AD1D6F9F7" box="[1099,1165,1612,1631]" captionStart="Figs 2428" captionStartId="8.[237,268,1896,1913]" captionTargetBox="[198,1386,277,1875]" captionTargetId="figure-104@8.[189,1398,272,1877]" captionTargetPageId="8" captionText="Figs 2428. Trigoniophthalmus tseyi sp. n., female, paratype. 24 urosternite and urocoxites IV; 25 urosternite and urocoxites VII; 26 ovipositor with urocoxite IX; 27 apex of anterior gonapophysis of ovipositor; 28 apex of posterior gonapophysis of ovipositor.Scale bars 0.1 mm. Рис. 2428. Trigoniophthalmus tseyi sp. n., самка, паратип. 24 стернит и кокситы IV сегмента брюшка; 25 стернит и кокситы VII сегмента брюшка; 26 яйцекΛаΑ с кокситом IX сегмента брюшка; 27 вершина переΑних створок яйцекΛаΑа; 28 вершина заΑних створок яйцекΛаΑа. Масштабные Λинейки 0.1 мм." figureDoi="http://doi.org/10.5281/zenodo.8145604" httpUri="https://zenodo.org/record/8145604/files/figure.png" pageId="4" pageNumber="28">Fig. 25</figureCitation>
); ratio of length to total width of these lobes about 0.7. Thoracic tergites, urotergites IIV, urosternites and urocoxites IVI without macrochaetae in both sexes. Distribution of sublateral macrochaetae on urotergites VX and spines on urocoxites VIIIX as shown in
<tableCitation id="C6D469CBC4405E39FDD63978D7DAFEDA" box="[578,641,350,370]" captionStart="Table 7" captionStartId="7.[237,279,1675,1692]" captionTargetId="graphics-806@7.[189,1396,1727,2024]" captionTargetPageId="7" captionText="Table 7. Distribution of sublateral macrochaetae on urotergites and spines on urocoxites of Trigoniophthalmus tseyi sp. n. ТабΛица 7. РаспреΑеΛение субΛатераΛьных макрохет на тергитах и игΛ на кокситах брюшка Trigoniophthalmus tseyi sp. n." pageId="7" pageNumber="31">Table 7</tableCitation>
.Urocoxites IX, respectively, with 2 and 78 (male) or 1 and 711 (female) outer and inner sublateral spines (
<figureCitation id="136D40F5C4405E39FE2339B4D4ACFE0D" box="[439,503,402,421]" captionStart="Figs 1623" captionStartId="6.[237,268,1876,1893]" captionTargetBox="[192,1394,273,1833]" captionTargetId="figure-1@6.[189,1398,272,1848]" captionTargetPageId="6" captionText="Figs 1623. Trigoniophthalmus tseyi sp. n., male, holotype. 16 distal chain of antennal flagellum; 17 apex of cercus; 18 eyes and paired ocelli; 19 maxillary palp; 20 labial palp;21 distal part of mandible; 22 fore leg (part); 23 male genitalia with urocoxite IX. Scale bars 0.1 mm. Рис. 1623. Trigoniophthalmus tseyi sp. n., самец, гоΛотип. 16 цепочка ΑистаΛьной части жгутика усиков; 17 вершина церки; 18 гΛаза и парные гΛазки; 19 нижнечеΛюстной щупик; 20 нижнегубной щупик; 21 ΑистаΛьная часть верхней чеΛюсти; 22 переΑняя нога (часть); 23 поΛовой аппарат самца с кокситом IX сегмента брюшка.Масштабные Λинейки 0.1 мм." figureDoi="http://doi.org/10.5281/zenodo.8145602" httpUri="https://zenodo.org/record/8145602/files/figure.png" pageId="7" pageNumber="31">Figs 23</figureCitation>
,
<figureCitation id="136D40F5C4405E39FE6B39B4D74CFE0D" box="[511,535,402,421]" captionStart="Figs 2428" captionStartId="8.[237,268,1896,1913]" captionTargetBox="[198,1386,277,1875]" captionTargetId="figure-104@8.[189,1398,272,1877]" captionTargetPageId="8" captionText="Figs 2428. Trigoniophthalmus tseyi sp. n., female, paratype. 24 urosternite and urocoxites IV; 25 urosternite and urocoxites VII; 26 ovipositor with urocoxite IX; 27 apex of anterior gonapophysis of ovipositor; 28 apex of posterior gonapophysis of ovipositor.Scale bars 0.1 mm. Рис. 2428. Trigoniophthalmus tseyi sp. n., самка, паратип. 24 стернит и кокситы IV сегмента брюшка; 25 стернит и кокситы VII сегмента брюшка; 26 яйцекΛаΑ с кокситом IX сегмента брюшка; 27 вершина переΑних створок яйцекΛаΑа; 28 вершина заΑних створок яйцекΛаΑа. Масштабные Λинейки 0.1 мм." figureDoi="http://doi.org/10.5281/zenodo.8145604" httpUri="https://zenodo.org/record/8145604/files/figure.png" pageId="7" pageNumber="31">26</figureCitation>
).
</paragraph>
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<paragraph id="8BE95C70C4435E3AFF793EB5D6C5F911" blockId="4.[237,926,1683,1722]" pageId="4" pageNumber="28">
Table 4. Ratios of length to width of leg segments of
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<emphasis id="B9228062C4435E3AFD153EB5D667F90B" box="[641,828,1683,1700]" italics="true" pageId="4" pageNumber="28">Trigoniophthalmus tseyi</emphasis>
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<emphasis id="B9228062C4435E3AFCD53EB5D637F90B" bold="true" box="[833,876,1683,1699]" pageId="4" pageNumber="28">
<taxonomicNameLabel id="A2113D19C4435E3AFCD53EB5D637F90B" box="[833,876,1683,1699]" pageId="4" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
ТабΛица 4. Отношения ΑΛины к ширине сегментов ног
<taxonomicName id="4C5627F3C4435E3AFD273E8FD635F911" authority="Kaplin, 2019" authorityName="Kaplin" authorityYear="2019" box="[691,878,1705,1722]" class="Insecta" family="Machilidae" genus="Trigoniophthalmus" kingdom="Animalia" order="Archaeognatha" pageId="4" pageNumber="28" phylum="Arthropoda" rank="species" species="tseyi" status="sp. n.">
<emphasis id="B9228062C4435E3AFD273E8FD635F911" box="[691,878,1705,1722]" italics="true" pageId="4" pageNumber="28">Trigoniophthalmus tseyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2113D19C4435E3AFCE63E8FD6C5F911" box="[882,926,1705,1721]" pageId="4" pageNumber="28" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BE95C70C4435E3AFD613EF4D06EF874" pageId="4" pageNumber="28">
<table id="F956AED0C443A1C1FF5C3EF7D018F875" box="[200,1347,1745,2013]" gridcols="7" gridrows="7" pageId="4" pageNumber="28">
<tr id="35665E32C443A1C1FF5C3EF7D018F94D" box="[200,1347,1745,1765]" gridrow="0" pageId="4" pageNumber="28">
<th id="76B7374EC443A1C1FF5C3EF7D018F94D" box="[200,1347,1745,1765]" colspan="7" colspanRight="6" gridcol="0" gridrow="0" pageId="4" pageNumber="28">Sex, pair of legs / ПоΛ, пара ног</th>
</tr>
<tr id="35665E32C443A1C1FF5C3ED0D018F8A2" box="[200,1347,1782,1802]" gridrow="1" pageId="4" pageNumber="28" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-6="1">
<th id="76B7374EC443A1C1FF5C3ED0D40DF8A2" box="[200,342,1782,1802]" gridcol="0" gridrow="1" pageId="4" pageNumber="28">Segments</th>
<td id="76B7374EC443A1C1FDC43ED0D79FF8A2" box="[592,708,1782,1802]" gridcol="2" gridrow="1" pageId="4" pageNumber="28">male / самец</td>
<td id="76B7374EC443A1C1FBAF3ED0D1E5F8A2" box="[1083,1214,1782,1802]" gridcol="5" gridrow="1" pageId="4" pageNumber="28">female / самка</td>
</tr>
<tr id="35665E32C443A1C1FF5C3F36D018F8E0" box="[200,1347,1808,1864]" gridrow="2" pageId="4" pageNumber="28">
<th id="76B7374EC443A1C1FF5C3F36D40DF8E0" box="[200,342,1808,1864]" gridcol="0" gridrow="2" pageId="4" pageNumber="28">Сегменты</th>
<td id="76B7374EC443A1C1FE2C3F36D74BF8E0" box="[440,528,1808,1864]" gridcol="1" gridrow="2" pageId="4" pageNumber="28">fore переΑняя</td>
<td id="76B7374EC443A1C1FDC43F36D79FF8E0" box="[592,708,1808,1864]" gridcol="2" gridrow="2" pageId="4" pageNumber="28">middle среΑняя</td>
<td id="76B7374EC443A1C1FC843F36D60BF8E0" box="[784,848,1808,1864]" gridcol="3" gridrow="2" pageId="4" pageNumber="28">hind заΑняя</td>
<td id="76B7374EC443A1C1FC3F3F36D158F8E0" box="[939,1027,1808,1864]" gridcol="4" gridrow="2" pageId="4" pageNumber="28">fore переΑняя</td>
<td id="76B7374EC443A1C1FBAF3F36D1E5F8E0" box="[1083,1214,1808,1864]" gridcol="5" gridrow="2" pageId="4" pageNumber="28">middle среΑняя</td>
<td id="76B7374EC443A1C1FA973F36D018F8E0" box="[1283,1347,1808,1864]" gridcol="6" gridrow="2" pageId="4" pageNumber="28">hind заΑняя</td>
</tr>
<tr id="35665E32C443A1C1FF5C3F7CD018F8C5" box="[200,1347,1882,1901]" gridrow="3" pageId="4" pageNumber="28">
<th id="76B7374EC443A1C1FF5C3F7CD40DF8C5" box="[200,342,1882,1901]" gridcol="0" gridrow="3" pageId="4" pageNumber="28">Tarsa / Αапка</th>
<td id="76B7374EC443A1C1FE2C3F7CD74BF8C5" box="[440,528,1882,1901]" gridcol="1" gridrow="3" pageId="4" pageNumber="28">4.47</td>
<td id="76B7374EC443A1C1FDC43F7CD79FF8C5" box="[592,708,1882,1901]" gridcol="2" gridrow="3" pageId="4" pageNumber="28">4.51</td>
<td id="76B7374EC443A1C1FC843F7CD60BF8C5" box="[784,848,1882,1901]" gridcol="3" gridrow="3" pageId="4" pageNumber="28">6.36</td>
<td id="76B7374EC443A1C1FC3F3F7CD158F8C5" box="[939,1027,1882,1901]" gridcol="4" gridrow="3" pageId="4" pageNumber="28">4.12</td>
<td id="76B7374EC443A1C1FBAF3F7CD1E5F8C5" box="[1083,1214,1882,1901]" gridcol="5" gridrow="3" pageId="4" pageNumber="28">4.91</td>
<td id="76B7374EC443A1C1FA973F7CD018F8C5" box="[1283,1347,1882,1901]" gridcol="6" gridrow="3" pageId="4" pageNumber="28">6.30</td>
</tr>
<tr id="35665E32C443A1C1FF5C3F59D018F83A" box="[200,1347,1919,1938]" gridrow="4" pageId="4" pageNumber="28">
<th id="76B7374EC443A1C1FF5C3F59D40DF83A" box="[200,342,1919,1938]" gridcol="0" gridrow="4" pageId="4" pageNumber="28">Tibia / ГоΛень</th>
<td id="76B7374EC443A1C1FE2C3F59D74BF83A" box="[440,528,1919,1938]" gridcol="1" gridrow="4" pageId="4" pageNumber="28">2.06</td>
<td id="76B7374EC443A1C1FDC43F59D79FF83A" box="[592,708,1919,1938]" gridcol="2" gridrow="4" pageId="4" pageNumber="28">2.05</td>
<td id="76B7374EC443A1C1FC843F59D60BF83A" box="[784,848,1919,1938]" gridcol="3" gridrow="4" pageId="4" pageNumber="28">2.95</td>
<td id="76B7374EC443A1C1FC3F3F59D158F83A" box="[939,1027,1919,1938]" gridcol="4" gridrow="4" pageId="4" pageNumber="28">2.08</td>
<td id="76B7374EC443A1C1FBAF3F59D1E5F83A" box="[1083,1214,1919,1938]" gridcol="5" gridrow="4" pageId="4" pageNumber="28">2.02</td>
<td id="76B7374EC443A1C1FA973F59D018F83A" box="[1283,1347,1919,1938]" gridcol="6" gridrow="4" pageId="4" pageNumber="28">2.69</td>
</tr>
<tr id="35665E32C443A1C1FF5C3F82D018F810" box="[200,1347,1956,1976]" gridrow="5" pageId="4" pageNumber="28">
<th id="76B7374EC443A1C1FF5C3F82D40DF810" box="[200,342,1956,1976]" gridcol="0" gridrow="5" pageId="4" pageNumber="28">Femur / БеΑро</th>
<td id="76B7374EC443A1C1FE2C3F82D74BF810" box="[440,528,1956,1976]" gridcol="1" gridrow="5" pageId="4" pageNumber="28">1.99</td>
<td id="76B7374EC443A1C1FDC43F82D79FF810" box="[592,708,1956,1976]" gridcol="2" gridrow="5" pageId="4" pageNumber="28">1.99</td>
<td id="76B7374EC443A1C1FC843F82D60BF810" box="[784,848,1956,1976]" gridcol="3" gridrow="5" pageId="4" pageNumber="28">2.42</td>
<td id="76B7374EC443A1C1FC3F3F82D158F810" box="[939,1027,1956,1976]" gridcol="4" gridrow="5" pageId="4" pageNumber="28">1.83</td>
<td id="76B7374EC443A1C1FBAF3F82D1E5F810" box="[1083,1214,1956,1976]" gridcol="5" gridrow="5" pageId="4" pageNumber="28">2.05</td>
<td id="76B7374EC443A1C1FA973F82D018F810" box="[1283,1347,1956,1976]" gridcol="6" gridrow="5" pageId="4" pageNumber="28">2.24</td>
</tr>
<tr id="35665E32C443A1C1FF5C3FEFD018F875" box="[200,1347,1993,2013]" gridrow="6" pageId="4" pageNumber="28">
<th id="76B7374EC443A1C1FF5C3FEFD40DF875" box="[200,342,1993,2013]" gridcol="0" gridrow="6" pageId="4" pageNumber="28">Coxa / Тазик</th>
<td id="76B7374EC443A1C1FE2C3FEFD74BF875" box="[440,528,1993,2013]" gridcol="1" gridrow="6" pageId="4" pageNumber="28">2.22</td>
<td id="76B7374EC443A1C1FDC43FEFD79FF875" box="[592,708,1993,2013]" gridcol="2" gridrow="6" pageId="4" pageNumber="28">2.32</td>
<td id="76B7374EC443A1C1FC843FEFD60BF875" box="[784,848,1993,2013]" gridcol="3" gridrow="6" pageId="4" pageNumber="28">2.44</td>
<td id="76B7374EC443A1C1FC3F3FEFD158F875" box="[939,1027,1993,2013]" gridcol="4" gridrow="6" pageId="4" pageNumber="28">2.23</td>
<td id="76B7374EC443A1C1FBAF3FEFD1E5F875" box="[1083,1214,1993,2013]" gridcol="5" gridrow="6" pageId="4" pageNumber="28">2.50</td>
<td id="76B7374EC443A1C1FA973FEFD018F875" box="[1283,1347,1993,2013]" gridcol="6" gridrow="6" pageId="4" pageNumber="28">2.62</td>
</tr>
</table>
</paragraph>
<caption id="DF290CF8C4425E3BFF793F13D6A2F84B" ID-DOI="http://doi.org/10.5281/zenodo.8145600" ID-Zenodo-Dep="8145600" httpUri="https://zenodo.org/record/8145600/files/figure.png" pageId="5" pageNumber="29" startId="5.[237,268,1845,1862]" targetBox="[193,1398,278,1756]" targetPageId="5" targetType="figure">
<paragraph id="8BE95C70C4425E3BFF793F13D767F8EE" blockId="5.[189,1400,1845,2019]" box="[237,572,1845,1862]" pageId="5" pageNumber="29">
Figs 915.
<taxonomicName id="4C5627F3C4425E3BFED43F13D757F8EE" authority="Kaplin, 2019" authorityName="Kaplin" authorityYear="2019" box="[320,524,1845,1862]" class="Insecta" family="Machilidae" genus="Trigoniophthalmus" kingdom="Animalia" order="Archaeognatha" pageId="5" pageNumber="29" phylum="Arthropoda" rank="species" species="kobani" status="sp. n.">
<emphasis id="B9228062C4425E3BFED43F13D757F8EE" box="[320,524,1845,1862]" italics="true" pageId="5" pageNumber="29">Trigoniophthalmus kobani</emphasis>
</taxonomicName>
<emphasis id="B9228062C4425E3BFD853F10D767F8EE" bold="true" box="[529,572,1846,1862]" pageId="5" pageNumber="29">
<taxonomicNameLabel id="A2113D19C4425E3BFD853F10D767F8EE" box="[529,572,1846,1862]" pageId="5" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BE95C70C4425E3BFF793F6AD4FDF821" blockId="5.[189,1400,1845,2019]" pageId="5" pageNumber="29">12 male, holotype; 911, 1315 female, paratype; 9 pronotum; 10 urosternite and urocoxites IV; 11 urosternite and urocoxites VII; 12 male genitalia with urocoxite IX; 13 ovipositor with urocoxite IX; 14 apex of anterior gonapophysis of ovipositor; 15 apex of posterior gonapophysis of ovipositor.Scale bars 0.1 mm.</paragraph>
<paragraph id="8BE95C70C4425E3BFF793FA9D764F837" blockId="5.[189,1400,1845,2019]" box="[237,575,1935,1952]" pageId="5" pageNumber="29">
Рис. 915.
<taxonomicName id="4C5627F3C4425E3BFED73FA9D754F808" authority="Kaplin, 2019" authorityName="Kaplin" authorityYear="2019" box="[323,527,1935,1952]" class="Insecta" family="Machilidae" genus="Trigoniophthalmus" kingdom="Animalia" order="Archaeognatha" pageId="5" pageNumber="29" phylum="Arthropoda" rank="species" species="kobani" status="sp. n.">
<emphasis id="B9228062C4425E3BFED73FA9D754F808" box="[323,527,1935,1952]" italics="true" pageId="5" pageNumber="29">Trigoniophthalmus kobani</emphasis>
</taxonomicName>
<emphasis id="B9228062C4425E3BFD803FA9D764F837" bold="true" box="[532,575,1935,1951]" pageId="5" pageNumber="29">
<taxonomicNameLabel id="A2113D19C4425E3BFD803FA9D764F837" box="[532,575,1935,1951]" pageId="5" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BE95C70C4425E3BFF793F80D6A2F84B" blockId="5.[189,1400,1845,2019]" pageId="5" pageNumber="29">12 самец, гоΛотип; 911, 1315 самка, паратип; 9 переΑнеспинка; 10 стернит и кокситы IV сегмента брюшка; 11 стернит и кокситы VII сегмента брюшка; 12 поΛовой аппарат самца с кокситом IX сегмента брюшка; 13 яйцекΛаΑ с кокситом IX сегмента брюшка; 14 вершина переΑней створки яйцекΛаΑа; 15 вершина заΑней створки яйцекΛаΑа.Масштабные Λинейки 0.1 мм.</paragraph>
</caption>
<caption id="DF290CF8C4415E38FF793F72D4A0F843" ID-DOI="http://doi.org/10.5281/zenodo.8145602" ID-Zenodo-Dep="8145602" httpUri="https://zenodo.org/record/8145602/files/figure.png" pageId="6" pageNumber="30" startId="6.[237,268,1876,1893]" targetBox="[192,1394,273,1833]" targetPageId="6" targetType="figure">
<paragraph id="8BE95C70C4415E38FF793F72D7E8F8CC" blockId="6.[188,1399,1876,2028]" box="[237,691,1876,1893]" pageId="6" pageNumber="30">
Figs 1623.
<taxonomicName id="4C5627F3C4415E38FEDD3F72D75FF8CD" authority="Kaplin, 2019" authorityName="Kaplin" authorityYear="2019" box="[329,516,1876,1893]" class="Insecta" family="Machilidae" genus="Trigoniophthalmus" kingdom="Animalia" order="Archaeognatha" pageId="6" pageNumber="30" phylum="Arthropoda" rank="species" species="tseyi" status="sp. n.">
<emphasis id="B9228062C4415E38FEDD3F72D75FF8CD" box="[329,516,1876,1893]" italics="true" pageId="6" pageNumber="30">Trigoniophthalmus tseyi</emphasis>
</taxonomicName>
<emphasis id="B9228062C4415E38FD9D3F72D76FF8CC" bold="true" box="[521,564,1876,1892]" pageId="6" pageNumber="30">
<taxonomicNameLabel id="A2113D19C4415E38FD9D3F72D76FF8CC" box="[521,564,1876,1892]" pageId="6" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male, holotype.
</paragraph>
<paragraph id="8BE95C70C4415E38FF793F4CD64AF839" blockId="6.[188,1399,1876,2028]" pageId="6" pageNumber="30">16 distal chain of antennal flagellum; 17 apex of cercus; 18 eyes and paired ocelli; 19 maxillary palp; 20 labial palp;21 distal part of mandible; 22 fore leg (part); 23 male genitalia with urocoxite IX. Scale bars 0.1 mm.</paragraph>
<paragraph id="8BE95C70C4415E38FF793FBED7E4F800" blockId="6.[188,1399,1876,2028]" box="[237,703,1943,1960]" pageId="6" pageNumber="30">
Рис. 1623.
<taxonomicName id="4C5627F3C4415E38FED83FB1D753F800" authority="Kaplin, 2019" authorityName="Kaplin" authorityYear="2019" box="[332,520,1943,1960]" class="Insecta" family="Machilidae" genus="Trigoniophthalmus" kingdom="Animalia" order="Archaeognatha" pageId="6" pageNumber="30" phylum="Arthropoda" rank="species" species="tseyi" status="sp. n.">
<emphasis id="B9228062C4415E38FED83FB1D753F800" box="[332,520,1943,1960]" italics="true" pageId="6" pageNumber="30">Trigoniophthalmus tseyi</emphasis>
</taxonomicName>
<emphasis id="B9228062C4415E38FD983FB1D76CF80F" bold="true" box="[524,567,1943,1959]" pageId="6" pageNumber="30">
<taxonomicNameLabel id="A2113D19C4415E38FD983FB1D76CF80F" box="[524,567,1943,1959]" pageId="6" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, самец, гоΛотип.
</paragraph>
<paragraph id="8BE95C70C4415E38FF793F88D4A0F843" blockId="6.[188,1399,1876,2028]" pageId="6" pageNumber="30">16 цепочка ΑистаΛьной части жгутика усиков; 17 вершина церки; 18 гΛаза и парные гΛазки; 19 нижнечеΛюстной щупик; 20 нижнегубной щупик; 21 ΑистаΛьная часть верхней чеΛюсти; 22 переΑняя нога (часть); 23 поΛовой аппарат самца с кокситом IX сегмента брюшка.Масштабные Λинейки 0.1 мм.</paragraph>
</caption>
<paragraph id="8BE95C70C4405E39FF79398DD7CBFD24" blockId="7.[189,776,273,729]" pageId="7" pageNumber="31">
Ovipositor slender, elongate (
<quantity id="4CAEF195C4405E39FD99398DD7D8FE16" box="[525,643,427,446]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.1" metricValueMin="1.6" pageId="7" pageNumber="31" unit="mm" value="1.85" valueMax="2.1" valueMin="1.6">1.62.1 mm</quantity>
), slightly not surpassing apex of styli IX (
<figureCitation id="136D40F5C4405E39FE4039E3D742FE70" box="[468,537,453,472]" captionStart="Figs 2428" captionStartId="8.[237,268,1896,1913]" captionTargetBox="[198,1386,277,1875]" captionTargetId="figure-104@8.[189,1398,272,1877]" captionTargetPageId="8" captionText="Figs 2428. Trigoniophthalmus tseyi sp. n., female, paratype. 24 urosternite and urocoxites IV; 25 urosternite and urocoxites VII; 26 ovipositor with urocoxite IX; 27 apex of anterior gonapophysis of ovipositor; 28 apex of posterior gonapophysis of ovipositor.Scale bars 0.1 mm. Рис. 2428. Trigoniophthalmus tseyi sp. n., самка, паратип. 24 стернит и кокситы IV сегмента брюшка; 25 стернит и кокситы VII сегмента брюшка; 26 яйцекΛаΑ с кокситом IX сегмента брюшка; 27 вершина переΑних створок яйцекΛаΑа; 28 вершина заΑних створок яйцекΛаΑа. Масштабные Λинейки 0.1 мм." figureDoi="http://doi.org/10.5281/zenodo.8145604" httpUri="https://zenodo.org/record/8145604/files/figure.png" pageId="7" pageNumber="31">Fig. 26</figureCitation>
). Anterior and posterior gonapophyses with approximately 47 and 50 divisions, respectively. Two basal divisions of anterior gonapophyses and about 27 basal divisions of posterior gonapophyses glabrous. Apical macrochaetae of gonapophyses as long as four apical divisions combined. Distal divisions of anterior gonapophyses with 46, posterior gonapophyses with 25 setae (not counting sensory setae and apical macrochaetae) (
<figureCitation id="136D40F5C4405E39FDB13A5FD7DDFD24" box="[549,646,633,652]" captionStart="Figs 2428" captionStartId="8.[237,268,1896,1913]" captionTargetBox="[198,1386,277,1875]" captionTargetId="figure-104@8.[189,1398,272,1877]" captionTargetPageId="8" captionText="Figs 2428. Trigoniophthalmus tseyi sp. n., female, paratype. 24 urosternite and urocoxites IV; 25 urosternite and urocoxites VII; 26 ovipositor with urocoxite IX; 27 apex of anterior gonapophysis of ovipositor; 28 apex of posterior gonapophysis of ovipositor.Scale bars 0.1 mm. Рис. 2428. Trigoniophthalmus tseyi sp. n., самка, паратип. 24 стернит и кокситы IV сегмента брюшка; 25 стернит и кокситы VII сегмента брюшка; 26 яйцекΛаΑ с кокситом IX сегмента брюшка; 27 вершина переΑних створок яйцекΛаΑа; 28 вершина заΑних створок яйцекΛаΑа. Масштабные Λинейки 0.1 мм." figureDoi="http://doi.org/10.5281/zenodo.8145604" httpUri="https://zenodo.org/record/8145604/files/figure.png" pageId="7" pageNumber="31">Figs 27, 28</figureCitation>
).
</paragraph>
<paragraph id="8BE95C70C4405E39FF793AB5D1B3FE96" blockId="7.[189,776,273,729]" lastBlockId="7.[813,1400,273,730]" pageId="7" pageNumber="31">
Male genitalia with one pair of parameres on abdominal segment IX. Parameres with 1 + 7 divisions, penis slightly not attaining apex of parameres (
<figureCitation id="136D40F5C4405E39FE743AE0D77CFD71" box="[480,551,710,729]" captionStart="Figs 1623" captionStartId="6.[237,268,1876,1893]" captionTargetBox="[192,1394,273,1833]" captionTargetId="figure-1@6.[189,1398,272,1848]" captionTargetPageId="6" captionText="Figs 1623. Trigoniophthalmus tseyi sp. n., male, holotype. 16 distal chain of antennal flagellum; 17 apex of cercus; 18 eyes and paired ocelli; 19 maxillary palp; 20 labial palp;21 distal part of mandible; 22 fore leg (part); 23 male genitalia with urocoxite IX. Scale bars 0.1 mm. Рис. 1623. Trigoniophthalmus tseyi sp. n., самец, гоΛотип. 16 цепочка ΑистаΛьной части жгутика усиков; 17 вершина церки; 18 гΛаза и парные гΛазки; 19 нижнечеΛюстной щупик; 20 нижнегубной щупик; 21 ΑистаΛьная часть верхней чеΛюсти; 22 переΑняя нога (часть); 23 поΛовой аппарат самца с кокситом IX сегмента брюшка.Масштабные Λинейки 0.1 мм." figureDoi="http://doi.org/10.5281/zenodo.8145602" httpUri="https://zenodo.org/record/8145602/files/figure.png" pageId="7" pageNumber="31">Fig. 23</figureCitation>
). Penis and parameres not attaining of apex of coxites IX for 4.6 of width of aedeagus. Phallobasis of penis 1.3 times as long as aedeagus.
</paragraph>
</subSubSection>
<subSubSection id="C34C0FFBC4405E39FCC83960D1F9FD37" pageId="7" pageNumber="31" type="diagnosis">
<paragraph id="8BE95C70C4405E39FCC83960D1D1FDEE" blockId="7.[813,1400,273,730]" pageId="7" pageNumber="31">
<emphasis id="B9228062C4405E39FCC83960D11CFEF4" bold="true" box="[860,1095,326,348]" pageId="7" pageNumber="31">Differential diagnosis.</emphasis>
<taxonomicName id="4C5627F3C4405E39FBD83960D060FEF4" authority="Kaplin, 2019" authorityName="Kaplin" authorityYear="2019" box="[1100,1339,326,348]" class="Insecta" family="Machilidae" genus="Trigoniophthalmus" kingdom="Animalia" order="Archaeognatha" pageId="7" pageNumber="31" phylum="Arthropoda" rank="species" species="tseyi" status="sp. n.">
<emphasis id="B9228062C4405E39FBD83960D060FEF4" box="[1100,1339,326,348]" italics="true" pageId="7" pageNumber="31">Trigoniophthalmus tseyi</emphasis>
</taxonomicName>
<emphasis id="B9228062C4405E39FAD43961D02DFEF4" bold="true" box="[1344,1398,327,348]" pageId="7" pageNumber="31">
<taxonomicNameLabel id="A2113D19C4405E39FAD43961D02DFEF4" box="[1344,1398,327,348]" pageId="7" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
belongs to the same species group as
<taxonomicName id="4C5627F3C4405E39FB583942D076FED1" authority="Kaplin, 2019" authorityName="Kaplin" authorityYear="2019" box="[1228,1325,356,377]" class="Insecta" family="Machilidae" genus="Trigoniophthalmus" kingdom="Animalia" order="Archaeognatha" pageId="7" pageNumber="31" phylum="Arthropoda" rank="species" species="kobani" status="sp. n.">
<emphasis id="B9228062C4405E39FB583942D076FED1" box="[1228,1325,356,377]" italics="true" pageId="7" pageNumber="31">T. kobani</emphasis>
</taxonomicName>
<emphasis id="B9228062C4405E39FAAD3942D02DFED1" bold="true" box="[1337,1398,356,377]" pageId="7" pageNumber="31">
<taxonomicNameLabel id="A2113D19C4405E39FAAD3942D02DFED1" box="[1337,1398,356,377]" pageId="7" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
described above.
<taxonomicName id="4C5627F3C4405E39FC7339A7D184FE3F" authority="Kaplin, 2019" authorityName="Kaplin" authorityYear="2019" box="[999,1247,385,407]" class="Insecta" family="Machilidae" genus="Trigoniophthalmus" kingdom="Animalia" order="Archaeognatha" pageId="7" pageNumber="31" phylum="Arthropoda" rank="species" species="tseyi" status="sp. n.">
<emphasis id="B9228062C4405E39FC7339A7D184FE3F" box="[999,1247,385,407]" italics="true" pageId="7" pageNumber="31">Trigoniophthalmus tseyi</emphasis>
</taxonomicName>
<emphasis id="B9228062C4405E39FB7839A7D070FE3E" bold="true" box="[1260,1323,385,406]" pageId="7" pageNumber="31">
<taxonomicNameLabel id="A2113D19C4405E39FB7839A7D070FE3E" box="[1260,1323,385,406]" pageId="7" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from other species of this group by ratio of distance between inner margins of paired ocelli to total width of eyes, presence of spine-like setae on legs, small posterior angle of urosternites IIV, large number of inner sublateral spines on urocoxite IX. Main differences between these species of this group are in
<tableCitation id="C6D469CBC4405E39FBA93A17D1DCFDEE" box="[1085,1159,561,583]" captionStart="Table 8" captionStartId="9.[237,279,273,290]" captionTargetBox="[200,1384,348,2017]" captionTargetId="graphics-0@9.[189,1396,327,2023]" captionText="Table 8. The main morphological differences between Trigoniophthalmus kobani sp. n., T. tseyi sp. n. and their closest congeners. ТабΛица 8. Основные морфоΛогические отΛичия межΑу Trigoniophthalmus kobani sp. n., T. tseyi sp. n. и схоΑными с ними виΑами." pageId="7" pageNumber="31">Table 8</tableCitation>
.
</paragraph>
<paragraph id="8BE95C70C4405E39FCC83A68D1F9FD37" blockId="7.[813,1400,273,730]" pageId="7" pageNumber="31">
<emphasis id="B9228062C4405E39FCC83A68D6E6FDCC" bold="true" box="[860,957,590,612]" pageId="7" pageNumber="31">Habitats.</emphasis>
Specimens of
<taxonomicName id="4C5627F3C4405E39FBDE3A68D060FDCC" authority="Kaplin, 2019" authorityName="Kaplin" authorityYear="2019" box="[1098,1339,590,612]" class="Insecta" family="Machilidae" genus="Trigoniophthalmus" kingdom="Animalia" order="Archaeognatha" pageId="7" pageNumber="31" phylum="Arthropoda" rank="species" species="tseyi" status="sp. n.">
<emphasis id="B9228062C4405E39FBDE3A68D060FDCC" box="[1098,1339,590,612]" italics="true" pageId="7" pageNumber="31">Trigoniophthalmus tseyi</emphasis>
</taxonomicName>
<emphasis id="B9228062C4405E39FAD43A69D02DFDCC" bold="true" box="[1344,1398,591,612]" pageId="7" pageNumber="31">
<taxonomicNameLabel id="A2113D19C4405E39FAD43A69D02DFDCC" box="[1344,1398,591,612]" pageId="7" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
were collected in mountain forest (Pinus, Fagus, shrubs) under stones,
<quantity id="4CAEF195C4405E39FC2D3AAFD158FD37" box="[953,1027,649,671]" metricMagnitude="3" metricUnit="m" metricValue="1.95" pageId="7" pageNumber="31" unit="m" value="1950.0">1950 m</quantity>
above sea level.
</paragraph>
</subSubSection>
<subSubSection id="C34C0FFBC4405E39FCC83A81D060FD72" pageId="7" pageNumber="31" type="etymology">
<paragraph id="8BE95C70C4405E39FCC83A81D060FD72" blockId="7.[813,1400,273,730]" pageId="7" pageNumber="31">
<emphasis id="B9228062C4405E39FCC83A81D688FD14" bold="true" box="[860,979,679,700]" pageId="7" pageNumber="31">Etymology.</emphasis>
The new species takes its name from the
<typeStatus id="54EDE2D2C4405E39FCBA3AE2D602FD71" box="[814,857,708,729]" pageId="7" pageNumber="31">type</typeStatus>
locality: Verkhniy Tsey,
<collectingRegion id="49929292C4405E39FBD83AE2D182FD72" box="[1100,1241,708,730]" country="Russia" name="North Ossetia" pageId="7" pageNumber="31">North Ossetia</collectingRegion>
<collectingRegion id="49929292C4405E39FB673AE2D06CFD72" box="[1267,1335,708,730]" country="Russia" name="North Ossetia" pageId="7" pageNumber="31">Alania</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>