338 lines
36 KiB
XML
338 lines
36 KiB
XML
<document id="BCD11621AF84BF0196809B8108E112E6" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFFFC734132FAF98BE622FD16" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium haackeri Wesener 2009, new species" docType="treatment" docUuid="CB1CAAF3-3BEC-48F5-95E4-F0CFED200AC6" docUuidSource="ZooBank" docVersion="5" lastPageNumber="18" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="16" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="D4342F0BB0A785B1EAC09552094133E1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="653333385F65D08CCB61D734069042FE">
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<mods:title id="6600BC1CB88FD417ED0DF81E31532082">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
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</mods:titleInfo>
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<mods:name id="D2A0BB4CE02583DF28E2DA74035C37C6" type="personal">
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<mods:role id="D50214378DA628484D9CD8C0906E02A5">
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<mods:roleTerm id="F0F7973A6F8E427AA931488F0490C20B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="29FE1C4AD43C5B85578B73E925AC87E2">Wesener, Thomas</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="104441BBCF782C058E4D13F5EB8942B7">text</mods:typeOfResource>
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<mods:titleInfo id="3053288EFBF0AA01726F0C1139337D1F">
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<mods:title id="F89D8FBCF818CBCBF1EC0435638AEE9C">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="9571F0C9EFAB953C80E4DC8EF11AD352">
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<mods:date id="17C9C62050C98D9F417717946DFD9CE9">2009</mods:date>
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<mods:detail id="5B6CAF4CB3D6E612DC5D9EFBA919756C" type="pubDate">
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<mods:number id="4B33C7AE50A666C9D6745D7A64C758D4">2009-05-11</mods:number>
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</mods:detail>
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<mods:detail id="FBBF665D382F286677AF5FD4128F4CEE" type="volume">
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<mods:number id="C33DABF7854C00F408792C0F1C4E4649">2097</mods:number>
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</mods:detail>
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<mods:detail id="FBF3595EB07B2C66E61037EC8A31AD1C" type="issue">
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<mods:number id="4EAA08A413A62090D170AA058A3F2156">1</mods:number>
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</mods:detail>
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<mods:extent id="393807B0744FAD12E7BEDB437B711156" unit="page">
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<mods:start id="04B5C24D1BC37F9CE6606073A396F2E6">1</mods:start>
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<mods:end id="862DDFBA9E25E587B821BE38A46F7283">134</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="D3CEF5ECF2722BB2CC214BA319C9B90F">
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<mods:url id="8FE1FB0CE6CF4E0E625C3225878CFC74">https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1</mods:url>
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</mods:location>
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<mods:classification id="8A186BF64DC4C059CD0FEBB7E3F8EBD5">journal article</mods:classification>
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<mods:identifier id="3F0174AE7A39F9EBA1551A5697862C51" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
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<mods:identifier id="2C93461DC1521696F4A7163DC136077B" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="0A432A8FC2804EF810301A72EB2CA108" type="Zenodo-Dep">5316536</mods:identifier>
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<mods:identifier id="B1FDD8B0276442CA761C899E20EC8555" type="ZooBank">0E18167A-78EB-4079-8BF4-972A4AC88C69</mods:identifier>
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</mods:mods>
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<treatment id="03FFBE1DFFFC734132FAF98BE622FD16" ID-DOI="http://doi.org/10.5281/zenodo.5325443" ID-Zenodo-Dep="5325443" LSID="urn:lsid:zoobank.org:act:CB1CAAF3-3BEC-48F5-95E4-F0CFED200AC6" httpUri="http://treatment.plazi.org/id/03FFBE1DFFFC734132FAF98BE622FD16" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16">
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<subSubSection id="C34C5C80FFFC735F32FAF98BE151F993" box="[151,598,1584,1611]" pageId="15" pageNumber="16" type="nomenclature">
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<paragraph id="8BE90F0BFFFC735F32FAF98BE151F993" blockId="15.[151,598,1584,1611]" box="[151,598,1584,1611]" pageId="15" pageNumber="16">
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<heading id="D0A1B867FFFC735F32FAF98BE151F993" bold="true" box="[151,598,1584,1611]" fontSize="11" level="1" pageId="15" pageNumber="16" reason="1">
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||
<emphasis id="B922D319FFFC735F32FAF98BE151F993" bold="true" box="[151,598,1584,1611]" pageId="15" pageNumber="16">
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<emphasis id="B922D319FFFC735F32FAF98BE2B8F992" bold="true" box="[151,447,1585,1611]" italics="true" pageId="15" pageNumber="16">
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||
<taxonomicName id="4C567488FFFC735F32FAF98BE2BEF992" ID-CoL="5D7H5" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,441,1585,1611]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="haackeri" status="sp. nov.">Zoosphaerium haackeri</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A2116E62FFFC735F33ABF98AE151F993" box="[454,598,1584,1610]" pageId="15" pageNumber="16" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34C5C80FFFC734032FAF9C0E3D0FF01" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="etymology">
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<paragraph id="8BE90F0BFFFC735F32FAF9C0E7D0F962" blockId="15.[151,1436,1658,2032]" pageId="15" pageNumber="16">
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<emphasis id="B922D319FFFC735F32FAF9C0E289F94D" bold="true" box="[151,398,1658,1684]" pageId="15" pageNumber="16">Derivatio nominis:</emphasis>
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<taxonomicName id="4C567488FFFC735F33CEF9C1E10DF94D" authorityName="Wesener" authorityYear="2009" box="[419,522,1659,1684]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="haackeri">
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<emphasis id="B922D319FFFC735F33CEF9C1E10DF94D" box="[419,522,1659,1684]" italics="true" pageId="15" pageNumber="16">haackeri</emphasis>
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</taxonomicName>
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, adjective, is named to honour myriapodologist Dr. Ulrich Haacker (1939–1972), who did excellent research on the behaviour of South African
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<taxonomicName id="4C567488FFFC735F367BF91BE7D4F962" box="[1046,1235,1697,1723]" class="Diplopoda" kingdom="Animalia" order="Sphaerotheriida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="order">Sphaerotheriida</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BE90F0BFFFC735F32ABF97DE71DF8F6" blockId="15.[151,1436,1658,2032]" pageId="15" pageNumber="16">
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<emphasis id="B922D319FFFC735F32ABF97DE298F938" bold="true" box="[198,415,1735,1761]" pageId="15" pageNumber="16">Studied material:</emphasis>
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<materialsCitation id="3B3E0556FFFC735F33CBF97DE71DF8F6" collectingDate="2002-02-11" collectingDateMax="2002-02-15" collectingDateMin="2002-02-11" collectionCode="CAS" collectorName="Fisher & Griswold" country="Madagascar" county="Locus" elevation="220" latitude="-25.581667" location="Reserve Speciale de Cap Sainte Marie" longLatPrecision="21" longitude="45.168335" municipality="Province" pageId="15" pageNumber="16" specimenCode="BLF 5499" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
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<emphasis id="B922D319FFFC735F33CBF97DE11FF938" bold="true" box="[422,536,1735,1761]" pageId="15" pageNumber="16">
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<typeStatus id="54EDB1A9FFFC735F33CBF97DE11FF938" box="[422,536,1735,1761]" pageId="15" pageNumber="16" type="holotype">Holotype</typeStatus>
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</emphasis>
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: 1 M (
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<quantity id="4CAEA2EEFFFC735F3008F972E1C8F93B" box="[613,719,1736,1762]" metricMagnitude="-2" metricUnit="m" metricValue="2.48" pageId="15" pageNumber="16" unit="mm" value="24.8">24.8 mm</quantity>
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long),
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<emphasis id="B922D319FFFC735F314CF97DE0DBF938" bold="true" box="[801,988,1735,1761]" pageId="15" pageNumber="16">
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<collectionCode id="ED4797CEFFFC735F314CF97DE05CF938" box="[801,859,1735,1761]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="15" pageNumber="16" type="Museum">CAS</collectionCode>
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<specimenCode id="DBF0A770FFFC735F310FF97DE0DBF938" box="[866,988,1735,1761]" collectionCode="BLF" pageId="15" pageNumber="16">BLF 5499</specimenCode>
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</emphasis>
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.
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<emphasis id="B922D319FFFC735F3186F972E78DF938" box="[1003,1162,1736,1761]" italics="true" pageId="15" pageNumber="16">
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<collectingCounty id="62887787FFFC735F3186F972E736F938" box="[1003,1073,1736,1761]" pageId="15" pageNumber="16">Locus</collectingCounty>
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typicus
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</emphasis>
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:
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<collectingCountry id="F3414F9BFFFC735F36F4F972E62EF93B" box="[1177,1321,1736,1762]" name="Madagascar" pageId="15" pageNumber="16">Madagascar</collectingCountry>
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,
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<collectingMunicipality id="6B8D9571FFFC735F3759F972E69BF93B" box="[1332,1436,1736,1762]" pageId="15" pageNumber="16">Province</collectingMunicipality>
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<collectingRegion id="4992C1E9FFFC735F32FAF954E3EAF8D1" box="[151,237,1774,1800]" country="Madagascar" name="Toliara" pageId="15" pageNumber="16">Toliara</collectingRegion>
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,
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||
<location id="8E8959D0FFFC735F3291F954E1D7F8D1" LSID="urn:lsid:plazi:treatment:03FFBE1DFFFC734132FAF98BE622FD16:8E8959D0FFFC735F3291F954E1D7F8D1" box="[252,720,1774,1800]" country="Madagascar" county="Locus" latitude="-25.581667" longLatPrecision="21" longitude="45.168335" municipality="Province" name="Reserve Speciale de Cap Sainte Marie" pageId="15" pageNumber="16" stateProvince="Toliara">Réserve Spéciale de Cap Sainte Marie</location>
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||
,
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<locationDeviation id="9A5B1629FFFC735F30B2F954E72BF8D1" box="[735,1068,1774,1800]" location="Marovato" pageId="15" pageNumber="16">
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||
<quantity id="4CAEA2EEFFFC735F30B2F954E045F8D1" box="[735,834,1774,1800]" metricMagnitude="4" metricUnit="m" metricValue="1.23" pageId="15" pageNumber="16" unit="km" value="12.3">12.3 km</quantity>
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262° W Marovato
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||
</locationDeviation>
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,
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<geoCoordinate id="EE6269CCFFFC735F3656F954E7C4F8D1" box="[1083,1219,1774,1800]" degrees="25" direction="south" minutes="34" orientation="latitude" pageId="15" pageNumber="16" precision="15" seconds="54" value="-25.581667">25°34'54"S</geoCoordinate>
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||
,
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<geoCoordinate id="EE6269CCFFFC735F36BFF954E64AF8D1" box="[1234,1357,1774,1800]" degrees="45" direction="east" minutes="10" orientation="longitude" pageId="15" pageNumber="16" precision="15" seconds="6" value="45.168335">45°10'6"E</geoCoordinate>
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||
, spiny forest thicket,
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||
<quantity id="4CAEA2EEFFFC735F3352F8AFE28CF8F6" box="[319,395,1813,1839]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="15" pageNumber="16" unit="m" value="220.0">
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||
<elevation id="007BE838FFFC735F3352F8AFE28CF8F6" box="[319,395,1813,1839]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="15" pageNumber="16" unit="m" value="220.0">220 m</elevation>
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||
</quantity>
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||
, coll.
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<collectorName id="26A36ADDFFFC735F33BFF8AFE11AF8F6" box="[466,541,1813,1839]" pageId="15" pageNumber="16">Fisher</collectorName>
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,
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<collectorName id="26A36ADDFFFC735F3044F8AFE191F8F6" box="[553,662,1813,1839]" pageId="15" pageNumber="16">Griswold</collectorName>
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<emphasis id="B922D319FFFC735F30F0F8AFE1DFF8F7" box="[669,728,1813,1838]" italics="true" pageId="15" pageNumber="16">et al.</emphasis>
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,
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<date id="FFE829CBFFFC735F308AF8AFE08CF8F6" box="[743,907,1813,1839]" pageId="15" pageNumber="16" value="2002-02-11" valueMax="2002-02-15" valueMin="2002-02-11">
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||
<collectingDate id="EFACD023FFFC735F308AF8AFE08CF8F6" box="[743,907,1813,1839]" pageId="15" pageNumber="16" value="2002-02-11" valueMax="2002-02-15" valueMin="2002-02-11">11–15.ii.2002</collectingDate>
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</date>
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,
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<collectingMethod id="5217771CFFFC735F31FAF8AFE711F8F6" box="[919,1046,1813,1839]" pageId="15" pageNumber="16">pitfall trap</collectingMethod>
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.
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</materialsCitation>
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||
</paragraph>
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<paragraph id="8BE90F0BFFFC735F32AAF881E183F88F" blockId="15.[151,1436,1658,2032]" box="[199,644,1851,1878]" pageId="15" pageNumber="16">
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<emphasis id="B922D319FFFC735F32AAF881E23EF88C" bold="true" box="[199,313,1851,1877]" pageId="15" pageNumber="16">
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<typeStatus id="54EDB1A9FFFC735F32AAF881E23EF88C" box="[199,313,1851,1877]" pageId="15" pageNumber="16" type="paratype">Paratype</typeStatus>
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</emphasis>
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: 1 F, same data as holotype.
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</paragraph>
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<paragraph id="8BE90F0BFFFC734032AAF8D8E3D0FF01" blockId="15.[151,1436,1658,2032]" lastBlockId="16.[151,1436,151,2034]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
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||
<emphasis id="B922D319FFFC735F32AAF8D8E288F8A5" bold="true" box="[199,399,1890,1916]" pageId="15" pageNumber="16">Other material:</emphasis>
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||
3 M, 1 F,
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||
<materialsCitation id="3B3E0556FFFC735F3079F8D8E00FF87A" collectingDate="2007-06-18" collectionCode="FMMC, T" collectorName="Wesener" elevation="27" latitude="-25.4234" location="Lavanono Beach" longLatPrecision="1" longitude="44.9404" pageId="15" pageNumber="16" specimenCount="2">
|
||
<specimenCount id="9D50C482FFFC735F3079F8D8E16DF8A5" box="[532,618,1890,1916]" pageId="15" pageNumber="16" type="generic">2 imm.</specimenCount>
|
||
<emphasis id="B922D319FFFC735F3019F8D8E1D5F8A5" bold="true" box="[628,722,1890,1916]" pageId="15" pageNumber="16">
|
||
<collectionCode id="ED4797CEFFFC735F3019F8D8E1D5F8A5" box="[628,722,1890,1916]" pageId="15" pageNumber="16">FMMC</collectionCode>
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||
</emphasis>
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||
,
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||
<location id="8E8959D0FFFC735F308EF8D8E0A9F8A5" LSID="urn:lsid:plazi:treatment:03FFBE1DFFFC734132FAF98BE622FD16:8E8959D0FFFC735F308EF8D8E0A9F8A5" box="[739,942,1890,1916]" latitude="-25.4234" longLatPrecision="1" longitude="44.9404" name="Lavanono Beach" pageId="15" pageNumber="16">Lavanono Beach</location>
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||
,
|
||
<geoCoordinate id="EE6269CCFFFC735F31D1F8D8E749F8A5" box="[956,1102,1890,1916]" degrees="25" direction="south" minutes="25.404" orientation="latitude" pageId="15" pageNumber="16" precision="1" value="-25.4234">25°25.404'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE6269CCFFFC735F3630F8D8E7F8F8A5" box="[1117,1279,1890,1916]" degrees="044" direction="east" minutes="56.424" orientation="longitude" pageId="15" pageNumber="16" precision="1" value="44.9404">044°56.424'E</geoCoordinate>
|
||
,
|
||
<quantity id="4CAEA2EEFFFC735F3760F8D8E64AF8A5" box="[1293,1357,1890,1916]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="15" pageNumber="16" unit="m" value="27.0">
|
||
<elevation id="007BE838FFFC735F3760F8D8E64AF8A5" box="[1293,1357,1890,1916]" metricMagnitude="1" metricUnit="m" metricValue="2.7" pageId="15" pageNumber="16" unit="m" value="27.0">27 m</elevation>
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||
</quantity>
|
||
, spiny bush close to the coast, coll.
|
||
<collectionCode id="ED4797CEFFFC735F3399F833E100F87A" box="[500,519,1929,1955]" name="Tavera, Department of Geology and Geophysics" pageId="15" pageNumber="16">T</collectionCode>
|
||
.
|
||
<collectorName id="26A36ADDFFFC735F3079F833E17DF87A" box="[532,634,1929,1955]" pageId="15" pageNumber="16">Wesener</collectorName>
|
||
,
|
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<date id="FFE829CBFFFC735F30E5F833E00FF87A" box="[648,776,1929,1955]" pageId="15" pageNumber="16" value="2007-06-18">
|
||
<collectingDate id="EFACD023FFFC735F30E5F833E00FF87A" box="[648,776,1929,1955]" pageId="15" pageNumber="16" value="2007-06-18">18.vi.2007</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
. 2 M, 1 F,
|
||
<materialsCitation id="3B3E0556FFFC734031E2F833E105FF6B" collectingDate="2002-02-21" collectingDateMax="2002-02-25" collectingDateMin="2002-02-21" collectionCode="CAS" collectorName="Fisher & Griswold" county="University" elevation="80" lastPageId="16" lastPageNumber="17" latitude="-24.65361" location="Mahafaly Plateau" longLatPrecision="21" longitude="43.996666" municipality="Province" pageId="15" pageNumber="16" specimenCode="BLF 5762" specimenCount="3" stateProvince="Antananarivo">
|
||
<specimenCount id="9D50C482FFFC735F31E2F833E0E0F87A" box="[911,999,1929,1955]" pageId="15" pageNumber="16" type="generic">3 imm.</specimenCount>
|
||
, voucher specimens deposited at the
|
||
<collectingCounty id="62887787FFFC735F32FAF80AE214F813" box="[151,275,1968,1994]" pageId="15" pageNumber="16">University</collectingCounty>
|
||
of
|
||
<collectingRegion id="4992C1E9FFFC735F332BF80AE2EFF813" box="[326,488,1968,1994]" country="Madagascar" name="Antananarivo" pageId="15" pageNumber="16">Antananarivo</collectingRegion>
|
||
, same collection data as previous. 2 M,
|
||
<emphasis id="B922D319FFFC735F3183F815E7B0F810" bold="true" box="[1006,1207,1967,1993]" pageId="15" pageNumber="16">
|
||
<collectionCode id="ED4797CEFFFC735F3183F815E72EF810" box="[1006,1065,1967,1993]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="15" pageNumber="16" type="Museum">CAS</collectionCode>
|
||
<specimenCode id="DBF0A770FFFC735F365BF815E7B0F810" box="[1078,1207,1967,1993]" collectionCode="BLF" pageId="15" pageNumber="16">BLF 5762</specimenCode>
|
||
</emphasis>
|
||
,
|
||
<collectingMunicipality id="6B8D9571FFFC735F36A1F80AE633F813" box="[1228,1332,1968,1994]" pageId="15" pageNumber="16">Province</collectingMunicipality>
|
||
<collectingRegion id="4992C1E9FFFC735F372FF80AE69FF813" box="[1346,1432,1968,1994]" country="Madagascar" name="Toliara" pageId="15" pageNumber="16">Toliara</collectingRegion>
|
||
,
|
||
<location id="8E8959D0FFFC735F32FAF86CE260F829" LSID="urn:lsid:plazi:treatment:03FFBE1DFFFC734132FAF98BE622FD16:8E8959D0FFFC735F32FAF86CE260F829" box="[151,359,2006,2032]" county="University" latitude="-24.65361" longLatPrecision="21" longitude="43.996666" municipality="Province" name="Mahafaly Plateau" pageId="15" pageNumber="16" stateProvince="Antananarivo">Mahafaly Plateau</location>
|
||
,
|
||
<locationDeviation id="9A5B1629FFFC735F331EF86CE1A0F829" box="[371,679,2006,2032]" location="Itampolo" pageId="15" pageNumber="16">
|
||
<quantity id="4CAEA2EEFFFC735F331EF86CE2C4F829" box="[371,451,2006,2032]" metricMagnitude="3" metricUnit="m" metricValue="6.2" pageId="15" pageNumber="16" unit="km" value="6.2">6.2 km</quantity>
|
||
74° ENE Itampolo
|
||
</locationDeviation>
|
||
,
|
||
<geoCoordinate id="EE6269CCFFFC735F30DEF86CE03DF829" box="[691,826,2006,2032]" degrees="24" direction="south" minutes="39" orientation="latitude" pageId="15" pageNumber="16" precision="15" seconds="13" value="-24.65361">24°39'13"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE6269CCFFFC735F312BF86CE0C8F829" box="[838,975,2006,2032]" degrees="43" direction="east" minutes="59" orientation="longitude" pageId="15" pageNumber="16" precision="15" seconds="48" value="43.996666">43°59'48"E</geoCoordinate>
|
||
, spiny forest thicket,
|
||
<quantity id="4CAEA2EEFFFC735F36A4F86CE601F829" box="[1225,1286,2006,2032]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="16" unit="m" value="80.0">
|
||
<elevation id="007BE838FFFC735F36A4F86CE601F829" box="[1225,1286,2006,2032]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="15" pageNumber="16" unit="m" value="80.0">80 m</elevation>
|
||
</quantity>
|
||
, coll.
|
||
<collectorName id="26A36ADDFFFC735F3720F86CE69FF829" box="[1357,1432,2006,2032]" pageId="15" pageNumber="16">Fisher</collectorName>
|
||
,
|
||
<collectorName id="26A36ADDFFE3734032FAFF22E203FF6B" box="[151,260,152,178]" pageId="16" pageNumber="17">Griswold</collectorName>
|
||
<emphasis id="B922D319FFE373403363FF22E24CFF68" box="[270,331,152,177]" italics="true" pageId="16" pageNumber="17">et al.</emphasis>
|
||
,
|
||
<date id="FFE829CBFFE373403331FF22E105FF6B" box="[348,514,152,178]" pageId="16" pageNumber="17" value="2002-02-21" valueMax="2002-02-25" valueMin="2002-02-21">
|
||
<collectingDate id="EFACD023FFE373403331FF22E105FF6B" box="[348,514,152,178]" pageId="16" pageNumber="17" value="2002-02-21" valueMax="2002-02-25" valueMin="2002-02-21">21–25.ii.2002</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
,
|
||
<materialsCitation id="3B3E0556FFE37340307DFF22E3D3FF01" collectingDate="2002-02-21" collectingDateMax="2002-02-25" collectingDateMin="2002-02-21" collectionCode="CAS" collectorName="Fisher & Griswold" county="University" elevation="80" latitude="-24.65361" location="Mahafaly Plateau" longLatPrecision="21" longitude="43.996666" municipality="Province" pageId="16" pageNumber="17" specimenCode="BLF 5763" specimenCount="1" stateProvince="Antananarivo">
|
||
general collecting; 1 M,
|
||
<emphasis id="B922D319FFE373403157FF2DE0FCFF68" bold="true" box="[826,1019,151,177]" pageId="16" pageNumber="17">
|
||
<collectionCode id="ED4797CEFFE373403157FF2DE073FF68" box="[826,884,151,177]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="16" pageNumber="17" type="Museum">CAS</collectionCode>
|
||
<specimenCode id="DBF0A770FFE373403113FF2DE0FCFF68" box="[894,1019,151,177]" collectionCode="BLF" pageId="16" pageNumber="17">BLF 5763</specimenCode>
|
||
</emphasis>
|
||
, same data as previous,
|
||
<collectingMethod id="5217771CFFE37340374BFF22E3D3FF01" pageId="16" pageNumber="17">but pitfall traps</collectingMethod>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C5C80FFE3734032ABFF5FE092FF26" box="[198,917,229,255]" pageId="16" pageNumber="17" type="distribution">
|
||
<paragraph id="8BE90F0BFFE3734032ABFF5FE092FF26" blockId="16.[151,1436,151,2034]" box="[198,917,229,255]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032ABFF5FE26EFF26" bold="true" box="[198,361,229,255]" pageId="16" pageNumber="17">Distribution:</emphasis>
|
||
Spiny forests in Southern
|
||
<collectingCountry id="F3414F9BFFE3734030CCFF5FE028FF26" box="[673,815,229,255]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
|
||
(Fig. 5).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C5C80FFE3734032ABFEB1E1E1FCC5" pageId="16" pageNumber="17" type="diagnosis">
|
||
<paragraph id="8BE90F0BFFE3734032ABFEB1E14DFD5B" blockId="16.[151,1436,151,2034]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032ABFEB1E24FFEFC" bold="true" box="[198,328,267,293]" pageId="16" pageNumber="17">Diagnosis:</emphasis>
|
||
Up to
|
||
<quantity id="4CAEA2EEFFE3734033C8FEB6E2FCFEFF" box="[421,507,268,294]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="17" unit="mm" value="28.0">28 mm</quantity>
|
||
long. Both sexes of same size. Body dark green, posterior margins of each tergite with yellow line (Fig. 6A). Texture of tergites smooth and glabrous. Tarsi of all legs broad, length 3 times larger than width (Fig. 7A). Antenna very short (Fig. 8A), without groove (Fig. 8B), disc with 9–15 apical cones (Fig. 9A). Fifth antennomere with
|
||
<emphasis id="B922D319FFE3734030B0FE3AE0C6FE40" box="[733,961,384,409]" italics="true" pageId="16" pageNumber="17">sensilla basiconica</emphasis>
|
||
(Fig. 8C). Movable finger of posterior telopods weakly curved, longer than fixed finger (Fig. 7G), on posterior side with
|
||
<emphasis id="B922D319FFE373403607FE1DE7A2FE19" box="[1130,1189,423,448]" italics="true" pageId="16" pageNumber="17">circa</emphasis>
|
||
20 large, sclerotized teeth, two large, non-sclerotized lobes and 3–5 thin spines (Fig. 7H). Fixed finger very wide, basally with large membranous lobe and small spine. Two stridulation ribs on each male harp (Fig. 7E) and two on each side of washboard. Anal shield without locking carinae. Operculum large, notched, dividing operculum into two, well-rounded, same-sized tips (Fig. 7C). Mandible with very small intermediate area between molar plate and pectinate lamella (Fig. 9C).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE3734032ABFD34E1E1FCC5" blockId="16.[151,1436,151,2034]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032ABFD34E28CFD71" bold="true" box="[198,395,654,680]" pageId="16" pageNumber="17">Similar species:</emphasis>
|
||
The species is in the anterior telopods very similar to
|
||
<taxonomicName id="4C567488FFE373403679FD35E7B9FD71" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[1044,1214,655,680]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="libidinosum">
|
||
<emphasis id="B922D319FFE373403679FD35E7B9FD71" box="[1044,1214,655,680]" italics="true" pageId="16" pageNumber="17">Z. libidinosum</emphasis>
|
||
</taxonomicName>
|
||
but
|
||
<taxonomicName id="4C567488FFE373403698FD35E67CFD71" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1269,1403,655,680]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="haackeri" status="sp. nov.">
|
||
<emphasis id="B922D319FFE373403698FD35E67CFD71" box="[1269,1403,655,680]" italics="true" pageId="16" pageNumber="17">Z. haackeri</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B922D319FFE3734037E9FD34E3BDFD16" bold="true" pageId="16" pageNumber="17">
|
||
<taxonomicNameLabel id="A2116E62FFE3734037E9FD34E3BDFD16" pageId="16" pageNumber="17" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from the latter in the following characters: the tarsi are of much greater width (Fig. 7A), the mandible of unique shape, without an intermediate area (Fig. 9C), the operculum is notched (Fig. 7C) and the posterior telopods are of different shape (Fig. 7G).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C5C80FFE3734132ABFC92E622FD16" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
|
||
<paragraph id="8BE90F0BFFE3734032ABFC92E004FC49" blockId="16.[151,1436,151,2034]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032ABFC92E267FC9B" bold="true" box="[198,352,808,834]" pageId="16" pageNumber="17">Description:</emphasis>
|
||
<emphasis id="B922D319FFE373403304FC93E2F0FC9B" box="[361,503,809,834]" italics="true" pageId="16" pageNumber="17">Body length</emphasis>
|
||
: Males (
|
||
<specimenCount id="9D50C482FFE37340300DFC93E1FFFC9A" box="[608,760,809,835]" pageId="16" pageNumber="17" type="generic">9 specimens</specimenCount>
|
||
): length up to
|
||
<quantity id="4CAEA2EEFFE3734031CFFC93E0FFFC9A" box="[930,1016,809,835]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="16" pageNumber="17" unit="mm" value="25.0">25 mm</quantity>
|
||
, width of thoracic shield up to
|
||
<quantity id="4CAEA2EEFFE373403704FC93E3CFFCB3" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="16" pageNumber="17" unit="mm" value="11.8">11.8 mm</quantity>
|
||
, height of thoracic shield up to
|
||
<quantity id="4CAEA2EEFFE373403025FCEAE1A0FCB3" box="[584,679,848,874]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="16" pageNumber="17" unit="mm" value="6.8">6.8 mm</quantity>
|
||
. Females (
|
||
<specimenCount id="9D50C482FFE373403144FCEAE0C6FCB3" box="[809,961,848,874]" pageId="16" pageNumber="17" type="generic">3 specimens</specimenCount>
|
||
): length up to
|
||
<quantity id="4CAEA2EEFFE37340361DFCEAE7C0FCB3" box="[1136,1223,848,874]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="16" pageNumber="17" unit="mm" value="26.0">26 mm</quantity>
|
||
, width up to
|
||
<quantity id="4CAEA2EEFFE373403704FCEAE3CFFC49" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="16" pageNumber="17" unit="mm" value="12.8">12.8 mm</quantity>
|
||
, height up to
|
||
<quantity id="4CAEA2EEFFE37340330BFCCCE2C4FC49" box="[358,451,886,912]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="16" pageNumber="17" unit="mm" value="7.2">7.2 mm</quantity>
|
||
. Both sexes of similar size.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE3734032AAFC27E732FC6E" blockId="16.[151,1436,151,2034]" box="[199,1077,925,951]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032AAFC27E223FC6F" box="[199,292,925,950]" italics="true" pageId="16" pageNumber="17">Habitus</emphasis>
|
||
: Anal shield well-rounded (Fig. 6A). Tergites smooth and hairless.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE3734032AAFC7EE018FBDD" blockId="16.[151,1436,151,2034]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032AAFC7EE24FFC04" box="[199,328,964,989]" italics="true" pageId="16" pageNumber="17">Coloration</emphasis>
|
||
: Body dark green, posterior end of each tergite (including collum and anal shield) with thin yellow stripe (Fig. 6A). Head, antennae and legs green.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE3734032AAFBABE12DFBF2" blockId="16.[151,1436,151,2034]" box="[199,554,1041,1067]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032AAFBABE201FBF3" box="[199,262,1041,1066]" italics="true" pageId="16" pageNumber="17">Head</emphasis>
|
||
: Eyes with 50–60 ocelli.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE3734032AAFB82E753FB1F" blockId="16.[151,1436,151,2034]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032AAFB82E231FB88" box="[199,310,1080,1105]" italics="true" pageId="16" pageNumber="17">Antennae</emphasis>
|
||
: Very short (Fig. 8A). Length of antennomeres: 1>2=3=4<5=6. Sixth antennomere cylindrical (Fig. 8A), disc with 9–15 apical cones (Fig. 9A). First antennomere broadest, short and cylindrical, apically with row of
|
||
<emphasis id="B922D319FFE37340334BFB3FE102FB47" box="[294,517,1157,1182]" italics="true" pageId="16" pageNumber="17">sensilla basiconica</emphasis>
|
||
, without groove (Fig. 8B). Sclerotized teeth at base of antennomeres 1–5 (Fig. 8A). Apical part of fifth antennomere also with few
|
||
<emphasis id="B922D319FFE373403096FB16E0DDFB1C" box="[763,986,1196,1221]" italics="true" pageId="16" pageNumber="17">sensilla basiconica</emphasis>
|
||
(Fig. 8C).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE3734032AAFB69E119FAE3" blockId="16.[151,1436,151,2034]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032AAFB69E230FB35" box="[199,311,1235,1260]" italics="true" pageId="16" pageNumber="17">Mandible</emphasis>
|
||
: Six rows of pectinate lamellae; number of teeth declining from apical to proximal. Intermediate area very small, pectinate lamellae almost directly touching molar plate (Fig. 9C). Condylus apically with two steps (Fig. 9C).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE3734032AAFAFDE1BFFA5E" blockId="16.[151,1436,151,2034]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032AAFAFDE296FAB9" box="[199,401,1351,1376]" italics="true" pageId="16" pageNumber="17">Gnathochilarium</emphasis>
|
||
: Laterally of palpi four sensory cones, located together (Fig. 9B). Depression on posterior side of palpi with few sensory cones.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE3734032AAFA2EE1B4FA77" blockId="16.[151,1436,151,2034]" box="[199,691,1428,1454]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032AAFA2EE21AFA74" box="[199,285,1428,1453]" italics="true" pageId="16" pageNumber="17">Collum</emphasis>
|
||
: Median parts of collum glabrous.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE3734032AAFA01E020FA0C" blockId="16.[151,1436,151,2034]" box="[199,807,1467,1493]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032AAFA01E279FA0D" box="[199,382,1467,1492]" italics="true" pageId="16" pageNumber="17">Thoracic shield</emphasis>
|
||
: Surface similar to those of tergites.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE3734032AAFA5BE72AFA22" blockId="16.[151,1436,151,2034]" box="[199,1069,1505,1531]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032AAFA5BE224FA23" box="[199,291,1505,1530]" italics="true" pageId="16" pageNumber="17">Tergites</emphasis>
|
||
: Smooth and glabrous. Paratergites slightly projecting posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE3734032AAF9B2E028F9B6" blockId="16.[151,1436,151,2034]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032AAF9B2E250F9F8" box="[199,343,1544,1569]" italics="true" pageId="16" pageNumber="17">Endotergum</emphasis>
|
||
: Inner section with short spines and very few, isolated bristles (Fig. 23B). Cuticular impressions absent (Fig. 23B). Externally three rows of marginal bristles, standing irregularly to one another. Bristles scaly, long, extending beyond tergite (Fig. 23B).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE3734032AAF9C6E1BEF964" blockId="16.[151,1436,151,2034]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032AAF9C6E24BF94C" box="[199,332,1660,1685]" italics="true" pageId="16" pageNumber="17">Anal shield</emphasis>
|
||
: Rounded, neither bell-shaped nor tapered (Fig. 6A). Slightly sexually dimorphic, in females glabrous. Ventral side without locking carinae.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE3734032AAF973E785F8E9" blockId="16.[151,1436,151,2034]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032AAF973E3F9F93B" box="[199,254,1737,1762]" italics="true" pageId="16" pageNumber="17">Legs</emphasis>
|
||
: Tarsi of first pair of legs with three, second with up to four ventral spines. First two leg pairs with only weakly curved claws and without apical spine. Tarsi of legs 3–21 with curved claws, 10–12 ventral spines and one apical spine. Femur 1.8 and tarsi 3.0 times longer than wide (Fig. 7A).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE3734032AAF887E7DAF88E" blockId="16.[151,1436,151,2034]" box="[199,1245,1853,1879]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032AAF887E271F88F" box="[199,374,1853,1878]" italics="true" pageId="16" pageNumber="17">Stigmatic plate</emphasis>
|
||
: First lobe well-rounded, long and weakly curved towards coxa (Fig. 7B).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE3734032AAF8DEE25DF82B" blockId="16.[151,1436,151,2034]" pageId="16" pageNumber="17">
|
||
<emphasis id="B922D319FFE3734032AAF8DEE100F8A4" box="[199,519,1892,1917]" italics="true" pageId="16" pageNumber="17">Female sexual characters</emphasis>
|
||
: Subanal plate with washboard, consisting of short, but well-developed stridulation ribs, two ribs on each half. Stridulation ribs symmetrical and strong. Vulva large, covering more than 4/5 of coxa. Apical margin of operculum weakly notched. Mesal plate reaching up to 2/3 of operculum's height (Fig. 7C).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE2734132AAFF22E2AFFF26" blockId="17.[151,1437,152,719]" pageId="17" pageNumber="18">
|
||
<emphasis id="B922D319FFE2734132AAFF22E2D8FF68" box="[199,479,152,177]" italics="true" pageId="17" pageNumber="18">Male sexual characters</emphasis>
|
||
: Male gonopore covered with sclerotized, undivided and rounded plate. Apical part of plate membranous (Fig. 7D). Anal shield not modified, but covered with numerous minute hairs around hindmost point.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE2734132AAFEB6E252FE3E" blockId="17.[151,1437,152,719]" pageId="17" pageNumber="18">
|
||
<emphasis id="B922D319FFE2734132AAFEB6E29FFEFC" box="[199,408,268,293]" italics="true" pageId="17" pageNumber="18">Anterior telopods</emphasis>
|
||
(Figs 7E, F): Harp with two stridulation ribs. Both ribs straight, mesal one slightly larger than lateral rib. Second podomere process lobe-like and little curved with rounded edges, reaching half of third podomeres height, apically with sclerotized spots. Third podomere on posterior side with cavity juxtaposed to second podomere process. Mesal margin towards cavity with numerous sclerotized spots and 3 or 4 small spines, lateral margin with 2–4 crenulated teeth. Tip of third podomere with one triangular, sclerotized spot.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFE2734132AAFE4EE622FD16" blockId="17.[151,1437,152,719]" pageId="17" pageNumber="18">
|
||
<emphasis id="B922D319FFE2734132AAFE4EE2A3FDD4" box="[199,420,500,525]" italics="true" pageId="17" pageNumber="18">Posterior telopods</emphasis>
|
||
(Figs 7G, H): Movable finger on both sides with few hairs. Finger slightly curved towards fixed finger, hollowed-out inner margin with two non-sclerotized lobes and four up to five smaller, sclerotized spines; posterior aspect with row of
|
||
<emphasis id="B922D319FFE2734130A8FDFBE007FD83" box="[709,768,577,602]" italics="true" pageId="17" pageNumber="18">circa</emphasis>
|
||
20 large, crenulated teeth. Fixed finger only on posterior side with few hairs, shorter but almost twice as wide as third podomere, with straight tip; anterior side basally with single membranous lobe and small, sclerotized spine, on inner margin covered with numerous sclerotized spots juxtaposed to third podomere sclerotized teeth. First podomere on both sides almost glabrous.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |