449 lines
45 KiB
XML
449 lines
45 KiB
XML
<document id="E3EB5386D9EC2845AE80DC1D5E1CED06" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFFCF731132FAF8ABE64BFEAA" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium tsingy Wesener 2009, new species" docType="treatment" docUuid="E0CB1946-D7C8-4C6F-BFA1-BECBB3F6401E" docUuidSource="ZooBank" docVersion="4" lastPageNumber="66" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="61" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="065C84AAAB0B2E52B728EC1A4FAAE8A4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="46872078CF59885D1A0D9C5B1FC23364">
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<mods:title id="E11522D6BFC481826489862348C9EF0C">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
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</mods:titleInfo>
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<mods:name id="E824909FE661D3622A4C39115FB8D02E" type="personal">
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<mods:role id="98E2C0F3D4C00E220519B80C532CFD88">
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<mods:roleTerm id="764469CE5E6103AA5E7C59982F4596DF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8367F8B31DDAE19F3CA41E757C7675F3">Wesener, Thomas</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="FF59566264A1C5FE86C44B9B1CD7A0A0">text</mods:typeOfResource>
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<mods:titleInfo id="3BFD9D4C3ABFF23B887D0107A803DEC9">
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<mods:title id="B44A5952BFD8C0B034BC458F6CE0C263">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="D25C1A2E0467BABF0BF7AC62FE1D8C5D">
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<mods:date id="D2D449877DA4D2EA82DE5FB8C902F2B3">2009</mods:date>
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<mods:detail id="88D4C6A8B963CB9B58A55C844C68479C" type="pubDate">
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<mods:number id="86ECD8D54CB915E81CCC0100F5A6BAD3">2009-05-11</mods:number>
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</mods:detail>
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<mods:detail id="9CC665BFF8FCD27DE4CA38B334D2DC87" type="volume">
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<mods:number id="D493BB5212400ABBC33F07542AF48723">2097</mods:number>
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</mods:detail>
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<mods:detail id="B29B2272BBB3286E5D913E2532480971" type="issue">
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<mods:number id="B858DE50FF616A9FF02639A9C5A666AD">1</mods:number>
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</mods:detail>
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<mods:extent id="1403326C204B964D2C9F9FE3F9A2C695" unit="page">
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<mods:start id="F48E8782C6DE6A3BA0C7396CCF18BB9C">1</mods:start>
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<mods:end id="315CCDDBD1296D05C0122A152417D9AD">134</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location id="D437C9CDA17485223CE4FFE6ED70343A">
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<mods:url id="9B5B0D6DBF026A9A19FE49AEB055BE8A">https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1</mods:url>
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</mods:location>
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<mods:classification id="B89B00AE7E216A93BFC8A874E8D73567">journal article</mods:classification>
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<mods:identifier id="E9DF7F712DE824EA6C54997B27202DB7" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
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<mods:identifier id="869D4351FB2FE489884E88AE32C61F9D" type="ISSN">11755334</mods:identifier>
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<mods:identifier id="44FC851A51A3F337C641159D708BCA1E" type="Zenodo-Dep">5316536</mods:identifier>
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<mods:identifier id="48BF00450518D1FE74D9D76D1C681779" type="ZooBank">0E18167A-78EB-4079-8BF4-972A4AC88C69</mods:identifier>
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</mods:mods>
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<treatment id="03FFBE1DFFCF731132FAF8ABE64BFEAA" LSID="urn:lsid:zoobank.org:act:E0CB1946-D7C8-4C6F-BFA1-BECBB3F6401E" httpUri="http://treatment.plazi.org/id/03FFBE1DFFCF731132FAF8ABE64BFEAA" lastPageId="65" lastPageNumber="66" pageId="60" pageNumber="61">
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<subSubSection id="C34C5C80FFCF736C32FAF8ABE135F8F2" box="[151,562,1809,1835]" pageId="60" pageNumber="61" type="nomenclature">
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<paragraph id="8BE90F0BFFCF736C32FAF8ABE135F8F2" blockId="60.[151,562,1809,1835]" box="[151,562,1809,1835]" pageId="60" pageNumber="61">
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<heading id="D0A1B867FFCF736C32FAF8ABE135F8F2" bold="true" box="[151,562,1809,1835]" fontSize="11" level="1" pageId="60" pageNumber="61" reason="1">
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||
<emphasis id="B922D319FFCF736C32FAF8ABE135F8F2" bold="true" box="[151,562,1809,1835]" pageId="60" pageNumber="61">
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<emphasis id="B922D319FFCF736C32FAF8ABE29CF8F2" bold="true" box="[151,411,1809,1835]" italics="true" pageId="60" pageNumber="61">
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<taxonomicName id="4C567488FFCF736C32FAF8ABE292F8F2" ID-CoL="5D7JC" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,405,1809,1835]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="tsingy" status="sp. nov.">Zoosphaerium tsingy</taxonomicName>
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,
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||
</emphasis>
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||
<taxonomicNameLabel id="A2116E62FFCF736C33CEF8ABE135F8F2" box="[419,562,1809,1835]" pageId="60" pageNumber="61" rank="species">new species</taxonomicNameLabel>
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||
</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34C5C80FFCF736C32FAF8E0E127F81B" pageId="60" pageNumber="61" type="etymology">
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||
<paragraph id="8BE90F0BFFCF736C32FAF8E0E22EF842" blockId="60.[151,1435,1882,2024]" pageId="60" pageNumber="61">
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||
<emphasis id="B922D319FFCF736C32FAF8E0E28EF8AD" bold="true" box="[151,393,1882,1908]" pageId="60" pageNumber="61">Derivatio nominis:</emphasis>
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<taxonomicName id="4C567488FFCF736C33F4F8E1E2DAF8AD" authorityName="Wesener" authorityYear="2009" box="[409,477,1883,1908]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="tsingy">
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<emphasis id="B922D319FFCF736C33F4F8E1E2DAF8AD" box="[409,477,1883,1908]" italics="true" pageId="60" pageNumber="61">tsingy</emphasis>
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</taxonomicName>
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||
, noun in apposition, after the
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<typeStatus id="54EDB1A9FFCF736C3102F8E1E0A6F8AC" box="[879,929,1883,1909]" pageId="60" pageNumber="61">type</typeStatus>
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locality, the world heritage site
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<taxonomicName id="4C567488FFCF736C3721F8E1E22EF842" authority="d'Bemaraha." class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="60" pageNumber="61" phylum="Arthropoda" rank="species" species="tsingy">Tsingy d'Bemaraha.</taxonomicName>
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||
</paragraph>
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<paragraph id="8BE90F0BFFCF736C32ABF81DE127F81B" blockId="60.[151,1435,1882,2024]" box="[198,544,1959,1986]" pageId="60" pageNumber="61">
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<emphasis id="B922D319FFCF736C32ABF81DE2A7F818" bold="true" box="[198,416,1959,1985]" pageId="60" pageNumber="61">Studied material:</emphasis>
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13 M,
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||
<specimenCount id="9D50C482FFCF736C3399F812E127F81B" box="[500,544,1960,1986]" pageId="60" pageNumber="61" type="generic">5 F.</specimenCount>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34C5C80FFCF736D32AAF874E638FEAA" lastPageId="61" lastPageNumber="62" pageId="60" pageNumber="61" type="materials_examined">
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||
<paragraph id="8BE90F0BFFCF736C32AAF874E69CF831" blockId="60.[151,1435,1882,2024]" box="[199,1435,1998,2024]" pageId="60" pageNumber="61">
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||
<materialsCitation id="3B3E0556FFCF736C32AAF874E69CF831" box="[199,1435,1998,2024]" collectionCode="FMMC" country="Madagascar" location="Parc National Tsingy" municipality="Province" pageId="60" pageNumber="61" specimenCode="FMMC 5455" specimenCount="1" stateProvince="Mahajanga" typeStatus="holotype">
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||
<emphasis id="B922D319FFCF736C32AAF874E23EF831" bold="true" box="[199,313,1998,2024]" pageId="60" pageNumber="61">
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||
<typeStatus id="54EDB1A9FFCF736C32AAF874E23EF831" box="[199,313,1998,2024]" pageId="60" pageNumber="61" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
: 1 M (
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||
<quantity id="4CAEA2EEFFCF736C33E7F874E2DAF831" box="[394,477,1998,2024]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="60" pageNumber="61" unit="mm" value="36.0">36 mm</quantity>
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||
long),
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||
<emphasis id="B922D319FFCF736C305EF874E1D4F831" bold="true" box="[563,723,1998,2024]" pageId="60" pageNumber="61">
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||
<specimenCode id="DBF0A770FFCF736C305EF874E1D4F831" box="[563,723,1998,2024]" collectionCode="FMMC" pageId="60" pageNumber="61">FMMC 5455</specimenCode>
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||
</emphasis>
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||
.
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||
<collectingCountry id="F3414F9BFFCF736C308EF874E075F831" box="[739,882,1998,2024]" name="Madagascar" pageId="60" pageNumber="61">Madagascar</collectingCountry>
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||
,
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||
<collectingMunicipality id="6B8D9571FFCF736C3112F874E0E0F831" box="[895,999,1998,2024]" pageId="60" pageNumber="61">Province</collectingMunicipality>
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||
<collectingRegion id="4992C1E9FFCF736C3182F874E773F831" box="[1007,1140,1998,2024]" country="Madagascar" name="Mahajanga" pageId="60" pageNumber="61">Mahajanga</collectingRegion>
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||
,
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||
<location id="8E8959D0FFCF736C36EDF874E67FF831" LSID="urn:lsid:plazi:treatment:03FFBE1DFFCF731132FAF8ABE64BFEAA:8E8959D0FFCF736C36EDF874E67FF831" box="[1152,1400,1998,2024]" country="Madagascar" municipality="Province" name="Parc National Tsingy" pageId="60" pageNumber="61" stateProvince="Mahajanga">
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Parc National
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<taxonomicName id="4C567488FFCF736C3744F874E67FF831" box="[1321,1400,1998,2024]" pageId="60" pageNumber="61">Tsingy</taxonomicName>
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||
</location>
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||
de
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="8BE90F0BFFCE736D32FAFF22E120FF01" blockId="61.[151,1436,152,2034]" pageId="61" pageNumber="62">
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||
Bemaraha, Ankidrodroa,
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||
<quantity id="4CAEA2EEFFCE736D33A5FF22E11AFF6B" box="[456,541,152,178]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="61" pageNumber="62" unit="km" value="2.5">2.5 km</quantity>
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NE Bekopaka, secondary dry forest,
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||
<quantity id="4CAEA2EEFFCE736D3187FF22E73DFF6B" box="[1002,1082,152,178]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="61" pageNumber="62" unit="m" value="100.0">100 m</quantity>
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||
,
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||
<geoCoordinate id="EE6269CCFFCE736D3626FF22E7B7FF6B" box="[1099,1200,152,178]" degrees="19" direction="south" minutes="7.9" orientation="latitude" pageId="61" pageNumber="62" precision="92" value="-19.131666">19°7.9'S</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE6269CCFFCE736D36ADFF22E631FF6B" box="[1216,1334,152,178]" degrees="44" direction="east" minutes="48.5" orientation="longitude" pageId="61" pageNumber="62" precision="92" value="44.808334">44°48.5'E</geoCoordinate>
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||
, coll. S. Goodman,
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<date id="FFE829CBFFCE736D3374FF04E29EFF01" box="[281,409,190,216]" pageId="61" pageNumber="62" value="2001-11-28">28.xi.2001</date>
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||
, pitfall trap.
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||
</paragraph>
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||
<paragraph id="8BE90F0BFFCE736D32AAFF5FE222FEFF" blockId="61.[151,1436,152,2034]" pageId="61" pageNumber="62">
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||
<materialsCitation id="3B3E0556FFCE736D32AAFF5FE07AFF26" box="[199,893,229,255]" collectionCode="FMMC" country="Madagascar" location="Parc National Tsingy" municipality="Province" pageId="61" pageNumber="62" specimenCode="FMMC 5455" specimenCount="1" stateProvince="Mahajanga" typeStatus="paratype">
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||
<emphasis id="B922D319FFCE736D32AAFF5FE243FF26" bold="true" box="[199,324,229,255]" pageId="61" pageNumber="62">
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||
<typeStatus id="54EDB1A9FFCE736D32AAFF5FE243FF26" box="[199,324,229,255]" pageId="61" pageNumber="62" type="paratype">Paratypes</typeStatus>
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||
</emphasis>
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||
: 8 M, 2 F,
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<emphasis id="B922D319FFCE736D33AAFF5FE160FF26" bold="true" box="[455,615,229,255]" pageId="61" pageNumber="62">
|
||
<specimenCode id="DBF0A770FFCE736D33AAFF5FE160FF26" box="[455,615,229,255]" collectionCode="FMMC" pageId="61" pageNumber="62">FMMC 5455</specimenCode>
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</emphasis>
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, same data as holotype
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</materialsCitation>
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. 5 M, 5 F,
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<materialsCitation id="3B3E0556FFCE736D3191FF5FE227FEFF" collectionCode="FMMC" country="Madagascar" location="Parc National Tsingy" municipality="Province" pageId="61" pageNumber="62" specimenCode="FMMC 5454" specimenCount="1" stateProvince="Mahajanga" typeStatus="paratype">
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||
<emphasis id="B922D319FFCE736D3191FF5FE79BFF26" bold="true" box="[1020,1180,229,255]" pageId="61" pageNumber="62">
|
||
<specimenCode id="DBF0A770FFCE736D3191FF5FE79BFF26" box="[1020,1180,229,255]" collectionCode="FMMC" pageId="61" pageNumber="62">FMMC 5454</specimenCode>
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||
</emphasis>
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||
, same collection data as holotype
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8BE90F0BFFCE736D32AAFE88E638FEAA" blockId="61.[151,1436,152,2034]" pageId="61" pageNumber="62">
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||
<emphasis id="B922D319FFCE736D32AAFE88E280FE95" bold="true" box="[199,391,306,332]" pageId="61" pageNumber="62">Other material</emphasis>
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||
: 1 F,
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||
<materialsCitation id="3B3E0556FFCE736D33B9FE88E638FEAA" collectingDate="2001-11-21" collectingDateMax="2001-11-25" collectingDateMin="2001-11-21" collectionCode="CAS" collectorName="de Tsimembo & Fisher & Griswold" elevation="20" latitude="-19.021389" location="Foret de Tsimembo" longLatPrecision="21" longitude="44.440556" municipality="Province" pageId="61" pageNumber="62" specimenCode="BLF 4509" specimenCount="1" stateProvince="Mahajanga">
|
||
<emphasis id="B922D319FFCE736D33B9FE88E19EFE95" bold="true" box="[468,665,306,332]" pageId="61" pageNumber="62">
|
||
<collectionCode id="ED4797CEFFCE736D33B9FE88E109FE95" box="[468,526,306,332]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="61" pageNumber="62" type="Museum">CAS</collectionCode>
|
||
<specimenCode id="DBF0A770FFCE736D3076FE88E19EFE95" box="[539,665,306,332]" collectionCode="BLF" pageId="61" pageNumber="62">BLF 4509</specimenCode>
|
||
</emphasis>
|
||
,
|
||
<collectingMunicipality id="6B8D9571FFCE736D30C1FE88E013FE95" box="[684,788,306,332]" pageId="61" pageNumber="62">Province</collectingMunicipality>
|
||
<collectingRegion id="4992C1E9FFCE736D3172FE88E0A3FE95" box="[799,932,306,332]" country="Madagascar" name="Mahajanga" pageId="61" pageNumber="62">Mahajanga</collectingRegion>
|
||
,
|
||
<location id="8E8959D0FFCE736D31D9FE88E7A5FE95" LSID="urn:lsid:plazi:treatment:03FFBE1DFFCF731132FAF8ABE64BFEAA:8E8959D0FFCE736D31D9FE88E7A5FE95" box="[948,1186,306,332]" latitude="-19.021389" longLatPrecision="21" longitude="44.440556" municipality="Province" name="Foret de Tsimembo" pageId="61" pageNumber="62" stateProvince="Mahajanga">
|
||
Forêt
|
||
<collectorName id="26A36ADDFFCE736D3190FE88E7A5FE95" box="[1021,1186,306,332]" pageId="61" pageNumber="62">de Tsimembo</collectorName>
|
||
</location>
|
||
,
|
||
<locationDeviation id="9A5B1629FFCE736D36DFFE88E3F0FEAA" location="Soatana" pageId="61" pageNumber="62">
|
||
<quantity id="4CAEA2EEFFCE736D36DFFE88E600FE95" box="[1202,1287,306,332]" metricMagnitude="3" metricUnit="m" metricValue="8.7" pageId="61" pageNumber="62" unit="km" value="8.7">8.7 km</quantity>
|
||
336° NNW Soatana
|
||
</locationDeviation>
|
||
, tropical dry forest,
|
||
<quantity id="4CAEA2EEFFCE736D338EFEE3E127FEAA" box="[483,544,345,371]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="61" pageNumber="62" unit="m" value="20.0">
|
||
<elevation id="007BE838FFCE736D338EFEE3E127FEAA" box="[483,544,345,371]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="61" pageNumber="62" unit="m" value="20.0">20 m</elevation>
|
||
</quantity>
|
||
,
|
||
<geoCoordinate id="EE6269CCFFCE736D3040FEE3E1A2FEAA" box="[557,677,345,371]" degrees="19" direction="south" minutes="1" orientation="latitude" pageId="61" pageNumber="62" precision="15" seconds="17" value="-19.021389">19°1'17"S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE6269CCFFCE736D30DFFEE3E03CFEAA" box="[690,827,345,371]" degrees="44" direction="east" minutes="26" orientation="longitude" pageId="61" pageNumber="62" precision="15" seconds="26" value="44.440556">44°26'26"E</geoCoordinate>
|
||
, coll.
|
||
<collectorName id="26A36ADDFFCE736D31EFFEE3E0CAFEAA" box="[898,973,345,371]" pageId="61" pageNumber="62">Fisher</collectorName>
|
||
,
|
||
<collectorName id="26A36ADDFFCE736D31B4FEE3E741FEAA" box="[985,1094,345,371]" pageId="61" pageNumber="62">Griswold</collectorName>
|
||
<emphasis id="B922D319FFCE736D3620FEE3E78FFEAB" box="[1101,1160,345,370]" italics="true" pageId="61" pageNumber="62">et al.</emphasis>
|
||
,
|
||
<date id="FFE829CBFFCE736D36FBFEE3E638FEAA" box="[1174,1343,345,371]" pageId="61" pageNumber="62" value="2001-11-21" valueMax="2001-11-25" valueMin="2001-11-21">
|
||
<collectingDate id="EFACD023FFCE736D36FBFEE3E638FEAA" box="[1174,1343,345,371]" pageId="61" pageNumber="62" value="2001-11-21" valueMax="2001-11-25" valueMin="2001-11-21">21–25.xi.2001</collectingDate>
|
||
</date>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C5C80FFCE736D32ABFEC5E7B3FE43" box="[198,1204,383,410]" pageId="61" pageNumber="62" type="distribution">
|
||
<paragraph id="8BE90F0BFFCE736D32ABFEC5E7B3FE43" blockId="61.[151,1436,152,2034]" box="[198,1204,383,410]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32ABFEC5E26EFE40" bold="true" box="[198,361,383,409]" pageId="61" pageNumber="62">Distribution:</emphasis>
|
||
Dry forests Bemaraha and Tsimembo in Western
|
||
<collectingCountry id="F3414F9BFFCE736D31DCFE3AE739FE43" box="[945,1086,384,410]" name="Madagascar" pageId="61" pageNumber="62">Madagascar</collectingCountry>
|
||
(Fig. 38).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C5C80FFCE736D32ABFE1CE637FCB3" pageId="61" pageNumber="62" type="diagnosis">
|
||
<paragraph id="8BE90F0BFFCE736D32ABFE1CE663FD2F" blockId="61.[151,1436,152,2034]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32ABFE1CE24FFE19" bold="true" box="[198,328,422,448]" pageId="61" pageNumber="62">Diagnosis:</emphasis>
|
||
Medium sized, up to
|
||
<quantity id="4CAEA2EEFFCE736D3039FE1CE1AFFE19" box="[596,680,422,448]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="61" pageNumber="62" unit="mm" value="50.0">50 mm</quantity>
|
||
long. Males slightly (
|
||
<emphasis id="B922D319FFCE736D31DEFE1DE0E9FE19" box="[947,1006,423,448]" italics="true" pageId="61" pageNumber="62">circa</emphasis>
|
||
<quantity id="4CAEA2EEFFCE736D3195FE1CE738FE19" box="[1016,1087,422,448]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="61" pageNumber="62" unit="mm" value="5.0">5 mm</quantity>
|
||
) smaller than females. Color olive-greenish (Fig. 39A). All legs without long hairs on inner margin. Texture of tergites smooth and glabrous. First antennomere with groove (Fig. 43C), antennomeres 1–4 with sclerotized teeth (Fig. 43B), disc with 16–31 apical cones (Fig. 43A). Movable finger of posterior telopods not curved, of almost cylindrical shape with two non-sclerotized lobes and three thin spines (Fig. 42E), posterior side with
|
||
<emphasis id="B922D319FFCE736D36AFFDFBE7FAFD83" box="[1218,1277,577,602]" italics="true" pageId="61" pageNumber="62">circa</emphasis>
|
||
9 sclerotized teeth, (Fig. 42F). Two stridulation ribs on each male harp (Fig. 42B) and 2 or 3 on each side of female washboard. Third podomere of anterior telopods large, elongated (Fig. 42C). Two locking carinae on the anal shield, anterior carina short and dot-like, posterior one straight and of medium length. Operculum of special shape, anterior margin excavated into two wide lobes. Lateral lobe twice as long as inner one (Fig. 42A).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCE736D32ABFCB8E637FCB3" blockId="61.[151,1436,152,2034]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32ABFCB8E28BFCC5" bold="true" box="[198,396,770,796]" pageId="61" pageNumber="62">Similar species:</emphasis>
|
||
<emphasis id="B922D319FFCE736D33FBFCB8E103FCC2" box="[406,516,770,795]" italics="true" pageId="61" pageNumber="62">
|
||
<taxonomicName id="4C567488FFCE736D33FBFCB8E2F9FCC2" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[406,510,770,795]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="tsingy" status="sp. nov.">Z. tsingy</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="B922D319FFCE736D3060FCB8E156FCC5" bold="true" box="[525,593,770,796]" pageId="61" pageNumber="62">
|
||
<taxonomicNameLabel id="A2116E62FFCE736D3060FCB8E156FCC5" box="[525,593,770,796]" pageId="61" pageNumber="62" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
telopods have almost no similarities to any other described giant pillmillipede species. The shape and position of the locking carinae and vulva place
|
||
<taxonomicName id="4C567488FFCE736D362BFC93E7ADFC9B" authorityName="Wesener" authorityYear="2009" box="[1094,1194,809,834]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="tsingy">
|
||
<emphasis id="B922D319FFCE736D362BFC93E7ADFC9B" box="[1094,1194,809,834]" italics="true" pageId="61" pageNumber="62">Z. tsingy</emphasis>
|
||
</taxonomicName>
|
||
in the
|
||
<taxonomicName id="4C567488FFCE736D3697FC93E69BFC9B" baseAuthorityName="de Saussure & Zehntner" baseAuthorityYear="1897" box="[1274,1436,809,834]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="platylabum">
|
||
<emphasis id="B922D319FFCE736D3697FC93E69BFC9B" box="[1274,1436,809,834]" italics="true" pageId="61" pageNumber="62">Z. platylabum</emphasis>
|
||
</taxonomicName>
|
||
species-group, but the special-shaped posterior telopods are so far unique for
|
||
<taxonomicName id="4C567488FFCE736D364EFCEAE637FCB3" box="[1059,1328,848,874]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="61" pageNumber="62" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B922D319FFCE736D364EFCEAE7CBFCB0" box="[1059,1228,848,873]" italics="true" pageId="61" pageNumber="62">Zoosphaerium</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34C5C80FFCE731132ABFCCCE64BFEAA" lastPageId="65" lastPageNumber="66" pageId="61" pageNumber="62" type="description">
|
||
<paragraph id="8BE90F0BFFCE736D32ABFCCCE7E6FC07" blockId="61.[151,1436,152,2034]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32ABFCCCE267FC49" bold="true" box="[198,352,886,912]" pageId="61" pageNumber="62">Description:</emphasis>
|
||
<emphasis id="B922D319FFCE736D3305FCCDE2F2FC49" box="[360,501,887,912]" italics="true" pageId="61" pageNumber="62">Body length</emphasis>
|
||
: Males (
|
||
<specimenCount id="9D50C482FFCE736D3031FCCCE005FC49" box="[604,770,886,912]" pageId="61" pageNumber="62" type="generic">13 specimens</specimenCount>
|
||
): length up to
|
||
<quantity id="4CAEA2EEFFCE736D31C5FCCCE0FAFC49" box="[936,1021,886,912]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="61" pageNumber="62" unit="mm" value="36.0">36 mm</quantity>
|
||
, width of thoracic shield up to
|
||
<quantity id="4CAEA2EEFFCE736D3705FCCCE3CFFC6E" metricMagnitude="-2" metricUnit="m" metricValue="1.93" pageId="61" pageNumber="62" unit="mm" value="19.3">19.3 mm</quantity>
|
||
, height of thoracic shield up to
|
||
<quantity id="4CAEA2EEFFCE736D302EFC27E1B7FC6E" box="[579,688,925,951]" metricMagnitude="-2" metricUnit="m" metricValue="1.0699999999999998" pageId="61" pageNumber="62" unit="mm" value="10.7">10.7 mm</quantity>
|
||
. Females (
|
||
<specimenCount id="9D50C482FFCE736D315DFC27E0CFFC6E" box="[816,968,925,951]" pageId="61" pageNumber="62" type="generic">6 specimens</specimenCount>
|
||
): length up to
|
||
<quantity id="4CAEA2EEFFCE736D3619FC27E7CCFC6E" box="[1140,1227,925,951]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="61" pageNumber="62" unit="mm" value="48.0">48 mm</quantity>
|
||
, width up to
|
||
<quantity id="4CAEA2EEFFCE736D3704FC27E3CFFC07" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="61" pageNumber="62" unit="mm" value="25.8">25.8 mm</quantity>
|
||
, height up to
|
||
<quantity id="4CAEA2EEFFCE736D330BFC7EE2D5FC07" box="[358,466,964,990]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="61" pageNumber="62" unit="mm" value="14.1">14.1 mm</quantity>
|
||
. Females
|
||
<quantity id="4CAEA2EEFFCE736D302BFC7EE1B1FC07" box="[582,694,964,990]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="61" pageNumber="62" unit="mm" value="7.5" valueMax="10.0" valueMin="5.0">5–10 mm</quantity>
|
||
larger than males of the same collection series.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCE736D32AAFC50E78BFBDD" blockId="61.[151,1436,152,2034]" box="[199,1164,1002,1028]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32AAFC50E229FBDA" box="[199,302,1002,1027]" italics="true" pageId="61" pageNumber="62">Habitus:</emphasis>
|
||
Anal shield well-rounded (Fig. 39A). Tergites very smooth and glabrous.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCE736D32AAFBABE0B9FB8B" blockId="61.[151,1436,152,2034]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32AAFBABE256FBF3" box="[199,337,1041,1066]" italics="true" pageId="61" pageNumber="62">Coloration:</emphasis>
|
||
Thoracic shield and tergites dirty olive-greenish, posterior margin and paratergite tips brown (Fig. 39A). Anal shield, collum, head, antennae and legs olive green.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCE736D32AAFBE5E17AFBA1" blockId="61.[151,1436,152,2034]" box="[199,637,1118,1144]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32AAFBE5E217FBA1" box="[199,272,1119,1144]" italics="true" pageId="61" pageNumber="62">Head:</emphasis>
|
||
Eyes with more than 75 ocelli.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCE736D32AAFB3FE03FFACA" blockId="61.[151,1436,152,2034]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32AAFB3FE238FB47" box="[199,319,1157,1182]" italics="true" pageId="61" pageNumber="62">Antennae:</emphasis>
|
||
Short, length of antennomeres: 1>2=3=4=5<6, sixth antennomere largest (Fig. 43B), disc with 16–31 apical cones (Fig. 43A). First antennomere of greatest width, short and cylindrical, with groove and one row of
|
||
<emphasis id="B922D319FFCE736D3376FB69E2FCFB35" box="[283,507,1235,1260]" italics="true" pageId="61" pageNumber="62">sensilla basiconica</emphasis>
|
||
(Fig. 43C). Fifth antennomere with two rows of
|
||
<emphasis id="B922D319FFCE736D365EFB69E614FB35" box="[1075,1299,1235,1260]" italics="true" pageId="61" pageNumber="62">sensilla basiconica</emphasis>
|
||
(Fig. 43D). Sclerotized teeth at base of antennomeres 1–4 (Fig. 43B).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCE736D32AAFA9AE0E9FAB9" blockId="61.[151,1436,152,2034]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32AAFA9AE247FAE0" box="[199,320,1312,1337]" italics="true" pageId="61" pageNumber="62">Mandible:</emphasis>
|
||
Six rows of pectinate lamellae and large intermediate area. Molar plate with one deep invagination. Condylus apically with strongly developed step (Fig. 44B).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCE736D32AAFAD7E074FA0C" blockId="61.[151,1436,152,2034]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32AAFAD7E296FA5F" box="[199,401,1389,1414]" italics="true" pageId="61" pageNumber="62">Gnathochilarium</emphasis>
|
||
: Lateral of palpi 6 or 7 sensory cones, located together (Fig. 43E). Depression on posterior side of palpi with some sensory cones (Fig. 43E).
|
||
<emphasis id="B922D319FFCE736D3115FA2EE0E8FA74" box="[888,1007,1428,1453]" italics="true" pageId="61" pageNumber="62">Tentorium</emphasis>
|
||
towards endochilarium with single large tooth and two rows of spines forming zippers (Fig. 43E).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCE736D32AAFA5BE7B1FA22" blockId="61.[151,1436,152,2034]" box="[199,1206,1505,1531]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32AAFA5BE220FA23" box="[199,295,1505,1530]" italics="true" pageId="61" pageNumber="62">Collum:</emphasis>
|
||
Margins covered with few short hairs, some hairs also standing on inner part.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCE736D32AAF9B2E1E3F9FB" blockId="61.[151,1436,152,2034]" box="[199,740,1544,1570]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32AAF9B2E279F9F8" box="[199,382,1544,1569]" italics="true" pageId="61" pageNumber="62">Thoracic shield</emphasis>
|
||
: Surface like those of tergites.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCE736D32AAF995E0C9F9B6" blockId="61.[151,1436,152,2034]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32AAF995E224F991" box="[199,291,1583,1608]" italics="true" pageId="61" pageNumber="62">Tergites</emphasis>
|
||
: Smooth and mostly glabrous. On high magnification (90x) very small, dot-like impressions become visible. Paratergites projecting slightly posteriorly (Fig. 39A).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCE736D32AAF9C6E275F8D3" blockId="61.[151,1436,152,2034]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32AAF9C6E267F94C" box="[199,352,1660,1685]" italics="true" pageId="61" pageNumber="62">Endotergum:</emphasis>
|
||
Inner section with numerous short spines and isolated bristles (Fig. 47C). Between marginal ridge and inner area single row of large, circular cuticular impressions (Fig. 47C). Externally 2 or 3 rows of marginal bristle, standing irregularly to each other. Bristles scaly, of medium length, reaching up to tergite margin (Fig. 47C).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCE736D32AAF8ADE107F8A7" blockId="61.[151,1436,152,2034]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32AAF8ADE25EF8E9" box="[199,345,1815,1840]" italics="true" pageId="61" pageNumber="62">Anal shield:</emphasis>
|
||
Rounded, neither bell-shaped nor tapered (Fig. 39A). Ventral side of anal shield with two black locking carinae, located closely to laterotergites. Anterior carina small and dot-like, posterior carina straight and of medium length.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCE736D32AAF831E667F82B" blockId="61.[151,1436,152,2034]" pageId="61" pageNumber="62">
|
||
<emphasis id="B922D319FFCE736D32AAF831E20FF87D" box="[199,264,1931,1956]" italics="true" pageId="61" pageNumber="62">Legs:</emphasis>
|
||
Tarsi of first pair of legs with four to seven, second with seven or eight and third with 8–10 ventral spines. First two leg pairs with only weakly curved claws and without apical spine. Tarsi of legs 3–21 with curved claws, 10–13 ventral spines and one apical spine. Femur 1.8 and tarsi 4.4 times longer than wide.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCD736E32AAFF22E705FF6B" blockId="62.[151,1436,152,448]" box="[199,1026,152,178]" pageId="62" pageNumber="63">
|
||
<emphasis id="B922D319FFCD736E32AAFF22E28BFF68" box="[199,396,152,177]" italics="true" pageId="62" pageNumber="63">Stigmatic plates:</emphasis>
|
||
First lobe short and triangular, with well-rounded tip.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCD736E32AAFF05E7E0FEAA" blockId="62.[151,1436,152,448]" pageId="62" pageNumber="63">
|
||
<emphasis id="B922D319FFCD736E32AAFF05E2FDFF01" box="[199,506,191,216]" italics="true" pageId="62" pageNumber="63">Female sexual characters</emphasis>
|
||
: Subanal plate with washboard, consisting of short, incontinuous stridulation ribs, with two or three ribs on each half. Vulva large, covering more than 3/4 of coxa. Operculum long, extending beyond coxa up to first third of prefemur. Apical margin of operculum medially deeply notched, with two large lobes. Lateral lobe of greater length and width than inner lobe. Both lobes with well-rounded tips (Fig. 42A). Mesal plate long, extending beyond coxa up to operculum's height (Fig. 42A).
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCD736E32AAFE3AE3D0FE19" blockId="62.[151,1436,152,448]" pageId="62" pageNumber="63">
|
||
<emphasis id="B922D319FFCD736E32AAFE3AE2D8FE40" box="[199,479,384,409]" italics="true" pageId="62" pageNumber="63">Male sexual characters</emphasis>
|
||
: Anal shield well-rounded, at hindmost point covered with dense field of short hairs.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCD736E32FAF9D7E013F8EB" blockId="62.[151,1436,1645,1842]" pageId="62" pageNumber="63">
|
||
<emphasis id="B922D319FFCD736E32FAF9D7E229F95C" bold="true" box="[151,302,1645,1669]" pageId="62" pageNumber="63">FIGURE 042</emphasis>
|
||
.
|
||
<taxonomicName id="4C567488FFCD736E3351F9D4E11AF95C" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[316,541,1646,1669]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="62" pageNumber="63" phylum="Arthropoda" rank="species" species="tsingy" status="sp. nov.">
|
||
<emphasis id="B922D319FFCD736E3351F9D4E11AF95C" box="[316,541,1646,1669]" italics="true" pageId="62" pageNumber="63">Zoosphaerium tsingy</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B922D319FFCD736E3049F9D7E166F95C" bold="true" box="[548,609,1645,1669]" pageId="62" pageNumber="63">
|
||
<taxonomicNameLabel id="A2116E62FFCD736E3049F9D7E166F95C" box="[548,609,1645,1669]" pageId="62" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, A female
|
||
<typeStatus id="54EDB1A9FFCD736E30B7F9D7E03EF95D" box="[730,825,1645,1668]" pageId="62" pageNumber="63" type="paratype">paratype</typeStatus>
|
||
, B–F
|
||
<typeStatus id="54EDB1A9FFCD736E3114F9D7E0DEF95D" box="[889,985,1645,1668]" pageId="62" pageNumber="63" type="holotype">holotype</typeStatus>
|
||
,
|
||
<emphasis id="B922D319FFCD736E3189F9D7E0F0F95C" bold="true" box="[996,1015,1645,1669]" pageId="62" pageNumber="63">A</emphasis>
|
||
: left second coxa with vulva, posterior view;
|
||
<emphasis id="B922D319FFCD736E32B4F92AE3ECF971" bold="true" box="[217,235,1680,1704]" pageId="62" pageNumber="63">B</emphasis>
|
||
: right half of anterior telopod, anterior view;
|
||
<emphasis id="B922D319FFCD736E30BEF92AE1E1F971" bold="true" box="[723,742,1680,1704]" pageId="62" pageNumber="63">C</emphasis>
|
||
: last three podomeres of anterior telopod, posterior view;
|
||
<emphasis id="B922D319FFCD736E3738F92AE66FF971" bold="true" box="[1365,1384,1680,1704]" pageId="62" pageNumber="63">D</emphasis>
|
||
: last two podomeres of anterior telopod, lateral view;
|
||
<emphasis id="B922D319FFCD736E30D2F908E1D6F913" bold="true" box="[703,721,1714,1738]" pageId="62" pageNumber="63">E</emphasis>
|
||
: posterior telopod, anterior view;
|
||
<emphasis id="B922D319FFCD736E3639F908E763F913" bold="true" box="[1108,1124,1714,1738]" pageId="62" pageNumber="63">F</emphasis>
|
||
: chela of posterior telopod, posterior view. Abbreviations:
|
||
<emphasis id="B922D319FFCD736E338CF96FE10FF934" bold="true" box="[481,520,1749,1773]" pageId="62" pageNumber="63">cav</emphasis>
|
||
= cavity;
|
||
<emphasis id="B922D319FFCD736E301EF96FE19BF934" bold="true" box="[627,668,1749,1773]" pageId="62" pageNumber="63">cr-t</emphasis>
|
||
= crenulated teeth;
|
||
<emphasis id="B922D319FFCD736E311DF96FE097F934" bold="true" box="[880,912,1749,1773]" pageId="62" pageNumber="63">Cx</emphasis>
|
||
= coxa;
|
||
<emphasis id="B922D319FFCD736E3180F96FE708F934" bold="true" box="[1005,1039,1749,1773]" pageId="62" pageNumber="63">EP</emphasis>
|
||
= external plate of vulva;
|
||
<emphasis id="B922D319FFCD736E3744F96FE64FF934" bold="true" box="[1321,1352,1749,1773]" pageId="62" pageNumber="63">IH</emphasis>
|
||
= inner horns;
|
||
<emphasis id="B922D319FFCD736E328DF942E3FDF8C9" bold="true" box="[224,250,1784,1808]" pageId="62" pageNumber="63">IP</emphasis>
|
||
= mesal plate of vulva;
|
||
<emphasis id="B922D319FFCD736E3396F942E11EF8C9" bold="true" box="[507,537,1784,1808]" pageId="62" pageNumber="63">ml</emphasis>
|
||
= membranous lobe;
|
||
<emphasis id="B922D319FFCD736E316CF942E011F8C9" bold="true" box="[769,790,1784,1808]" pageId="62" pageNumber="63">O</emphasis>
|
||
= operculum;
|
||
<emphasis id="B922D319FFCD736E31DFF942E0DEF8C9" bold="true" box="[946,985,1784,1808]" pageId="62" pageNumber="63">Pre</emphasis>
|
||
= prefemur;
|
||
<emphasis id="B922D319FFCD736E3609F942E781F8C9" bold="true" box="[1124,1158,1784,1808]" pageId="62" pageNumber="63">s-p</emphasis>
|
||
= sclerotized spot;
|
||
<emphasis id="B922D319FFCD736E373AF942E681F8C9" bold="true" box="[1367,1414,1784,1808]" pageId="62" pageNumber="63">scl-t</emphasis>
|
||
= sclerotized teeth;
|
||
<emphasis id="B922D319FFCD736E333EF8A0E272F8EB" bold="true" box="[339,373,1818,1842]" pageId="62" pageNumber="63">SR</emphasis>
|
||
= stridulation ribs. Scale bars =
|
||
<quantity id="4CAEA2EEFFCD736E30BDF8A1E016F8EB" box="[720,785,1819,1842]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="62" pageNumber="63" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCD736F32AAF8DEE0E6FF01" blockId="62.[151,1436,1892,2034]" lastBlockId="63.[151,1436,152,216]" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
|
||
<emphasis id="B922D319FFCD736E32AAF8DEE29EF8A4" box="[199,409,1892,1917]" italics="true" pageId="62" pageNumber="63">Anterior telopods</emphasis>
|
||
(Figs 42B–D): Podomeres 1–3 covered on both sides with numerous long, isolated hairs. Harp with two stridulation ribs, mesal rib smaller than lateral one, both ribs well-developed. Second podomere process lobe-like and little curved with rounded edges, protruding up to half of third podomere height, apically with sclerotized spots. Third podomere long, almost as long as first and second podomere combined. Cavity mesally with numerous sclerotized spots and five thin, sclerotized spines, laterally with three crenulated, small teeth. One sclerotized spot located closely to tip.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFCC736F32FAF8D1E0F0F832" blockId="63.[151,1436,1899,2027]" pageId="63" pageNumber="64">
|
||
<emphasis id="B922D319FFCC736F32FAF8D1E22BF85A" bold="true" box="[151,300,1899,1923]" pageId="63" pageNumber="64">FIGURE 043</emphasis>
|
||
.
|
||
<taxonomicName id="4C567488FFCC736F3354F8D6E11EF85A" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[313,537,1900,1923]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="63" pageNumber="64" phylum="Arthropoda" rank="species" species="tsingy" status="sp. nov.">
|
||
<emphasis id="B922D319FFCC736F3354F8D6E11EF85A" box="[313,537,1900,1923]" italics="true" pageId="63" pageNumber="64">Zoosphaerium tsingy</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B922D319FFCC736F3072F8D1E15DF85A" bold="true" box="[543,602,1899,1923]" pageId="63" pageNumber="64">
|
||
<taxonomicNameLabel id="A2116E62FFCC736F3072F8D1E15DF85A" box="[543,602,1899,1923]" pageId="63" pageNumber="64" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<typeStatus id="54EDB1A9FFCC736F300AF8D6E1C1F85A" box="[615,710,1900,1923]" pageId="63" pageNumber="64" type="paratype">paratype</typeStatus>
|
||
, SEM, A–D antenna,
|
||
<emphasis id="B922D319FFCC736F31C0F8D1E0C7F85A" bold="true" box="[941,960,1899,1923]" pageId="63" pageNumber="64">A</emphasis>
|
||
: apical view on disc;
|
||
<emphasis id="B922D319FFCC736F36CAF8D1E7BEF85A" bold="true" box="[1191,1209,1899,1923]" pageId="63" pageNumber="64">B</emphasis>
|
||
: lateral view;
|
||
<emphasis id="B922D319FFCC736F373DF8D1E664F85A" bold="true" box="[1360,1379,1899,1923]" pageId="63" pageNumber="64">C</emphasis>
|
||
: first antennomere, lateral view;
|
||
<emphasis id="B922D319FFCC736F33D7F834E2CAF87F" bold="true" box="[442,461,1934,1958]" pageId="63" pageNumber="64">D</emphasis>
|
||
: margin between fifth and sixth antennomere, lateral view;
|
||
<emphasis id="B922D319FFCC736F3622F834E766F87F" bold="true" box="[1103,1121,1934,1958]" pageId="63" pageNumber="64">E</emphasis>
|
||
: gnathochilarium, right inner palpus and rudimentary lateral palpus with five sensory cones. Abbreviations:
|
||
<emphasis id="B922D319FFCC736F366DF80BE71FF810" bold="true" box="[1024,1048,1969,1993]" pageId="63" pageNumber="64">iP</emphasis>
|
||
= inner palpus;
|
||
<emphasis id="B922D319FFCC736F36B9F80BE7F1F810" bold="true" box="[1236,1270,1969,1993]" pageId="63" pageNumber="64">rlp</emphasis>
|
||
= rudimentary lateral palpus;
|
||
<emphasis id="B922D319FFCC736F3359F869E24AF832" bold="true" box="[308,333,2003,2027]" pageId="63" pageNumber="64">sb</emphasis>
|
||
=
|
||
<emphasis id="B922D319FFCC736F3304F86EE132F832" box="[361,565,2004,2027]" italics="true" pageId="63" pageNumber="64">sensilla basiconica</emphasis>
|
||
;
|
||
<emphasis id="B922D319FFCC736F302EF869E157F832" bold="true" box="[579,592,2003,2027]" pageId="63" pageNumber="64">v</emphasis>
|
||
= depression on posterior side of palpi.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFB3731032FAF991E15EF951" blockId="64.[151,1436,1579,1673]" pageId="64" pageNumber="65">
|
||
<emphasis id="B922D319FFB3731032FAF991E228F99A" bold="true" box="[151,303,1579,1603]" pageId="64" pageNumber="65">FIGURE 044</emphasis>
|
||
.
|
||
<taxonomicName id="4C567488FFB373103350F996E118F99A" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[317,543,1580,1603]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="64" pageNumber="65" phylum="Arthropoda" rank="species" species="tsingy" status="sp. nov.">
|
||
<emphasis id="B922D319FFB373103350F996E118F99A" box="[317,543,1580,1603]" italics="true" pageId="64" pageNumber="65">Zoosphaerium tsingy</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B922D319FFB37310304AF991E162F99A" bold="true" box="[551,613,1579,1603]" pageId="64" pageNumber="65">
|
||
<taxonomicNameLabel id="A2116E62FFB37310304AF991E162F99A" box="[551,613,1579,1603]" pageId="64" pageNumber="65" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<typeStatus id="54EDB1A9FFB373103019F996E1D4F99A" box="[628,723,1580,1603]" pageId="64" pageNumber="65" type="paratype">paratype</typeStatus>
|
||
, SEM.
|
||
<emphasis id="B922D319FFB373103149F991E030F99A" bold="true" box="[804,823,1579,1603]" pageId="64" pageNumber="65">A</emphasis>
|
||
: tentorium towards endochilarium, dorsal view;
|
||
<emphasis id="B922D319FFB373103725F991E65DF99A" bold="true" box="[1352,1370,1579,1603]" pageId="64" pageNumber="65">B</emphasis>
|
||
: right mandible, mesal view. Abbreviations:
|
||
<emphasis id="B922D319FFB373103058F9F4E15BF9BF" bold="true" box="[565,604,1614,1638]" pageId="64" pageNumber="65">3iT</emphasis>
|
||
= 3 inner teeth;
|
||
<emphasis id="B922D319FFB37310317CF9F4E036F9BF" bold="true" box="[785,817,1614,1638]" pageId="64" pageNumber="65">Co</emphasis>
|
||
= condylus;
|
||
<emphasis id="B922D319FFB3731031D2F9F4E0FBF9BF" bold="true" box="[959,1020,1614,1638]" pageId="64" pageNumber="65">Endo</emphasis>
|
||
= endochilarium;
|
||
<emphasis id="B922D319FFB3731036A9F9F4E7E5F9BF" bold="true" box="[1220,1250,1614,1638]" pageId="64" pageNumber="65">eT</emphasis>
|
||
= external tooth;
|
||
<emphasis id="B922D319FFB3731032FAF9CBE3BBF950" bold="true" box="[151,188,1649,1673]" pageId="64" pageNumber="65">mp</emphasis>
|
||
= molar plate;
|
||
<emphasis id="B922D319FFB373103332F9CBE278F950" bold="true" box="[351,383,1649,1673]" pageId="64" pageNumber="65">pL</emphasis>
|
||
= pectinate lamella.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFB3731032FAF90DE21EF852" blockId="64.[151,1436,1718,2007]" pageId="64" pageNumber="65">
|
||
<emphasis id="B922D319FFB3731032FAF90DE276F909" box="[151,369,1719,1744]" italics="true" pageId="64" pageNumber="65">Posterior telopods</emphasis>
|
||
(Figs 42E, F): Movable finger with well-rounded tip, apically not tapering; hollowed-out inner margin with two non-sclerotized lobes and 3 or 4 sclerotized spines, posterior aspect with
|
||
<emphasis id="B922D319FFB373103749F966E658F92C" box="[1316,1375,1756,1781]" italics="true" pageId="64" pageNumber="65">circa</emphasis>
|
||
nine small, sclerotized teeth. Movable finger slightly longer than fixed finger, the latter straight, as wide as third podomere. Fixed finger at margin with numerous sclerotized spots. All podomeres covered on both sides with numerous, long isolated hairs, only tips of chela glabrous (Figs 42E, F). Inner horns of syncoxite with pointed, slender tip.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFB3731132AAF82CE1C7FF01" blockId="64.[151,1436,1718,2007]" lastBlockId="65.[151,1436,152,371]" lastPageId="65" lastPageNumber="66" pageId="64" pageNumber="65">
|
||
<emphasis id="B922D319FFB3731032AAF82CE2E9F869" bold="true" box="[199,494,1942,1968]" pageId="64" pageNumber="65">Intraspecific variation:</emphasis>
|
||
Two males of the type series which are in all other aspects similar to the
|
||
<typeStatus id="54EDB1A9FFB3731032FAF807E3F9F80E" box="[151,254,1981,2007]" pageId="64" pageNumber="65" type="holotype">holotype</typeStatus>
|
||
have only one stridulation rib on each harp. One male of the type series has only four apical sensory cones on both antennae. The shape of the female operculum varies slightly in different sized females. The operculum is not notched in immature females.
|
||
</paragraph>
|
||
<paragraph id="8BE90F0BFFB2731132AAFF5FE64BFEAA" blockId="65.[151,1436,152,371]" pageId="65" pageNumber="66">
|
||
The (only female) specimen from Tsimembo differs in the number of antennal cones (31, maximum
|
||
<quantity id="4CAEA2EEFFB27311370DFF5FE69BFF26" box="[1376,1436,229,255]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" pageId="65" pageNumber="66" unit="in" value="27.0">27 in</quantity>
|
||
Bemaraha specimens) and number of tarsal spines (first leg with 3 or 4, while
|
||
<quantity id="4CAEA2EEFFB273113629FEB6E788FEFF" box="[1092,1167,268,294]" metricMagnitude="-1" metricUnit="m" metricValue="1.397" metricValueMax="1.778" metricValueMin="1.016" pageId="65" pageNumber="66" unit="in" value="5.5" valueMax="7.0" valueMin="4.0">4–7 in</quantity>
|
||
Bemaraha specimens) from the
|
||
<typeStatus id="54EDB1A9FFB27311336BFE88E23FFE95" box="[262,312,306,332]" pageId="65" pageNumber="66">type</typeStatus>
|
||
series. Those differences are treated as intraspecific variation. When male specimens become available or molecular studies are undertaken, this treatment might be viewed as an oversimplification.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |