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<document id="950F252703AD82E36F6809E10C4C1BA9" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFF98733D32FAFF2DE728FC49" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium denticulatum Wesener 2009, new species" docType="treatment" docUuid="B2B882E6-7D9B-4CAB-AEC2-51F2ACEF9A8D" docUuidSource="ZooBank" docVersion="5" lastPageNumber="110" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="108" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DA11DB0D67A821B1989DBF85699FCD6B">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
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<mods:name id="D5EACBF79D791263B894B563D776C046" type="personal">
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<mods:namePart id="43D08DC668E2FC4009045F3721341496">Wesener, Thomas</mods:namePart>
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<mods:typeOfResource id="7169625B3B7F70B16773C03F3A158689">text</mods:typeOfResource>
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<mods:title id="0425362B935DA1375D4FC194DD942855">Zootaxa</mods:title>
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<mods:part id="1B54C3D498731AA55A42A7B3679F33E8">
<mods:date id="B26D581C81B52FF019E304814064EB30">2009</mods:date>
<mods:detail id="A7F5D2C2A125D62D075FD6D10DE390CC" type="pubDate">
<mods:number id="68F6EB40B77D4AF89332F7E0A9388E35">2009-05-11</mods:number>
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<mods:number id="6AC9B67010173BE82E8D999DAD9242B7">2097</mods:number>
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<mods:classification id="536B37D80A71A18CC931DBAC792BA264">journal article</mods:classification>
<mods:identifier id="F89D97B95C62CC1DA02A2D8B9C263573" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
<mods:identifier id="61E21074B90DFEB4CA0C646F7D32A32F" type="ISSN">1175­5334</mods:identifier>
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<treatment id="03FFBE1DFF98733D32FAFF2DE728FC49" ID-DOI="http://doi.org/10.5281/zenodo.5325525" ID-Zenodo-Dep="5325525" LSID="urn:lsid:zoobank.org:act:B2B882E6-7D9B-4CAB-AEC2-51F2ACEF9A8D" httpUri="http://treatment.plazi.org/id/03FFBE1DFF98733D32FAFF2DE728FC49" lastPageId="109" lastPageNumber="110" pageId="107" pageNumber="108">
<subSubSection id="C34C5C80FF98733B32FAFF2DE18BFF68" box="[151,652,151,177]" pageId="107" pageNumber="108" type="nomenclature">
<paragraph id="8BE90F0BFF98733B32FAFF2DE18BFF68" blockId="107.[151,652,151,177]" box="[151,652,151,177]" pageId="107" pageNumber="108">
<heading id="D0A1B867FF98733B32FAFF2DE18BFF68" bold="true" box="[151,652,151,177]" fontSize="11" level="1" pageId="107" pageNumber="108" reason="1">
<emphasis id="B922D319FF98733B32FAFF2DE18BFF68" bold="true" box="[151,652,151,177]" pageId="107" pageNumber="108">
<emphasis id="B922D319FF98733B32FAFF2DE2F3FF68" bold="true" box="[151,500,151,177]" italics="true" pageId="107" pageNumber="108">
<taxonomicName id="4C567488FF98733B32FAFF2DE2E8FF68" ID-CoL="7GGPZ" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,495,151,177]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="107" pageNumber="108" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">Zoosphaerium denticulatum</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A2116E62FF98733B3391FF2DE18BFF68" box="[508,652,151,177]" pageId="107" pageNumber="108" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF98733B32FAFF5AE124FE4F" pageId="107" pageNumber="108" type="etymology">
<paragraph id="8BE90F0BFF98733B32FAFF5AE3D0FEFB" blockId="107.[151,1436,224,715]" pageId="107" pageNumber="108">
<emphasis id="B922D319FF98733B32FAFF5AE287FF23" bold="true" box="[151,384,224,250]" pageId="107" pageNumber="108">Derivatio nominis:</emphasis>
<taxonomicName id="4C567488FF98733B33EAFF5BE127FF23" authorityName="Wesener" authorityYear="2009" box="[391,544,225,250]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="107" pageNumber="108" phylum="Arthropoda" rank="species" species="denticulatum">
<emphasis id="B922D319FF98733B33EAFF5BE127FF23" box="[391,544,225,250]" italics="true" pageId="107" pageNumber="108">denticulatum</emphasis>
</taxonomicName>
, adjective, refers to the numerous small teeth on the posterior telopods (Fig. 72E).
</paragraph>
<paragraph id="8BE90F0BFF98733B32ABFE94E124FE4F" blockId="107.[151,1436,224,715]" pageId="107" pageNumber="108">
<emphasis id="B922D319FF98733B32ABFE94E2A5FE91" bold="true" box="[198,418,302,328]" pageId="107" pageNumber="108">Studied material:</emphasis>
<materialsCitation id="3B3E0556FF98733B33C0FE94E124FE4F" collectingDate="2000-11-12" collectingDateMax="2000-11-19" collectingDateMin="2000-11-12" collectionCode="FMMC" collectorName="S. Goodman" country="Madagascar" county="Locus" elevation="1075" latitude="-21.511667" location="Andrombovata" longLatPrecision="126" longitude="47.41" municipality="Province" pageId="107" pageNumber="108" specimenCode="FMMC 5446" specimenCount="1" stateProvince="Fianarantsoa" typeStatus="holotype">
<emphasis id="B922D319FF98733B33C0FE94E12FFE91" bold="true" box="[429,552,302,328]" pageId="107" pageNumber="108">
<typeStatus id="54EDB1A9FF98733B33C0FE94E124FE91" box="[429,547,302,328]" pageId="107" pageNumber="108" type="holotype">Holotype</typeStatus>
:
</emphasis>
1 M (
<quantity id="4CAEA2EEFF98733B3014FE94E1D6FE91" box="[633,721,302,328]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="107" pageNumber="108" unit="mm" value="30.0">30 mm</quantity>
long),
<emphasis id="B922D319FF98733B3147FE94E0CAFE91" bold="true" box="[810,973,302,328]" pageId="107" pageNumber="108">
<specimenCode id="DBF0A770FF98733B3147FE94E0CAFE91" box="[810,973,302,328]" collectionCode="FMMC" pageId="107" pageNumber="108">FMMC 5446</specimenCode>
</emphasis>
,
<emphasis id="B922D319FF98733B31B2FE95E784FE91" box="[991,1155,303,328]" italics="true" pageId="107" pageNumber="108">
<collectingCounty id="62887787FF98733B31B2FE95E722FE91" box="[991,1061,303,328]" pageId="107" pageNumber="108">Locus</collectingCounty>
typicus
</emphasis>
:
<collectingCountry id="F3414F9BFF98733B36F8FE94E623FE91" box="[1173,1316,302,328]" name="Madagascar" pageId="107" pageNumber="108">Madagascar</collectingCountry>
,
<collectingMunicipality id="6B8D9571FF98733B3759FE94E69BFE91" box="[1332,1436,302,328]" pageId="107" pageNumber="108">Province</collectingMunicipality>
<collectingRegion id="4992C1E9FF98733B32FAFEEFE237FEB6" box="[151,304,341,367]" country="Madagascar" name="Fianarantsoa" pageId="107" pageNumber="108">Fianarantsoa</collectingRegion>
,
<location id="8E8959D0FF98733B3351FEEFE2E8FEB6" LSID="urn:lsid:plazi:treatment:03FFBE1DFF98733D32FAFF2DE728FC49:8E8959D0FF98733B3351FEEFE2E8FEB6" box="[316,495,341,367]" country="Madagascar" county="Locus" latitude="-21.511667" longLatPrecision="126" longitude="47.41" municipality="Province" name="Andrombovata" pageId="107" pageNumber="108" stateProvince="Fianarantsoa">Andrombovata</location>
,
<locationDeviation id="9A5B1629FF98733B3391FEEFE01EFEB6" box="[508,793,341,367]" location="Andrombovata" pageId="107" pageNumber="108">
<quantity id="4CAEA2EEFF98733B3391FEEFE13EFEB6" box="[508,569,341,367]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="107" pageNumber="108" unit="km" value="2.0">2 km</quantity>
W Andrombovata
</locationDeviation>
, along
<location id="8E8959D0FF98733B311DFEEFE72FFEB6" LSID="urn:lsid:plazi:treatment:03FFBE1DFF98733D32FAFF2DE728FC49:8E8959D0FF98733B311DFEEFE72FFEB6" box="[880,1064,341,367]" country="Madagascar" county="Locus" latitude="-21.511667" longLatPrecision="126" longitude="47.41" municipality="Province" name="Tatamaly River" pageId="107" pageNumber="108" stateProvince="Fianarantsoa">Tatamaly River</location>
,
<geoCoordinate id="EE6269CCFF98733B3658FEEFE7ADFEB6" box="[1077,1194,341,367]" degrees="21" direction="south" minutes="30.7" orientation="latitude" pageId="107" pageNumber="108" precision="92" value="-21.511667">21°30.7'S</geoCoordinate>
,
<geoCoordinate id="EE6269CCFF98733B36DAFEEFE62BFEB6" box="[1207,1324,341,367]" degrees="47" direction="east" minutes="24.6" orientation="longitude" pageId="107" pageNumber="108" precision="92" value="47.41">47°24.6'E</geoCoordinate>
,
<quantity id="4CAEA2EEFF98733B3757FEEFE691FEB6" box="[1338,1430,341,367]" metricMagnitude="3" metricUnit="m" metricValue="1.075" pageId="107" pageNumber="108" unit="m" value="1075.0">
<elevation id="007BE838FF98733B3757FEEFE691FEB6" box="[1338,1430,341,367]" metricMagnitude="3" metricUnit="m" metricValue="1.075" pageId="107" pageNumber="108" unit="m" value="1075.0">1075 m</elevation>
</quantity>
, coll.
<collectorName id="26A36ADDFF98733B32BCFEC6E261FE4F" box="[209,358,380,406]" pageId="107" pageNumber="108">S. Goodman</collectorName>
,
<date id="FFE829CBFF98733B331EFEC6E118FE4F" box="[371,543,380,406]" pageId="107" pageNumber="108" value="2000-11-12" valueMax="2000-11-19" valueMin="2000-11-12">
<collectingDate id="EFACD023FF98733B331EFEC6E118FE4F" box="[371,543,380,406]" pageId="107" pageNumber="108" value="2000-11-12" valueMax="2000-11-19" valueMin="2000-11-12">1219.xi.2000</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF98733B32ABFE18E664FE65" box="[198,1379,418,444]" pageId="107" pageNumber="108" type="distribution">
<paragraph id="8BE90F0BFF98733B32ABFE18E664FE65" blockId="107.[151,1436,224,715]" box="[198,1379,418,444]" pageId="107" pageNumber="108">
<emphasis id="B922D319FF98733B32ABFE18E2F4FE65" bold="true" box="[198,499,418,444]" pageId="107" pageNumber="108">Distribution &amp; Ecology:</emphasis>
Only known from the
<typeStatus id="54EDB1A9FF98733B3092FE18E036FE65" box="[767,817,418,444]" pageId="107" pageNumber="108">type</typeStatus>
locality,
<quantity id="4CAEA2EEFF98733B31F0FE18E0E0FE65" box="[925,999,418,444]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="107" pageNumber="108" unit="km" value="25.0">25 km</quantity>
South of Ranomafana (Fig. 50).
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF98733B32ABFE72E12CF8FF" pageId="107" pageNumber="108" type="diagnosis">
<paragraph id="8BE90F0BFF98733B32ABFE72E747FD12" blockId="107.[151,1436,224,715]" pageId="107" pageNumber="108">
<emphasis id="B922D319FF98733B32ABFE72E24FFE3B" bold="true" box="[198,328,456,482]" pageId="107" pageNumber="108">Diagnosis:</emphasis>
Up to
<quantity id="4CAEA2EEFF98733B33F0FE73E2F7FE3A" box="[413,496,457,483]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="107" pageNumber="108" unit="mm" value="30.0">30 mm</quantity>
long. Color blackish with dark green legs. Tergites glabrous. Anal shield wellrounded, covered with dense field of minute setae. Disc of antenna with
<emphasis id="B922D319FF98733B3660FE4AE74FFDD0" box="[1037,1096,496,521]" italics="true" pageId="107" pageNumber="108">circa</emphasis>
26 apical cones (Fig. 73B). Movable finger of posterior telopods almost glabrous, broad, with well-rounded tip, with three sclerotized spines (Fig. 72D); posterior side with&gt;25 small, sclerotized teeth (Fig. 72E). Fixed finger long and slender, basally with one small spine, distally at inner margin with numerous sclerotized spots (Fig. 72D). Fixed finger shorter than movable finger. One stridulation rib on each male harp (Fig. 72A). Anal shield with two locking carinae, anterior small, posterior&gt; 2.5 times longer than first. Female unknown.
</paragraph>
<paragraph id="8BE90F0BFF98733B32FAF9E8E092F955" blockId="107.[151,1435,1618,1677]" pageId="107" pageNumber="108">
<emphasis id="B922D319FF98733B32FAF9E8E22AF9B3" bold="true" box="[151,301,1618,1642]" pageId="107" pageNumber="108">FIGURE 073</emphasis>
.
<taxonomicName id="4C567488FF98733B3354F9E9E162F9B3" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[313,613,1619,1642]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="107" pageNumber="108" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
<emphasis id="B922D319FF98733B3354F9E9E162F9B3" box="[313,613,1619,1642]" italics="true" pageId="107" pageNumber="108">Zoosphaerium denticulatum</emphasis>
</taxonomicName>
<emphasis id="B922D319FF98733B3006F9E8E1A1F9B3" bold="true" box="[619,678,1618,1642]" pageId="107" pageNumber="108">
<taxonomicNameLabel id="A2116E62FF98733B3006F9E8E1A1F9B3" box="[619,678,1618,1642]" pageId="107" pageNumber="108" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FF98733B30DEF9E9E014F9B3" box="[691,787,1619,1642]" pageId="107" pageNumber="108" type="holotype">holotype</typeStatus>
, SEM, antenna
<emphasis id="B922D319FF98733B31D7F9E8E0CAF9B3" bold="true" box="[954,973,1618,1642]" pageId="107" pageNumber="108">A</emphasis>
: first antennomere with
<emphasis id="B922D319FF98733B36BCF9E9E69CF9B3" box="[1233,1435,1619,1642]" italics="true" pageId="107" pageNumber="108">sensilla basiconica</emphasis>
(arrow);
<emphasis id="B922D319FF98733B3298F9CFE200F954" bold="true" box="[245,263,1653,1677]" pageId="107" pageNumber="108">B</emphasis>
: apical view on sixth antennomere with disc;
<emphasis id="B922D319FF98733B309CF9CFE003F954" bold="true" box="[753,772,1653,1677]" pageId="107" pageNumber="108">C</emphasis>
: lateral view.
</paragraph>
<paragraph id="8BE90F0BFF98733B32ABF904E12CF8FF" blockId="107.[151,1436,1726,2023]" pageId="107" pageNumber="108">
<emphasis id="B922D319FF98733B32ABF904E28BF901" bold="true" box="[198,396,1726,1752]" pageId="107" pageNumber="108">Similar species:</emphasis>
<emphasis id="B922D319FF98733B33F9F905E152F901" box="[404,597,1727,1752]" italics="true" pageId="107" pageNumber="108">
<taxonomicName id="4C567488FF98733B33F9F905E156F901" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[404,593,1727,1752]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="107" pageNumber="108" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">Z. denticulatum</taxonomicName>
,
</emphasis>
<emphasis id="B922D319FF98733B3033F904E1A5F901" bold="true" box="[606,674,1726,1752]" pageId="107" pageNumber="108">
<taxonomicNameLabel id="A2116E62FF98733B3033F904E1A5F901" box="[606,674,1726,1752]" pageId="107" pageNumber="108" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
features a unique combination of characters, as one stridulation rib on the harp, more than 20 apical cones on the antennae and a row of crenulated teeth on the third podomere of anterior telopods (Figs 72B, C).
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF98733D32ABF888E728FC49" lastPageId="109" lastPageNumber="110" pageId="107" pageNumber="108" type="description">
<paragraph id="8BE90F0BFF98733B32ABF888E10AF8AA" blockId="107.[151,1436,1726,2023]" pageId="107" pageNumber="108">
<emphasis id="B922D319FF98733B32ABF888E267F895" bold="true" box="[198,352,1842,1868]" pageId="107" pageNumber="108">Description:</emphasis>
<emphasis id="B922D319FF98733B3304F889E2F1F895" box="[361,502,1843,1868]" italics="true" pageId="107" pageNumber="108">Body length</emphasis>
:
<typeStatus id="54EDB1A9FF98733B306BF889E174F894" box="[518,627,1843,1869]" pageId="107" pageNumber="108" type="holotype">Holotype</typeStatus>
male: length
<quantity id="4CAEA2EEFF98733B3178F889E06DF894" box="[789,874,1843,1869]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="107" pageNumber="108" unit="mm" value="30.0">30 mm</quantity>
, width of thoracic shield up to
<quantity id="4CAEA2EEFF98733B36B7F889E641F894" box="[1242,1350,1843,1869]" metricMagnitude="-2" metricUnit="m" metricValue="1.53" pageId="107" pageNumber="108" unit="mm" value="15.3">15.3 mm</quantity>
, height of thoracic shield up to
<quantity id="4CAEA2EEFF98733B33C1F8E3E10EF8AA" box="[428,521,1881,1907]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="107" pageNumber="108" unit="mm" value="8.5">8.5 mm</quantity>
.
</paragraph>
<paragraph id="8BE90F0BFF98733B32AAF83AE04EF843" blockId="107.[151,1436,1726,2023]" box="[199,841,1920,1946]" pageId="107" pageNumber="108">
<emphasis id="B922D319FF98733B32AAF83AE229F840" box="[199,302,1920,1945]" italics="true" pageId="107" pageNumber="108">Habitus:</emphasis>
Medium sized, black, glabrous pill millipede.
</paragraph>
<paragraph id="8BE90F0BFF98733B32AAF81DE284F83E" blockId="107.[151,1436,1726,2023]" pageId="107" pageNumber="108">
<emphasis id="B922D319FF98733B32AAF81DE256F819" box="[199,337,1959,1984]" italics="true" pageId="107" pageNumber="108">Coloration:</emphasis>
Head, collum, thoracic shield, tergites and anal shield colored completely blackish, antennae and legs dark green.
</paragraph>
<paragraph id="8BE90F0BFF9F733C32AAFF22E116FF6B" blockId="108.[151,1436,152,564]" box="[199,529,152,178]" pageId="108" pageNumber="109">
<emphasis id="B922D319FF9F733C32AAFF22E217FF68" box="[199,272,152,177]" italics="true" pageId="108" pageNumber="109">Head:</emphasis>
Eyes with&gt;75 ocelli.
</paragraph>
<paragraph id="8BE90F0BFF9F733C32AAFF05E63FFEFF" blockId="108.[151,1436,152,564]" pageId="108" pageNumber="109">
<emphasis id="B922D319FF9F733C32AAFF05E238FF01" box="[199,319,191,216]" italics="true" pageId="108" pageNumber="109">Antennae:</emphasis>
Length of antennomeres: 1&gt;2=3&gt;4=5&lt;6 (Fig. 73C), sixth antennomere bearing disc with 2627 sensory cones (Fig. 73B). First antennomere remarkably broader than others, short, with groove and 1 or 2 rows of
<emphasis id="B922D319FF9F733C3341FEB6E10BFEFC" box="[300,524,268,293]" italics="true" pageId="108" pageNumber="109">sensilla basiconica</emphasis>
(Fig.73A). Sclerotized teeth at base of antennomeres 13 (Fig. 73C).
</paragraph>
<paragraph id="8BE90F0BFF9F733C32AAFE88E2F0FE95" blockId="108.[151,1436,152,564]" box="[199,503,306,332]" pageId="108" pageNumber="109">
<emphasis id="B922D319FF9F733C32AAFE88E257FE92" box="[199,336,306,331]" italics="true" pageId="108" pageNumber="109">Mouthparts</emphasis>
not dissected.
</paragraph>
<paragraph id="8BE90F0BFF9F733C32AAFEE3E1AEFEAA" blockId="108.[151,1436,152,564]" box="[199,681,345,371]" pageId="108" pageNumber="109">
<emphasis id="B922D319FF9F733C32AAFEE3E220FEAB" box="[199,295,345,370]" italics="true" pageId="108" pageNumber="109">Collum:</emphasis>
Median part of collum glabrous.
</paragraph>
<paragraph id="8BE90F0BFF9F733C32AAFE3AE17AFE43" blockId="108.[151,1436,152,564]" box="[199,637,384,410]" pageId="108" pageNumber="109">
<emphasis id="B922D319FF9F733C32AAFE3AE279FE40" box="[199,382,384,409]" italics="true" pageId="108" pageNumber="109">Thoracic shield</emphasis>
without peculiarities.
</paragraph>
<paragraph id="8BE90F0BFF9F733C32AAFE1DE021FE19" blockId="108.[151,1436,152,564]" box="[199,806,422,448]" pageId="108" pageNumber="109">
<emphasis id="B922D319FF9F733C32AAFE1DE224FE19" box="[199,291,423,448]" italics="true" pageId="108" pageNumber="109">Tergites</emphasis>
: Paratergites projecting slightly posteriorly.
</paragraph>
<paragraph id="8BE90F0BFF9F733C32FAFE77E210FDED" blockId="108.[151,1436,152,564]" pageId="108" pageNumber="109">
<emphasis id="B922D319FF9F733C32FAFE77E237FE3F" box="[151,304,461,486]" italics="true" pageId="108" pageNumber="109">Endotergum:</emphasis>
Inner area with broad conical spines and long hair. Two rows of small, circular cuticular impression, regularly distributed (Fig. 74E). Single row of marginal bristles, reaching 1/2 towards margin (Fig. 74E).
</paragraph>
<paragraph id="8BE90F0BFF9F733C32FAF838E1F6F83A" blockId="108.[151,1436,1922,2019]" pageId="108" pageNumber="109">
<emphasis id="B922D319FF9F733C32FAF838E236F843" bold="true" box="[151,305,1922,1946]" pageId="108" pageNumber="109">FIGURE 074</emphasis>
. SEM, endoterga of midbody tergites,
<emphasis id="B922D319FF9F733C3085F838E1FCF843" bold="true" box="[744,763,1922,1946]" pageId="108" pageNumber="109">A</emphasis>
:
<taxonomicName id="4C567488FF9F733C3160F838E70EF840" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[781,1033,1922,1945]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="tampolo" status="sp. nov.">
<emphasis id="B922D319FF9F733C3160F838E70EF840" box="[781,1033,1922,1945]" italics="true" pageId="108" pageNumber="109">Zoosphaerium tampolo</emphasis>
</taxonomicName>
<emphasis id="B922D319FF9F733C3679F838E753F843" bold="true" box="[1044,1108,1922,1946]" pageId="108" pageNumber="109">
<taxonomicNameLabel id="A2116E62FF9F733C3679F838E753F843" box="[1044,1108,1922,1946]" pageId="108" pageNumber="109" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B922D319FF9F733C360AF838E785F843" bold="true" box="[1127,1154,1922,1946]" pageId="108" pageNumber="109">B:</emphasis>
<taxonomicName id="4C567488FF9F733C36E1F838E604F840" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[1164,1283,1922,1945]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="ignotum" status="sp. nov.">
<emphasis id="B922D319FF9F733C36E1F838E604F840" box="[1164,1283,1922,1945]" italics="true" pageId="108" pageNumber="109">Z. ignotum</emphasis>
</taxonomicName>
<emphasis id="B922D319FF9F733C3762F838E648F843" bold="true" box="[1295,1359,1922,1946]" pageId="108" pageNumber="109">
<taxonomicNameLabel id="A2116E62FF9F733C3762F838E648F843" box="[1295,1359,1922,1946]" pageId="108" pageNumber="109" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B922D319FF9F733C370CF838E673F843" bold="true" box="[1377,1396,1922,1946]" pageId="108" pageNumber="109">C</emphasis>
:
<taxonomicName id="4C567488FF9F733C37EBF838E217F864" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="tainkintana" status="sp. nov.">
<emphasis id="B922D319FF9F733C37EBF838E217F864" italics="true" pageId="108" pageNumber="109">Z. tainkintana</emphasis>
</taxonomicName>
<emphasis id="B922D319FF9F733C3375F81CE252F867" bold="true" box="[280,341,1958,1982]" pageId="108" pageNumber="109">
<taxonomicNameLabel id="A2116E62FF9F733C3375F81CE252F867" box="[280,341,1958,1982]" pageId="108" pageNumber="109" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B922D319FF9F733C3308F81CE27FF867" bold="true" box="[357,376,1958,1982]" pageId="108" pageNumber="109">D</emphasis>
:
<taxonomicName id="4C567488FF9F733C33EAF81CE128F864" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[391,559,1958,1981]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="viridissimum" status="sp. nov.">
<emphasis id="B922D319FF9F733C33EAF81CE128F864" box="[391,559,1958,1981]" italics="true" pageId="108" pageNumber="109">Z. viridissimum</emphasis>
</taxonomicName>
<emphasis id="B922D319FF9F733C305AF81CE173F867" bold="true" box="[567,628,1958,1982]" pageId="108" pageNumber="109">
<taxonomicNameLabel id="A2116E62FF9F733C305AF81CE173F867" box="[567,628,1958,1982]" pageId="108" pageNumber="109" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B922D319FF9F733C30E9F81CE191F867" bold="true" box="[644,662,1958,1982]" pageId="108" pageNumber="109">E</emphasis>
:
<taxonomicName id="4C567488FF9F733C30C9F81CE04AF864" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[676,845,1958,1981]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="108" pageNumber="109" phylum="Arthropoda" rank="species" species="denticulatum" status="sp. nov.">
<emphasis id="B922D319FF9F733C30C9F81CE04AF864" box="[676,845,1958,1981]" italics="true" pageId="108" pageNumber="109">Z. denticulatum</emphasis>
</taxonomicName>
<emphasis id="B922D319FF9F733C3138F81CE095F867" bold="true" box="[853,914,1958,1982]" pageId="108" pageNumber="109">
<taxonomicNameLabel id="A2116E62FF9F733C3138F81CE095F867" box="[853,914,1958,1982]" pageId="108" pageNumber="109" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Abbreviations:
<emphasis id="B922D319FF9F733C362CF81CE753F867" bold="true" box="[1089,1108,1958,1982]" pageId="108" pageNumber="109">ci</emphasis>
= cuticular impressions;
<emphasis id="B922D319FF9F733C370AF81CE683F867" bold="true" box="[1383,1412,1958,1982]" pageId="108" pageNumber="109">IA</emphasis>
= inner area;
<emphasis id="B922D319FF9F733C3362F871E233F83A" bold="true" box="[271,308,1995,2019]" pageId="108" pageNumber="109">mb</emphasis>
= marginal bristles;
<emphasis id="B922D319FF9F733C307DF871E135F83A" bold="true" box="[528,562,1995,2019]" pageId="108" pageNumber="109">mr</emphasis>
= marginal ridge.
</paragraph>
<paragraph id="8BE90F0BFF9E733D32ABFF22E1FDFF26" blockId="109.[151,1437,152,912]" pageId="109" pageNumber="110">
<emphasis id="B922D319FF9E733D32ABFF22E252FF68" box="[198,341,152,177]" italics="true" pageId="109" pageNumber="110">Anal shield:</emphasis>
Well-rounded, neither bell-shaped nor tapered. Completely covered by dense field of minute setae. Underside carries two well-developed black locking carinae, anterior one small, but well-developed. Posterior carina&gt; 2.5 times longer than anterior one
</paragraph>
<paragraph id="8BE90F0BFF9E733D32AAFEB6E642FEAA" blockId="109.[151,1437,152,912]" pageId="109" pageNumber="110">
<emphasis id="B922D319FF9E733D32AAFEB6E20FFEFC" box="[199,264,268,293]" italics="true" pageId="109" pageNumber="110">Legs:</emphasis>
Tarsi of first pair of legs with four, second with five and third pair with seven ventral spines. First two leg pairs with only weakly curved claws and without apical spine. Tarsi of legs 321 with curved claws, 1113 ventral spines and one apical spine. In 9th leg femur 2.2 times, tarsi 5.5 times longer than wide.
</paragraph>
<paragraph id="8BE90F0BFF9E733D32AAFE3AE09FFE43" blockId="109.[151,1437,152,912]" box="[199,920,384,410]" pageId="109" pageNumber="110">
<emphasis id="B922D319FF9E733D32AAFE3AE28BFE40" box="[199,396,384,409]" italics="true" pageId="109" pageNumber="110">Stigmatic plates:</emphasis>
First plate lobe short, with well-rounded tip.
</paragraph>
<paragraph id="8BE90F0BFF9E733D32AAFE1CE296FE19" blockId="109.[151,1437,152,912]" box="[199,401,422,448]" pageId="109" pageNumber="110">Female unknown</paragraph>
<paragraph id="8BE90F0BFF9E733D32AAFE77E0EFFE3E" blockId="109.[151,1437,152,912]" box="[199,1000,461,487]" pageId="109" pageNumber="110">
<emphasis id="B922D319FF9E733D32AAFE77E2DDFE3F" box="[199,474,461,486]" italics="true" pageId="109" pageNumber="110">Male sexual characters</emphasis>
: Anal shield covered by dense field of setae.
</paragraph>
<paragraph id="8BE90F0BFF9E733D32AAFE4EE27BFD71" blockId="109.[151,1437,152,912]" pageId="109" pageNumber="110">
<emphasis id="B922D319FF9E733D32AAFE4EE29BFDD4" box="[199,412,500,525]" italics="true" pageId="109" pageNumber="110">Anterior telopods</emphasis>
(Figs 72AC): Harp with single stridulation rib, rib straight, very long. Second podomere process lobe-like and little curved with rounded edges, reaching only up to 1/3 of third podomere height, apically with sclerotized spots. Third podomere long, cavity mesally with numerous sclerotized spots and three sclerotized spines, laterally with row of 1518 crenulated, large teeth. One sclerotized teeth located closely towards tip.
</paragraph>
<paragraph id="8BE90F0BFF9E733D32AAFD0FE728FC49" blockId="109.[151,1437,152,912]" pageId="109" pageNumber="110">
<emphasis id="B922D319FF9E733D32AAFD0FE2A3FD17" box="[199,420,693,718]" italics="true" pageId="109" pageNumber="110">Posterior telopods</emphasis>
(Figs 72D, E): Movable finger of chela thick, 2.4 times longer than wide, weakly curved, apically slightly tapering. Hollowed-out inner margin with three sclerotized spines, posterior aspect with&gt;25 small, sclerotized teeth. Fixed finger shorter and more slender than movable finger. Fixed finger tip weakly curved, anterior side basally with one short sclerotized spine, apically with numerous sclerotized spots. Finger on both sides covered with few long hairs. Movable finger on posterior side medially with few hairs, remaining parts of posterior telopods glabrous. Inner horns of syncoxite.
</paragraph>
</subSubSection>
</treatment>
</document>