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<document id="E8ADABCA1925532D21C55C9BC934CDBC" ID-DOI="10.11646/zootaxa.2097.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5316536" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629621849155" checkinUser="felipe" docAuthor="Wesener, Thomas" docDate="2009" docId="03FFBE1DFF81732532FAFC36E78AFACA" docLanguage="en" docName="zt02097p134.pdf" docOrigin="Zootaxa 2097 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Zoosphaerium micropiligerum Wesener 2009, new species" docType="treatment" docUuid="18F56A3C-358B-44FC-A9EC-2FB108476FFB" docUuidSource="ZooBank" docVersion="5" lastPageNumber="118" masterDocId="FFC6C665FFF37350326DFFBAE307FFD9" masterDocTitle="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)" masterLastPageNumber="134" masterPageNumber="1" pageNumber="115" updateTime="1699084836552" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B60716C7958D55AD984471A8461BAEB6">Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on microendemism (Diplopoda, Sphaerotheriida)</mods:title>
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<mods:namePart id="68F6A8BA398067A3BB94A402C326F063">Wesener, Thomas</mods:namePart>
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<mods:title id="AD030EC232398630FD093D879E79A361">Zootaxa</mods:title>
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<mods:date id="FED905954FB8DF2C34311C54EB042949">2009</mods:date>
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<mods:number id="5EDB02C99DAF00FDC54934320A93A5E9">2009-05-11</mods:number>
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<mods:number id="E02C59A9F551A37F61ADD42BA783D837">2097</mods:number>
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<mods:url id="35554F61DC16516651206FA81EAD2B59">https://biotaxa.org/Zootaxa/article/view/zootaxa.2097.1.1</mods:url>
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<mods:classification id="2FEA404BD979014D612CB0D367725FBD">journal article</mods:classification>
<mods:identifier id="683DAAEE69C04D60B726C33441FC6651" type="DOI">10.11646/zootaxa.2097.1.1</mods:identifier>
<mods:identifier id="291385FE3BFF2E25D8A76C3CDFDDFD14" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="B9B90972C0EE5462E1D7EF7BD0AA1B96" type="Zenodo-Dep">5316536</mods:identifier>
<mods:identifier id="B602D46B4448D1D2C9C91D6FD05AEA4E" type="ZooBank">0E18167A-78EB-4079-8BF4-972A4AC88C69</mods:identifier>
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<treatment id="03FFBE1DFF81732532FAFC36E78AFACA" ID-DOI="http://doi.org/10.5281/zenodo.5325533" ID-Zenodo-Dep="5325533" LSID="urn:lsid:zoobank.org:act:18F56A3C-358B-44FC-A9EC-2FB108476FFB" httpUri="http://treatment.plazi.org/id/03FFBE1DFF81732532FAFC36E78AFACA" lastPageId="117" lastPageNumber="118" pageId="114" pageNumber="115">
<subSubSection id="C34C5C80FF81732232FAFC36E1A0FC7C" box="[151,679,907,934]" pageId="114" pageNumber="115" type="nomenclature">
<paragraph id="8BE90F0BFF81732232FAFC36E1A0FC7C" blockId="114.[151,679,907,934]" box="[151,679,907,934]" pageId="114" pageNumber="115">
<heading id="D0A1B867FF81732232FAFC36E1A0FC7C" bold="true" box="[151,679,907,934]" fontSize="11" level="1" pageId="114" pageNumber="115" reason="1">
<emphasis id="B922D319FF81732232FAFC36E1A0FC7C" bold="true" box="[151,679,907,934]" pageId="114" pageNumber="115">
<taxonomicName id="4C567488FF81732232FAFC36E10EFC7F" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[151,521,908,934]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="micropiligerum" status="sp. nov.">
<emphasis id="B922D319FF81732232FAFC36E10EFC7F" bold="true" box="[151,521,908,934]" italics="true" pageId="114" pageNumber="115">Zoosphaerium micropiligerum</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A2116E62FF817322307AFC31E1A0FC7C" box="[535,679,907,933]" pageId="114" pageNumber="115" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF81732232FAFC6FE185FAAB" pageId="114" pageNumber="115" type="etymology">
<paragraph id="8BE90F0BFF81732232FAFC6FE73FFBCF" blockId="114.[151,1436,981,2013]" pageId="114" pageNumber="115">
<emphasis id="B922D319FF81732232FAFC6FE284FC36" bold="true" box="[151,387,981,1007]" pageId="114" pageNumber="115">Derivatio nominis:</emphasis>
<taxonomicName id="4C567488FF81732233E1FC6FE146FC37" authorityName="Wesener" authorityYear="2009" box="[396,577,981,1006]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="micropiligerum">
<emphasis id="B922D319FF81732233E1FC6FE146FC37" box="[396,577,981,1006]" italics="true" pageId="114" pageNumber="115">micropiligerum</emphasis>
</taxonomicName>
, adjective. Consisting out of micro = small, and piligerum, referring to the similarity of this species with
<taxonomicName id="4C567488FF817322304FFC46E733FBCF" authority="(de Saussure &amp; Zehntner, 1897)" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" box="[546,1076,1020,1046]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="piligerum">
<emphasis id="B922D319FF817322304FFC46E1B4FBCC" box="[546,691,1020,1045]" italics="true" pageId="114" pageNumber="115">Z. piligerum</emphasis>
(
<bibRefCitation id="EFC772FAFF81732230AEFC46E72CFBCF" author="de Saussure, H. &amp; Zehntner, L." box="[707,1067,1020,1046]" pageId="114" pageNumber="115" pagination="1 - 12" refId="ref60128" refString="de Saussure, H. &amp; Zehntner, L. (1897) Atlas de l'histoire naturelle des Myriapodes, In: Grandidier (Ed.) Histoire physique, naturelle et politique de Madagascar. 27 (53), Alfred Grandidier, Paris, France, plates 1 - 12." type="book chapter" year="1897">de Saussure &amp; Zehntner, 1897</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<paragraph id="8BE90F0BFF81732232ABFB98E039FB53" blockId="114.[151,1436,981,2013]" pageId="114" pageNumber="115">
<emphasis id="B922D319FF81732232ABFB98E2A7FBE5" bold="true" box="[198,416,1058,1084]" pageId="114" pageNumber="115">Studied material:</emphasis>
<materialsCitation id="3B3E0556FF81732233C5FB98E039FB53" collectingDate="1995-11-27" collectingDateMax="1995-12-05" collectingDateMin="1995-11-27" collectionCode="FMMC" collectorName="SE Andranodambo &amp; S. Goodman" country="Madagascar" elevation="1875" latitude="-24.561666" location="Andohahela" longLatPrecision="125" longitude="46.72167" municipality="Locus" pageId="114" pageNumber="115" specimenCode="FMMC 5399" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<emphasis id="B922D319FF81732233C5FB98E11DFBE5" bold="true" box="[424,538,1058,1084]" pageId="114" pageNumber="115">
<typeStatus id="54EDB1A9FF81732233C5FB98E11DFBE5" box="[424,538,1058,1084]" pageId="114" pageNumber="115" type="holotype">Holotype</typeStatus>
</emphasis>
: 1 M (
<quantity id="4CAEA2EEFF8173223001FB98E1B8FBE5" box="[620,703,1058,1084]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="114" pageNumber="115" unit="mm" value="18.0">18 mm</quantity>
long),
<emphasis id="B922D319FF8173223179FB98E0BBFBE5" bold="true" box="[788,956,1058,1084]" pageId="114" pageNumber="115">
<specimenCode id="DBF0A770FF8173223179FB98E0B0FBE5" box="[788,951,1058,1084]" collectionCode="FMMC" pageId="114" pageNumber="115">FMMC 5399</specimenCode>
.
</emphasis>
<emphasis id="B922D319FF81732231A9FB98E762FBE2" box="[964,1125,1058,1083]" italics="true" pageId="114" pageNumber="115">
<collectingMunicipality id="6B8D9571FF81732231A9FB98E70DFBE2" box="[964,1034,1058,1083]" pageId="114" pageNumber="115">Locus</collectingMunicipality>
typicus
</emphasis>
:
<collectingCountry id="F3414F9BFF8173223619FB98E604FBE5" box="[1140,1283,1058,1084]" name="Madagascar" pageId="114" pageNumber="115">Madagascar</collectingCountry>
,
<collectingRegion id="4992C1E9FF817322377DFB98E3EAFBBA" country="Madagascar" name="Toliara" pageId="114" pageNumber="115">Province de Toliara</collectingRegion>
, RNI
<location id="8E8959D0FF8173223355FBF3E2CFFBBA" LSID="urn:lsid:plazi:treatment:03FFBE1DFF81732532FAFC36E78AFACA:8E8959D0FF8173223355FBF3E2CFFBBA" box="[312,456,1097,1123]" country="Madagascar" latitude="-24.561666" longLatPrecision="125" longitude="46.72167" municipality="Locus" name="Andohahela" pageId="114" pageNumber="115" stateProvince="Toliara">Andohahela</location>
, parcel 1, 20.0 km
<collectorName id="26A36ADDFF81732230DAFBF3E09AFBBA" box="[695,925,1097,1123]" pageId="114" pageNumber="115">SE Andranodambo</collectorName>
,
<geoCoordinate id="EE6269CCFF81732231C1FBF3E727FBBA" box="[940,1056,1097,1123]" degrees="24" direction="south" minutes="33.7" orientation="latitude" pageId="114" pageNumber="115" precision="92" value="-24.561666">24°33.7'S</geoCoordinate>
,
<geoCoordinate id="EE6269CCFF817322365DFBF3E7A1FBBA" box="[1072,1190,1097,1123]" degrees="46" direction="east" minutes="43.3" orientation="longitude" pageId="114" pageNumber="115" precision="92" value="46.72167">46°43.3'E</geoCoordinate>
, montane rainforest,
<quantity id="4CAEA2EEFF81732232FAFBCAE3F5FB53" box="[151,242,1136,1162]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="114" pageNumber="115" unit="m" value="1875.0">
<elevation id="007BE838FF81732232FAFBCAE3F5FB53" box="[151,242,1136,1162]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="114" pageNumber="115" unit="m" value="1875.0">1875 m</elevation>
</quantity>
, coll.
<collectorName id="26A36ADDFF8173223354FBCAE2C8FB53" box="[313,463,1136,1162]" pageId="114" pageNumber="115">S. Goodman</collectorName>
,
<collectingMethod id="5217771CFF81732233B7FBCAE165FB53" box="[474,610,1136,1162]" pageId="114" pageNumber="115">pitfall traps</collectingMethod>
,
<date id="FFE829CBFF8173223002FBCAE03DFB53" box="[623,826,1136,1162]" pageId="114" pageNumber="115" value="1995-11-27" valueMax="1995-12-05" valueMin="1995-11-27">
<collectingDate id="EFACD023FF8173223002FBCAE03DFB53" box="[623,826,1136,1162]" pageId="114" pageNumber="115" value="1995-11-27" valueMax="1995-12-05" valueMin="1995-11-27">27.xi.5.xii.1995</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE90F0BFF81732232AAFB2CE248FB27" blockId="114.[151,1436,981,2013]" pageId="114" pageNumber="115">
<materialsCitation id="3B3E0556FF81732232AAFB2CE0C6FB69" box="[199,961,1174,1200]" collectingDate="1995-11-27" collectingDateMax="1995-12-05" collectingDateMin="1995-11-27" collectionCode="FMMC" collectorName="SE Andranodambo &amp; S. Goodman" country="Madagascar" elevation="1875" latitude="-24.561666" location="Andohahela" longLatPrecision="125" longitude="46.72167" municipality="Locus" pageId="114" pageNumber="115" specimenCode="FMMC 5399" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
<emphasis id="B922D319FF81732232AAFB2CE24AFB69" bold="true" box="[199,333,1174,1200]" pageId="114" pageNumber="115">
<typeStatus id="54EDB1A9FF81732232AAFB2CE24EFB69" box="[199,329,1174,1200]" pageId="114" pageNumber="115" type="paratype">Paratypes</typeStatus>
:
</emphasis>
1 M,
<emphasis id="B922D319FF81732233F8FB2CE131FB69" bold="true" box="[405,566,1174,1200]" pageId="114" pageNumber="115">
<specimenCode id="DBF0A770FF81732233F8FB2CE131FB69" box="[405,566,1174,1200]" collectionCode="FMMC" pageId="114" pageNumber="115">FMMC 5399</specimenCode>
</emphasis>
, same data as holotype; 8 M, 2 F
</materialsCitation>
,
<materialsCitation id="3B3E0556FF81732231A0FB2CE24CFB27" collectingDate="1995-11-17" collectingDateMax="1995-11-27" collectingDateMin="1995-11-17" collectionCode="FMMC" collectorName="Andohahela &amp; NW Eminiminy &amp; S. Goodman" country="Madagascar" elevation="1500" latitude="-24.57" longLatPrecision="125" longitude="46.731667" pageId="114" pageNumber="115" specimenCode="FMMC 5416" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
<emphasis id="B922D319FF81732231A0FB2CE76AFB69" bold="true" box="[973,1133,1174,1200]" pageId="114" pageNumber="115">
<specimenCode id="DBF0A770FF81732231A0FB2CE76AFB69" box="[973,1133,1174,1200]" collectionCode="FMMC" pageId="114" pageNumber="115">FMMC 5416</specimenCode>
</emphasis>
, RNI
<collectorName id="26A36ADDFF81732236DAFB2CE64FFB69" box="[1207,1352,1174,1200]" pageId="114" pageNumber="115">Andohahela</collectorName>
, parcel 1, camp 4, 15.0 km
<collectorName id="26A36ADDFF81732233D5FB07E187FB0E" box="[440,640,1213,1239]" pageId="114" pageNumber="115">NW Eminiminy</collectorName>
,
<geoCoordinate id="EE6269CCFF81732230FAFB07E00BFB0E" box="[663,780,1213,1239]" degrees="24" direction="south" minutes="34.2" orientation="latitude" pageId="114" pageNumber="115" precision="92" value="-24.57">24°34.2'S</geoCoordinate>
,
<geoCoordinate id="EE6269CCFF817322314EFB07E09EFB0E" box="[803,921,1213,1239]" degrees="46" direction="east" minutes="43.9" orientation="longitude" pageId="114" pageNumber="115" precision="92" value="46.731667">46°43.9'E</geoCoordinate>
, rainforest,
<quantity id="4CAEA2EEFF8173223657FB07E7A6FB0E" box="[1082,1185,1213,1239]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="114" pageNumber="115" unit="m" value="1500.0">
<elevation id="007BE838FF8173223657FB07E7A6FB0E" box="[1082,1185,1213,1239]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="114" pageNumber="115" unit="m" value="1500.0">1500 m</elevation>
</quantity>
, coll.
<collectorName id="26A36ADDFF8173223693FB07E69CFB0E" box="[1278,1435,1213,1239]" pageId="114" pageNumber="115">S. Goodman</collectorName>
<date id="FFE829CBFF81732232FAFB5EE24CFB27" box="[151,331,1252,1278]" pageId="114" pageNumber="115" value="1995-11-17" valueMax="1995-11-27" valueMin="1995-11-17">
<collectingDate id="EFACD023FF81732232FAFB5EE24CFB27" box="[151,331,1252,1278]" pageId="114" pageNumber="115" value="1995-11-17" valueMax="1995-11-27" valueMin="1995-11-17">1727.XI.1995</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE90F0BFF81732232ABFAB0E185FAAB" blockId="114.[151,1436,981,2013]" pageId="114" pageNumber="115">
<emphasis id="B922D319FF81732232ABFAB0E284FAFD" bold="true" box="[198,387,1290,1316]" pageId="114" pageNumber="115">Other material</emphasis>
: 5 M, 1 F
<materialsCitation id="3B3E0556FF817322306DFAB0E179FAAB" collectingDate="2002-01-21" collectingDateMax="2002-01-25" collectingDateMin="2002-01-21" collectionCode="CAS" collectorName="Col du Sedro &amp; Fisher &amp; Griswold" elevation="900" latitude="-24.76389" location="Parc National d'Andohahela" longLatPrecision="21" longitude="46.751667" municipality="Col du Sedro" pageId="114" pageNumber="115" specimenCode="BLF 5013" specimenCount="1">
<emphasis id="B922D319FF817322306DFAB0E1BAFAFD" bold="true" box="[512,701,1290,1316]" pageId="114" pageNumber="115">
<collectionCode id="ED4797CEFF817322306DFAB0E13CFAFD" box="[512,571,1290,1316]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="114" pageNumber="115" type="Museum">CAS</collectionCode>
<specimenCode id="DBF0A770FF817322302FFAB0E1BAFAFD" box="[578,701,1290,1316]" collectionCode="BLF" pageId="114" pageNumber="115">BLF 5013</specimenCode>
</emphasis>
,
<location id="8E8959D0FF81732230A0FAB0E71DFAFD" LSID="urn:lsid:plazi:treatment:03FFBE1DFF81732532FAFC36E78AFACA:8E8959D0FF81732230A0FAB0E71DFAFD" box="[717,1050,1290,1316]" latitude="-24.76389" longLatPrecision="21" longitude="46.751667" municipality="Col du Sedro" name="Parc National d'Andohahela" pageId="114" pageNumber="115">Parc National d'Andohahela</location>
,
<collectorName id="26A36ADDFF817322364BFAB0E7C3FAFD" box="[1062,1220,1290,1316]" pageId="114" pageNumber="115">
<collectingMunicipality id="6B8D9571FF817322364BFAB0E7C3FAFD" box="[1062,1220,1290,1316]" pageId="114" pageNumber="115">Col du Sedro</collectingMunicipality>
</collectorName>
,
<locationDeviation id="9A5B1629FF81732236BDFAB0E219FA92" location="Mahamavo" pageId="114" pageNumber="115">
<quantity id="4CAEA2EEFF81732236BDFAB0E626FAFD" box="[1232,1313,1290,1316]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="114" pageNumber="115" unit="km" value="3.8">3.8 km</quantity>
113° ESE Mahamavo
</locationDeviation>
,
<locationDeviation id="9A5B1629FF8173223346FA8BE19AFA92" box="[299,669,1329,1355]" location="Tolagnaro" pageId="114" pageNumber="115">
<quantity id="4CAEA2EEFF8173223346FA8BE28BFA92" box="[299,396,1329,1355]" metricMagnitude="4" metricUnit="m" metricValue="3.76" pageId="114" pageNumber="115" unit="km" value="37.6">37.6 km</quantity>
341° NNW Tolagnaro
</locationDeviation>
,
<geoCoordinate id="EE6269CCFF81732230C7FA8BE035FA92" box="[682,818,1329,1355]" degrees="24" direction="south" minutes="45" orientation="latitude" pageId="114" pageNumber="115" precision="15" seconds="50" value="-24.76389">24°45'50&quot;S</geoCoordinate>
,
<geoCoordinate id="EE6269CCFF8173223152FA8BE0BDFA92" box="[831,954,1329,1355]" degrees="46" direction="east" minutes="45" orientation="longitude" pageId="114" pageNumber="115" precision="15" seconds="6" value="46.751667">46°45'6&quot;E</geoCoordinate>
,
<quantity id="4CAEA2EEFF81732231A5FA8BE712FA92" box="[968,1045,1329,1355]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="114" pageNumber="115" unit="m" value="900.0">
<elevation id="007BE838FF81732231A5FA8BE712FA92" box="[968,1045,1329,1355]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="114" pageNumber="115" unit="m" value="900.0">900 m</elevation>
</quantity>
, montane rainforest, coll.
<collectorName id="26A36ADDFF8173223720FA8BE69FFA92" box="[1357,1432,1329,1355]" pageId="114" pageNumber="115">Fisher</collectorName>
,
<collectorName id="26A36ADDFF81732232FAFAE2E203FAAB" box="[151,260,1368,1394]" pageId="114" pageNumber="115">Griswold</collectorName>
<emphasis id="B922D319FF8173223366FAE2E241FAA8" box="[267,326,1368,1393]" italics="true" pageId="114" pageNumber="115">et al.</emphasis>
,
<collectingMethod id="5217771CFF8173223338FAE2E2D5FAAB" box="[341,466,1368,1394]" pageId="114" pageNumber="115">pitfall trap</collectingMethod>
,
<date id="FFE829CBFF81732233B3FAE2E179FAAB" box="[478,638,1368,1394]" pageId="114" pageNumber="115" value="2002-01-21" valueMax="2002-01-25" valueMin="2002-01-21">
<collectingDate id="EFACD023FF81732233B3FAE2E179FAAB" box="[478,638,1368,1394]" pageId="114" pageNumber="115" value="2002-01-21" valueMax="2002-01-25" valueMin="2002-01-21">2125.I.2002</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF81732232ABFAC4E624FA41" box="[198,1315,1406,1432]" pageId="114" pageNumber="115" type="distribution">
<paragraph id="8BE90F0BFF81732232ABFAC4E624FA41" blockId="114.[151,1436,981,2013]" box="[198,1315,1406,1432]" pageId="114" pageNumber="115">
<emphasis id="B922D319FF81732232ABFAC4E26EFA41" bold="true" box="[198,361,1406,1432]" pageId="114" pageNumber="115">Distribution:</emphasis>
This species is only known from the montane rainforest at Andohahela (Fig. 77).
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF81732232ABFA1FE663F8B1" pageId="114" pageNumber="115" type="diagnosis">
<paragraph id="8BE90F0BFF81732232ABFA1FE27FF92D" blockId="114.[151,1436,981,2013]" pageId="114" pageNumber="115">
<emphasis id="B922D319FF81732232ABFA1FE24FFA66" bold="true" box="[198,328,1445,1471]" pageId="114" pageNumber="115">Diagnosis:</emphasis>
Small and elongated,
<emphasis id="B922D319FF8173223037FA1FE192FA67" box="[602,661,1445,1470]" italics="true" pageId="114" pageNumber="115">circa</emphasis>
<quantity id="4CAEA2EEFF81732230F2FA1FE1F3FA66" box="[671,756,1445,1471]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="114" pageNumber="115" unit="mm" value="20.0">20 mm</quantity>
long. Color strongly faded in alcohol, shiny. Texture of tergites with small grooves and hairs. Anal shield in males covered completely with very short hairs. Disc of antenna with four apical cones (Fig. 79A). Antenna unusual long, reaching up to fifth tergite. Antennomeres 1 and 2 with sclerotized teeth. All antennomeres without groove (Fig. 79B). Movable finger of posterior telopods strongly curved, with five sclerotized spines (Fig. 78E); posterior side with up to eight sclerotized teeth (Fig. 78F). Fixed finger strongly curved, long and slender, basally with large membranous lobe and sclerotized spine (Fig. 78E). Two long stridulation ribs on male harp (Fig. 78C) and two on each half of female washboard. Anal shield with two locking carinae, first short, second 3 or 4 times longer than first. Females immature.
</paragraph>
<paragraph id="8BE90F0BFF81732232ABF8BAE663F8B1" blockId="114.[151,1436,981,2013]" pageId="114" pageNumber="115">
<emphasis id="B922D319FF81732232ABF8BAE296F8C3" bold="true" box="[198,401,1792,1818]" pageId="114" pageNumber="115">Similar species:</emphasis>
The general shape of telopods is similar to those of
<taxonomicName id="4C567488FF8173223652F8BBE2E1F89B" authority="(de Saussure &amp; Zehntner, 1897)" baseAuthorityName="de Saussure &amp; Zehntner" baseAuthorityYear="1897" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="piligerum">
<emphasis id="B922D319FF8173223652F8BBE66FF8C3" box="[1087,1384,1793,1818]" italics="true" pageId="114" pageNumber="115">Zoosphaerium piligerum</emphasis>
(
<bibRefCitation id="EFC772FAFF8173223712F8BBE2D8F89B" author="de Saussure, H. &amp; Zehntner, L." pageId="114" pageNumber="115" pagination="1 - 12" refId="ref60128" refString="de Saussure, H. &amp; Zehntner, L. (1897) Atlas de l'histoire naturelle des Myriapodes, In: Grandidier (Ed.) Histoire physique, naturelle et politique de Madagascar. 27 (53), Alfred Grandidier, Paris, France, plates 1 - 12." type="book chapter" year="1897">de Saussure &amp; Zehntner, 1897</bibRefCitation>
)
</taxonomicName>
and
<emphasis id="B922D319FF8173223049F892E016F898" box="[548,785,1832,1857]" italics="true" pageId="114" pageNumber="115">
<taxonomicName id="4C567488FF8173223049F892E009F898" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[548,782,1832,1857]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="pseudopiligerum" status="sp. nov.">Z. pseudopiligerum</taxonomicName>
,
</emphasis>
<emphasis id="B922D319FF8173223171F89DE067F898" bold="true" box="[796,864,1831,1857]" pageId="114" pageNumber="115">
<taxonomicNameLabel id="A2116E62FF8173223171F89DE067F898" box="[796,864,1831,1857]" pageId="114" pageNumber="115" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
. The much smaller size, unusual long antennae with only four apical cones and differences in the telopods identifies
<taxonomicName id="4C567488FF81732231ADF8F5E794F8B1" authorityName="Wesener" authorityYear="2009" box="[960,1171,1871,1896]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="114" pageNumber="115" phylum="Arthropoda" rank="species" species="micropiligerum">
<emphasis id="B922D319FF81732231ADF8F5E794F8B1" box="[960,1171,1871,1896]" italics="true" pageId="114" pageNumber="115">Z. micropiligerum</emphasis>
</taxonomicName>
as a new species.
</paragraph>
</subSubSection>
<subSubSection id="C34C5C80FF81732532ABF8CFE78AFACA" lastPageId="117" lastPageNumber="118" pageId="114" pageNumber="115" type="description">
<paragraph id="8BE90F0BFF81732232ABF8CFE3CBF804" blockId="114.[151,1436,981,2013]" pageId="114" pageNumber="115">
<emphasis id="B922D319FF81732232ABF8CFE267F856" bold="true" box="[198,352,1909,1935]" pageId="114" pageNumber="115">Description:</emphasis>
<emphasis id="B922D319FF8173223304F8CFE2F1F857" box="[361,502,1909,1934]" italics="true" pageId="114" pageNumber="115">Body length</emphasis>
: Males (
<specimenCount id="9D50C482FF817322300DF8CFE001F856" box="[608,774,1909,1935]" pageId="114" pageNumber="115" type="generic">10 specimens</specimenCount>
): length up to
<quantity id="4CAEA2EEFF81732231DDF8CFE702F856" box="[944,1029,1909,1935]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="114" pageNumber="115" unit="mm" value="18.0">18 mm</quantity>
, width of thoracic shield up to
<quantity id="4CAEA2EEFF817322371AF8CFE3CFF86F" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="114" pageNumber="115" unit="mm" value="7.1">7.1 mm</quantity>
, height of thoracic shield up to
<quantity id="4CAEA2EEFF817322302EF826E1A4F86F" box="[579,675,1948,1974]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="114" pageNumber="115" unit="mm" value="4.9">4.9 mm</quantity>
. Female (
<specimenCount id="9D50C482FF8173223174F826E0A3F86F" box="[793,932,1948,1974]" pageId="114" pageNumber="115" type="generic">1 specimen</specimenCount>
): length
<quantity id="4CAEA2EEFF8173223667F826E767F86F" box="[1034,1120,1948,1974]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="114" pageNumber="115" unit="mm" value="16.0">16 mm</quantity>
, width
<quantity id="4CAEA2EEFF81732236D7F826E61FF86F" box="[1210,1304,1948,1974]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="114" pageNumber="115" unit="mm" value="7.9">7.9 mm</quantity>
, height
<quantity id="4CAEA2EEFF817322371AF826E3CFF804" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="114" pageNumber="115" unit="mm" value="4.6">4.6 mm</quantity>
.
</paragraph>
<paragraph id="8BE90F0BFF80732332FAF8BEE103F87F" blockId="115.[151,1437,1796,1958]" pageId="115" pageNumber="116">
<emphasis id="B922D319FF80732332FAF8BEE228F8C5" bold="true" box="[151,303,1796,1820]" pageId="115" pageNumber="116">FIGURE 078</emphasis>
.
<taxonomicName id="4C567488FF8073233353F8BFE182F8C5" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[318,645,1797,1820]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="115" pageNumber="116" phylum="Arthropoda" rank="species" species="micropiligerum" status="sp. nov.">
<emphasis id="B922D319FF8073233353F8BFE182F8C5" box="[318,645,1797,1820]" italics="true" pageId="115" pageNumber="116">Zoosphaerium micropiligerum</emphasis>
</taxonomicName>
<emphasis id="B922D319FF80732330E0F8BEE1CCF8C5" bold="true" box="[653,715,1796,1820]" pageId="115" pageNumber="116">
<taxonomicNameLabel id="A2116E62FF80732330E0F8BEE1CCF8C5" box="[653,715,1796,1820]" pageId="115" pageNumber="116" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FF80732330B7F8BEE03DF8C2" box="[730,826,1796,1819]" pageId="115" pageNumber="116" type="holotype">holotype</typeStatus>
,
<emphasis id="B922D319FF807323312AF8BEE05DF8C5" bold="true" box="[839,858,1796,1820]" pageId="115" pageNumber="116">A</emphasis>
: first left coxa and prefemur with stigmatic plate;
<emphasis id="B922D319FF80732337EEF8BEE692F8C5" bold="true" box="[1411,1429,1796,1820]" pageId="115" pageNumber="116">B</emphasis>
: second left coxa with gonopore, posterior view;
<emphasis id="B922D319FF80732330F4F89DE1ABF8E6" bold="true" box="[665,684,1831,1855]" pageId="115" pageNumber="116">C</emphasis>
: left half of anterior telopod, anterior view;
<emphasis id="B922D319FF80732336EFF89DE792F8E6" bold="true" box="[1154,1173,1831,1855]" pageId="115" pageNumber="116">D</emphasis>
: last three podomeres of anterior telopod, posterior view;
<emphasis id="B922D319FF807323339BF8F0E10FF8BB" bold="true" box="[502,520,1866,1890]" pageId="115" pageNumber="116">E</emphasis>
: posterior telopod, anterior view;
<emphasis id="B922D319FF807323311BF8F0E081F8BB" bold="true" box="[886,902,1866,1890]" pageId="115" pageNumber="116">F</emphasis>
: posterior telopod, posterior view. Abbreviations:
<emphasis id="B922D319FF80732332FAF8D6E3B9F85D" bold="true" box="[151,190,1900,1924]" pageId="115" pageNumber="116">cav</emphasis>
= cavity;
<emphasis id="B922D319FF8073233342F8D6E248F85D" bold="true" box="[303,335,1900,1924]" pageId="115" pageNumber="116">Cx</emphasis>
= coxa;
<emphasis id="B922D319FF80732333D9F8D6E2DCF85D" bold="true" box="[436,475,1900,1924]" pageId="115" pageNumber="116">Pre</emphasis>
= prefemur;
<emphasis id="B922D319FF8073233003F8D6E197F85D" bold="true" box="[622,656,1900,1924]" pageId="115" pageNumber="116">s-p</emphasis>
= sclerotized spot;
<emphasis id="B922D319FF8073233107F8D6E09EF85D" bold="true" box="[874,921,1900,1924]" pageId="115" pageNumber="116">scl-t</emphasis>
= sclerotized teeth;
<emphasis id="B922D319FF8073233617F8D6E79BF85D" bold="true" box="[1146,1180,1900,1924]" pageId="115" pageNumber="116">SR</emphasis>
= stridulation rib;
<emphasis id="B922D319FF8073233700F8D6E683F85D" bold="true" box="[1389,1412,1900,1924]" pageId="115" pageNumber="116">St</emphasis>
= stigmatic plate. Scale bar =
<quantity id="4CAEA2EEFF80732333D2F835E107F87F" box="[447,512,1935,1958]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="115" pageNumber="116" unit="mm" value="1.0">1 mm</quantity>
.
</paragraph>
<paragraph id="8BE90F0BFF87732432FAFA49E2D5F989" blockId="116.[151,1436,1523,1616]" pageId="116" pageNumber="117">
<emphasis id="B922D319FF87732432FAFA49E22AF9D2" bold="true" box="[151,301,1523,1547]" pageId="116" pageNumber="117">FIGURE 079</emphasis>
.
<taxonomicName id="4C567488FF8773243357FA49E187F9D3" authority="Wesener, 2009" authorityName="Wesener" authorityYear="2009" box="[314,640,1523,1546]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="116" pageNumber="117" phylum="Arthropoda" rank="species" species="micropiligerum" status="sp. nov.">
<emphasis id="B922D319FF8773243357FA49E187F9D3" box="[314,640,1523,1546]" italics="true" pageId="116" pageNumber="117">Zoosphaerium micropiligerum</emphasis>
</taxonomicName>
<emphasis id="B922D319FF87732430EBFA49E1C5F9D2" bold="true" box="[646,706,1523,1547]" pageId="116" pageNumber="117">
<taxonomicNameLabel id="A2116E62FF87732430EBFA49E1C5F9D2" box="[646,706,1523,1547]" pageId="116" pageNumber="117" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
,
<typeStatus id="54EDB1A9FF87732430A2FA49E029F9D3" box="[719,814,1523,1546]" pageId="116" pageNumber="117" type="paratype">paratype</typeStatus>
, SEM,
<emphasis id="B922D319FF8773243110FA49E097F9D2" bold="true" box="[893,912,1523,1547]" pageId="116" pageNumber="117">A</emphasis>
: antenna, apical view on disc;
<emphasis id="B922D319FF87732436B4FA49E7ECF9D2" bold="true" box="[1241,1259,1523,1547]" pageId="116" pageNumber="117">B</emphasis>
: antenna, lateral view;
<emphasis id="B922D319FF87732432B5F9ACE3ECF9F7" bold="true" box="[216,235,1558,1582]" pageId="116" pageNumber="117">C</emphasis>
: right mandible, mesal view. Abbreviations:
<emphasis id="B922D319FF87732430A5F9ACE1E8F9F7" bold="true" box="[712,751,1558,1582]" pageId="116" pageNumber="117">3iT</emphasis>
= 3 inner teeth;
<emphasis id="B922D319FF87732431F1F9ACE0BBF9F7" bold="true" box="[924,956,1558,1582]" pageId="116" pageNumber="117">Co</emphasis>
= condylus;
<emphasis id="B922D319FF8773243628F9ACE764F9F7" bold="true" box="[1093,1123,1558,1582]" pageId="116" pageNumber="117">eT</emphasis>
= external tooth;
<emphasis id="B922D319FF8773243770F9ACE645F9F7" bold="true" box="[1309,1346,1558,1582]" pageId="116" pageNumber="117">mp</emphasis>
= molar plate;
<emphasis id="B922D319FF87732432B4F982E3FEF989" bold="true" box="[217,249,1592,1616]" pageId="116" pageNumber="117">pL</emphasis>
= pectinate lamella.
</paragraph>
<paragraph id="8BE90F0BFF87732432AAF9C4E086F941" blockId="116.[151,1436,1662,2031]" box="[199,897,1662,1688]" pageId="116" pageNumber="117">
<emphasis id="B922D319FF87732432AAF9C4E229F94E" box="[199,302,1662,1687]" italics="true" pageId="116" pageNumber="117">Habitus:</emphasis>
Especially males are remarkably long and slender.
</paragraph>
<paragraph id="8BE90F0BFF87732432AAF91FE14AF93C" blockId="116.[151,1436,1662,2031]" pageId="116" pageNumber="117">
<emphasis id="B922D319FF87732432AAF91FE256F967" box="[199,337,1701,1726]" italics="true" pageId="116" pageNumber="117">Coloration:</emphasis>
Faded in alcohol. Body with traces of pitchblack coloration. Appendages whitish, sometimes with traces of olive green.
</paragraph>
<paragraph id="8BE90F0BFF87732432AAF948E17AF8D5" blockId="116.[151,1436,1662,2031]" box="[199,637,1778,1804]" pageId="116" pageNumber="117">
<emphasis id="B922D319FF87732432AAF948E217F8D2" box="[199,272,1778,1803]" italics="true" pageId="116" pageNumber="117">Head:</emphasis>
Eyes with more than 50 ocelli.
</paragraph>
<paragraph id="8BE90F0BFF87732432FAF8A3E3DEF87A" blockId="116.[151,1436,1662,2031]" pageId="116" pageNumber="117">
<emphasis id="B922D319FF87732432FAF8A3E208F8EB" box="[151,271,1817,1842]" italics="true" pageId="116" pageNumber="117">Antennae:</emphasis>
Unusually long, reaching back to 5th tergite. Length of antennomeres: 1=2=3&gt;4=5&lt;6 (Fig. 79B), sixth antennomere longest, cylindrical, bearing disc with four sensory cones (Fig. 79A). First antennomere with neither groove nor
<emphasis id="B922D319FF87732433DDF8DEE188F8A4" box="[432,655,1892,1917]" italics="true" pageId="116" pageNumber="117">sensilla basiconica</emphasis>
(Fig. 79B). Sclerotized teeth at base of antennomeres 1 and 2 (Fig. 79B).
</paragraph>
<paragraph id="8BE90F0BFF87732432AAF814E089F836" blockId="116.[151,1436,1662,2031]" pageId="116" pageNumber="117">
<emphasis id="B922D319FF87732432AAF814E247F81E" box="[199,320,1966,1991]" italics="true" pageId="116" pageNumber="117">Mandible:</emphasis>
Six rows of pectinate lamellae, number of teeth declining proximally (Fig. 79C). Molar plate with deep groove (Fig. 79C). Condylus well-rounded (Fig. 79C).
</paragraph>
<paragraph id="8BE90F0BFF86732532AAFF22E103FF01" blockId="117.[151,1436,152,1299]" pageId="117" pageNumber="118">
<emphasis id="B922D319FF86732532AAFF22E296FF68" box="[199,401,152,177]" italics="true" pageId="117" pageNumber="118">Gnathochilarium</emphasis>
: Lateral of palpi
<emphasis id="B922D319FF8673253031FF22E190FF68" box="[604,663,152,177]" italics="true" pageId="117" pageNumber="118">circa</emphasis>
five sensory cones, located together. Depression on posterior side of palpi without sensory cones.
</paragraph>
<paragraph id="8BE90F0BFF86732532AAFF5FE1AEFF26" blockId="117.[151,1436,152,1299]" box="[199,681,229,255]" pageId="117" pageNumber="118">
<emphasis id="B922D319FF86732532AAFF5FE220FF27" box="[199,295,229,254]" italics="true" pageId="117" pageNumber="118">Collum:</emphasis>
Median part of collum glabrous.
</paragraph>
<paragraph id="8BE90F0BFF86732532AAFEB6E1C4FEFF" blockId="117.[151,1436,152,1299]" box="[199,707,268,294]" pageId="117" pageNumber="118">
<emphasis id="B922D319FF86732532AAFEB6E279FEFC" box="[199,382,268,293]" italics="true" pageId="117" pageNumber="118">Thoracic shield</emphasis>
: Surface similar to tergites.
</paragraph>
<paragraph id="8BE90F0BFF86732532AAFE88E692FE95" blockId="117.[151,1436,152,1299]" box="[199,1429,306,332]" pageId="117" pageNumber="118">
<emphasis id="B922D319FF86732532AAFE88E224FE92" box="[199,291,306,331]" italics="true" pageId="117" pageNumber="118">Tergites</emphasis>
: Posterior margin of paratergites projecting posteriorly. Tergites covered with small pits and hair.
</paragraph>
<paragraph id="8BE90F0BFF86732532AAFEE3E083FE19" blockId="117.[151,1436,152,1299]" pageId="117" pageNumber="118">
<emphasis id="B922D319FF86732532AAFEE3E267FEAB" box="[199,352,345,370]" italics="true" pageId="117" pageNumber="118">Endotergum:</emphasis>
Inner section with broad spines and few, isolated bristles (Fig. 88A). Between marginal ridge and inner area without special structures. Externally single row of very short marginal bristles, protruding up to 1/3 of space towards tergite margin (Fig. 88A).
</paragraph>
<paragraph id="8BE90F0BFF86732532AAFE77E13EFDED" blockId="117.[151,1436,152,1299]" pageId="117" pageNumber="118">
<emphasis id="B922D319FF86732532AAFE77E250FE3F" box="[199,343,461,486]" italics="true" pageId="117" pageNumber="118">Anal shield:</emphasis>
Neither bell-shaped nor tapered. At least in males covered with numerous hairs standing in minute impressions. Underside carries two well-developed black locking carinae, anterior short, posterior carina 3.54 times longer than first.
</paragraph>
<paragraph id="8BE90F0BFF86732532AAFDFBE703FD71" blockId="117.[151,1436,152,1299]" pageId="117" pageNumber="118">
<emphasis id="B922D319FF86732532AAFDFBE20FFD83" box="[199,264,577,602]" italics="true" pageId="117" pageNumber="118">Legs:</emphasis>
First tarsi with up to four, second with up to seven, third with eight long ventral spines. First two leg pairs with only weakly curved claws and without apical spine. Tarsi of legs 321 with curved claws, eight ventral spines and apical spine. In 9th leg, tarsi 4.2 times longer than wide.
</paragraph>
<paragraph id="8BE90F0BFF86732532AAFD0FE745FD16" blockId="117.[151,1436,152,1299]" box="[199,1090,693,719]" pageId="117" pageNumber="118">
<emphasis id="B922D319FF86732532AAFD0FE28BFD17" box="[199,396,693,718]" italics="true" pageId="117" pageNumber="118">Stigmatic plates:</emphasis>
First plate, lobe weakly curved, short and stout (Fig. 78A).
</paragraph>
<paragraph id="8BE90F0BFF86732532AAFD66E71BFD2F" blockId="117.[151,1436,152,1299]" box="[199,1052,732,758]" pageId="117" pageNumber="118">
<emphasis id="B922D319FF86732532AAFD66E218FD2C" box="[199,287,732,757]" italics="true" pageId="117" pageNumber="118">Female</emphasis>
: Immature, with two stridulation ribs on each side of washboard.
</paragraph>
<paragraph id="8BE90F0BFF86732532AAFCB8E713FC9A" blockId="117.[151,1436,152,1299]" pageId="117" pageNumber="118">
<emphasis id="B922D319FF86732532AAFCB8E2E6FCC2" box="[199,481,770,795]" italics="true" pageId="117" pageNumber="118">Male sexual characters</emphasis>
: Male gonopore large, covered with huge, sclerotized, undivided and rounded plate (Fig. 78B). Anal shield completely covered with numerous short hairs.
</paragraph>
<paragraph id="8BE90F0BFF86732532AAFCEAE782FBF2" blockId="117.[151,1436,152,1299]" pageId="117" pageNumber="118">
<emphasis id="B922D319FF86732532AAFCEAE293FCB0" box="[199,404,848,873]" italics="true" pageId="117" pageNumber="118">Anterior telopods</emphasis>
(Figs 78C, D): Harp with two stridulation ribs, both straight and long. Lateral rib longer than inner rib, reaching from basal to apical part of first podomere. Second podomere process lobe-like and little curved with rounded edges, reaching almost as high as third podomere, apically with sclerotized spots and basally with one spine. Third podomere cylindrical, almost as wide as second podomere process. Cavity mesally with numerous sclerotized spots and five small spines, spines located basally, medially and largest two apically closely towards tip. Tip of third podomere with circular, sclerotized spot.
</paragraph>
<paragraph id="8BE90F0BFF86732532AAFB82E78AFACA" blockId="117.[151,1436,152,1299]" pageId="117" pageNumber="118">
<emphasis id="B922D319FF86732532AAFB82E2A6FB88" box="[199,417,1080,1105]" italics="true" pageId="117" pageNumber="118">Posterior telopods</emphasis>
: Movable finger strongly curved. Hollowed-out inner margin with up to five smaller, sclerotized spines, two located at its tip; posterior aspect with 8 sclerotized spots. Fixed finger slender, far shorter than movable finger. Fixed finger tip strongly curved, anterior side basally with single, large membranous lobe and sclerotized spine, around its tip with numerous small, sclerotized spots. Movable finger glabrous, first and second podomere almost glabrous, on both sides with some isolated hair. Inner horns of syncoxite apically curved. Tips sharp, whole horns covered with numerous short hairs.
</paragraph>
</subSubSection>
</treatment>
</document>