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<document id="6464F4A60C5C960E1E1D2C878CF20899" ID="10.11646/zootaxa.4263.2.4SLASH10992" ID-CLB-Dataset="33087" ID-DOI="10.11646/zootaxa.4263.2.4" ID-GBIF-Dataset="aeb6b624-fa15-4767-9d42-7963502cd209" ID-ISSN="1175-5326" ID-Zenodo-Dep="573095" ID-ZooBank="6A54FA8D-213C-4776-B5CF-84949BE621D1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1494316787886" checkinUser="plazi" docAuthor="Sagorny, Christina &amp; Wesener, Thomas" docDate="2017" docId="03FF9C6CFFE2142889F4D6C2E3B0FD81" docLanguage="en" docName="zootaxa.4263.2.4SLASH10992.pdf" docOrigin="Zootaxa 4263 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zoosphaerium bemanevika Sagorny &amp; Wesener, n. sp." docType="treatment" docVersion="9" lastPageNumber="284" masterDocId="FFC6E414FFE714238963D059E078FF99" masterDocTitle="Two new giant pill-millipede species of the genus Zoosphaerium endemic to the Bemanevika area in northern Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae)" masterLastPageNumber="294" masterPageNumber="273" pageNumber="278" updateTime="1698439385235" updateUser="ExternalLinkService">
<mods:mods id="309B31660BFBE5B6D29C78A096DD375F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C8A22F6E23CA919FFF0CD49DB8DD7985">
<mods:title id="5E9D8806B6E3C925649243DD84FC3079">Two new giant pill-millipede species of the genus Zoosphaerium endemic to the Bemanevika area in northern Madagascar (Diplopoda, Sphaerotheriida, Arthrosphaeridae)</mods:title>
</mods:titleInfo>
<mods:name id="155158D57AA81EE44EF948096A7F01F5" type="personal">
<mods:role id="9455C2475AAF038B4867F61F4ACBAD0F">
<mods:roleTerm id="847C43C817FC42E957BD13D9F11D22EC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3FDACB1D8065B4B5BF59672936515067">Sagorny, Christina</mods:namePart>
</mods:name>
<mods:name id="6416C439783928469F21286FFDFAD18A" type="personal">
<mods:role id="0B75AA226C60A9E23A83C98C5F1A8631">
<mods:roleTerm id="A906016C2BB6E631F3E5B911E1493C23">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E0B06C0D7CACA9B1EF5EC3E731713011">Wesener, Thomas</mods:namePart>
</mods:name>
<mods:typeOfResource id="68E4CE586AB45A77A9974EA5E0FFE88F">text</mods:typeOfResource>
<mods:relatedItem id="D4176A2F4D4C1A203AB201B4F558D600" type="host">
<mods:titleInfo id="78AD7C61AD20AF18DBEB59D741CDFB47">
<mods:title id="18951421BEAA4116234B0D87C4CDAAD3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="46EE3FF5BDD91DC7E7DFEAD015301E06">
<mods:date id="F80364187F7A97C9C66E816924636ED7">2017</mods:date>
<mods:detail id="A2029D1CC527EA20CE4B73D16EDB70EB" type="volume">
<mods:number id="C91481B229F97FB952B4A7CB21596E51">4263</mods:number>
</mods:detail>
<mods:detail id="AE4DE756E73D078CCCF741F9C2DADB9C" type="issue">
<mods:number id="B39712DDF4BBA2227AD90ED133622B3D">2</mods:number>
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<mods:start id="DAAE57C8AFC1CD3DA8DC0FE3182AC1DE">273</mods:start>
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</mods:extent>
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<mods:classification id="280B540A11EF70777E45F81F31B8BEF9">journal article</mods:classification>
<mods:identifier id="76AB2BFF439081E34604AF7871DB0B3B" type="CLB-Dataset">33087</mods:identifier>
<mods:identifier id="C5ABB404668C76D120EF66FE7B10B2F9" type="DOI">10.11646/zootaxa.4263.2.4</mods:identifier>
<mods:identifier id="177432F7CB42C2CD240C1E8FBA863AF6" type="GBIF-Dataset">aeb6b624-fa15-4767-9d42-7963502cd209</mods:identifier>
<mods:identifier id="FD9B7E0C899232A1E3CDB3B09A5C7BCF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D7BD403BC4A4A98FA1B96502A26E45BC" type="Zenodo-Dep">573095</mods:identifier>
<mods:identifier id="7CDD020E5F723B239826D0DA5A4C5A5E" type="ZooBank">6A54FA8D-213C-4776-B5CF-84949BE621D1</mods:identifier>
</mods:mods>
<treatment id="03FF9C6CFFE2142889F4D6C2E3B0FD81" ID-DOI="http://doi.org/10.5281/zenodo.5690092" ID-GBIF-Taxon="130497681" ID-Zenodo-Dep="5690092" LSID="urn:lsid:plazi:treatment:03FF9C6CFFE2142889F4D6C2E3B0FD81" httpUri="http://treatment.plazi.org/id/03FF9C6CFFE2142889F4D6C2E3B0FD81" lastPageId="11" lastPageNumber="284" pageId="5" pageNumber="278">
<subSubSection id="C34C7EF1FFE2142689F4D6C2E15AF94E" pageId="5" pageNumber="278" type="nomenclature">
<paragraph id="8BE92D7AFFE2142689F4D6C2E357F92F" blockId="5.[151,815,1691,1751]" box="[151,815,1691,1718]" pageId="5" pageNumber="278">
<heading id="D0A19A16FFE2142689F4D6C2E357F92F" bold="true" box="[151,815,1691,1718]" fontSize="11" level="1" pageId="5" pageNumber="278" reason="1">
<emphasis id="B922F168FFE2142689F4D6C2E357F92F" bold="true" box="[151,815,1691,1718]" pageId="5" pageNumber="278">
<taxonomicName id="4C5656F9FFE2142689F4D6C2E298F92F" ID-CoL="84F5T" authority="Sagorny &amp; Wesener" authorityName="Sagorny &amp; Wesener" box="[151,736,1691,1718]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="278" phylum="Arthropoda" rank="species" species="bemanevika" status="sp. nov.">
<emphasis id="B922F168FFE2142689F4D6C2E1A7F92C" bold="true" box="[151,479,1691,1717]" italics="true" pageId="5" pageNumber="278">Zoosphaerium bemanevika</emphasis>
Sagorny &amp; Wesener
</taxonomicName>
,
<taxonomicNameLabel id="A2114C13FFE214268B8ED6C5E357F92F" box="[749,815,1692,1718]" pageId="5" pageNumber="278" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE92D7AFFE2142689F4D6E7E15AF94E" blockId="5.[151,815,1691,1751]" box="[151,290,1726,1751]" pageId="5" pageNumber="278">
<figureCitation id="136D31FFFFE2142689F4D6E7E083F94E" box="[151,251,1726,1751]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="6.[151,250,1801,1823]" captionStartId-1="7.[151,250,700,722]" captionStartId-2="7.[151,250,1892,1914]" captionStartId-3="8.[151,250,1054,1076]" captionStartId-4="9.[151,250,1889,1911]" captionStartId-5="10.[151,250,849,871]" captionTargetBox-0="[168,1414,336,1783]" captionTargetBox-1="[175,1398,193,689]" captionTargetBox-2="[176,1389,821,1861]" captionTargetBox-3="[150,1436,194,1027]" captionTargetBox-4="[152,1433,664,1861]" captionTargetBox-5="[151,1435,194,824]" captionTargetId-0="figure@6.[168,1414,336,1783]" captionTargetId-1="figure@7.[175,1398,193,689]" captionTargetId-2="figure@7.[176,1390,821,1862]" captionTargetId-3="figure@8.[150,1437,193,1033]" captionTargetId-4="figure@9.[152,1435,663,1867]" captionTargetId-5="figure@10.[151,1435,193,828]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionText-0="FIGURE 2. Zoosphaerium bemanevika n. sp.., A, B: female (ZFMK MYR 6144); C-H: holotype male (FMNH-INS 2858686 - C); A: coxa of first left leg with first stigmatic plate; B: 9 th right leg; C coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view G: right posterior telopod, posterior view; H: right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-p = sclerotized spot; s-ps = field of sclerotized spots; sr = stridulation ribs; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. A, B Scale bars = 1 mm, C – H Scale bars = 0.5 mm." captionText-1="FIGURE 3. Zoosphaerium bemanevika n. sp., female (ZFMK MYR 6144), A: coxa of second left leg with vulva; B: female subanal plate with washboard. Abbreviations: cx = coxa; ep = external plate of vulva; ip = internal plate of vulva; o = operculum; sr = stridulation ribs. Scale bars = 1 mm." captionText-2="FIGURE 4. Zoosphaerium bemanevika n. sp., female (ZFMK MYR 6144), head, µCT, A: frontal view; B: lateral view; C: internal morphology. Abbreviations: aG = antennal groove; aO = aberrant ocellus; c = condyles; cp = central pad; Gn = gnathochilarium; gu = gula; il = incisura lateralis; ip = inner palpus; lb = labrum; LL = lamellae linguales; mdb = mandible base; mdt = mandible tendon; Me = mentum; oc = ocelli; Ph = pharynx; PM = paramentum; St = stipites; t = tentorium; TO = Organ of Tömosváry. Images not to scale." captionText-3="FIGURE 5. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), left antenna, SEM, A: lateral view; B: sensilla basiconica on 1 st antennomere; C: apical disc with sensory cones. Abbreviations: aD = apical disc; sb = sensilla basiconica." captionText-4="FIGURE 6. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), SEM, A: gnathochilarium, ventral view; B: gnathochilarium, apical view of dorsal side; C: gnathochilarium, rudimentary lateral palpus with sensory cones; D: gnathochilarium, sensory cones on central pad; E: left mandible, mesal view; F: left mandible, pectinate lamellae. Abbreviations: c = condyles; cp = central pad; eT = external tooth; iP = inner palpus; iT = internal tooth; LL = lamellae linguales; LP = lateral palpus; mp = molar plate; pL = pectinate lamellae; St = stipites." captionText-5="FIGURE 7. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), endotergum of mid-body tergite, SEM, A: ventral view; B: bristles on apical margin." httpUri-0="https://zenodo.org/record/584046/files/figure.png" httpUri-1="https://zenodo.org/record/584047/files/figure.png" httpUri-2="https://zenodo.org/record/584048/files/figure.png" httpUri-3="https://zenodo.org/record/584049/files/figure.png" httpUri-4="https://zenodo.org/record/584050/files/figure.png" httpUri-5="https://zenodo.org/record/584051/files/figure.png" pageId="5" pageNumber="278">Figs 2–7</figureCitation>
,
<figureCitation id="136D31FFFFE214268865D6E6E15AF94E" box="[262,290,1727,1751]" captionStart="FIGURE 12" captionStartId="16.[151,250,1864,1886]" captionTargetBox="[163,1436,1100,1835]" captionTargetId="figure@16.[150,1436,1094,1842]" captionTargetPageId="16" captionText="FIGURE 12. Maximum likelihood tree inferred from the COI dataset with 1000 bootstrap pseudoreplicates implementing the GTR + I + G model. Habitus photograph shows Z. bemanevika (FMNH-INS 3196510). Colors used to separate species. Green = Z. bemanevika; Orange = Z. minutus. Circles indicating sister group relationships, as mapped in figure 1." httpUri="https://zenodo.org/record/584057/files/figure.png" pageId="5" pageNumber="278">12</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34C7EF1FFE2142689F4D758E548F883" box="[151,1328,1793,1818]" pageId="5" pageNumber="278" type="etymology">
<paragraph id="8BE92D7AFFE2142689F4D758E548F883" blockId="5.[151,1328,1793,1818]" box="[151,1328,1793,1818]" pageId="5" pageNumber="278">
<emphasis id="B922F168FFE2142689F4D758E166F883" bold="true" box="[151,286,1793,1818]" pageId="5" pageNumber="278">Etymology.</emphasis>
<taxonomicName id="4C5656F9FFE214268847D758E1CBF883" box="[292,435,1793,1818]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="bemanevika" status="sp. nov.">Bemanevika</taxonomicName>
, noun in apposition. After the name of the village located near the
<typeStatus id="54ED93D8FFE214268DFCD75BE4B7F883" box="[1183,1231,1794,1818]" pageId="5" pageNumber="278">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C34C7EF1FFE2142589F4D710E55EFE84" lastPageId="6" lastPageNumber="279" pageId="5" pageNumber="278" type="materials_examined">
<paragraph id="8BE92D7AFFE2142689F4D710E526F830" blockId="5.[151,1436,1865,2034]" pageId="5" pageNumber="278">
<materialsCitation id="3B3E2727FFE2142689F4D710E526F830" ID-GBIF-Occurrence="1500393085" collectingDate="2013-10" collectionCode="FMNH" collectorName="Pitfalls PF &amp; Steven M." country="Madagascar" county="Material" elevation="1570" latitude="-14.346472" location="Amberivery Forest" longLatPrecision="1" longitude="48.579803" municipality="Sofia" pageId="5" pageNumber="278" specimenCount="1" stateProvince="Mahajanga" typeStatus="holotype">
<emphasis id="B922F168FFE2142689F4D710E272F8FB" bold="true" box="[151,522,1865,1890]" pageId="5" pageNumber="278">
<collectingCounty id="628855F6FFE2142689F4D710E087F8FB" box="[151,255,1865,1890]" pageId="5" pageNumber="278">Material</collectingCounty>
examined.
<typeStatus id="54ED93D8FFE2142688FED710E272F8FB" box="[413,522,1865,1890]" pageId="5" pageNumber="278" type="holotype">Holotype</typeStatus>
</emphasis>
: 1 M,
<emphasis id="B922F168FFE214268B15D710E3EDF8FB" bold="true" box="[630,917,1865,1890]" pageId="5" pageNumber="278">FMNH-INS 2858686-C</emphasis>
(
<collectionCode id="ED47B5BFFFE214268ACCD710E47EF8F8" box="[943,1030,1865,1889]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="5" pageNumber="278">FMNH</collectionCode>
),
<collectingCountry id="F3416DEAFFE214268D40D710E4D7F8FB" box="[1059,1199,1865,1890]" name="Madagascar" pageId="5" pageNumber="278">Madagascar</collectingCountry>
:
<collectingRegion id="4992E398FFE214268DA6D710E53CF8FB" box="[1221,1348,1865,1890]" country="Madagascar" name="Mahajanga" pageId="5" pageNumber="278">Mahajanga</collectingRegion>
,
<collectingMunicipality id="6B8DB700FFE214268C39D710E5E0F8FB" box="[1370,1432,1865,1890]" pageId="5" pageNumber="278">Sofia</collectingMunicipality>
,
<location id="8E897BA1FFE2142689F4D734E10AF81C" LSID="urn:lsid:plazi:treatment:03FF9C6CFFE2142889F4D6C2E3B0FD81:8E897BA1FFE2142689F4D734E10AF81C" box="[151,370,1901,1926]" country="Madagascar" county="Material" latitude="-14.346472" longLatPrecision="1" longitude="48.579803" municipality="Sofia" name="Amberivery Forest" pageId="5" pageNumber="278" stateProvince="Mahajanga">Amberivery Forest</location>
,
<location id="8E897BA1FFE2142688E3D737E2ACF81F" LSID="urn:lsid:plazi:treatment:03FF9C6CFFE2142889F4D6C2E3B0FD81:8E897BA1FFE2142688E3D737E2ACF81F" box="[384,724,1901,1926]" country="Madagascar" county="Material" latitude="-14.346472" longLatPrecision="1" longitude="48.579803" municipality="Sofia" name="Paysage Harmonieux Protege" pageId="5" pageNumber="278" stateProvince="Mahajanga">Paysage Harmonieux Protégé</location>
de
<taxonomicName id="4C5656F9FFE214268A60D734E3EAF81F" box="[771,914,1901,1926]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="bemanevika" status="sp. nov.">Bemanevika</taxonomicName>
, site 2,
<geoCoordinate id="EE624BBDFFE214268A91D737E4F0F81F" box="[1010,1160,1902,1926]" direction="south" orientation="latitude" pageId="5" pageNumber="278" precision="1" value="-14.346472">14°20'47.3''S</geoCoordinate>
,
<geoCoordinate id="EE624BBDFFE214268DF4D737E555F81F" box="[1175,1325,1902,1926]" direction="east" orientation="longitude" pageId="5" pageNumber="278" precision="1" value="48.579803">48°34'47.3''E</geoCoordinate>
,
<quantity id="4CAE809FFFE214268C5ED734E5EFF81C" box="[1341,1431,1901,1926]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="5" pageNumber="278" unit="m" value="1570.0">
<elevation id="007BCA49FFE214268C5ED734E5EFF81C" box="[1341,1431,1901,1926]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="5" pageNumber="278" unit="m" value="1570.0">1570 m</elevation>
</quantity>
, slightly disturbed montane forest,
<collectorName id="26A348ACFFE214268B75D7C8E2F7F833" box="[534,655,1937,1962]" pageId="5" pageNumber="278">Pitfalls PF</collectorName>
#5, coll.
<collectorName id="26A348ACFFE214268B8DD7C8E319F830" box="[750,865,1937,1961]" pageId="5" pageNumber="278">Steven M.</collectorName>
Goodman and Toky Randriamora,
<date id="FFE80BBAFFE214268D8ED7C8E522F830" box="[1261,1370,1937,1961]" pageId="5" pageNumber="278" value="2013-10">
<collectingDate id="EFACF252FFE214268D8ED7C8E522F830" box="[1261,1370,1937,1961]" pageId="5" pageNumber="278" value="2013-10">Oct-2013</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE92D7AFFE2142589A4D7ECE55EFE84" blockId="5.[151,1436,1865,2034]" lastBlockId="6.[151,1437,151,285]" lastPageId="6" lastPageNumber="279" pageId="5" pageNumber="278">
<materialsCitation id="3B3E2727FFE2142589A4D7ECE51FFF29" ID-GBIF-Occurrence="1500393080" collectingDate="2013-10" collectorName="Pitfalls, PF &amp; Steven M." country="Madagascar" county="Other" elevation="1620" lastPageId="6" lastPageNumber="279" latitude="-14.34075" location="Along" longLatPrecision="1" longitude="48.585636" municipality="Sofia" pageId="5" pageNumber="278" specimenCount="1" stateProvince="Mahajanga">
<emphasis id="B922F168FFE2142689A4D7ECE279F857" bold="true" box="[199,513,1973,1998]" pageId="5" pageNumber="278">
<collectingCounty id="628855F6FFE2142689A4D7ECE177F857" box="[199,271,1973,1998]" pageId="5" pageNumber="278">Other</collectingCounty>
material examined.
</emphasis>
1 M (immature),
<emphasis id="B922F168FFE214268BBFD7ECE3ADF857" bold="true" box="[732,981,1973,1998]" pageId="5" pageNumber="278">FMNH-INS 2858678</emphasis>
,
<collectingCountry id="F3416DEAFFE214268A86D7ECE409F857" box="[997,1137,1973,1998]" name="Madagascar" pageId="5" pageNumber="278">Madagascar</collectingCountry>
:
<collectingRegion id="4992E398FFE214268D1ED7ECE485F857" box="[1149,1277,1973,1998]" country="Madagascar" name="Mahajanga" pageId="5" pageNumber="278">Mahajanga</collectingRegion>
,
<collectingMunicipality id="6B8DB700FFE214268C69D7ECE530F857" box="[1290,1352,1973,1998]" pageId="5" pageNumber="278">Sofia</collectingMunicipality>
,
<location id="8E897BA1FFE214268C35D7ECE5E4F857" LSID="urn:lsid:plazi:treatment:03FF9C6CFFE2142889F4D6C2E3B0FD81:8E897BA1FFE214268C35D7ECE5E4F857" box="[1366,1436,1973,1998]" country="Madagascar" county="Other" latitude="-14.34075" longLatPrecision="1" longitude="48.585636" municipality="Sofia" name="Along" pageId="5" pageNumber="278" stateProvince="Mahajanga">Along</location>
trail to
<location id="8E897BA1FFE21426898BD780E1E1F86B" LSID="urn:lsid:plazi:treatment:03FF9C6CFFE2142889F4D6C2E3B0FD81:8E897BA1FFE21426898BD780E1E1F86B" box="[232,409,2009,2034]" country="Madagascar" county="Other" latitude="-14.34075" longLatPrecision="1" longitude="48.585636" municipality="Sofia" name="Matsaborimena" pageId="5" pageNumber="278" stateProvince="Mahajanga">Matsaborimena</location>
,
<location id="8E897BA1FFE2142688C7D783E297F86B" LSID="urn:lsid:plazi:treatment:03FF9C6CFFE2142889F4D6C2E3B0FD81:8E897BA1FFE2142688C7D783E297F86B" box="[420,751,2009,2034]" country="Madagascar" county="Other" latitude="-14.34075" longLatPrecision="1" longitude="48.585636" municipality="Sofia" name="Paysage Harmonieux Protege" pageId="5" pageNumber="278" stateProvince="Mahajanga">Paysage Harmonieux Protégé</location>
de
<taxonomicName id="4C5656F9FFE214268A7BD780E3DFF86B" box="[792,935,2009,2034]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="5" pageNumber="291" phylum="Arthropoda" rank="species" species="bemanevika" status="sp. nov.">Bemanevika</taxonomicName>
, site 1,
<geoCoordinate id="EE624BBDFFE214268A9ED780E4EAF86B" box="[1021,1170,2009,2034]" direction="south" orientation="latitude" pageId="5" pageNumber="278" precision="1" value="-14.34075">14°20'26.7''S</geoCoordinate>
,
<geoCoordinate id="EE624BBDFFE214268DFDD780E54DF86B" box="[1182,1333,2009,2034]" direction="east" orientation="longitude" pageId="5" pageNumber="278" precision="1" value="48.585636">48°35'08.3''E</geoCoordinate>
,
<quantity id="4CAE809FFFE214268C23D780E5EFF868" box="[1344,1431,2009,2034]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="5" pageNumber="278" unit="m" value="1620.0">
<elevation id="007BCA49FFE214268C23D780E5EFF868" box="[1344,1431,2009,2034]" metricMagnitude="3" metricUnit="m" metricValue="1.6199999999999999" pageId="5" pageNumber="278" unit="m" value="1620.0">1620 m</elevation>
</quantity>
, slightly disturbed montane forest,
<collectorName id="26A348ACFFE114258B7BD0CEE2E1FF29" box="[536,665,151,176]" pageId="6" pageNumber="279">Pitfalls, PF</collectorName>
#3, coll.
<collectorName id="26A348ACFFE114258B9BD0C1E313FF36" box="[760,875,152,176]" pageId="6" pageNumber="279">Steven M.</collectorName>
Goodman and Toky Randriamora,
<date id="FFE80BBAFFE114258D9AD0C1E51FFF29" box="[1273,1383,152,176]" pageId="6" pageNumber="279" value="2013-10">
<collectingDate id="EFACF252FFE114258D9AD0C1E51FFF29" box="[1273,1383,152,176]" pageId="6" pageNumber="279" value="2013-10">Oct-2013</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E2727FFE114258C11D0C1E55EFE84" ID-GBIF-Occurrence="1500393088" collectingDate="2013-10" collectionCode="ZFMK, MYR" collectorName="Pitfalls, PF &amp; Steven M." country="Madagascar" county="Sofia" elevation="1570" latitude="-14.346472" location="Paysage Harmonieux Protege" longLatPrecision="1" longitude="48.579803" municipality="Amberivery Forest" pageId="6" pageNumber="279" specimenCode="MYR 6144" specimenCount="1" stateProvince="Mahajanga">
1 F,
<emphasis id="B922F168FFE1142589F4D0E5E1E3FF4C" bold="true" box="[151,411,188,213]" pageId="6" pageNumber="279">FMNH-INS 3196510,</emphasis>
<collectingCountry id="F3416DEAFFE1142588C9D0E5E24EFF4C" box="[426,566,188,213]" name="Madagascar" pageId="6" pageNumber="279">Madagascar</collectingCountry>
:
<collectingRegion id="4992E398FFE114258B24D0E5E2BEFF4C" box="[583,710,188,213]" country="Madagascar" name="Mahajanga" pageId="6" pageNumber="279">Mahajanga</collectingRegion>
,
<collectingCounty id="628855F6FFE114258BBAD0E5E36FFF4C" box="[729,791,188,213]" pageId="6" pageNumber="279">Sofia</collectingCounty>
,
<collectingMunicipality id="6B8DB700FFE114258A4AD0E5E470FF4D" box="[809,1032,188,213]" pageId="6" pageNumber="279">Amberivery Forest</collectingMunicipality>
,
<location id="8E897BA1FFE114258D79D0E4E50BFF4D" LSID="urn:lsid:plazi:treatment:03FF9C6CFFE2142889F4D6C2E3B0FD81:8E897BA1FFE114258D79D0E4E50BFF4D" box="[1050,1395,188,213]" country="Madagascar" county="Sofia" latitude="-14.346472" longLatPrecision="1" longitude="48.579803" municipality="Amberivery Forest" name="Paysage Harmonieux Protege" pageId="6" pageNumber="279" stateProvince="Mahajanga">Paysage Harmonieux Protégé</location>
de
<taxonomicName id="4C5656F9FFE1142589F4D086E15EFF61" box="[151,294,223,248]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="6" pageNumber="291" phylum="Arthropoda" rank="species" species="bemanevika" status="sp. nov.">Bemanevika</taxonomicName>
, site 2,
<geoCoordinate id="EE624BBDFFE1142588E0D0B9E261FF61" box="[387,537,224,248]" direction="south" orientation="latitude" pageId="6" pageNumber="279" precision="1" value="-14.346472">14°20'47.3''S</geoCoordinate>
,
<geoCoordinate id="EE624BBDFFE114258B44D0B9E2C6FF61" box="[551,702,224,248]" direction="east" orientation="longitude" pageId="6" pageNumber="279" precision="1" value="48.579803">48°34'47.3''E</geoCoordinate>
,
<quantity id="4CAE809FFFE114258BAFD086E35EFF6E" box="[716,806,223,248]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="6" pageNumber="279" unit="m" value="1570.0">
<elevation id="007BCA49FFE114258BAFD086E35EFF6E" box="[716,806,223,248]" metricMagnitude="3" metricUnit="m" metricValue="1.5699999999999998" pageId="6" pageNumber="279" unit="m" value="1570.0">1570 m</elevation>
</quantity>
, slightly disturbed montane forest,
<collectorName id="26A348ACFFE114258DDCD086E539FF61" box="[1215,1345,223,248]" pageId="6" pageNumber="279">Pitfalls, PF</collectorName>
#6, coll.
<collectorName id="26A348ACFFE1142589F4D15DE172FE85" box="[151,266,260,284]" pageId="6" pageNumber="279">Steven M.</collectorName>
Goodman and Toky Randriamora,
<date id="FFE80BBAFFE114258BF5D15DE37BFE85" box="[662,771,260,284]" pageId="6" pageNumber="279" value="2013-10">
<collectingDate id="EFACF252FFE114258BF5D15DE37BFE85" box="[662,771,260,284]" pageId="6" pageNumber="279" value="2013-10">Oct-2013</collectingDate>
</date>
; 1 F,
<emphasis id="B922F168FFE114258A5DD15DE463FE84" bold="true" box="[830,1051,260,285]" pageId="6" pageNumber="279">
<collectionCode id="ED47B5BFFFE114258A5DD15DE3EAFE84" box="[830,914,260,285]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="6" pageNumber="279">ZFMK</collectionCode>
<specimenCode id="DBF08501FFE114258AFAD15DE463FE84" box="[921,1051,260,285]" collectionCode="MYR" pageId="6" pageNumber="279">
<collectionCode id="ED47B5BFFFE114258AFAD15DE3A4FE84" box="[921,988,260,285]" pageId="6" pageNumber="279">MYR</collectionCode>
6144
</specimenCode>
</emphasis>
, same data as previous.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF297DF2FFE1142589F4D750E1DFF84E" httpUri="https://zenodo.org/record/584046/files/figure.png" pageId="6" pageNumber="279" targetBox="[168,1414,336,1783]" targetPageId="6">
<paragraph id="8BE92D7AFFE1142589F4D750E1DFF84E" blockId="6.[151,1437,1801,2007]" pageId="6" pageNumber="279">
<emphasis id="B922F168FFE1142589F4D750E174F886" bold="true" box="[151,268,1801,1823]" pageId="6" pageNumber="279">FIGURE 2</emphasis>
.
<taxonomicName id="4C5656F9FFE11425887BD750E252F886" box="[280,554,1801,1823]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="6" pageNumber="279" phylum="Arthropoda" rank="species" species="bemanevika" status="sp. nov.">
<emphasis id="B922F168FFE11425887BD750E252F886" box="[280,554,1801,1823]" italics="true" pageId="6" pageNumber="279">Zoosphaerium bemanevika</emphasis>
</taxonomicName>
<emphasis id="B922F168FFE114258B53D750E211F886" bold="true" box="[560,617,1801,1823]" pageId="6" pageNumber="279">
<taxonomicNameLabel id="A2114C13FFE114258B53D750E211F886" box="[560,617,1801,1823]" pageId="6" pageNumber="279" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
.,
<emphasis id="B922F168FFE114258B1FD750E2D3F886" bold="true" box="[636,683,1801,1823]" pageId="6" pageNumber="279">A, B</emphasis>
: female (ZFMK MYR 6144);
<emphasis id="B922F168FFE114258A83D750E476F886" bold="true" box="[992,1038,1801,1823]" pageId="6" pageNumber="279">C-H</emphasis>
: holotype male (FMNH-INS 2858686- C);
<emphasis id="B922F168FFE1142589DDD771E0A8F8A7" bold="true" box="[190,208,1832,1854]" pageId="6" pageNumber="279">A</emphasis>
: coxa of first left leg with first stigmatic plate;
<emphasis id="B922F168FFE114258BD7D771E2BDF8A7" bold="true" box="[692,709,1832,1854]" pageId="6" pageNumber="279">B</emphasis>
: 9th right leg;
<emphasis id="B922F168FFE114258A3ED771E317F8A7" bold="true" box="[861,879,1832,1854]" pageId="6" pageNumber="279">C</emphasis>
coxa of second left leg with gonopore;
<emphasis id="B922F168FFE114258C67D771E56EF8A7" bold="true" box="[1284,1302,1832,1854]" pageId="6" pageNumber="279">D</emphasis>
: left anterior telopod, anterior view;
<emphasis id="B922F168FFE1142588E6D71FE1EEF8C5" bold="true" box="[389,406,1862,1884]" pageId="6" pageNumber="279">E</emphasis>
: left anterior telopod, posterior view;
<emphasis id="B922F168FFE114258A7FD71FE353F8C5" bold="true" box="[796,811,1862,1884]" pageId="6" pageNumber="279">F</emphasis>
: left anterior telopod, lateral view
<emphasis id="B922F168FFE114258DF3D71FE4DCF8C5" bold="true" box="[1168,1188,1862,1884]" pageId="6" pageNumber="279">G</emphasis>
: right posterior telopod, posterior view;
<emphasis id="B922F168FFE114258856D73CE130F8E2" bold="true" box="[309,328,1893,1915]" pageId="6" pageNumber="279">H</emphasis>
: right posterior telopod, anterior view. Abbreviations:
<emphasis id="B922F168FFE114258A17D73CE3F2F8E2" bold="true" box="[884,906,1893,1915]" pageId="6" pageNumber="279">as</emphasis>
= apical spine;
<emphasis id="B922F168FFE114258D49D73CE444F8E2" bold="true" box="[1066,1084,1893,1915]" pageId="6" pageNumber="279">cl</emphasis>
= claw;
<emphasis id="B922F168FFE114258DF7D73CE4C2F8E2" bold="true" box="[1172,1210,1893,1915]" pageId="6" pageNumber="279">cr-t</emphasis>
= crenulated teeth;
<emphasis id="B922F168FFE114258CE7D73CE5E5F8E2" bold="true" box="[1412,1437,1893,1915]" pageId="6" pageNumber="279">cx</emphasis>
= coxa;
<emphasis id="B922F168FFE11425898AD7DDE085F803" bold="true" box="[233,253,1924,1946]" pageId="6" pageNumber="279">fe</emphasis>
= femur;
<emphasis id="B922F168FFE114258800D7DDE106F803" bold="true" box="[355,382,1924,1946]" pageId="6" pageNumber="279">gp</emphasis>
= gonopore;
<emphasis id="B922F168FFE114258B64D7DDE252F803" bold="true" box="[519,554,1924,1946]" pageId="6" pageNumber="279">pof</emphasis>
= postfemur;
<emphasis id="B922F168FFE114258BD9D7DDE29FF803" bold="true" box="[698,743,1924,1946]" pageId="6" pageNumber="279">pref</emphasis>
= prefemur;
<emphasis id="B922F168FFE114258A0ED7DDE3F5F803" bold="true" box="[877,909,1924,1946]" pageId="6" pageNumber="279">s-p</emphasis>
= sclerotized spot;
<emphasis id="B922F168FFE114258D36D7DDE407F803" bold="true" box="[1109,1151,1924,1946]" pageId="6" pageNumber="279">s-ps</emphasis>
= field of sclerotized spots;
<emphasis id="B922F168FFE1142589F4D7FAE0D4F820" bold="true" box="[151,172,1955,1977]" pageId="6" pageNumber="279">sr</emphasis>
= stridulation ribs;
<emphasis id="B922F168FFE114258816D7FAE1FFF820" bold="true" box="[373,391,1955,1977]" pageId="6" pageNumber="279">st</emphasis>
= stigmatic plate;
<emphasis id="B922F168FFE114258B25D7FAE212F820" bold="true" box="[582,618,1955,1977]" pageId="6" pageNumber="279">syn</emphasis>
= syncoxite;
<emphasis id="B922F168FFE114258B96D7FAE372F820" bold="true" box="[757,778,1955,1977]" pageId="6" pageNumber="279">ta</emphasis>
= tarsus;
<emphasis id="B922F168FFE114258A0CD7FAE306F820" bold="true" box="[879,894,1955,1977]" pageId="6" pageNumber="279">ti</emphasis>
= tibia;
<emphasis id="B922F168FFE114258AB6D7FAE393F820" bold="true" box="[981,1003,1955,1977]" pageId="6" pageNumber="279">vs</emphasis>
= ventral spines.
<emphasis id="B922F168FFE114258DC3D7FAE4A9F820" bold="true" box="[1184,1233,1955,1977]" pageId="6" pageNumber="279">A, B</emphasis>
Scale bars = 1 mm,
<emphasis id="B922F168FFE1142589F4D798E0B1F84E" bold="true" box="[151,201,1985,2007]" pageId="6" pageNumber="279">C–H</emphasis>
Scale bars = 0.5 mm.
</paragraph>
</caption>
<caption id="DF297DF2FFE0142489F4D2E5E2C8FC89" httpUri="https://zenodo.org/record/584047/files/figure.png" pageId="7" pageNumber="280" targetBox="[175,1398,193,689]" targetPageId="7">
<paragraph id="8BE92D7AFFE0142489F4D2E5E2C8FC89" blockId="7.[151,1436,700,784]" pageId="7" pageNumber="280">
<emphasis id="B922F168FFE0142489F4D2E5E16CFD4B" bold="true" box="[151,276,700,722]" pageId="7" pageNumber="280">FIGURE 3.</emphasis>
<taxonomicName id="4C5656F9FFE014248878D2E5E256FD4B" box="[283,558,700,722]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="280" phylum="Arthropoda" rank="species" species="bemanevika" status="sp. nov.">
<emphasis id="B922F168FFE014248878D2E5E256FD4B" box="[283,558,700,722]" italics="true" pageId="7" pageNumber="280">Zoosphaerium bemanevika</emphasis>
</taxonomicName>
<emphasis id="B922F168FFE014248B56D2E5E208FD4B" bold="true" box="[565,624,700,722]" pageId="7" pageNumber="280">
<taxonomicNameLabel id="A2114C13FFE014248B56D2E5E208FD4B" box="[565,624,700,722]" pageId="7" pageNumber="280" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female (ZFMK MYR 6144),
<emphasis id="B922F168FFE014248AC9D2E5E3C4FD4B" bold="true" box="[938,956,700,722]" pageId="7" pageNumber="280">A</emphasis>
: coxa of second left leg with vulva;
<emphasis id="B922F168FFE014248C5BD2E5E531FD4B" bold="true" box="[1336,1353,700,722]" pageId="7" pageNumber="280">B</emphasis>
: female subanal plate with washboard. Abbreviations:
<emphasis id="B922F168FFE014248BE0D282E2E3FD68" bold="true" box="[643,667,731,753]" pageId="7" pageNumber="280">cx</emphasis>
= coxa;
<emphasis id="B922F168FFE014248A63D282E361FD68" bold="true" box="[768,793,731,753]" pageId="7" pageNumber="280">ep</emphasis>
= external plate of vulva;
<emphasis id="B922F168FFE014248D5CD282E42CFD68" bold="true" box="[1087,1108,731,753]" pageId="7" pageNumber="280">ip</emphasis>
= internal plate of vulva;
<emphasis id="B922F168FFE014248C15D282E5FBFD68" bold="true" box="[1398,1411,731,753]" pageId="7" pageNumber="280">o</emphasis>
= operculum;
<emphasis id="B922F168FFE014248873D2A3E15DFC89" bold="true" box="[272,293,762,784]" pageId="7" pageNumber="280">sr</emphasis>
= stridulation ribs. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="DF297DF2FFE0142489F4D73DE247F86C" httpUri="https://zenodo.org/record/584048/files/figure.png" pageId="7" pageNumber="280" targetBox="[176,1389,821,1861]" targetPageId="7">
<paragraph id="8BE92D7AFFE0142489F4D73DE247F86C" blockId="7.[151,1436,1892,2037]" pageId="7" pageNumber="280">
<emphasis id="B922F168FFE0142489F4D73DE177F8E3" bold="true" box="[151,271,1892,1914]" pageId="7" pageNumber="280">FIGURE 4</emphasis>
.
<taxonomicName id="4C5656F9FFE01424887ED73DE248F8E3" box="[285,560,1892,1914]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="7" pageNumber="280" phylum="Arthropoda" rank="species" species="bemanevika" status="sp. nov.">
<emphasis id="B922F168FFE01424887ED73DE248F8E3" box="[285,560,1892,1914]" italics="true" pageId="7" pageNumber="280">Zoosphaerium bemanevika</emphasis>
</taxonomicName>
<emphasis id="B922F168FFE014248B5AD73DE20CF8E3" bold="true" box="[569,628,1892,1914]" pageId="7" pageNumber="280">
<taxonomicNameLabel id="A2114C13FFE014248B5AD73DE20CF8E3" box="[569,628,1892,1914]" pageId="7" pageNumber="280" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female (ZFMK MYR 6144), head, µCT,
<emphasis id="B922F168FFE014248D4FD73DE446F8E3" bold="true" box="[1068,1086,1892,1914]" pageId="7" pageNumber="280">A</emphasis>
: frontal view;
<emphasis id="B922F168FFE014248DB9D73DE493F8E3" bold="true" box="[1242,1259,1892,1914]" pageId="7" pageNumber="280">B</emphasis>
: lateral view;
<emphasis id="B922F168FFE014248CE0D73DE5EDF8E3" bold="true" box="[1411,1429,1892,1914]" pageId="7" pageNumber="280">C</emphasis>
: internal morphology. Abbreviations:
<emphasis id="B922F168FFE014248B7AD7DAE241F800" bold="true" box="[537,569,1923,1945]" pageId="7" pageNumber="280">aG</emphasis>
= antennal groove;
<emphasis id="B922F168FFE014248A71D7DAE34AF800" bold="true" box="[786,818,1923,1945]" pageId="7" pageNumber="280">aO</emphasis>
= aberrant ocellus;
<emphasis id="B922F168FFE014248D69D7DAE455F800" bold="true" box="[1034,1069,1923,1945]" pageId="7" pageNumber="280">c =</emphasis>
condyles;
<emphasis id="B922F168FFE014248DC1D7DAE4C3F800" bold="true" box="[1186,1211,1923,1945]" pageId="7" pageNumber="280">cp</emphasis>
= central pad;
<emphasis id="B922F168FFE014248C01D7DAE5FCF800" bold="true" box="[1378,1412,1923,1945]" pageId="7" pageNumber="280">Gn</emphasis>
= gnathochilarium;
<emphasis id="B922F168FFE014248828D7FBE11EF821" bold="true" box="[331,358,1954,1976]" pageId="7" pageNumber="280">gu</emphasis>
= gula;
<emphasis id="B922F168FFE0142488DDD7FBE1B4F821" bold="true" box="[446,460,1954,1976]" pageId="7" pageNumber="280">il</emphasis>
= incisura lateralis;
<emphasis id="B922F168FFE014248BC3D7FBE2CDF821" bold="true" box="[672,693,1954,1976]" pageId="7" pageNumber="280">ip</emphasis>
= inner palpus;
<emphasis id="B922F168FFE014248A3CD7FBE30CF821" bold="true" box="[863,884,1954,1976]" pageId="7" pageNumber="280">lb</emphasis>
= labrum;
<emphasis id="B922F168FFE014248A8BD7FBE472F821" bold="true" box="[1000,1034,1954,1976]" pageId="7" pageNumber="280">LL</emphasis>
= lamellae linguales;
<emphasis id="B922F168FFE014248D8CD7FBE558F821" bold="true" box="[1263,1312,1954,1976]" pageId="7" pageNumber="280">mdb</emphasis>
= mandible base;
<emphasis id="B922F168FFE0142489B2D799E085F84F" bold="true" box="[209,253,1984,2006]" pageId="7" pageNumber="280">mdt</emphasis>
= mandible tendon;
<emphasis id="B922F168FFE0142488ADD799E189F84F" bold="true" box="[462,497,1984,2006]" pageId="7" pageNumber="280">Me</emphasis>
= mentum;
<emphasis id="B922F168FFE014248B0FD799E2FCF84F" bold="true" box="[620,644,1984,2006]" pageId="7" pageNumber="280">oc</emphasis>
= ocelli;
<emphasis id="B922F168FFE014248B87D799E379F84F" bold="true" box="[740,769,1984,2006]" pageId="7" pageNumber="280">Ph</emphasis>
= pharynx;
<emphasis id="B922F168FFE014248A1ED799E3C0F84F" bold="true" box="[893,952,1984,2006]" pageId="7" pageNumber="280">PM =</emphasis>
paramentum;
<emphasis id="B922F168FFE014248D28D799E418F84F" bold="true" box="[1099,1120,1984,2006]" pageId="7" pageNumber="280">St</emphasis>
= stipites;
<emphasis id="B922F168FFE014248DB3D799E495F84F" bold="true" box="[1232,1261,1984,2006]" pageId="7" pageNumber="280">t =</emphasis>
tentorium;
<emphasis id="B922F168FFE014248C00D799E5FFF84F" bold="true" box="[1379,1415,1984,2006]" pageId="7" pageNumber="280">TO</emphasis>
= Organ of Tömosváry. Images not to scale.
</paragraph>
</caption>
<caption id="DF297DF2FFEF142B89F4D447E171FBE8" httpUri="https://zenodo.org/record/584049/files/figure.png" pageId="8" pageNumber="281" targetBox="[150,1436,194,1027]" targetPageId="8">
<paragraph id="8BE92D7AFFEF142B89F4D447E171FBE8" blockId="8.[151,1436,1054,1137]" pageId="8" pageNumber="281">
<emphasis id="B922F168FFEF142B89F4D447E16EFBAD" bold="true" box="[151,278,1054,1076]" pageId="8" pageNumber="281">FIGURE 5.</emphasis>
<taxonomicName id="4C5656F9FFEF142B8843D447E24DFBAD" box="[288,565,1054,1076]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="281" phylum="Arthropoda" rank="species" species="bemanevika" status="sp. nov.">
<emphasis id="B922F168FFEF142B8843D447E24DFBAD" box="[288,565,1054,1076]" italics="true" pageId="8" pageNumber="281">Zoosphaerium bemanevika</emphasis>
</taxonomicName>
<emphasis id="B922F168FFEF142B8B5CD447E204FBAD" bold="true" box="[575,636,1054,1076]" pageId="8" pageNumber="281">
<taxonomicNameLabel id="A2114C13FFEF142B8B5CD447E204FBAD" box="[575,636,1054,1076]" pageId="8" pageNumber="281" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype (FMNH-INS 2858686-C), left antenna, SEM,
<emphasis id="B922F168FFEF142B8DB6D447E49FFBAD" bold="true" box="[1237,1255,1054,1076]" pageId="8" pageNumber="281">A</emphasis>
: lateral view;
<emphasis id="B922F168FFEF142B8CE7D447E5EDFBAD" bold="true" box="[1412,1429,1054,1076]" pageId="8" pageNumber="281">B</emphasis>
: sensilla basiconica on 1st antennomere;
<emphasis id="B922F168FFEF142B8B23D465E22AFBCB" bold="true" box="[576,594,1084,1106]" pageId="8" pageNumber="281">C</emphasis>
: apical disc with sensory cones. Abbreviations:
<emphasis id="B922F168FFEF142B8D37D465E40AFBCB" bold="true" box="[1108,1138,1084,1106]" pageId="8" pageNumber="281">aD</emphasis>
= apical disc;
<emphasis id="B922F168FFEF142B8C75D465E556FBCB" bold="true" box="[1302,1326,1084,1106]" pageId="8" pageNumber="281">sb</emphasis>
= sensilla basiconica.
</paragraph>
</caption>
<subSubSection id="C34C7EF1FFEF142B89A4D4C0E40FFA13" pageId="8" pageNumber="281" type="diagnosis">
<paragraph id="8BE92D7AFFEF142B89A4D4C0E40FFA13" blockId="8.[151,1436,1177,1418]" pageId="8" pageNumber="281">
<emphasis id="B922F168FFEF142B89A4D4C0E139FB2B" bold="true" box="[199,321,1177,1202]" pageId="8" pageNumber="281">Diagnosis.</emphasis>
Females very large,&gt;
<quantity id="4CAE809FFFEF142B8B20D4C0E2EDFB28" box="[579,661,1177,1202]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="281" unit="mm" value="50.0">50 mm</quantity>
(
<figureCitation id="136D31FFFFEF142B8BC5D4C0E284FB28" box="[678,764,1177,1202]" captionStart="FIGURE 12" captionStartId="16.[151,250,1864,1886]" captionTargetBox="[163,1436,1100,1835]" captionTargetId="figure@16.[150,1436,1094,1842]" captionTargetPageId="16" captionText="FIGURE 12. Maximum likelihood tree inferred from the COI dataset with 1000 bootstrap pseudoreplicates implementing the GTR + I + G model. Habitus photograph shows Z. bemanevika (FMNH-INS 3196510). Colors used to separate species. Green = Z. bemanevika; Orange = Z. minutus. Circles indicating sister group relationships, as mapped in figure 1." httpUri="https://zenodo.org/record/584057/files/figure.png" pageId="8" pageNumber="281">Fig. 12</figureCitation>
). Posterior telopods (
<figureCitation id="136D31FFFFEF142B8A92D4C0E441FB28" box="[1009,1081,1177,1202]" captionStart="FIGURE 2" captionStartId="6.[151,250,1801,1823]" captionTargetBox="[168,1414,336,1783]" captionTargetId="figure@6.[168,1414,336,1783]" captionTargetPageId="6" captionText="FIGURE 2. Zoosphaerium bemanevika n. sp.., A, B: female (ZFMK MYR 6144); C-H: holotype male (FMNH-INS 2858686 - C); A: coxa of first left leg with first stigmatic plate; B: 9 th right leg; C coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view G: right posterior telopod, posterior view; H: right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-p = sclerotized spot; s-ps = field of sclerotized spots; sr = stridulation ribs; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. A, B Scale bars = 1 mm, C – H Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/584046/files/figure.png" pageId="8" pageNumber="281">Figs 2</figureCitation>
G, H) identify this species as a member of the
<taxonomicName id="4C5656F9FFEF142B8826D4E7E273FB4C" box="[325,523,1212,1237]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="281" phylum="Arthropoda" rank="species" species="coquerelianum">
<emphasis id="B922F168FFEF142B8826D4E7E273FB4C" box="[325,523,1212,1237]" italics="true" pageId="8" pageNumber="281">Z. coquerelianum</emphasis>
</taxonomicName>
species-group.
<taxonomicName id="4C5656F9FFEF142B8BDDD4E5E396FB4C" box="[702,1006,1212,1237]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="281" phylum="Arthropoda" rank="species" species="bemanevika" status="sp. nov.">
<emphasis id="B922F168FFEF142B8BDDD4E5E396FB4C" box="[702,1006,1212,1237]" italics="true" pageId="8" pageNumber="281">Zoosphaerium bemanevika</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2114C13FFEF142B8A95D4E7E44BFB4C" box="[1014,1075,1214,1237]" pageId="8" pageNumber="281" rank="species">n. sp.</taxonomicNameLabel>
has two stridulation ribs on the male harp, more than 20 apical cones on the antenna, and more than 10 ventral spines on the tarsi of walking legs, characters only shared with
<taxonomicName id="4C5656F9FFEF142B88BCD55FE227FA87" box="[479,607,1285,1310]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="281" phylum="Arthropoda" rank="species" species="villosum">
<emphasis id="B922F168FFEF142B88BCD55FE227FA87" box="[479,607,1285,1310]" italics="true" pageId="8" pageNumber="281">Z. villosum</emphasis>
</taxonomicName>
.
<taxonomicName id="4C5656F9FFEF142B8B12D55CE3DCFA87" box="[625,932,1285,1310]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="8" pageNumber="281" phylum="Arthropoda" rank="species" species="bemanevika" status="sp. nov.">
<emphasis id="B922F168FFEF142B8B12D55CE3DCFA87" box="[625,932,1285,1310]" italics="true" pageId="8" pageNumber="281">Zoosphaerium bemanevika</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2114C13FFEF142B8ACDD55FE396FA84" box="[942,1006,1286,1309]" pageId="8" pageNumber="281" rank="species">n. sp.</taxonomicNameLabel>
differs from the latter species in the presence of only one row of marginal bristles on the endotergum (
<figureCitation id="136D31FFFFEF142B8A14D570E3C1FAD8" box="[887,953,1321,1346]" captionStart="FIGURE 7" captionStartId="10.[151,250,849,871]" captionTargetBox="[151,1435,194,824]" captionTargetId="figure@10.[151,1435,193,828]" captionTargetPageId="10" captionText="FIGURE 7. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), endotergum of mid-body tergite, SEM, A: ventral view; B: bristles on apical margin." httpUri="https://zenodo.org/record/584051/files/figure.png" pageId="8" pageNumber="281">Fig. 7</figureCitation>
A), only 20–30 apical cones on the antenna (
<figureCitation id="136D31FFFFEF142B89FCD514E09BFAFF" box="[159,227,1357,1382]" captionStart="FIGURE 5" captionStartId="8.[151,250,1054,1076]" captionTargetBox="[150,1436,194,1027]" captionTargetId="figure@8.[150,1437,193,1033]" captionTargetPageId="8" captionText="FIGURE 5. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), left antenna, SEM, A: lateral view; B: sensilla basiconica on 1 st antennomere; C: apical disc with sensory cones. Abbreviations: aD = apical disc; sb = sensilla basiconica." httpUri="https://zenodo.org/record/584049/files/figure.png" pageId="8" pageNumber="281">Fig. 5</figureCitation>
C), the presence of sclerotized teeth on antennomere 4 (
<figureCitation id="136D31FFFFEF142B8A04D514E3D2FAFF" box="[871,938,1357,1382]" captionStart="FIGURE 5" captionStartId="8.[151,250,1054,1076]" captionTargetBox="[150,1436,194,1027]" captionTargetId="figure@8.[150,1437,193,1033]" captionTargetPageId="8" captionText="FIGURE 5. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), left antenna, SEM, A: lateral view; B: sensilla basiconica on 1 st antennomere; C: apical disc with sensory cones. Abbreviations: aD = apical disc; sb = sensilla basiconica." httpUri="https://zenodo.org/record/584049/files/figure.png" pageId="8" pageNumber="281">Fig. 5</figureCitation>
A) and the absence of pits and setae on the tergites. Both species also differ by an uncorrected p-distance of the COI gene of 10.6%.
</paragraph>
</subSubSection>
<subSubSection id="C34C7EF1FFEF142889F4D5E0E1EFFE35" lastPageId="11" lastPageNumber="284" pageId="8" pageNumber="281" type="description">
<paragraph id="8BE92D7AFFEF142B89F4D5E0E23EF983" blockId="8.[151,1436,1465,2030]" pageId="8" pageNumber="281">
<emphasis id="B922F168FFEF142B89F4D5E0E151FA4B" bold="true" box="[151,297,1465,1490]" pageId="8" pageNumber="281">Description.</emphasis>
<emphasis id="B922F168FFEF142B884CD5E0E1CEFA4B" box="[303,438,1465,1490]" italics="true" pageId="8" pageNumber="281">Body length</emphasis>
:
<typeStatus id="54ED93D8FFEF142B88A6D5E0E25FFA4B" box="[453,551,1465,1490]" pageId="8" pageNumber="281" type="holotype">holotype</typeStatus>
male: length ca.
<quantity id="4CAE809FFFEF142B8B84D5E0E340FA48" box="[743,824,1465,1489]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="8" pageNumber="281" unit="mm" value="22.0">22 mm</quantity>
, width of thoracic shield
<quantity id="4CAE809FFFEF142B8D37D5E0E4CFFA48" box="[1108,1207,1465,1489]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="8" pageNumber="281" unit="mm" value="11.1">11.1 mm</quantity>
(=widest), height of thoracic shield
<quantity id="4CAE809FFFEF142B8827D584E1E3FA6C" box="[324,411,1501,1526]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="8" pageNumber="281" unit="mm" value="6.2">6.2 mm</quantity>
(=highest). Female: length ca.
<quantity id="4CAE809FFFEF142B8B9ED584E336FA6C" box="[765,846,1501,1526]" metricMagnitude="-2" metricUnit="m" metricValue="5.4" pageId="8" pageNumber="281" unit="mm" value="54.0">54 mm</quantity>
, width of thoracic shield
<quantity id="4CAE809FFFEF142B8D0CD584E4ACFA6C" box="[1135,1236,1501,1525]" metricMagnitude="-2" metricUnit="m" metricValue="2.88" pageId="8" pageNumber="281" unit="mm" value="28.8">28.8 mm</quantity>
(=widest), height of thoracic shield
<quantity id="4CAE809FFFEF142B8803D658E1BCF980" box="[352,452,1537,1562]" metricMagnitude="-2" metricUnit="m" metricValue="1.67" pageId="8" pageNumber="281" unit="mm" value="16.7">16.7 mm</quantity>
(=highest).
</paragraph>
<paragraph id="8BE92D7AFFEF142B89A4D67CE2CDF933" blockId="8.[151,1436,1465,2030]" pageId="8" pageNumber="281">
<emphasis id="B922F168FFEF142B89A4D67CE13AF9A7" box="[199,322,1573,1598]" italics="true" pageId="8" pageNumber="281">Coloration</emphasis>
: some discoloration may have occurred because of preservation in ethanol. Tergites orange to lightbrown with a slightly darker rim at posterior edge of tergite. Anal shield uniform solid color, more orange than tergites. Collum and thoracic shield darker brown to olive in color (
<figureCitation id="136D31FFFFEF142B8AF7D634E390F91C" box="[916,1000,1645,1670]" captionStart="FIGURE 12" captionStartId="16.[151,250,1864,1886]" captionTargetBox="[163,1436,1100,1835]" captionTargetId="figure@16.[150,1436,1094,1842]" captionTargetPageId="16" captionText="FIGURE 12. Maximum likelihood tree inferred from the COI dataset with 1000 bootstrap pseudoreplicates implementing the GTR + I + G model. Habitus photograph shows Z. bemanevika (FMNH-INS 3196510). Colors used to separate species. Green = Z. bemanevika; Orange = Z. minutus. Circles indicating sister group relationships, as mapped in figure 1." httpUri="https://zenodo.org/record/584057/files/figure.png" pageId="8" pageNumber="281">Fig. 12</figureCitation>
). Head and antennae light-brown, legs light-brown with slightly green apical segments.
</paragraph>
<paragraph id="8BE92D7AFFEF142B89A4D6ECE5E4F850" blockId="8.[151,1436,1465,2030]" pageId="8" pageNumber="281">
<emphasis id="B922F168FFEF142B89A4D6ECE17CF957" box="[199,260,1717,1742]" italics="true" pageId="8" pageNumber="281">Head</emphasis>
: wide and short (
<figureCitation id="136D31FFFFEF142B88A8D6ECE268F954" box="[459,528,1717,1742]" captionStart="FIGURE 4" captionStartId="7.[151,250,1892,1914]" captionTargetBox="[176,1389,821,1861]" captionTargetId="figure@7.[176,1390,821,1862]" captionTargetPageId="7" captionText="FIGURE 4. Zoosphaerium bemanevika n. sp., female (ZFMK MYR 6144), head, µCT, A: frontal view; B: lateral view; C: internal morphology. Abbreviations: aG = antennal groove; aO = aberrant ocellus; c = condyles; cp = central pad; Gn = gnathochilarium; gu = gula; il = incisura lateralis; ip = inner palpus; lb = labrum; LL = lamellae linguales; mdb = mandible base; mdt = mandible tendon; Me = mentum; oc = ocelli; Ph = pharynx; PM = paramentum; St = stipites; t = tentorium; TO = Organ of Tömosváry. Images not to scale." httpUri="https://zenodo.org/record/584048/files/figure.png" pageId="8" pageNumber="281">Fig. 4</figureCitation>
A). Eyes with&gt;80 ocelli. Aberrant ocellus located in antennal groove (
<figureCitation id="136D31FFFFEF142B8C55D6ECE503F954" box="[1334,1403,1717,1742]" captionStart="FIGURE 4" captionStartId="7.[151,250,1892,1914]" captionTargetBox="[176,1389,821,1861]" captionTargetId="figure@7.[176,1390,821,1862]" captionTargetPageId="7" captionText="FIGURE 4. Zoosphaerium bemanevika n. sp., female (ZFMK MYR 6144), head, µCT, A: frontal view; B: lateral view; C: internal morphology. Abbreviations: aG = antennal groove; aO = aberrant ocellus; c = condyles; cp = central pad; Gn = gnathochilarium; gu = gula; il = incisura lateralis; ip = inner palpus; lb = labrum; LL = lamellae linguales; mdb = mandible base; mdt = mandible tendon; Me = mentum; oc = ocelli; Ph = pharynx; PM = paramentum; St = stipites; t = tentorium; TO = Organ of Tömosváry. Images not to scale." httpUri="https://zenodo.org/record/584048/files/figure.png" pageId="8" pageNumber="281">Fig. 4</figureCitation>
B). Antennae short, with cylindrical joints, protruding posteriorly to leg pair 3. Size of antennomeres 1&gt;2=3&gt;4&lt;5&lt;6 (
<figureCitation id="136D31FFFFEF142B89FCD6A4E09AF88F" box="[159,226,1789,1814]" captionStart="FIGURE 5" captionStartId="8.[151,250,1054,1076]" captionTargetBox="[150,1436,194,1027]" captionTargetId="figure@8.[150,1437,193,1033]" captionTargetPageId="8" captionText="FIGURE 5. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), left antenna, SEM, A: lateral view; B: sensilla basiconica on 1 st antennomere; C: apical disc with sensory cones. Abbreviations: aD = apical disc; sb = sensilla basiconica." httpUri="https://zenodo.org/record/584049/files/figure.png" pageId="8" pageNumber="281">Fig. 5</figureCitation>
A). Lateral base of antennomeres 1–4 with small scale-like teeth, numerous teeth present on antennomere 1 protruding to base of antennomere 2, only very few teeth present on antennomeres 3 and 4, restricted to base. One apical row of sensilla basiconica and a shallow groove present on antennomere 1 (
<figureCitation id="136D31FFFFEF142B8D0BD71CE4D7F8C7" box="[1128,1199,1861,1886]" captionStart="FIGURE 5" captionStartId="8.[151,250,1054,1076]" captionTargetBox="[150,1436,194,1027]" captionTargetId="figure@8.[150,1437,193,1033]" captionTargetPageId="8" captionText="FIGURE 5. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), left antenna, SEM, A: lateral view; B: sensilla basiconica on 1 st antennomere; C: apical disc with sensory cones. Abbreviations: aD = apical disc; sb = sensilla basiconica." httpUri="https://zenodo.org/record/584049/files/figure.png" pageId="8" pageNumber="281">Fig. 5</figureCitation>
B). Few larger setae present on antennomeres 1–4, more setae present on antennomere 5, antennomere 6 densely pubescent, possessing single row of sensilla basiconica surrounding apical disc. Male apical disc with 21/28 apical sensory cones (
<figureCitation id="136D31FFFFEF142B8C0CD7D4E0DDF853" captionStart="FIGURE 5" captionStartId="8.[151,250,1054,1076]" captionTargetBox="[150,1436,194,1027]" captionTargetId="figure@8.[150,1437,193,1033]" captionTargetPageId="8" captionText="FIGURE 5. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), left antenna, SEM, A: lateral view; B: sensilla basiconica on 1 st antennomere; C: apical disc with sensory cones. Abbreviations: aD = apical disc; sb = sensilla basiconica." httpUri="https://zenodo.org/record/584049/files/figure.png" pageId="8" pageNumber="281">Fig. 5</figureCitation>
C), disc of much larger female with 32/33 apical cones. Organ of Tömösváry located in antennal groove (
<figureCitation id="136D31FFFFEF142B8C55D7E8E501F850" box="[1334,1401,1969,1994]" captionStart="FIGURE 4" captionStartId="7.[151,250,1892,1914]" captionTargetBox="[176,1389,821,1861]" captionTargetId="figure@7.[176,1390,821,1862]" captionTargetPageId="7" captionText="FIGURE 4. Zoosphaerium bemanevika n. sp., female (ZFMK MYR 6144), head, µCT, A: frontal view; B: lateral view; C: internal morphology. Abbreviations: aG = antennal groove; aO = aberrant ocellus; c = condyles; cp = central pad; Gn = gnathochilarium; gu = gula; il = incisura lateralis; ip = inner palpus; lb = labrum; LL = lamellae linguales; mdb = mandible base; mdt = mandible tendon; Me = mentum; oc = ocelli; Ph = pharynx; PM = paramentum; St = stipites; t = tentorium; TO = Organ of Tömosváry. Images not to scale." httpUri="https://zenodo.org/record/584048/files/figure.png" pageId="8" pageNumber="281">Fig. 4</figureCitation>
A).
</paragraph>
<paragraph id="8BE92D7AFFEF142A89A4D78CE24DFED9" blockId="8.[151,1436,1465,2030]" lastBlockId="9.[151,1436,151,608]" lastPageId="9" lastPageNumber="282" pageId="8" pageNumber="281">
<emphasis id="B922F168FFEF142B89A4D78CE1E9F877" box="[199,401,2005,2030]" italics="true" pageId="8" pageNumber="281">Gnathochilarium:</emphasis>
structure typical of the order (
<figureCitation id="136D31FFFFEF142B8B9FD78CE33BF877" box="[764,835,2005,2030]" captionStart="FIGURE 6" captionStartId="9.[151,250,1889,1911]" captionTargetBox="[152,1433,664,1861]" captionTargetId="figure@9.[152,1435,663,1867]" captionTargetPageId="9" captionText="FIGURE 6. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), SEM, A: gnathochilarium, ventral view; B: gnathochilarium, apical view of dorsal side; C: gnathochilarium, rudimentary lateral palpus with sensory cones; D: gnathochilarium, sensory cones on central pad; E: left mandible, mesal view; F: left mandible, pectinate lamellae. Abbreviations: c = condyles; cp = central pad; eT = external tooth; iP = inner palpus; iT = internal tooth; LL = lamellae linguales; LP = lateral palpus; mp = molar plate; pL = pectinate lamellae; St = stipites." httpUri="https://zenodo.org/record/584050/files/figure.png" pageId="8" pageNumber="281">Fig. 6</figureCitation>
A). Lateral stipites and paramentum bearing several long setae (
<figureCitation id="136D31FFFFEE142A8875D0CEE120FF29" box="[278,344,151,176]" captionStart="FIGURE 6" captionStartId="9.[151,250,1889,1911]" captionTargetBox="[152,1433,664,1861]" captionTargetId="figure@9.[152,1435,663,1867]" captionTargetPageId="9" captionText="FIGURE 6. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), SEM, A: gnathochilarium, ventral view; B: gnathochilarium, apical view of dorsal side; C: gnathochilarium, rudimentary lateral palpus with sensory cones; D: gnathochilarium, sensory cones on central pad; E: left mandible, mesal view; F: left mandible, pectinate lamellae. Abbreviations: c = condyles; cp = central pad; eT = external tooth; iP = inner palpus; iT = internal tooth; LL = lamellae linguales; LP = lateral palpus; mp = molar plate; pL = pectinate lamellae; St = stipites." httpUri="https://zenodo.org/record/584050/files/figure.png" pageId="9" pageNumber="282">Fig. 6</figureCitation>
B). No setae present at center of lamellae linguales (
<figureCitation id="136D31FFFFEE142A8AF8D0CEE3A5FF29" box="[923,989,151,176]" captionStart="FIGURE 6" captionStartId="9.[151,250,1889,1911]" captionTargetBox="[152,1433,664,1861]" captionTargetId="figure@9.[152,1435,663,1867]" captionTargetPageId="9" captionText="FIGURE 6. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), SEM, A: gnathochilarium, ventral view; B: gnathochilarium, apical view of dorsal side; C: gnathochilarium, rudimentary lateral palpus with sensory cones; D: gnathochilarium, sensory cones on central pad; E: left mandible, mesal view; F: left mandible, pectinate lamellae. Abbreviations: c = condyles; cp = central pad; eT = external tooth; iP = inner palpus; iT = internal tooth; LL = lamellae linguales; LP = lateral palpus; mp = molar plate; pL = pectinate lamellae; St = stipites." httpUri="https://zenodo.org/record/584050/files/figure.png" pageId="9" pageNumber="282">Fig. 6</figureCitation>
A). Inner palpi on stipites bearing single field of large sensory cones interspersed with smaller scale-like structures (
<figureCitation id="136D31FFFFEE142A8D63D0E5E43EFF4C" box="[1024,1094,188,213]" captionStart="FIGURE 6" captionStartId="9.[151,250,1889,1911]" captionTargetBox="[152,1433,664,1861]" captionTargetId="figure@9.[152,1435,663,1867]" captionTargetPageId="9" captionText="FIGURE 6. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), SEM, A: gnathochilarium, ventral view; B: gnathochilarium, apical view of dorsal side; C: gnathochilarium, rudimentary lateral palpus with sensory cones; D: gnathochilarium, sensory cones on central pad; E: left mandible, mesal view; F: left mandible, pectinate lamellae. Abbreviations: c = condyles; cp = central pad; eT = external tooth; iP = inner palpus; iT = internal tooth; LL = lamellae linguales; LP = lateral palpus; mp = molar plate; pL = pectinate lamellae; St = stipites." httpUri="https://zenodo.org/record/584050/files/figure.png" pageId="9" pageNumber="282">Fig. 6</figureCitation>
B). Lateral palpi rudimentary, consisting of four sensory cones (
<figureCitation id="136D31FFFFEE142A8B4ED086E20EFF61" box="[557,630,223,248]" captionStart="FIGURE 6" captionStartId="9.[151,250,1889,1911]" captionTargetBox="[152,1433,664,1861]" captionTargetId="figure@9.[152,1435,663,1867]" captionTargetPageId="9" captionText="FIGURE 6. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), SEM, A: gnathochilarium, ventral view; B: gnathochilarium, apical view of dorsal side; C: gnathochilarium, rudimentary lateral palpus with sensory cones; D: gnathochilarium, sensory cones on central pad; E: left mandible, mesal view; F: left mandible, pectinate lamellae. Abbreviations: c = condyles; cp = central pad; eT = external tooth; iP = inner palpus; iT = internal tooth; LL = lamellae linguales; LP = lateral palpus; mp = molar plate; pL = pectinate lamellae; St = stipites." httpUri="https://zenodo.org/record/584050/files/figure.png" pageId="9" pageNumber="282">Fig. 6</figureCitation>
C).
<collectingRegion id="4992E398FFEE142A8BC6D086E28FFF61" box="[677,759,223,248]" country="Fiji" name="Central" pageId="9" pageNumber="282">Central</collectingRegion>
pads as protuberance of endochilarium projecting above lamellae linguales. Each pad with a broad, flat incision and a single field of different
<typeStatus id="54ED93D8FFEE142A8D10D15CE4D6FE84" box="[1139,1198,261,285]" pageId="9" pageNumber="282">types</typeStatus>
of sensory cones on underside of posterior half (
<figureCitation id="136D31FFFFEE142A88ADD17EE26AFED9" box="[462,530,295,320]" captionStart="FIGURE 6" captionStartId="9.[151,250,1889,1911]" captionTargetBox="[152,1433,664,1861]" captionTargetId="figure@9.[152,1435,663,1867]" captionTargetPageId="9" captionText="FIGURE 6. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), SEM, A: gnathochilarium, ventral view; B: gnathochilarium, apical view of dorsal side; C: gnathochilarium, rudimentary lateral palpus with sensory cones; D: gnathochilarium, sensory cones on central pad; E: left mandible, mesal view; F: left mandible, pectinate lamellae. Abbreviations: c = condyles; cp = central pad; eT = external tooth; iP = inner palpus; iT = internal tooth; LL = lamellae linguales; LP = lateral palpus; mp = molar plate; pL = pectinate lamellae; St = stipites." httpUri="https://zenodo.org/record/584050/files/figure.png" pageId="9" pageNumber="282">Fig. 6</figureCitation>
D).
</paragraph>
<paragraph id="8BE92D7AFFEE142A89A4D115E3BBFE35" blockId="9.[151,1436,151,608]" pageId="9" pageNumber="282">
<emphasis id="B922F168FFEE142A89A4D115E143FEFC" box="[199,315,332,357]" italics="true" pageId="9" pageNumber="282">Mandible:</emphasis>
typical shape for the order, inner tooth three-combed (
<figureCitation id="136D31FFFFEE142A8AD9D115E478FEFC" box="[954,1024,332,357]" captionStart="FIGURE 6" captionStartId="9.[151,250,1889,1911]" captionTargetBox="[152,1433,664,1861]" captionTargetId="figure@9.[152,1435,663,1867]" captionTargetPageId="9" captionText="FIGURE 6. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), SEM, A: gnathochilarium, ventral view; B: gnathochilarium, apical view of dorsal side; C: gnathochilarium, rudimentary lateral palpus with sensory cones; D: gnathochilarium, sensory cones on central pad; E: left mandible, mesal view; F: left mandible, pectinate lamellae. Abbreviations: c = condyles; cp = central pad; eT = external tooth; iP = inner palpus; iT = internal tooth; LL = lamellae linguales; LP = lateral palpus; mp = molar plate; pL = pectinate lamellae; St = stipites." httpUri="https://zenodo.org/record/584050/files/figure.png" pageId="9" pageNumber="282">Fig. 6</figureCitation>
E), pectinate lamellae arranged in 5 rows (
<figureCitation id="136D31FFFFEE142A89BFD136E167FE11" box="[220,287,367,392]" captionStart="FIGURE 6" captionStartId="9.[151,250,1889,1911]" captionTargetBox="[152,1433,664,1861]" captionTargetId="figure@9.[152,1435,663,1867]" captionTargetPageId="9" captionText="FIGURE 6. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), SEM, A: gnathochilarium, ventral view; B: gnathochilarium, apical view of dorsal side; C: gnathochilarium, rudimentary lateral palpus with sensory cones; D: gnathochilarium, sensory cones on central pad; E: left mandible, mesal view; F: left mandible, pectinate lamellae. Abbreviations: c = condyles; cp = central pad; eT = external tooth; iP = inner palpus; iT = internal tooth; LL = lamellae linguales; LP = lateral palpus; mp = molar plate; pL = pectinate lamellae; St = stipites." httpUri="https://zenodo.org/record/584050/files/figure.png" pageId="9" pageNumber="282">Fig. 6</figureCitation>
F). Condylus pointing posteriorly, apically with a sharp-edged step (
<figureCitation id="136D31FFFFEE142A8D76D136E421FE11" box="[1045,1113,367,392]" captionStart="FIGURE 4" captionStartId="7.[151,250,1892,1914]" captionTargetBox="[176,1389,821,1861]" captionTargetId="figure@7.[176,1390,821,1862]" captionTargetPageId="7" captionText="FIGURE 4. Zoosphaerium bemanevika n. sp., female (ZFMK MYR 6144), head, µCT, A: frontal view; B: lateral view; C: internal morphology. Abbreviations: aG = antennal groove; aO = aberrant ocellus; c = condyles; cp = central pad; Gn = gnathochilarium; gu = gula; il = incisura lateralis; ip = inner palpus; lb = labrum; LL = lamellae linguales; mdb = mandible base; mdt = mandible tendon; Me = mentum; oc = ocelli; Ph = pharynx; PM = paramentum; St = stipites; t = tentorium; TO = Organ of Tömosváry. Images not to scale." httpUri="https://zenodo.org/record/584048/files/figure.png" pageId="9" pageNumber="282">Fig. 4</figureCitation>
C). Large ventral mandibular base (
<figureCitation id="136D31FFFFEE142A89BBD1CDE163FE35" box="[216,283,404,429]" captionStart="FIGURE 4" captionStartId="7.[151,250,1892,1914]" captionTargetBox="[176,1389,821,1861]" captionTargetId="figure@7.[176,1390,821,1862]" captionTargetPageId="7" captionText="FIGURE 4. Zoosphaerium bemanevika n. sp., female (ZFMK MYR 6144), head, µCT, A: frontal view; B: lateral view; C: internal morphology. Abbreviations: aG = antennal groove; aO = aberrant ocellus; c = condyles; cp = central pad; Gn = gnathochilarium; gu = gula; il = incisura lateralis; ip = inner palpus; lb = labrum; LL = lamellae linguales; mdb = mandible base; mdt = mandible tendon; Me = mentum; oc = ocelli; Ph = pharynx; PM = paramentum; St = stipites; t = tentorium; TO = Organ of Tömosváry. Images not to scale." httpUri="https://zenodo.org/record/584048/files/figure.png" pageId="9" pageNumber="282">Fig. 4</figureCitation>
A). Mentum of gnathochilarium connected to gula (
<figureCitation id="136D31FFFFEE142A8A3ED1CDE3DAFE35" box="[861,930,404,429]" captionStart="FIGURE 4" captionStartId="7.[151,250,1892,1914]" captionTargetBox="[176,1389,821,1861]" captionTargetId="figure@7.[176,1390,821,1862]" captionTargetPageId="7" captionText="FIGURE 4. Zoosphaerium bemanevika n. sp., female (ZFMK MYR 6144), head, µCT, A: frontal view; B: lateral view; C: internal morphology. Abbreviations: aG = antennal groove; aO = aberrant ocellus; c = condyles; cp = central pad; Gn = gnathochilarium; gu = gula; il = incisura lateralis; ip = inner palpus; lb = labrum; LL = lamellae linguales; mdb = mandible base; mdt = mandible tendon; Me = mentum; oc = ocelli; Ph = pharynx; PM = paramentum; St = stipites; t = tentorium; TO = Organ of Tömosváry. Images not to scale." httpUri="https://zenodo.org/record/584048/files/figure.png" pageId="9" pageNumber="282">Fig. 4</figureCitation>
C).
</paragraph>
<paragraph id="8BE92D7AFFEE142A89A4D1E1E45EFE6D" blockId="9.[151,1436,151,608]" pageId="9" pageNumber="282">
<emphasis id="B922F168FFEE142A89A4D1E1E107FE49" box="[199,383,439,464]" italics="true" pageId="9" pageNumber="282">Stigmatic plates</emphasis>
: first stigmatic plate triangular, apex well rounded, nearly as wide as at base. Hair most abundant on apical margin, very few hairs on basal and lateral margins (
<figureCitation id="136D31FFFFEE142A8ADDD185E47AFE6D" box="[958,1026,476,501]" captionStart="FIGURE 2" captionStartId="6.[151,250,1801,1823]" captionTargetBox="[168,1414,336,1783]" captionTargetId="figure@6.[168,1414,336,1783]" captionTargetPageId="6" captionText="FIGURE 2. Zoosphaerium bemanevika n. sp.., A, B: female (ZFMK MYR 6144); C-H: holotype male (FMNH-INS 2858686 - C); A: coxa of first left leg with first stigmatic plate; B: 9 th right leg; C coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view G: right posterior telopod, posterior view; H: right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-p = sclerotized spot; s-ps = field of sclerotized spots; sr = stridulation ribs; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. A, B Scale bars = 1 mm, C – H Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/584046/files/figure.png" pageId="9" pageNumber="282">Fig. 2</figureCitation>
A).
</paragraph>
<paragraph id="8BE92D7AFFEE142A89A4D1A6E50FFD8E" blockId="9.[151,1436,151,608]" box="[199,1399,511,536]" pageId="9" pageNumber="282">
<emphasis id="B922F168FFEE142A89A4D1A6E14DFD81" box="[199,309,511,536]" italics="true" pageId="9" pageNumber="282">Pleurites:</emphasis>
first pleurite laterally modified and extending backwards with a broad, apically rounded process.
</paragraph>
<paragraph id="8BE92D7AFFEE142A89A4D27DE51AFDA4" blockId="9.[151,1436,151,608]" box="[199,1378,548,573]" pageId="9" pageNumber="282">
<emphasis id="B922F168FFEE142A89A4D27DE162FDA4" box="[199,282,548,573]" italics="true" pageId="9" pageNumber="282">Collum</emphasis>
: glabrous with very sparse setae around margins, mostly located at corners on either side of head.
</paragraph>
<paragraph id="8BE92D7AFFEE142A89A4D21EE305FDF9" blockId="9.[151,1436,151,608]" box="[199,893,583,608]" pageId="9" pageNumber="282">
<emphasis id="B922F168FFEE142A89A4D21EE10EFDF9" box="[199,374,583,608]" italics="true" pageId="9" pageNumber="282">Thoracic shield</emphasis>
: smooth and glabrous, without protuberances.
</paragraph>
<caption id="DF297DF2FFEE142A89F4D738E474F86B" httpUri="https://zenodo.org/record/584050/files/figure.png" pageId="9" pageNumber="282" targetBox="[152,1433,664,1861]" targetPageId="9">
<paragraph id="8BE92D7AFFEE142A89F4D738E474F86B" blockId="9.[151,1437,1889,2034]" pageId="9" pageNumber="282">
<emphasis id="B922F168FFEE142A89F4D738E16AF8EE" bold="true" box="[151,274,1889,1911]" pageId="9" pageNumber="282">FIGURE 6.</emphasis>
<taxonomicName id="4C5656F9FFEE142A887AD738E253F8EE" box="[281,555,1889,1911]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="9" pageNumber="282" phylum="Arthropoda" rank="species" species="bemanevika" status="sp. nov.">
<emphasis id="B922F168FFEE142A887AD738E253F8EE" box="[281,555,1889,1911]" italics="true" pageId="9" pageNumber="282">Zoosphaerium bemanevika</emphasis>
</taxonomicName>
<emphasis id="B922F168FFEE142A8B52D738E212F8EE" bold="true" box="[561,618,1889,1911]" pageId="9" pageNumber="282">
<taxonomicNameLabel id="A2114C13FFEE142A8B52D738E212F8EE" box="[561,618,1889,1911]" pageId="9" pageNumber="282" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype (FMNH-INS 2858686-C), SEM,
<emphasis id="B922F168FFEE142A8D45D738E440F8EE" bold="true" box="[1062,1080,1889,1911]" pageId="9" pageNumber="282">A</emphasis>
: gnathochilarium, ventral view;
<emphasis id="B922F168FFEE142A8CE7D738E5EDF8EE" bold="true" box="[1412,1429,1889,1911]" pageId="9" pageNumber="282">B</emphasis>
: gnathochilarium, apical view of dorsal side;
<emphasis id="B922F168FFEE142A8BEED7D9E2E7F80F" bold="true" box="[653,671,1920,1942]" pageId="9" pageNumber="282">C</emphasis>
: gnathochilarium, rudimentary lateral palpus with sensory cones;
<emphasis id="B922F168FFEE142A8CE0D7D9E5EDF80F" bold="true" box="[1411,1429,1920,1942]" pageId="9" pageNumber="282">D</emphasis>
: gnathochilarium, sensory cones on central pad;
<emphasis id="B922F168FFEE142A8BCDD7C6E2C7F82C" bold="true" box="[686,703,1951,1973]" pageId="9" pageNumber="282">E</emphasis>
: left mandible, mesal view;
<emphasis id="B922F168FFEE142A8D68D7C6E462F82C" bold="true" box="[1035,1050,1951,1973]" pageId="9" pageNumber="282">F</emphasis>
: left mandible, pectinate lamellae. Abbreviations:
<emphasis id="B922F168FFEE142A8854D7E4E13AF84A" bold="true" box="[311,322,1981,2003]" pageId="9" pageNumber="282">c</emphasis>
= condyles;
<emphasis id="B922F168FFEE142A88AED7E4E19EF84A" bold="true" box="[461,486,1981,2003]" pageId="9" pageNumber="282">cp</emphasis>
= central pad;
<emphasis id="B922F168FFEE142A8BE8D7E4E2DFF84A" bold="true" box="[651,679,1981,2003]" pageId="9" pageNumber="282">eT</emphasis>
= external tooth;
<emphasis id="B922F168FFEE142A8A0BD7E4E306F84A" bold="true" box="[872,894,1981,2003]" pageId="9" pageNumber="282">iP</emphasis>
= inner palpus;
<emphasis id="B922F168FFEE142A8D4CD7E4E43FF84A" bold="true" box="[1071,1095,1981,2003]" pageId="9" pageNumber="282">iT</emphasis>
= internal tooth;
<emphasis id="B922F168FFEE142A8C60D7E4E55DF84A" bold="true" box="[1283,1317,1981,2003]" pageId="9" pageNumber="282">LL</emphasis>
= lamellae linguales;
<emphasis id="B922F168FFEE142A899CD785E167F86B" bold="true" box="[255,287,2012,2034]" pageId="9" pageNumber="282">LP</emphasis>
= lateral palpus;
<emphasis id="B922F168FFEE142A88B3D785E18BF86B" bold="true" box="[464,499,2012,2034]" pageId="9" pageNumber="282">mp</emphasis>
= molar plate;
<emphasis id="B922F168FFEE142A8BEDD785E2D5F86B" bold="true" box="[654,685,2012,2034]" pageId="9" pageNumber="282">pL</emphasis>
= pectinate lamellae;
<emphasis id="B922F168FFEE142A8AEFD785E3D9F86B" bold="true" box="[908,929,2012,2034]" pageId="9" pageNumber="282">St</emphasis>
= stipites.
</paragraph>
</caption>
<caption id="DF297DF2FFED142989F4D308E239FC1F" httpUri="https://zenodo.org/record/584051/files/figure.png" pageId="10" pageNumber="283" targetBox="[151,1435,194,824]" targetPageId="10">
<paragraph id="8BE92D7AFFED142989F4D308E239FC1F" blockId="10.[151,1436,849,902]" pageId="10" pageNumber="283">
<emphasis id="B922F168FFED142989F4D308E16BFCFE" bold="true" box="[151,275,849,871]" pageId="10" pageNumber="283">FIGURE 7.</emphasis>
<taxonomicName id="4C5656F9FFED14298878D308E255FCFE" box="[283,557,849,871]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="10" pageNumber="283" phylum="Arthropoda" rank="species" species="bemanevika" status="sp. nov.">
<emphasis id="B922F168FFED14298878D308E255FCFE" box="[283,557,849,871]" italics="true" pageId="10" pageNumber="283">Zoosphaerium bemanevika</emphasis>
</taxonomicName>
<emphasis id="B922F168FFED14298B57D308E216FCFE" bold="true" box="[564,622,849,871]" pageId="10" pageNumber="283">
<taxonomicNameLabel id="A2114C13FFED14298B57D308E216FCFE" box="[564,622,849,871]" pageId="10" pageNumber="283" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype (FMNH-INS 2858686-C), endotergum of mid-body tergite, SEM,
<emphasis id="B922F168FFED14298CE0D308E5EDFCFE" bold="true" box="[1411,1429,849,871]" pageId="10" pageNumber="283">A</emphasis>
: ventral view;
<emphasis id="B922F168FFED14298841D329E14BFC1F" bold="true" box="[290,307,880,902]" pageId="10" pageNumber="283">B</emphasis>
: bristles on apical margin.
</paragraph>
</caption>
<paragraph id="8BE92D7AFFED142989A4D3EDE16EFC76" blockId="10.[151,1436,946,2015]" pageId="10" pageNumber="283">
<emphasis id="B922F168FFED142989A4D3EDE165FC52" box="[199,285,948,971]" italics="true" pageId="10" pageNumber="283">Tergites</emphasis>
: surface glabrous and shiny, lacking small pits. Tips of paratergites of midbody tergites projecting posteriorly.
</paragraph>
<paragraph id="8BE92D7AFFED142989A4D3A3E12AFBE6" blockId="10.[151,1436,946,2015]" pageId="10" pageNumber="283">
<emphasis id="B922F168FFED142989A4D3A3E128FB8A" box="[199,336,1018,1043]" italics="true" pageId="10" pageNumber="283">Endotergum</emphasis>
: inner section with numerous short triangular spines and very few longer setae. Middle area above spines with single sparse row of circular cuticular impressions slightly varying in size. Apically single row of short marginal bristles, protruding to half distance of tergite margin (
<figureCitation id="136D31FFFFED14298A38D41BE3E6FBC2" box="[859,926,1090,1115]" captionStart="FIGURE 7" captionStartId="10.[151,250,849,871]" captionTargetBox="[151,1435,194,824]" captionTargetId="figure@10.[151,1435,193,828]" captionTargetPageId="10" captionText="FIGURE 7. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), endotergum of mid-body tergite, SEM, A: ventral view; B: bristles on apical margin." httpUri="https://zenodo.org/record/584051/files/figure.png" pageId="10" pageNumber="283">Fig. 7</figureCitation>
A). Bristles not smooth, with numerous small spines (
<figureCitation id="136D31FFFFED1429898FD43FE157FBE6" box="[236,303,1126,1151]" captionStart="FIGURE 7" captionStartId="10.[151,250,849,871]" captionTargetBox="[151,1435,194,824]" captionTargetId="figure@10.[151,1435,193,828]" captionTargetPageId="10" captionText="FIGURE 7. Zoosphaerium bemanevika n. sp., holotype (FMNH-INS 2858686 - C), endotergum of mid-body tergite, SEM, A: ventral view; B: bristles on apical margin." httpUri="https://zenodo.org/record/584051/files/figure.png" pageId="10" pageNumber="283">Fig. 7</figureCitation>
B).
</paragraph>
<paragraph id="8BE92D7AFFED142989A4D4D3E4BEFB72" blockId="10.[151,1436,946,2015]" pageId="10" pageNumber="283">
<emphasis id="B922F168FFED142989A4D4D3E131FB3A" box="[199,329,1162,1187]" italics="true" pageId="10" pageNumber="283">Anal Shield</emphasis>
: large and well rounded. Female anal shield with slightly elongated tip (
<figureCitation id="136D31FFFFED14298D1FD4D3E4A8FB3A" box="[1148,1232,1162,1187]" captionStart="FIGURE 12" captionStartId="16.[151,250,1864,1886]" captionTargetBox="[163,1436,1100,1835]" captionTargetId="figure@16.[150,1436,1094,1842]" captionTargetPageId="16" captionText="FIGURE 12. Maximum likelihood tree inferred from the COI dataset with 1000 bootstrap pseudoreplicates implementing the GTR + I + G model. Habitus photograph shows Z. bemanevika (FMNH-INS 3196510). Colors used to separate species. Green = Z. bemanevika; Orange = Z. minutus. Circles indicating sister group relationships, as mapped in figure 1." httpUri="https://zenodo.org/record/584057/files/figure.png" pageId="10" pageNumber="283">Fig. 12</figureCitation>
). No hair on male anal shield but with numerous small pits. Female anal shield covered with hair. Underside carries two black locking carinae, posterior one 4 times longer than anterior one, both located close to anal shield margin.
</paragraph>
<paragraph id="8BE92D7AFFED142989A4D4A1E0B0FAE2" blockId="10.[151,1436,946,2015]" pageId="10" pageNumber="283">
<emphasis id="B922F168FFED142989A4D4A1E084FA96" box="[199,252,1272,1295]" italics="true" pageId="10" pageNumber="283">Legs</emphasis>
: leg 1 with 4 or 5 ventral spines, leg 2 with 5–7, leg 3 with 8–11. First two leg pairs without an apical spine. Leg pairs 4–21 with 9–12 ventral spines and an apical spine. In leg 9 femur 1.8, tarsus 5.1 times longer than wide. All podomeres with setae. Coxa with field of tiny spines laterally close to articulation with prefemur (
<figureCitation id="136D31FFFFED14298C0CD567E0DFFAE2" captionStart="FIGURE 2" captionStartId="6.[151,250,1801,1823]" captionTargetBox="[168,1414,336,1783]" captionTargetId="figure@6.[168,1414,336,1783]" captionTargetPageId="6" captionText="FIGURE 2. Zoosphaerium bemanevika n. sp.., A, B: female (ZFMK MYR 6144); C-H: holotype male (FMNH-INS 2858686 - C); A: coxa of first left leg with first stigmatic plate; B: 9 th right leg; C coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view G: right posterior telopod, posterior view; H: right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-p = sclerotized spot; s-ps = field of sclerotized spots; sr = stridulation ribs; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. A, B Scale bars = 1 mm, C – H Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/584046/files/figure.png" pageId="10" pageNumber="283">Fig. 2</figureCitation>
B).
</paragraph>
<paragraph id="8BE92D7AFFED142989A4D5DFE1C5FA5A" blockId="10.[151,1436,946,2015]" pageId="10" pageNumber="283">
<emphasis id="B922F168FFED142989A4D5DFE1A7FA06" box="[199,479,1414,1439]" italics="true" pageId="10" pageNumber="283">Male sexual characters:</emphasis>
gonopore covered with a single undivided, rounded membranous plate. No setae located on plate (
<figureCitation id="136D31FFFFED14298834D5F3E1E3FA5A" box="[343,411,1450,1475]" captionStart="FIGURE 2" captionStartId="6.[151,250,1801,1823]" captionTargetBox="[168,1414,336,1783]" captionTargetId="figure@6.[168,1414,336,1783]" captionTargetPageId="6" captionText="FIGURE 2. Zoosphaerium bemanevika n. sp.., A, B: female (ZFMK MYR 6144); C-H: holotype male (FMNH-INS 2858686 - C); A: coxa of first left leg with first stigmatic plate; B: 9 th right leg; C coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view G: right posterior telopod, posterior view; H: right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-p = sclerotized spot; s-ps = field of sclerotized spots; sr = stridulation ribs; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. A, B Scale bars = 1 mm, C – H Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/584046/files/figure.png" pageId="10" pageNumber="283">Fig. 2</figureCitation>
C).
</paragraph>
<paragraph id="8BE92D7AFFED142989A4D596E474F926" blockId="10.[151,1436,946,2015]" pageId="10" pageNumber="283">
<emphasis id="B922F168FFED142989A4D596E1F4FA7E" box="[199,396,1486,1511]" italics="true" pageId="10" pageNumber="283">Anterior telopods</emphasis>
: harp with two stridulation ribs—one long and well pronounced rib located mesally of a less pronounced and shorter lateral rib (
<figureCitation id="136D31FFFFED14298B41D5ABE21EF992" box="[546,614,1522,1547]" captionStart="FIGURE 2" captionStartId="6.[151,250,1801,1823]" captionTargetBox="[168,1414,336,1783]" captionTargetId="figure@6.[168,1414,336,1783]" captionTargetPageId="6" captionText="FIGURE 2. Zoosphaerium bemanevika n. sp.., A, B: female (ZFMK MYR 6144); C-H: holotype male (FMNH-INS 2858686 - C); A: coxa of first left leg with first stigmatic plate; B: 9 th right leg; C coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view G: right posterior telopod, posterior view; H: right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-p = sclerotized spot; s-ps = field of sclerotized spots; sr = stridulation ribs; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. A, B Scale bars = 1 mm, C – H Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/584046/files/figure.png" pageId="10" pageNumber="283">Fig. 2</figureCitation>
D). First podomere with only few setae restricted to apical parts of lateral margins. Process of second podomere nearly as long as third podomere, laterally visible in anterior view, and with an apical mound of few sclerotized spots juxtaposed to third podomere. Third podomere with an apical brown sclerotized spot and few elevated sclerotized spots at inner margin below brown spot. Two small spines located below sclerotized spot (
<figureCitation id="136D31FFFFED142988CAD6DBE197F902" box="[425,495,1666,1691]" captionStart="FIGURE 2" captionStartId="6.[151,250,1801,1823]" captionTargetBox="[168,1414,336,1783]" captionTargetId="figure@6.[168,1414,336,1783]" captionTargetPageId="6" captionText="FIGURE 2. Zoosphaerium bemanevika n. sp.., A, B: female (ZFMK MYR 6144); C-H: holotype male (FMNH-INS 2858686 - C); A: coxa of first left leg with first stigmatic plate; B: 9 th right leg; C coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view G: right posterior telopod, posterior view; H: right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-p = sclerotized spot; s-ps = field of sclerotized spots; sr = stridulation ribs; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. A, B Scale bars = 1 mm, C – H Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/584046/files/figure.png" pageId="10" pageNumber="283">Fig. 2</figureCitation>
E). Depression lateral to field of spots with one small sclerotized spine (
<figureCitation id="136D31FFFFED14298C54D6DBE505F902" box="[1335,1405,1666,1691]" captionStart="FIGURE 2" captionStartId="6.[151,250,1801,1823]" captionTargetBox="[168,1414,336,1783]" captionTargetId="figure@6.[168,1414,336,1783]" captionTargetPageId="6" captionText="FIGURE 2. Zoosphaerium bemanevika n. sp.., A, B: female (ZFMK MYR 6144); C-H: holotype male (FMNH-INS 2858686 - C); A: coxa of first left leg with first stigmatic plate; B: 9 th right leg; C coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view G: right posterior telopod, posterior view; H: right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-p = sclerotized spot; s-ps = field of sclerotized spots; sr = stridulation ribs; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. A, B Scale bars = 1 mm, C – H Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/584046/files/figure.png" pageId="10" pageNumber="283">Fig. 2</figureCitation>
F). Second and third podomere with setae mainly apically, no hair at inner margin.
</paragraph>
<paragraph id="8BE92D7AFFED142989A4D695E395F846" blockId="10.[151,1436,946,2015]" pageId="10" pageNumber="283">
<emphasis id="B922F168FFED142989A4D695E1E7F97A" box="[199,415,1738,1763]" italics="true" pageId="10" pageNumber="283">Posterior telopods</emphasis>
: movable finger (podomere 3) 2.6 times longer than wide, weakly curved towards immovable finger (process of podomere 2). Shallowly hollowed out inner margin with two sclerotized spines, one positioned apically and one in middle of cavity. Additional sclerotized spine at base of movable finger. Posterior aspect with 7 or 8 small, crenulated teeth (
<figureCitation id="136D31FFFFED14298BF1D76FE2A2F8D6" box="[658,730,1846,1871]" captionStart="FIGURE 2" captionStartId="6.[151,250,1801,1823]" captionTargetBox="[168,1414,336,1783]" captionTargetId="figure@6.[168,1414,336,1783]" captionTargetPageId="6" captionText="FIGURE 2. Zoosphaerium bemanevika n. sp.., A, B: female (ZFMK MYR 6144); C-H: holotype male (FMNH-INS 2858686 - C); A: coxa of first left leg with first stigmatic plate; B: 9 th right leg; C coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view G: right posterior telopod, posterior view; H: right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-p = sclerotized spot; s-ps = field of sclerotized spots; sr = stridulation ribs; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. A, B Scale bars = 1 mm, C – H Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/584046/files/figure.png" pageId="10" pageNumber="283">Fig. 2</figureCitation>
G). Immovable finger almost as long as and curved towards movable finger. Sclerotized spine located on anterior side near base of immovable finger. Inner part of fixed finger apically covered with few sclerotized spots (
<figureCitation id="136D31FFFFED14298BE4D727E2B2F80E" box="[647,714,1918,1943]" captionStart="FIGURE 2" captionStartId="6.[151,250,1801,1823]" captionTargetBox="[168,1414,336,1783]" captionTargetId="figure@6.[168,1414,336,1783]" captionTargetPageId="6" captionText="FIGURE 2. Zoosphaerium bemanevika n. sp.., A, B: female (ZFMK MYR 6144); C-H: holotype male (FMNH-INS 2858686 - C); A: coxa of first left leg with first stigmatic plate; B: 9 th right leg; C coxa of second left leg with gonopore; D: left anterior telopod, anterior view; E: left anterior telopod, posterior view; F: left anterior telopod, lateral view G: right posterior telopod, posterior view; H: right posterior telopod, anterior view. Abbreviations: as = apical spine; cl = claw; cr-t = crenulated teeth; cx = coxa; fe = femur; gp = gonopore; pof = postfemur; pref = prefemur; s-p = sclerotized spot; s-ps = field of sclerotized spots; sr = stridulation ribs; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; vs = ventral spines. A, B Scale bars = 1 mm, C – H Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/584046/files/figure.png" pageId="10" pageNumber="283">Fig. 2</figureCitation>
H). Movable finger completely glabrous on both sides except for a few hairs baso-laterally. Second podomere with few hairs baso-laterally and on central part of fixed finger, remaining parts of podomere glabrous. First podomere completely glabrous.
</paragraph>
<paragraph id="8BE92D7AFFEC142889A4D0CEE1E6FED9" blockId="11.[151,1437,151,429]" pageId="11" pageNumber="284">
<emphasis id="B922F168FFEC142889A4D0CEE192FF29" box="[199,490,151,176]" italics="true" pageId="11" pageNumber="284">Female sexual characters</emphasis>
: vulva large, covering 3/4 of coxa, not extending to prefemur. Operculum rounded, medially invaginated, apical margin extended into two well-rounded lobes. Inner mesal plate long and slender and extending to apex of coxa and operculum. Lateral margin covered by hair. External mesal plate broader and only extending to base of operculum, lateral margins also covered by hair. Suture between inner and external plate membranous (
<figureCitation id="136D31FFFFEC14288854D17EE101FED9" box="[311,377,295,320]" captionStart="FIGURE 3" captionStartId="7.[151,250,700,722]" captionTargetBox="[175,1398,193,689]" captionTargetId="figure@7.[175,1398,193,689]" captionTargetPageId="7" captionText="FIGURE 3. Zoosphaerium bemanevika n. sp., female (ZFMK MYR 6144), A: coxa of second left leg with vulva; B: female subanal plate with washboard. Abbreviations: cx = coxa; ep = external plate of vulva; ip = internal plate of vulva; o = operculum; sr = stridulation ribs. Scale bars = 1 mm." httpUri="https://zenodo.org/record/584047/files/figure.png" pageId="11" pageNumber="284">Fig. 3</figureCitation>
A).
</paragraph>
<paragraph id="8BE92D7AFFEC142889A4D115E1EFFE35" blockId="11.[151,1437,151,429]" pageId="11" pageNumber="284">
<emphasis id="B922F168FFEC142889A4D115E111FEFC" box="[199,361,332,357]" italics="true" pageId="11" pageNumber="284">Subanal plate</emphasis>
with shallow invagination at apical margin. Washboard with 2 short but well-developed stridulation ribs on each side. Margins and median part densely covered with hair, lateral parts with only few shorter hairs (
<figureCitation id="136D31FFFFEC14288852D1CDE10CFE35" box="[305,372,404,429]" captionStart="FIGURE 3" captionStartId="7.[151,250,700,722]" captionTargetBox="[175,1398,193,689]" captionTargetId="figure@7.[175,1398,193,689]" captionTargetPageId="7" captionText="FIGURE 3. Zoosphaerium bemanevika n. sp., female (ZFMK MYR 6144), A: coxa of second left leg with vulva; B: female subanal plate with washboard. Abbreviations: cx = coxa; ep = external plate of vulva; ip = internal plate of vulva; o = operculum; sr = stridulation ribs. Scale bars = 1 mm." httpUri="https://zenodo.org/record/584047/files/figure.png" pageId="11" pageNumber="284">Fig. 3</figureCitation>
B).
</paragraph>
</subSubSection>
<subSubSection id="C34C7EF1FFEC142889F4D185E3B0FD81" pageId="11" pageNumber="284" type="distribution">
<paragraph id="8BE92D7AFFEC142889F4D185E3B0FD81" blockId="11.[151,1436,476,536]" pageId="11" pageNumber="284">
<emphasis id="B922F168FFEC142889F4D185E1BAFE6C" bold="true" box="[151,450,476,501]" pageId="11" pageNumber="284">Distribution and ecology.</emphasis>
<taxonomicName id="4C5656F9FFEC142888AAD185E28FFE6C" box="[457,759,476,501]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="284" phylum="Arthropoda" rank="species" species="bemanevika">
<emphasis id="B922F168FFEC142888AAD185E28FFE6C" box="[457,759,476,501]" italics="true" pageId="11" pageNumber="284">Zoosphaerium bemanevika</emphasis>
</taxonomicName>
is currently only known from two collection sites in patches of mountainous rainforest (
<figureCitation id="136D31FFFFEC142888A4D1A6E260FD81" box="[455,536,511,536]" captionStart="FIGURE 13" captionStartId="17.[151,250,1820,1842]" captionTargetBox="[150,1435,224,1798]" captionTargetId="figure@17.[150,1436,193,1799]" captionTargetPageId="17" captionText="FIGURE 13. Habitat of Zoosphaerium bemanevika n. sp. and Zoosphaerium minutus n. sp. in patches of mountainous rainforest in the Paysage Harmonieux Protégé de Bemanevika. Surroundings of remaining rainforest are prone to habitat destruction and deforestation. A: shows the grazing of Zebu cattle in the surrounding pseudosteppe; B: indicating the destruction of forest edges due to the yearly burning of the pseudosteppe. Each fire lets the forest fragments shrink a few meters in diameter. Photos are the courtesy of Achille P. Raselimanana." httpUri="https://zenodo.org/record/584058/files/figure.png" pageId="11" pageNumber="284">Fig. 13</figureCitation>
) in
<taxonomicName id="4C5656F9FFEC14288B26D1A6E2ACFD81" box="[581,724,511,536]" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" kingdom="Animalia" order="Sphaerotheriida" pageId="11" pageNumber="291" phylum="Arthropoda" rank="species" species="bemanevika" status="sp. nov.">Bemanevika</taxonomicName>
,
<collectingCountry id="F3416DEAFFEC14288BBCD1A6E31EFD81" box="[735,870,511,536]" name="Madagascar" pageId="11" pageNumber="284">Madagascar</collectingCountry>
(
<figureCitation id="136D31FFFFEC14288A15D1A6E3CFFD81" box="[886,951,511,536]" captionStart="FIGURE 1" captionStartId="5.[151,250,1538,1560]" captionTargetBox="[209,1377,432,1508]" captionTargetId="figure@5.[199,1387,427,1517]" captionTargetPageId="5" captionText="FIGURE 1. Distribution map of Zoosphaerium species in Madagascar, including Zoosphaerium bemanevika n. sp., and Zoosphaerium minutus n. sp. Red points refer to other Zoosphaerium species (See Table 1). Circles refer to clades (see Figure 12). Vegetation mapped after literature (DuPuy &amp; Moat 2007)." httpUri="https://zenodo.org/record/584045/files/figure.png" pageId="11" pageNumber="284">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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