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<mods:title id="431E81B5813662DB9AB21E89ACC72282">Integrative taxonomy study of brackish water crabs of the genus Ptychognathus Stimpson, 1858 (Crustacea: Brachyura: Varunidae) from Polynesia, with description of two new species</mods:title>
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<emphasis id="B922EAF7E30BFFF61CBEFA0CFE4AF9C8" bold="true" box="[151,491,1534,1561]" italics="true" pageId="5" pageNumber="272">Ptychognathus crassimanus</emphasis>
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<treatmentCitation id="0AF710F4E30BFFF61DF9F972FDDBF949" author="Poupin, J." box="[464,634,1664,1688]" page="69" pageId="5" pageNumber="272" year="1996">
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<bibRefCitation id="EFC74B14E30BFFF61EADF972FD7FF948" author="Poupin, J." box="[644,734,1664,1689]" pageId="5" pageNumber="272" refId="ref19132" refString="Poupin, J. (2005) Systematique et ecologie des Crustaces Decapodes et Stomatopodes de Polynesie Francaise. Ecologie, Environnement. Universite de Perpignan, 115 pp. [https: // tel. archives-ouvertes. fr / tel- 00085049]" type="book" year="2005">2005: 28</bibRefCitation>
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.—
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only).—
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Salvat
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2016: 244
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[
<collectingRegion id="4992F807E30BFFF61D73F94CFE6CF907" box="[346,461,1726,1750]" country="French Polynesia" name="Iles Marquises" pageId="5" pageNumber="272">Marquesas</collectingRegion>
].—All not
<taxonomicName id="4C564D66E30BFFF61E69F94DFCC3F907" ID-CoL="6WNM3" authority="Rathbun, 1907" authorityName="Rathbun" authorityYear="1907" box="[576,866,1726,1750]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="272" phylum="Arthropoda" rank="species" species="easteranus">
<emphasis id="B922EAF7E30BFFF61E69F94DFD67F907" box="[576,710,1727,1750]" italics="true" pageId="5" pageNumber="272">P. easteranus</emphasis>
<bibRefCitation id="EFC74B14E30BFFF61EE4F94CFCC3F907" author="Rathbun, M. J." box="[717,866,1726,1750]" pageId="5" pageNumber="272" pagination="23 - 74" refId="ref19375" refString="Rathbun, M. J. (1907) Reports on the scientific results of the expedition to the Tropical Pacific, in charge of Alexander Agassiz, by the U. S. Fish Commission steamer Albatross, from August, 1899, to March, 1900, Commander Jefferson F. Moser, U. S. N., commanding - IX. ibidem. from October, 1904, to March 1905, lieut. - commander L. M. Garrett, U. S. N., commanding - X. The Brachyura. Memoirs of the Museum of Comparative Zoology at Harvard College, 35 (2), 23 - 74." type="journal article" year="1907">Rathbun, 1907</bibRefCitation>
</taxonomicName>
but
<taxonomicName id="4C564D66E30BFFF61FA6F94DFB6FF907" ID-CoL="6WNLP" authority="Finnegan, 1931" authorityName="Finnegan" authorityYear="1931" box="[911,1230,1726,1750]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="272" phylum="Arthropoda" rank="species" species="crassimanus">
<emphasis id="B922EAF7E30BFFF61FA6F94DFB86F907" box="[911,1063,1727,1750]" italics="true" pageId="5" pageNumber="272">P. crassimanus</emphasis>
<bibRefCitation id="EFC74B14E30BFFF61804F94CFB6FF907" author="Finnegan, S." box="[1069,1230,1726,1750]" pageId="5" pageNumber="272" pagination="607 - 673" refId="ref17489" refString="Finnegan, S. (1931) Report on the brachyura collected in Central America, the Gorgona and Galapagos islands, by Dr. Crossland on the St George Expedition to the Pacific, 1924 - 25. Linnean Society's Journal, Zoology, 37 (255), 607 - 673. https: // doi. org / 10.1111 / j. 1096 - 3642.1931. tb 02367. x" type="journal article" year="1931">Finnegan, 1931</bibRefCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34C656EE30BFFF51CBEF8F6FC2EFE58" lastPageId="6" lastPageNumber="273" pageId="5" pageNumber="272" type="materials_examined">
<paragraph id="8BE936E5E30BFFF61CBEF8F6FA3DF827" blockId="5.[151,1437,1796,2039]" pageId="5" pageNumber="272">
<emphasis id="B922EAF7E30BFFF61CBEF8F6FEDEF8CF" bold="true" box="[151,383,1796,1822]" pageId="5" pageNumber="272">Material examined.</emphasis>
<materialsCitation id="3B3E3CB8E30BFFF61DADF8F6FC8CF8B7" ID-GBIF-Occurrence="4905160314" collectingDate="2012-01-17" collectingDateMax="2012-01-18" collectingDateMin="2012-01-17" collectionCode="TE, MOANA" country="French Polynesia" county="Fatu Hiva" latitude="-10.464" location="Stn" longLatPrecision="13" longitude="-138.66617" municipality="Baie des Vierges" pageId="5" pageNumber="272" specimenCode="CA2646" specimenCount="1" specimenCount-female="1" stateProvince="Iles Marquises">
<emphasis id="B922EAF7E30BFFF61DADF8F6FC49F8CF" box="[388,1000,1796,1822]" italics="true" pageId="5" pageNumber="272">
<collectingCountry id="F3417675E30BFFF61DADF8F6FD28F8CF" box="[388,649,1796,1822]" name="French Polynesia" pageId="5" pageNumber="272">FRENCH POLYNESIA</collectingCountry>
.
<collectingRegion id="4992F807E30BFFF61EBBF8F7FCC7F8CF" box="[658,870,1796,1822]" country="French Polynesia" name="Iles Marquises" pageId="5" pageNumber="272">Marquesas Islands</collectingRegion>
.
<collectingCounty id="62884E69E30BFFF61F46F8F7FC43F8CF" box="[879,994,1797,1822]" pageId="5" pageNumber="272">Fatu Hiva</collectingCounty>
:
</emphasis>
MNHN-IU-2011-9062 (DNA voucher:
<specimenCode id="DBF09E9EE30BFFF61CBEF8DAFF5AF893" box="[151,251,1832,1858]" collectionCode="CA" country="USA" name="Chicago Academy of Sciences" pageId="5" pageNumber="272" type="Museum">CA2646</specimenCode>
),
<specimenCount id="9D50FD6CE30BFFF61D22F8DAFE92F892" box="[267,307,1832,1859]" count="1" pageId="5" pageNumber="272" type="female">1 ♀</specimenCount>
, 14.0 mm ×
<quantity id="4CAE9B00E30BFFF61D90F8DAFDBDF892" box="[441,540,1832,1859]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="5" pageNumber="272" unit="mm" value="11.8">11.8 mm</quantity>
,
<collectingMunicipality id="6B8DAC9FE30BFFF61E0CF8DAFD7EF893" box="[549,735,1832,1858]" pageId="5" pageNumber="272">Baie des Vierges</collectingMunicipality>
,
<location id="8E89603EE30BFFF61ECEF8DAFCADF893" LSID="urn:lsid:plazi:treatment:03FF87F3E30BFFFB1CBEFA0CFDD0F859:8E89603EE30BFFF61ECEF8DAFCADF893" box="[743,780,1832,1858]" country="French Polynesia" county="Fatu Hiva" latitude="-10.464" longLatPrecision="13" longitude="-138.66617" municipality="Baie des Vierges" name="Stn" pageId="5" pageNumber="272" stateProvince="Iles Marquises">Stn</location>
MQ16-M,
<geoCoordinate id="EE625022E30BFFF61FA0F8DAFBA7F893" box="[905,1030,1832,1858]" degrees="10" direction="south" minutes="27.84" orientation="latitude" pageId="5" pageNumber="272" precision="9" value="-10.464">10°27.84'S</geoCoordinate>
,
<geoCoordinate id="EE625022E30BFFF61826F8DAFB02F893" box="[1039,1187,1832,1858]" degrees="138" direction="west" minutes="39.97" orientation="longitude" pageId="5" pageNumber="272" precision="9" value="-138.66617">138°39.97'W</geoCoordinate>
,
<location id="8E89603EE30BFFF61885F8DAFA86F893" LSID="urn:lsid:plazi:treatment:03FF87F3E30BFFFB1CBEFA0CFDD0F859:8E89603EE30BFFF61885F8DAFA86F893" box="[1196,1319,1832,1858]" country="French Polynesia" county="Fatu Hiva" latitude="-10.464" longLatPrecision="13" longitude="-138.66617" municipality="Baie des Vierges" name="Expedition" pageId="5" pageNumber="272" stateProvince="Iles Marquises">Expedition</location>
PAKAIHI I
<collectionCode id="ED47AE20E30BFFF61C80F8BFFEE1F8B7" box="[169,320,1868,1895]" pageId="5" pageNumber="272">TE MOANA</collectionCode>
, coll. MNHN-AAMP,
<date id="FFE81025E30BFFF61E6AF8BEFC8CF8B7" box="[579,813,1868,1895]" pageId="5" pageNumber="272" value="2012-01-17" valueMax="2012-01-18" valueMin="2012-01-17">
<collectingDate id="EFACE9CDE30BFFF61E6AF8BEFC8CF8B7" box="[579,813,1868,1895]" pageId="5" pageNumber="272" value="2012-01-17" valueMax="2012-01-18" valueMin="2012-01-17">1718 January 2012</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E3CB8E30BFFF61F10F8BEFA95F87F" ID-GBIF-Occurrence="4905160353" collectingDate="2000-02-13" collectingDateMax="2012-01-18" collectingDateMin="2012-01-17" collectionCode="CA" country="French Polynesia" county="Fatu Hiva" latitude="-10.464" location="Ta'aoa River" longLatPrecision="13" longitude="-138.66617" municipality="Hiva Oa" pageId="5" pageNumber="272" specimenCode="CA2647, CA2668" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Marquesas Islands">
MNHN-IU-2022-4082 (DNA voucher:
<specimenCode id="DBF09E9EE30BFFF618D0F8BEFAFDF8B7" box="[1273,1372,1868,1894]" collectionCode="CA" country="USA" name="Chicago Academy of Sciences" pageId="5" pageNumber="272" type="Museum">CA2647</specimenCode>
),
<specimenCount id="9D50FD6CE30BFFF61958F8BEFA3DF8B6" box="[1393,1436,1868,1895]" count="1" pageId="5" pageNumber="272" type="female">1 ♀</specimenCount>
ovig.,
<quantity id="4CAE9B00E30BFFF61CC8F882FEE9F85A" box="[225,328,1904,1931]" metricMagnitude="-2" metricUnit="m" metricValue="1.3699999999999999" pageId="5" pageNumber="272" unit="mm" value="13.7">13.7 mm</quantity>
×
<quantity id="4CAE9B00E30BFFF61D45F882FE74F85A" box="[364,469,1904,1931]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="5" pageNumber="272" unit="mm" value="12.4">12.4 mm</quantity>
, same collection data as previous.
<emphasis id="B922EAF7E30BFFF61F45F883FC79F85B" box="[876,984,1904,1930]" italics="true" pageId="5" pageNumber="272">
<collectingMunicipality id="6B8DAC9FE30BFFF61F45F883FC73F85B" box="[876,978,1904,1930]" pageId="5" pageNumber="272">Hiva Oa</collectingMunicipality>
:
</emphasis>
MNHN-IU-2022-4083 (DNA voucher:
<specimenCode id="DBF09E9EE30BFFF61CBEF866FF5BF87F" box="[151,250,1940,1966]" collectionCode="CA" country="USA" name="Chicago Academy of Sciences" pageId="5" pageNumber="272" type="Museum">CA2668</specimenCode>
),
<specimenCount id="9D50FD6CE30BFFF61D22F866FE94F87E" box="[267,309,1940,1967]" count="1" pageId="5" pageNumber="272" type="male">1 ♂</specimenCount>
, 13.0 mm ×
<quantity id="4CAE9B00E30BFFF61D93F866FDBFF87E" box="[442,542,1940,1967]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="5" pageNumber="272" unit="mm" value="12.5">12.5 mm</quantity>
,
<location id="8E89603EE30BFFF61E0EF866FD15F87F" LSID="urn:lsid:plazi:treatment:03FF87F3E30BFFFB1CBEFA0CFDD0F859:8E89603EE30BFFF61E0EF866FD15F87F" box="[551,692,1940,1966]" country="French Polynesia" municipality="Hiva Oa" name="Ta'aoa River" pageId="5" pageNumber="272" stateProvince="Marquesas Islands">Ta'aoa River</location>
, coll.
<collectorName id="26A35333E30BFFF61EDAF867FCDAF87E" box="[755,891,1941,1967]" pageId="5" pageNumber="272">E. Feunteun</collectorName>
,
<collectorName id="26A35333E30BFFF61FADF867FC7CF87F" box="[900,989,1940,1967]" pageId="5" pageNumber="272">P. Keith</collectorName>
,
<collectorName id="26A35333E30BFFF61FCFF867FBC4F87F" box="[998,1125,1940,1967]" pageId="5" pageNumber="272">E. Vigneux</collectorName>
,
<date id="FFE81025E30BFFF61847F866FA95F87F" box="[1134,1332,1940,1966]" pageId="5" pageNumber="272" value="2000-02-13">
<collectingDate id="EFACE9CDE30BFFF61847F866FA95F87F" box="[1134,1332,1940,1966]" pageId="5" pageNumber="272" value="2000-02-13">13 February 2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E3CB8E30BFFF61914F867FE6EF827" ID-GBIF-Occurrence="4905160303" collectingDate="2000-02-13" collectionCode="MNHN-" country="French Polynesia" location="Ta'aoa River" pageId="5" pageNumber="272" specimenCode="CA2669" specimenCount="1" specimenCount-male="1" stateProvince="Marquesas Islands">
<collectionCode id="ED47AE20E30BFFF61914F867FA3DF87E" box="[1341,1436,1941,1967]" pageId="5" pageNumber="272">MNHNIU-</collectionCode>
2022-4084 (DNA voucher:
<specimenCode id="DBF09E9EE30BFFF61DDFF84AFDFBF803" box="[502,602,1976,2002]" collectionCode="CA" country="USA" name="Chicago Academy of Sciences" pageId="5" pageNumber="272" type="Museum">CA2669</specimenCode>
),
<specimenCount id="9D50FD6CE30BFFF61E44F84AFD3AF802" box="[621,667,1976,2003]" count="1" pageId="5" pageNumber="272" type="male">1 ♂</specimenCount>
, 14.0 mm ×
<quantity id="4CAE9B00E30BFFF61F05F84AFC32F802" box="[812,915,1976,2003]" metricMagnitude="-2" metricUnit="m" metricValue="1.3199999999999998" pageId="5" pageNumber="272" unit="mm" value="13.2">13.2 mm</quantity>
,
<location id="8E89603EE30BFFF61F89F84AFB8EF803" LSID="urn:lsid:plazi:treatment:03FF87F3E30BFFFB1CBEFA0CFDD0F859:8E89603EE30BFFF61F89F84AFB8EF803" box="[928,1071,1976,2002]" country="French Polynesia" name="Ta'aoa River" pageId="5" pageNumber="272" stateProvince="Marquesas Islands">Ta'aoa River</location>
,, coll.
<collectorName id="26A35333E30BFFF618ADF84BFAAEF802" box="[1156,1295,1977,2003]" pageId="5" pageNumber="272">E. Feunteun</collectorName>
,
<collectorName id="26A35333E30BFFF61932F84BFAD9F803" box="[1307,1400,1976,2003]" pageId="5" pageNumber="272">P. Keith</collectorName>
,
<collectorName id="26A35333E30BFFF619ADF84BFF58F827" pageId="5" pageNumber="272">E. Vigneux</collectorName>
,
<date id="FFE81025E30BFFF61D2DF82EFE6EF827" box="[260,463,2012,2038]" pageId="5" pageNumber="272" value="2000-02-13">
<collectingDate id="EFACE9CDE30BFFF61D2DF82EFE6EF827" box="[260,463,2012,2038]" pageId="5" pageNumber="272" value="2000-02-13">13 February 2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E3CB8E30BFFF61DF0F82EFA39F827" ID-GBIF-Occurrence="4905160367" box="[473,1432,2012,2039]" collectionCode="CA" country="French Polynesia" location="Ta'aoa River" pageId="5" pageNumber="272" specimenCode="CA2465" specimenCount="1" specimenCount-male="1" stateProvince="Marquesas Islands">
MNHN-IU-2019-180 (DNA voucher:
<specimenCode id="DBF09E9EE30BFFF61FAFF82EFC48F826" box="[902,1001,2012,2039]" collectionCode="CA" country="USA" name="Chicago Academy of Sciences" pageId="5" pageNumber="272" type="Museum">CA2465</specimenCode>
),
<specimenCount id="9D50FD6CE30BFFF61FD5F82EFB89F826" box="[1020,1064,2012,2039]" count="1" pageId="5" pageNumber="272" type="male">1 ♂</specimenCount>
,
<quantity id="4CAE9B00E30BFFF6181DF82EFB2BF826" box="[1076,1162,2012,2039]" metricMagnitude="-3" metricUnit="m" metricValue="9.4" pageId="5" pageNumber="272" unit="mm" value="9.4">9.4 mm</quantity>
×
<quantity id="4CAE9B00E30BFFF6188EF82EFB5EF826" box="[1191,1279,2012,2039]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="5" pageNumber="272" unit="mm" value="8.4">8.4 mm</quantity>
,
<location id="8E89603EE30BFFF61923F82EFA39F827" LSID="urn:lsid:plazi:treatment:03FF87F3E30BFFFB1CBEFA0CFDD0F859:8E89603EE30BFFF61923F82EFA39F827" box="[1290,1432,2012,2038]" country="French Polynesia" name="Ta'aoa River" pageId="5" pageNumber="272" stateProvince="Marquesas Islands">Ta'aoa River</location>
</materialsCitation>
,
</paragraph>
<paragraph id="8BE936E5E308FFF51CBEFF64FC2EFE58" blockId="6.[151,1437,150,1977]" pageId="6" pageNumber="273">
<geoCoordinate id="EE625022E308FFF51CBEFF64FEB4FF60" box="[151,277,150,177]" degrees="9" direction="south" minutes="50.101" orientation="latitude" pageId="6" pageNumber="273" precision="1" value="-9.835016">9°50.101'S</geoCoordinate>
,
<geoCoordinate id="EE625022E308FFF51D09FF64FE60FF60" box="[288,449,150,177]" degrees="139" direction="west" minutes="03.725" orientation="longitude" pageId="6" pageNumber="273" precision="1" value="-139.06209">139°03.725'W</geoCoordinate>
, coll. E. Feunteun, P. Gerbeaux, G. Marquet, P. Tiberghien,
<date id="FFE81025E308FFF51872FF65FAAAFF60" box="[1115,1291,151,177]" pageId="6" pageNumber="273" value="2017-10-09">9 October 2017</date>
;
<materialsCitation id="3B3E3CB8E308FFF5193FFF65FC5EFF04" ID-GBIF-Occurrence="4905160354" collectionCode="CA" location="Faakuaa River" pageId="6" pageNumber="273" specimenCode="CA2464" specimenCount="1" specimenCount-male="1">
MNHN-IU-2019-183 (DNA voucher:
<specimenCode id="DBF09E9EE308FFF51D94FF49FD81FF04" box="[445,544,187,213]" collectionCode="CA" country="USA" name="Chicago Academy of Sciences" pageId="6" pageNumber="273" type="Museum">CA2464</specimenCode>
),
<specimenCount id="9D50FD6CE308FFF51E1AFF49FDFFFF04" box="[563,606,187,213]" count="1" pageId="6" pageNumber="273" type="male">1 ♂</specimenCount>
,
<quantity id="4CAE9B00E308FFF51E43FF49FD6CFF04" box="[618,717,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="6" pageNumber="273" unit="mm" value="18.2">18.2 mm</quantity>
×
<quantity id="4CAE9B00E308FFF51EC3FF48FCF1FF04" box="[746,848,186,213]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="6" pageNumber="273" unit="mm" value="15.4">15.4 mm</quantity>
,
<location id="8E89603EE308FFF51F72FF49FC5EFF04" LSID="urn:lsid:plazi:treatment:03FF87F3E30BFFFB1CBEFA0CFDD0F859:8E89603EE308FFF51F72FF49FC5EFF04" box="[859,1023,187,213]" name="Faakuaa River" pageId="6" pageNumber="273">Faakuaa River</location>
</materialsCitation>
,
<geoCoordinate id="EE625022E308FFF51820FF49FB34FF04" box="[1033,1173,187,213]" degrees="09" direction="south" minutes="46.097" orientation="latitude" pageId="6" pageNumber="273" precision="1" value="-9.768283">09°46.097'S</geoCoordinate>
,
<geoCoordinate id="EE625022E308FFF51889FF49FAE0FF04" box="[1184,1345,187,213]" degrees="139" direction="west" minutes="01.768" orientation="longitude" pageId="6" pageNumber="273" precision="1" value="-139.02946">139°01.768'W</geoCoordinate>
, coll. E. Feunteun, P. Gerbeaux, G. Marquet, P. Tiberghien,
<date id="FFE81025E308FFF51EE5FF2DFC2AFF28" box="[716,907,223,249]" pageId="6" pageNumber="273" value="2017-10-10">10 October 2017</date>
.
<materialsCitation id="3B3E3CB8E308FFF51FBDFF2DFD55FECC" ID-GBIF-Occurrence="4905160352" collectionCode="CA" collectorName="Ua Pou" location="Hakahetau River" pageId="6" pageNumber="273" specimenCode="CA2466" specimenCount="1" specimenCount-male="1">
<emphasis id="B922EAF7E308FFF51FBDFF2DFC52FF29" box="[916,1011,223,248]" italics="true" pageId="6" pageNumber="273">
<collectorName id="26A35333E308FFF51FBDFF2DFC4DFF29" box="[916,1004,223,248]" pageId="6" pageNumber="273">Ua Pou</collectorName>
:
</emphasis>
MNHN-IU-2019-182 (DNA voucher:
<specimenCode id="DBF09E9EE308FFF51CBEFEF1FF5AFECC" box="[151,251,259,285]" collectionCode="CA" country="USA" name="Chicago Academy of Sciences" pageId="6" pageNumber="273" type="Museum">CA2466</specimenCode>
),
<specimenCount id="9D50FD6CE308FFF51D24FEF1FE98FECC" box="[269,313,259,285]" count="1" pageId="6" pageNumber="273" type="male">1 ♂</specimenCount>
,
<quantity id="4CAE9B00E308FFF51D6DFEF1FE09FECC" box="[324,424,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.6199999999999999" pageId="6" pageNumber="273" unit="mm" value="16.2">16.2 mm</quantity>
×
<quantity id="4CAE9B00E308FFF51DECFEF1FD8BFECC" box="[453,554,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="1.38" pageId="6" pageNumber="273" unit="mm" value="13.8">13.8 mm</quantity>
,
<location id="8E89603EE308FFF51E1FFEF1FD55FECC" LSID="urn:lsid:plazi:treatment:03FF87F3E30BFFFB1CBEFA0CFDD0F859:8E89603EE308FFF51E1FFEF1FD55FECC" box="[566,756,259,285]" name="Hakahetau River" pageId="6" pageNumber="273">Hakahetau River</location>
</materialsCitation>
,
<geoCoordinate id="EE625022E308FFF51ED6FEF1FC2BFECC" box="[767,906,259,285]" degrees="09" direction="south" minutes="21.600" orientation="latitude" pageId="6" pageNumber="273" precision="1" value="-9.36">09°21.600'S</geoCoordinate>
,
<geoCoordinate id="EE625022E308FFF51FBCFEF1FB96FECC" box="[917,1079,259,285]" degrees="140" direction="west" minutes="06.313" orientation="longitude" pageId="6" pageNumber="273" precision="1" value="-140.10521">140°06.313'W</geoCoordinate>
, coll. E. Feunteun, P. Gerbeaux, G. Marquet, P. Tiberghien,
<date id="FFE81025E308FFF51DE3FED5FD2AFE90" box="[458,651,295,321]" pageId="6" pageNumber="273" value="2017-10-16">16 October 2017</date>
;
<materialsCitation id="3B3E3CB8E308FFF51EBEFED4FD4FFEB4" ID-GBIF-Occurrence="4905160358" collectingDate="1996-02-12" collectionCode="CA" collectorName="J. Poupin" location="Hakahau Bay" pageId="6" pageNumber="273" specimenCode="CA2665" specimenCount="1" specimenCount-male="1">
MNHN-IU-2022-4085 (DNA voucher:
<specimenCode id="DBF09E9EE308FFF5187DFED4FB16FE90" box="[1108,1207,294,321]" collectionCode="CA" country="USA" name="Chicago Academy of Sciences" pageId="6" pageNumber="273" type="Museum">CA2665</specimenCode>
),
<specimenCount id="9D50FD6CE308FFF518E2FED5FB56FE90" box="[1227,1271,295,321]" count="1" pageId="6" pageNumber="273" type="male">1 ♂</specimenCount>
,
<quantity id="4CAE9B00E308FFF5192DFED5FAFAFE90" box="[1284,1371,295,321]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="6" pageNumber="273" unit="mm" value="9.6">9.6 mm</quantity>
×
<quantity id="4CAE9B00E308FFF51950FED5FF64FEB4" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="6" pageNumber="273" unit="mm" value="8.4">8.4 mm</quantity>
,
<location id="8E89603EE308FFF51CF8FEB9FECBFEB4" LSID="urn:lsid:plazi:treatment:03FF87F3E30BFFFB1CBEFA0CFDD0F859:8E89603EE308FFF51CF8FEB9FECBFEB4" box="[209,362,331,357]" name="Hakahau Bay" pageId="6" pageNumber="273">Hakahau Bay</location>
, coll.
<collectorName id="26A35333E308FFF51D84FEB9FDB9FEB4" box="[429,536,331,357]" pageId="6" pageNumber="273">J. Poupin</collectorName>
,
<date id="FFE81025E308FFF51E0DFEB9FD4FFEB4" box="[548,750,331,357]" pageId="6" pageNumber="273" value="1996-02-12">
<collectingDate id="EFACE9CDE308FFF51E0DFEB9FD4FFEB4" box="[548,750,331,357]" pageId="6" pageNumber="273" value="1996-02-12">12 February 1996</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3E3CB8E308FFF51ED3FEB9FC2BFE58" ID-GBIF-Occurrence="4905160334" collectingDate="1996-02-12" collectionCode="CA" collectorName="J. Poupin" location="Hakahau Bay" pageId="6" pageNumber="273" specimenCode="CA2666" specimenCount="1" specimenCount-female="1">
MNHN-IU-2022-4086 (DNA voucher:
<specimenCode id="DBF09E9EE308FFF5189CFEB9FAB9FEB4" box="[1205,1304,331,357]" collectionCode="CA" country="USA" name="Chicago Academy of Sciences" pageId="6" pageNumber="273" type="Museum">CA2666</specimenCode>
),
<specimenCount id="9D50FD6CE308FFF51902FEB9FAF4FEB4" box="[1323,1365,331,357]" count="1" pageId="6" pageNumber="273" type="female">1 ♀</specimenCount>
ovig.,
<quantity id="4CAE9B00E308FFF51CBEFE9DFF4CFE58" box="[151,237,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="9.2" pageId="6" pageNumber="273" unit="mm" value="9.2">9.2 mm</quantity>
×
<quantity id="4CAE9B00E308FFF51D22FE9CFEC2FE58" box="[267,355,366,393]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="6" pageNumber="273" unit="mm" value="8.5">8.5 mm</quantity>
,
<location id="8E89603EE308FFF51D47FE9DFDA6FE58" LSID="urn:lsid:plazi:treatment:03FF87F3E30BFFFB1CBEFA0CFDD0F859:8E89603EE308FFF51D47FE9DFDA6FE58" box="[366,519,367,393]" name="Hakahau Bay" pageId="6" pageNumber="273">Hakahau Bay</location>
, coll.
<collectorName id="26A35333E308FFF51E63FE9DFD14FE58" box="[586,693,367,393]" pageId="6" pageNumber="273">J. Poupin</collectorName>
,
<date id="FFE81025E308FFF51EE9FE9DFC2BFE58" box="[704,906,367,393]" pageId="6" pageNumber="273" value="1996-02-12">
<collectingDate id="EFACE9CDE308FFF51EE9FE9DFC2BFE58" box="[704,906,367,393]" pageId="6" pageNumber="273" value="1996-02-12">12 February 1996</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34C656EE308FFF51CEEFE60FCE2FDB0" pageId="6" pageNumber="273" type="diagnosis">
<paragraph id="8BE936E5E308FFF51CEEFE60FCE2FDB0" blockId="6.[151,1437,150,1977]" pageId="6" pageNumber="273">
<emphasis id="B922EAF7E308FFF51CEEFE60FEE3FE7D" bold="true" box="[199,322,402,428]" pageId="6" pageNumber="273">Diagnosis.</emphasis>
Carapace quadrate, dorsal surface smooth and flat. Front broad, anterior margin straight to faintly sinuous, frontal width 0.370.42 times CW; anterolateral margin with 3 teeth including orbital tooth. Third maxillipeds with auriculiform merus; exopod broad, inflated, greatest width 1.31.6 times of median width of ischium. Chelipeds stout, symmetrical in males; inner angle of carpus with single blunt spine only in females; fingers glabrous. Ambulatory legs slender, somewhat compressed laterally; merus with single spine on anterior margins near carpus in P24, spine indistinct or absent in P5.
</paragraph>
</subSubSection>
<subSubSection id="C34C656EE308FFFB1CEEFD98FBA0FA64" lastPageId="8" lastPageNumber="275" pageId="6" pageNumber="273" type="description">
<paragraph id="8BE936E5E308FFF51CEEFD98FB68FC8C" blockId="6.[151,1437,150,1977]" pageId="6" pageNumber="273">
<emphasis id="B922EAF7E308FFF51CEEFD98FEF8FD55" bold="true" box="[199,345,618,644]" pageId="6" pageNumber="273">Description.</emphasis>
Carapace (
<figureCitation id="136D2A60E308FFF51DF2FD99FD94FD54" box="[475,565,619,645]" captionStart="FIGURE 4" captionStartId="7.[151,250,1909,1933]" captionTargetBox="[194,1393,183,1885]" captionTargetId="figure-19@7.[191,1396,181,1887]" captionTargetPageId="7" captionText="FIGURE 4. Ptychognathus crassimanus Finnegan, 1931: A, carapace; B, left third maxilliped; C, male cheliped; D, E, female cheliped; F, third pereiopod with inner view of dactylus; G, fifth pereiopod; H, male pleon; I, female pleon; J, male right first gonopod. MNHN-IU-2019-182 (A, B, C, F, G, H) male from Ua Pou Island; MNHN-IU-2019-183 (J) male from Hiva Oa Island; MNHN-IU-2011-9062 (E, D, I) female from Fatu Hiva Island." figureDoi="http://doi.org/10.5281/zenodo.12681756" httpUri="https://zenodo.org/record/12681756/files/figure.png" pageId="6" pageNumber="273">Fig. 4A</figureCitation>
) quadrate, slightly broader than long, 1.1 times as wide as long, flat, dorsal surface smooth and glabrous; regions poorly defined but with central H-shaped grooves clearly discernible. Front broad, anterior margin faintly sinuous; frontal width 0.370.42 times CW, fronto-orbital width 0.910.96 times CW. Upper border of orbit S-shaped, inner angle with distinct slit. Anterolateral margins and supraorbital margins lined with small granules, anterolateral margins with 3 teeth including orbital tooth, first tooth most distinct and acute, slightly sloping forward, second and third teeth, clearly visible, blunt. Posterolateral margins divergent posteriorly, moderately sloping outwards. Lateral and posterolateral margins furnished with short soft setae.
</paragraph>
<paragraph id="8BE936E5E308FFF51CEEFC95FD34FC3C" blockId="6.[151,1437,150,1977]" pageId="6" pageNumber="273">
Third maxillipeds (
<figureCitation id="136D2A60E308FFF51DB2FC95FE51FC50" box="[411,496,871,897]" captionStart="FIGURE 4" captionStartId="7.[151,250,1909,1933]" captionTargetBox="[194,1393,183,1885]" captionTargetId="figure-19@7.[191,1396,181,1887]" captionTargetPageId="7" captionText="FIGURE 4. Ptychognathus crassimanus Finnegan, 1931: A, carapace; B, left third maxilliped; C, male cheliped; D, E, female cheliped; F, third pereiopod with inner view of dactylus; G, fifth pereiopod; H, male pleon; I, female pleon; J, male right first gonopod. MNHN-IU-2019-182 (A, B, C, F, G, H) male from Ua Pou Island; MNHN-IU-2019-183 (J) male from Hiva Oa Island; MNHN-IU-2011-9062 (E, D, I) female from Fatu Hiva Island." figureDoi="http://doi.org/10.5281/zenodo.12681756" httpUri="https://zenodo.org/record/12681756/files/figure.png" pageId="6" pageNumber="273">Fig. 4B</figureCitation>
) with ischium subequal in proximal and distal widths, lateral margin concave, ventral surface with longitudinal median ridge. Merus very broad, with anterolateral angle strongly inflated, auriculiform; ventral surface with oblique ridge on mesial part. Exopod broad, inflated, greatest width 1.41.6 times (male), 1.31.4 (female) of median width of ischium.
</paragraph>
<paragraph id="8BE936E5E308FFF51CEEFC05FB0EFB38" blockId="6.[151,1437,150,1977]" pageId="6" pageNumber="273">
Male chelipeds (
<figureCitation id="136D2A60E308FFF51DABFC05FE79FBC0" box="[386,472,1015,1041]" captionStart="FIGURE 4" captionStartId="7.[151,250,1909,1933]" captionTargetBox="[194,1393,183,1885]" captionTargetId="figure-19@7.[191,1396,181,1887]" captionTargetPageId="7" captionText="FIGURE 4. Ptychognathus crassimanus Finnegan, 1931: A, carapace; B, left third maxilliped; C, male cheliped; D, E, female cheliped; F, third pereiopod with inner view of dactylus; G, fifth pereiopod; H, male pleon; I, female pleon; J, male right first gonopod. MNHN-IU-2019-182 (A, B, C, F, G, H) male from Ua Pou Island; MNHN-IU-2019-183 (J) male from Hiva Oa Island; MNHN-IU-2011-9062 (E, D, I) female from Fatu Hiva Island." figureDoi="http://doi.org/10.5281/zenodo.12681756" httpUri="https://zenodo.org/record/12681756/files/figure.png" pageId="6" pageNumber="273">Fig. 4C</figureCitation>
) symmetrical, not granular at all. Merus longer than broad (length to width ratio 1.8 2.1, anterior and posterior proximal margins with long soft setae, posterior distal margins bluntly carinate. Carpus strongly inflated, slightly longer than wide (length to width ratio 1.01.2); anterior distal angle with single blunt spine only in females. Palm very much inflated (
<taxonomicName id="4C564D66E308FFF51EE4FB91FCF8FBAD" authorityName="Finnegan" authorityYear="1931" box="[717,857,1123,1148]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="273" phylum="Arthropoda" rank="species" species="crassimanus">
<emphasis id="B922EAF7E308FFF51EE4FB91FCF8FBAD" box="[717,857,1123,1148]" italics="true" pageId="6" pageNumber="273">crassimanus</emphasis>
</taxonomicName>
means stout palm), outer surface smooth, without distinct granules; inner surface glabrous. Fingers glabrous. Dactylus about as long as palm, terminating in small, hoof-shaped corneous claw, bearing 812 small teeth, pollex with only faint indication of longitudinal groove, with 48 teeth, more conical and larger than those of dactylus. Without pulvinus at base of fingers.
</paragraph>
<paragraph id="8BE936E5E308FFF51CEEFB01FAEDFA84" blockId="6.[151,1437,150,1977]" pageId="6" pageNumber="273">
Female chelipeds (
<figureCitation id="136D2A60E308FFF51DB5FB01FD82FADC" box="[412,547,1267,1293]" captionStart="FIGURE 4" captionStartId="7.[151,250,1909,1933]" captionTargetBox="[194,1393,183,1885]" captionTargetId="figure-19@7.[191,1396,181,1887]" captionTargetPageId="7" captionText="FIGURE 4. Ptychognathus crassimanus Finnegan, 1931: A, carapace; B, left third maxilliped; C, male cheliped; D, E, female cheliped; F, third pereiopod with inner view of dactylus; G, fifth pereiopod; H, male pleon; I, female pleon; J, male right first gonopod. MNHN-IU-2019-182 (A, B, C, F, G, H) male from Ua Pou Island; MNHN-IU-2019-183 (J) male from Hiva Oa Island; MNHN-IU-2011-9062 (E, D, I) female from Fatu Hiva Island." figureDoi="http://doi.org/10.5281/zenodo.12681756" httpUri="https://zenodo.org/record/12681756/files/figure.png" pageId="6" pageNumber="273">Fig. 4D, 4E</figureCitation>
) distinctly smaller than those of male, chela much narrower than that of male. Palm not strongly inflated; movable finger distinctly shorter than palm. Carpus with small blunt spine at inner distal angle (
<figureCitation id="136D2A60E308FFF51CCAFAC9FE9AFA84" box="[227,315,1339,1365]" captionStart="FIGURE 4" captionStartId="7.[151,250,1909,1933]" captionTargetBox="[194,1393,183,1885]" captionTargetId="figure-19@7.[191,1396,181,1887]" captionTargetPageId="7" captionText="FIGURE 4. Ptychognathus crassimanus Finnegan, 1931: A, carapace; B, left third maxilliped; C, male cheliped; D, E, female cheliped; F, third pereiopod with inner view of dactylus; G, fifth pereiopod; H, male pleon; I, female pleon; J, male right first gonopod. MNHN-IU-2019-182 (A, B, C, F, G, H) male from Ua Pou Island; MNHN-IU-2019-183 (J) male from Hiva Oa Island; MNHN-IU-2011-9062 (E, D, I) female from Fatu Hiva Island." figureDoi="http://doi.org/10.5281/zenodo.12681756" httpUri="https://zenodo.org/record/12681756/files/figure.png" pageId="6" pageNumber="273">Fig. 4E</figureCitation>
). Narrow hiatus between fingers. Pollex with ridge consisting of large granules toward palm.
</paragraph>
<paragraph id="8BE936E5E308FFF51CEEFAADFADCF96C" blockId="6.[151,1437,150,1977]" pageId="6" pageNumber="273">
Ambulatory legs (P25), slender, somewhat compressed laterally; P4 longest (male: length 1.41.9 times carapace width, female: length 1.21.4 times carapace width); P5 shortest. Merus elongated, subrectangular in lateral view, with single spine on anterior margins near carpus in P2P4, spine indistinct or absent in P5. Anterior margins of P25 meri with long soft setae; posterior margins glabrous. Anterior margin of carpus with dense short setae, posterior margins glabrous. Propodus with dense short setae on anterior margins and dense short setae with sparse long setae on posterior margins. Dactylus terminating in subacute corneous claw, with dense short setae on anterior and posterior margins; ventral surface with rows of short setae. P3 (
<figureCitation id="136D2A60E308FFF51FC5F9C5FB9EF980" box="[1004,1087,1591,1617]" captionStart="FIGURE 4" captionStartId="7.[151,250,1909,1933]" captionTargetBox="[194,1393,183,1885]" captionTargetId="figure-19@7.[191,1396,181,1887]" captionTargetPageId="7" captionText="FIGURE 4. Ptychognathus crassimanus Finnegan, 1931: A, carapace; B, left third maxilliped; C, male cheliped; D, E, female cheliped; F, third pereiopod with inner view of dactylus; G, fifth pereiopod; H, male pleon; I, female pleon; J, male right first gonopod. MNHN-IU-2019-182 (A, B, C, F, G, H) male from Ua Pou Island; MNHN-IU-2019-183 (J) male from Hiva Oa Island; MNHN-IU-2011-9062 (E, D, I) female from Fatu Hiva Island." figureDoi="http://doi.org/10.5281/zenodo.12681756" httpUri="https://zenodo.org/record/12681756/files/figure.png" pageId="6" pageNumber="273">Fig. 4F</figureCitation>
) with merus 3.23.9 as long as wide, carpus 1.52.1 as long as wide, propodus 2.12.3 as long as wide, dactylus 3.75.2 as long as wide, dactylus 1.01.2 times as long as propodus. P5 (
<figureCitation id="136D2A60E308FFF51E60F98DFD01F948" box="[585,672,1663,1689]" captionStart="FIGURE 4" captionStartId="7.[151,250,1909,1933]" captionTargetBox="[194,1393,183,1885]" captionTargetId="figure-19@7.[191,1396,181,1887]" captionTargetPageId="7" captionText="FIGURE 4. Ptychognathus crassimanus Finnegan, 1931: A, carapace; B, left third maxilliped; C, male cheliped; D, E, female cheliped; F, third pereiopod with inner view of dactylus; G, fifth pereiopod; H, male pleon; I, female pleon; J, male right first gonopod. MNHN-IU-2019-182 (A, B, C, F, G, H) male from Ua Pou Island; MNHN-IU-2019-183 (J) male from Hiva Oa Island; MNHN-IU-2011-9062 (E, D, I) female from Fatu Hiva Island." figureDoi="http://doi.org/10.5281/zenodo.12681756" httpUri="https://zenodo.org/record/12681756/files/figure.png" pageId="6" pageNumber="273">Fig. 4G</figureCitation>
) with merus 2.22.8 as long as wide, carpus 1.62.5 as long as wide, propodus 1.21.6 as long as wide, dactylus 2.53.0 as long as wide, dactylus 0.70.9 times as long as propodus.
</paragraph>
<paragraph id="8BE936E5E308FFF51CEEF935FF4BF89C" blockId="6.[151,1437,150,1977]" pageId="6" pageNumber="273">
Male pleon (
<figureCitation id="136D2A60E308FFF51D7FF935FE0FF930" box="[342,430,1735,1761]" captionStart="FIGURE 4" captionStartId="7.[151,250,1909,1933]" captionTargetBox="[194,1393,183,1885]" captionTargetId="figure-19@7.[191,1396,181,1887]" captionTargetPageId="7" captionText="FIGURE 4. Ptychognathus crassimanus Finnegan, 1931: A, carapace; B, left third maxilliped; C, male cheliped; D, E, female cheliped; F, third pereiopod with inner view of dactylus; G, fifth pereiopod; H, male pleon; I, female pleon; J, male right first gonopod. MNHN-IU-2019-182 (A, B, C, F, G, H) male from Ua Pou Island; MNHN-IU-2019-183 (J) male from Hiva Oa Island; MNHN-IU-2011-9062 (E, D, I) female from Fatu Hiva Island." figureDoi="http://doi.org/10.5281/zenodo.12681756" httpUri="https://zenodo.org/record/12681756/files/figure.png" pageId="6" pageNumber="273">Fig. 4H</figureCitation>
) narrow, broadly triangular, nearly smooth, distinctly narrowed at base of telson; margins covered with short setae, setae on margins of telson longer. Telson linguiform, terminal margin broadly rounded; length 1.01.2 times basal width, 1.11.3 times as long as sixth somite, sixth somite 1.11.4 times longer than fifth somite.
</paragraph>
<paragraph id="8BE936E5E308FFF51CEEF8A5FB42F868" blockId="6.[151,1437,150,1977]" pageId="6" pageNumber="273">
Female pleon (
<figureCitation id="136D2A60E308FFF51D43F8A5FE17F8A0" box="[362,438,1879,1905]" captionStart="FIGURE 4" captionStartId="7.[151,250,1909,1933]" captionTargetBox="[194,1393,183,1885]" captionTargetId="figure-19@7.[191,1396,181,1887]" captionTargetPageId="7" captionText="FIGURE 4. Ptychognathus crassimanus Finnegan, 1931: A, carapace; B, left third maxilliped; C, male cheliped; D, E, female cheliped; F, third pereiopod with inner view of dactylus; G, fifth pereiopod; H, male pleon; I, female pleon; J, male right first gonopod. MNHN-IU-2019-182 (A, B, C, F, G, H) male from Ua Pou Island; MNHN-IU-2019-183 (J) male from Hiva Oa Island; MNHN-IU-2011-9062 (E, D, I) female from Fatu Hiva Island." figureDoi="http://doi.org/10.5281/zenodo.12681756" httpUri="https://zenodo.org/record/12681756/files/figure.png" pageId="6" pageNumber="273">Fig. 4I</figureCitation>
,
<figureCitation id="136D2A60E308FFF51DE9F8A4FE43F8A0" box="[448,482,1878,1905]" captionStart="FIGURE 5" captionStartId="8.[151,250,1076,1100]" captionTargetBox="[151,1435,181,1052]" captionTargetId="figure-26@8.[151,1435,181,1052]" captionTargetPageId="8" captionText="FIGURE 5. Ptychognathus crassimanus Finnegan, 1931:A, dorsal view of a living specimen; B, front view of a living specimen; C, ventral view; D, typical habitat of the species, Faakuaa River, Hiva Oa Island. Photo credits: É. Vigneux (A, B), J. Poupin (C) and G. Marquet (D)." figureDoi="http://doi.org/10.5281/zenodo.12681758" httpUri="https://zenodo.org/record/12681758/files/figure.png" pageId="6" pageNumber="273">5C</figureCitation>
) very broad, covering most of thoracic sternum. Telson broadly subtriangular, terminal margin broadly rounded, length 0.25 times basal width, 0.80.9 times longer than sixth somite, sixth somite 1.01.2 times longer than fifth somite, measured on midline. Small round eggs
<quantity id="4CAE9B00E308FFF51F9DF86CFBF6F868" box="[948,1111,1950,1977]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" metricValueMax="3.1" metricValueMin="2.5" pageId="6" pageNumber="273" unit="mm" value="0.28" valueMax="0.31" valueMin="0.25">0.250.31 mm</quantity>
in diameter.
</paragraph>
<caption id="DF29666DE309FFF41CBEF887FCFDF828" ID-DOI="http://doi.org/10.5281/zenodo.12681756" ID-Zenodo-Dep="12681756" httpUri="https://zenodo.org/record/12681756/files/figure.png" pageId="7" pageNumber="274" startId="7.[151,250,1909,1933]" targetBox="[194,1393,183,1885]" targetPageId="7" targetType="figure">
<paragraph id="8BE936E5E309FFF41CBEF887FCFDF828" blockId="7.[151,1437,1909,2041]" pageId="7" pageNumber="274">
<emphasis id="B922EAF7E309FFF41CBEF887FEB2F85C" bold="true" box="[151,275,1909,1933]" pageId="7" pageNumber="274">FIGURE 4.</emphasis>
<taxonomicName id="4C564D66E309FFF41D33F887FD7FF85C" authority="Finnegan, 1931" authorityName="Finnegan" authorityYear="1931" box="[282,734,1909,1933]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="274" phylum="Arthropoda" rank="species" species="crassimanus">
<emphasis id="B922EAF7E309FFF41D33F887FD97F85C" box="[282,566,1909,1933]" italics="true" pageId="7" pageNumber="274">Ptychognathus crassimanus</emphasis>
<bibRefCitation id="EFC74B14E309FFF41E15F887FD7FF85C" author="Finnegan, S." box="[572,734,1909,1933]" pageId="7" pageNumber="274" pagination="607 - 673" refId="ref17489" refString="Finnegan, S. (1931) Report on the brachyura collected in Central America, the Gorgona and Galapagos islands, by Dr. Crossland on the St George Expedition to the Pacific, 1924 - 25. Linnean Society's Journal, Zoology, 37 (255), 607 - 673. https: // doi. org / 10.1111 / j. 1096 - 3642.1931. tb 02367. x" type="journal article" year="1931">Finnegan, 1931</bibRefCitation>
</taxonomicName>
: A, carapace; B, left third maxilliped; C, male cheliped; D, E, female cheliped; F, third pereiopod with inner view of dactylus; G, fifth pereiopod; H, male pleon; I, female pleon; J, male right first gonopod. MNHN-IU-2019-182 (A, B, C, F, G, H) male from Ua Pou Island; MNHN-IU-2019-183 (J) male from Hiva Oa Island; MNHN-IU-2011-9062 (E, D, I) female from Fatu Hiva Island.
</paragraph>
</caption>
<caption id="DF29666DE306FFFB1CBEFBC6FE2EFB45" ID-DOI="http://doi.org/10.5281/zenodo.12681758" ID-Zenodo-Dep="12681758" httpUri="https://zenodo.org/record/12681758/files/figure.png" pageId="8" pageNumber="275" startId="8.[151,250,1076,1100]" targetBox="[151,1435,181,1052]" targetPageId="8" targetType="figure">
<paragraph id="8BE936E5E306FFFB1CBEFBC6FE2EFB45" blockId="8.[151,1437,1076,1172]" pageId="8" pageNumber="275">
<emphasis id="B922EAF7E306FFFB1CBEFBC6FEB0FB9D" bold="true" box="[151,273,1076,1100]" pageId="8" pageNumber="275">FIGURE 5.</emphasis>
<taxonomicName id="4C564D66E306FFFB1D3FFBC6FD75FB9D" authority="Finnegan, 1931" authorityName="Finnegan" authorityYear="1931" box="[278,724,1076,1100]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="275" phylum="Arthropoda" rank="species" species="crassimanus">
<emphasis id="B922EAF7E306FFFB1D3FFBC6FD91FB9D" box="[278,560,1076,1100]" italics="true" pageId="8" pageNumber="275">Ptychognathus crassimanus</emphasis>
<bibRefCitation id="EFC74B14E306FFFB1E1DFBC6FD75FB9D" author="Finnegan, S." box="[564,724,1076,1100]" pageId="8" pageNumber="275" pagination="607 - 673" refId="ref17489" refString="Finnegan, S. (1931) Report on the brachyura collected in Central America, the Gorgona and Galapagos islands, by Dr. Crossland on the St George Expedition to the Pacific, 1924 - 25. Linnean Society's Journal, Zoology, 37 (255), 607 - 673. https: // doi. org / 10.1111 / j. 1096 - 3642.1931. tb 02367. x" type="journal article" year="1931">Finnegan, 1931</bibRefCitation>
</taxonomicName>
:A, dorsal view of a living specimen; B, front view of a living specimen; C, ventral view; D, typical habitat of the species, Faakuaa River, Hiva Oa Island. Photo credits: É. Vigneux (A, B), J. Poupin (C) and G. Marquet (D).
</paragraph>
</caption>
<paragraph id="8BE936E5E306FFFB1CEEFB30FC74FAD0" blockId="8.[151,1437,1218,1929]" pageId="8" pageNumber="275">
Male G1 (
<figureCitation id="136D2A60E306FFFB1D11FB30FE27FB0D" box="[312,390,1218,1244]" captionStart="FIGURE 4" captionStartId="7.[151,250,1909,1933]" captionTargetBox="[194,1393,183,1885]" captionTargetId="figure-19@7.[191,1396,181,1887]" captionTargetPageId="7" captionText="FIGURE 4. Ptychognathus crassimanus Finnegan, 1931: A, carapace; B, left third maxilliped; C, male cheliped; D, E, female cheliped; F, third pereiopod with inner view of dactylus; G, fifth pereiopod; H, male pleon; I, female pleon; J, male right first gonopod. MNHN-IU-2019-182 (A, B, C, F, G, H) male from Ua Pou Island; MNHN-IU-2019-183 (J) male from Hiva Oa Island; MNHN-IU-2011-9062 (E, D, I) female from Fatu Hiva Island." figureDoi="http://doi.org/10.5281/zenodo.12681756" httpUri="https://zenodo.org/record/12681756/files/figure.png" pageId="8" pageNumber="275">Fig. 4J</figureCitation>
) slender, 8 times as long as wide, slightly curving dorsally; tip bluntly rounded, forming short chitinous structure; subdistal lobe obtuse, partially obscured by stiff setae.
</paragraph>
<paragraph id="8BE936E5E306FFFB1CEEFAF8FBC5FA98" blockId="8.[151,1437,1218,1929]" pageId="8" pageNumber="275">
<emphasis id="B922EAF7E306FFFB1CEEFAF8FE0AFAF5" bold="true" box="[199,427,1290,1316]" pageId="8" pageNumber="275">Colouration in life.</emphasis>
Carapace (
<figureCitation id="136D2A60E306FFFB1E02FAF8FD22FAF4" box="[555,643,1290,1317]" captionStart="FIGURE 5" captionStartId="8.[151,250,1076,1100]" captionTargetBox="[151,1435,181,1052]" captionTargetId="figure-26@8.[151,1435,181,1052]" captionTargetPageId="8" captionText="FIGURE 5. Ptychognathus crassimanus Finnegan, 1931:A, dorsal view of a living specimen; B, front view of a living specimen; C, ventral view; D, typical habitat of the species, Faakuaa River, Hiva Oa Island. Photo credits: É. Vigneux (A, B), J. Poupin (C) and G. Marquet (D)." figureDoi="http://doi.org/10.5281/zenodo.12681758" httpUri="https://zenodo.org/record/12681758/files/figure.png" pageId="8" pageNumber="275">Fig. 5A</figureCitation>
) bluish brownish. Chelipeds (
<figureCitation id="136D2A60E306FFFB1FFEFAF8FB8FFAF4" box="[983,1070,1290,1317]" captionStart="FIGURE 5" captionStartId="8.[151,250,1076,1100]" captionTargetBox="[151,1435,181,1052]" captionTargetId="figure-26@8.[151,1435,181,1052]" captionTargetPageId="8" captionText="FIGURE 5. Ptychognathus crassimanus Finnegan, 1931:A, dorsal view of a living specimen; B, front view of a living specimen; C, ventral view; D, typical habitat of the species, Faakuaa River, Hiva Oa Island. Photo credits: É. Vigneux (A, B), J. Poupin (C) and G. Marquet (D)." figureDoi="http://doi.org/10.5281/zenodo.12681758" httpUri="https://zenodo.org/record/12681758/files/figure.png" pageId="8" pageNumber="275">Fig. 5B</figureCitation>
) bluish brownish superiorly and orange inferiorly; finger tips whitish. Ambulatory legs bluish-brown with orange parts.
</paragraph>
<paragraph id="8BE936E5E306FFFB1CEEFAA0FEEAFA40" blockId="8.[151,1437,1218,1929]" pageId="8" pageNumber="275">
<emphasis id="B922EAF7E306FFFB1CEEFAA0FF5FFABD" bold="true" box="[199,254,1362,1388]" pageId="8" pageNumber="275">Size.</emphasis>
Medium-sized. the largest male specimen examined is
<quantity id="4CAE9B00E306FFFB1F5FFAA0FC7AFABC" box="[886,987,1362,1389]" metricMagnitude="-2" metricUnit="m" metricValue="1.8199999999999998" pageId="8" pageNumber="275" unit="mm" value="18.2">18.2 mm</quantity>
×
<quantity id="4CAE9B00E306FFFB1FD5FAA0FBC0FABC" box="[1020,1121,1362,1389]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" pageId="8" pageNumber="275" unit="mm" value="15.4">15.4 mm</quantity>
and the largest female 14.0 mm ×
<quantity id="4CAE9B00E306FFFB1CC8FA84FEE7FA40" box="[225,326,1398,1425]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="8" pageNumber="275" unit="mm" value="11.8">11.8 mm</quantity>
.
</paragraph>
<paragraph id="8BE936E5E306FFFB1CEEFA68FBA0FA64" blockId="8.[151,1437,1218,1929]" box="[199,1025,1434,1461]" pageId="8" pageNumber="275">
<emphasis id="B922EAF7E306FFFB1CEEFA68FE8BFA65" bold="true" box="[199,298,1434,1460]" pageId="8" pageNumber="275">Habitat.</emphasis>
This species is found in brackish water, near estuaries (
<figureCitation id="136D2A60E306FFFB1FB3FA68FC52FA64" box="[922,1011,1434,1461]" captionStart="FIGURE 5" captionStartId="8.[151,250,1076,1100]" captionTargetBox="[151,1435,181,1052]" captionTargetId="figure-26@8.[151,1435,181,1052]" captionTargetPageId="8" captionText="FIGURE 5. Ptychognathus crassimanus Finnegan, 1931:A, dorsal view of a living specimen; B, front view of a living specimen; C, ventral view; D, typical habitat of the species, Faakuaa River, Hiva Oa Island. Photo credits: É. Vigneux (A, B), J. Poupin (C) and G. Marquet (D)." figureDoi="http://doi.org/10.5281/zenodo.12681758" httpUri="https://zenodo.org/record/12681758/files/figure.png" pageId="8" pageNumber="275">Fig. 5D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34C656EE306FFFB1CEEFA4CFEB4FA2D" pageId="8" pageNumber="275" type="distribution">
<paragraph id="8BE936E5E306FFFB1CEEFA4CFEB4FA2D" blockId="8.[151,1437,1218,1929]" pageId="8" pageNumber="275">
<emphasis id="B922EAF7E306FFFB1CEEFA4CFEC0FA09" bold="true" box="[199,353,1470,1496]" pageId="8" pageNumber="275">Distribution.</emphasis>
<taxonomicName id="4C564D66E306FFFB1D4FFA4CFCF0FA09" authority="Finnegan, 1931" authorityName="Finnegan" authorityYear="1931" box="[358,849,1470,1496]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="275" phylum="Arthropoda" rank="species" species="crassimanus">
<emphasis id="B922EAF7E306FFFB1D4FFA4CFD3EFA09" box="[358,671,1470,1496]" italics="true" pageId="8" pageNumber="275">Ptychognathus crassimanus</emphasis>
<bibRefCitation id="EFC74B14E306FFFB1E8DFA4CFCF0FA09" author="Finnegan, S." box="[676,849,1470,1496]" pageId="8" pageNumber="275" pagination="607 - 673" refId="ref17489" refString="Finnegan, S. (1931) Report on the brachyura collected in Central America, the Gorgona and Galapagos islands, by Dr. Crossland on the St George Expedition to the Pacific, 1924 - 25. Linnean Society's Journal, Zoology, 37 (255), 607 - 673. https: // doi. org / 10.1111 / j. 1096 - 3642.1931. tb 02367. x" type="journal article" year="1931">Finnegan, 1931</bibRefCitation>
</taxonomicName>
(
<figureCitation id="136D2A60E306FFFB1F49FA4CFC03FA09" box="[864,930,1470,1496]" captionStart="FIGURE 6" captionStartId="9.[151,250,1462,1486]" captionTargetBox="[154,1434,183,1438]" captionTargetId="figure-19@9.[151,1436,181,1441]" captionTargetPageId="9" captionText="FIGURE 6. Distribution map of the species studied. Stars indicate type localities." figureDoi="http://doi.org/10.5281/zenodo.12681762" httpUri="https://zenodo.org/record/12681762/files/figure.png" pageId="8" pageNumber="275">Fig. 6</figureCitation>
) is endemic to the
<collectingRegion id="4992F807E306FFFB1858FA4DFA9EFA09" box="[1137,1343,1470,1497]" country="French Polynesia" name="Iles Marquises" pageId="8" pageNumber="275">Marquesas Islands</collectingRegion>
(
<collectingCountry id="F3417675E306FFFB1964FA4CFEA8FA2D" name="French Polynesia" pageId="8" pageNumber="275">French Polynesia</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C34C656EE306FFFB1CEEF9F4FDD0F859" pageId="8" pageNumber="275" type="discussion">
<paragraph id="8BE936E5E306FFFB1CEEF9F4FC63F891" blockId="8.[151,1437,1218,1929]" pageId="8" pageNumber="275">
<emphasis id="B922EAF7E306FFFB1CEEF9F4FE9AF9F1" bold="true" box="[199,315,1542,1568]" pageId="8" pageNumber="275">Remarks.</emphasis>
This species resembles
<taxonomicName id="4C564D66E306FFFB1E62F9F5FD41F9F1" authorityName="Rathbun" authorityYear="1907" box="[587,736,1543,1568]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="275" phylum="Arthropoda" rank="species" species="easteranus">
<emphasis id="B922EAF7E306FFFB1E62F9F5FD41F9F1" box="[587,736,1543,1568]" italics="true" pageId="8" pageNumber="275">P. easteranus</emphasis>
</taxonomicName>
,
<emphasis id="B922EAF7E306FFFB1EC6F9F5FCE8F9F1" box="[751,841,1542,1568]" italics="true" pageId="8" pageNumber="275">P. makii</emphasis>
<bibRefCitation id="EFC74B14E306FFFB1F79F9F5FBBAF9F1" author="Hsu, J. - W. &amp; Shih, H. - T." box="[848,1051,1542,1569]" pageId="8" pageNumber="275" pagination="1 - 13" refId="ref17713" refString="Hsu, J. - W. &amp; Shih, H. - T. (2020) Diversity of Taiwanese brackish crabs Genus Ptychognathus Stimpson, 1958 (Crustacea: Brachyura: Varunidae) based on DNA barcode, with descriptions of two new species. Zoological Studies, 59, 1 - 13." type="journal article" year="2020">Hsu &amp; Shih, 2020</bibRefCitation>
and
<emphasis id="B922EAF7E306FFFB187AF9F5FB7DF9F1" box="[1107,1244,1542,1568]" italics="true" pageId="8" pageNumber="275">P. altimanus</emphasis>
(
<bibRefCitation id="EFC74B14E306FFFB18C5F9F4FA35F9F1" author="Rathbun, M. J." box="[1260,1428,1542,1568]" pageId="8" pageNumber="275" pagination="69 - 85" refId="ref19496" refString="Rathbun, M. J. (1914) New species of crabs of the families Grapsidae and Ocypodidae. (Scientific results of the Philippine cruise of the fisheries steamer &quot; Albatross. &quot; 1907 - 1910. No. 31). Proceedings of the United States National Museum, 47, 69 - 85. https: // doi. org / 10.5479 / si. 00963801.2044.69" type="journal article" year="1914">Rathbun, 1914</bibRefCitation>
) by its broad front, anterolateral margins of carapace with three teeth clearly visible, its P24 merus with a single small spine on anterior margin near the carpus.
<taxonomicName id="4C564D66E306FFFB1E9AF9BCFC4FF9B9" authorityName="Finnegan" authorityYear="1931" box="[691,1006,1614,1640]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="275" phylum="Arthropoda" rank="species" species="crassimanus">
<emphasis id="B922EAF7E306FFFB1E9AF9BCFC4FF9B9" box="[691,1006,1614,1640]" italics="true" pageId="8" pageNumber="275">Ptychognathus crassimanus</emphasis>
</taxonomicName>
differs from
<emphasis id="B922EAF7E306FFFB18ACF9BDFAAEF9B9" box="[1157,1295,1614,1640]" italics="true" pageId="8" pageNumber="275">P. altimanus</emphasis>
by having a shorter frontal region (vs longer) and posterolateral margins of carapace distinctly divergent posteriorly (vs almost parallel, not divergent posteriorly) (
<bibRefCitation id="EFC74B14E306FFFB1E03F964FCA0F960" author="Naruse, T. &amp; Shih, H. T. &amp; Ng, N. K. &amp; Hsu, H. - L." box="[554,769,1686,1713]" pageId="8" pageNumber="275" pagination="69 - 79" refId="ref18749" refString="Naruse, T., Shih, H. T., Ng, N. K. &amp; Hsu, H. - L. (2005) On two new records of varunid crabs (Crustacea: Brachyura: Varunidae) from Southern Taiwan. Collection and Research, 18, 69 - 79." type="journal article" year="2005">
Naruse
<emphasis id="B922EAF7E306FFFB1EAAF965FD1CF961" box="[643,701,1686,1712]" italics="true" pageId="8" pageNumber="275">et al.</emphasis>
2005
</bibRefCitation>
: fig. 1A). It differs from
<emphasis id="B922EAF7E306FFFB1837F965FBD8F961" box="[1054,1145,1686,1712]" italics="true" pageId="8" pageNumber="275">P. makii</emphasis>
by having fewer teeth on the dactylus 812 (vs 15) and on the pollex 48 (vs 10) (
<bibRefCitation id="EFC74B14E306FFFB1F3EF948FC40F905" author="Hsu, J. - W. &amp; Shih, H. - T." box="[791,993,1722,1748]" pageId="8" pageNumber="275" pagination="1 - 13" refId="ref17713" refString="Hsu, J. - W. &amp; Shih, H. - T. (2020) Diversity of Taiwanese brackish crabs Genus Ptychognathus Stimpson, 1958 (Crustacea: Brachyura: Varunidae) based on DNA barcode, with descriptions of two new species. Zoological Studies, 59, 1 - 13." type="journal article" year="2020">Hsu &amp; Shih 2020</bibRefCitation>
: fig. 3D) and merus of ambulatory legs with their posterior margins glabrous (vs with short setae densely and long setae sparsely (
<bibRefCitation id="EFC74B14E306FFFB18B4F92CFACBF929" author="Hsu, J. - W. &amp; Shih, H. - T." box="[1181,1386,1758,1784]" pageId="8" pageNumber="275" pagination="1 - 13" refId="ref17713" refString="Hsu, J. - W. &amp; Shih, H. - T. (2020) Diversity of Taiwanese brackish crabs Genus Ptychognathus Stimpson, 1958 (Crustacea: Brachyura: Varunidae) based on DNA barcode, with descriptions of two new species. Zoological Studies, 59, 1 - 13." type="journal article" year="2020">Hsu &amp; Shih 2020</bibRefCitation>
: fig. 3F).
<taxonomicName id="4C564D66E306FFFB1CE4F8F0FDA9F8CD" authorityName="Finnegan" authorityYear="1931" box="[205,520,1794,1820]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="275" phylum="Arthropoda" rank="species" species="crassimanus">
<emphasis id="B922EAF7E306FFFB1CE4F8F0FDA9F8CD" box="[205,520,1794,1820]" italics="true" pageId="8" pageNumber="275">Ptychognathus crassimanus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C564D66E306FFFB1E89F8F1FC97F8CD" authorityName="Rathbun" authorityYear="1907" box="[672,822,1795,1820]" class="Malacostraca" family="Varunidae" genus="Ptychognathus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="275" phylum="Arthropoda" rank="species" species="easteranus">
<emphasis id="B922EAF7E306FFFB1E89F8F1FC97F8CD" box="[672,822,1795,1820]" italics="true" pageId="8" pageNumber="275">P. easteranus</emphasis>
</taxonomicName>
by its stouter P5 dactylus (2.53.0 as long as wide vs 3.03.9) and its propodus (0.71.1 times as long as dactylus vs 0.91.4).
</paragraph>
<paragraph id="8BE936E5E306FFFB1CEEF8B9FDD0F859" blockId="8.[151,1437,1218,1929]" pageId="8" pageNumber="275">
For the present study, the
<typeStatus id="54ED8847E306FFFB1DDFF8B9FD87F8B4" box="[502,550,1867,1893]" pageId="8" pageNumber="275">type</typeStatus>
specimen preserved in the Natural History Museum, London (Catalogue no. NHM 1932.12.19.221) was not examined.
</paragraph>
</subSubSection>
</treatment>
</document>