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<document id="7C3C062D775F3EBC46142DB839006B83" ID-CLB-Dataset="25263" ID-DOI="10.11646/zootaxa.4683.4.1" ID-GBIF-Dataset="48496621-de1a-4490-b866-1ae69d81ecbc" ID-ISSN="1175-5326" ID-Zenodo-Dep="3479609" ID-ZooBank="982FE3FB-5610-44A1-AAE3-659B28F45307" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1570688900916" checkinUser="plazi" docAuthor="Lee, Chi-Feng &amp; Bezdĕk, Jan" docDate="2019" docId="03FF87D6FF8E4451FF6BAD5AFCD9F836" docLanguage="en" docName="zootaxa.4683.4.1.pdf" docOrigin="Zootaxa 4683 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Theopea sabahensis Lee &amp; Bezdĕk 2019, sp. nov." docType="treatment" docVersion="15" lastPageNumber="476" masterDocId="FFC6FFAEFF974448FFFCAF12FFAEFFA1" masterDocTitle="Revision of the genus Theopea Baly (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland and the Philippines: Redefinition of the genus" masterLastPageNumber="507" masterPageNumber="451" pageNumber="476" updateTime="1698758705207" updateUser="ExternalLinkService">
<mods:mods id="B3F373F91C1B9A1E9E5D81571E129B1E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="77EC64804DC09FDBC068E403EFD494AB">
<mods:title id="F6414ED899DA142F73E44D75D547982C">Revision of the genus Theopea Baly (Coleoptera: Chrysomelidae: Galerucinae) in Sundaland and the Philippines: Redefinition of the genus</mods:title>
</mods:titleInfo>
<mods:name id="DA68DED64307555BAC2B3B751EB87504" type="personal">
<mods:role id="89E8D3CF6A0A02EFF86254E3EBFAAF5A">
<mods:roleTerm id="957F5B74B8325C546EBBDB21BFA5A369">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9173D04C50460581951A787EAD2B21B8">Lee, Chi-Feng</mods:namePart>
</mods:name>
<mods:name id="80C563AD865C1AF4168C5E9CFEB79D8F" type="personal">
<mods:role id="E07EA341C242C052CA57540607412D12">
<mods:roleTerm id="295DFA63FB6C9F57AE8F445D543D48D9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4F8547F170891EF3FD7722DC99D26EF9">Bezdĕk, Jan</mods:namePart>
</mods:name>
<mods:typeOfResource id="610F6157982E74B67223731B83808BCB">text</mods:typeOfResource>
<mods:relatedItem id="E588E3F8CB3C14E9260E98470AA350E8" type="host">
<mods:titleInfo id="09BF0633D4EE70052EA91F3B133A7127">
<mods:title id="F48531A167BD5E52471A943ED6DA0E93">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DA261060717914ED5F9D193914C185A8">
<mods:date id="85C65F746C500177E14EDDB815632752">2019</mods:date>
<mods:detail id="18F0489A159B742ED24CCE144EEA6169" type="pubDate">
<mods:number id="B2FF3E1466B0923CCB714B7BDF8B4BBD">2019-10-10</mods:number>
</mods:detail>
<mods:detail id="8B25534D470EA465DE1E82CA219C8DE5" type="volume">
<mods:number id="C946D559329B787B1DCF047154D0185B">4683</mods:number>
</mods:detail>
<mods:detail id="12E676B1F9DC36327E7193C7B71639A5" type="issue">
<mods:number id="52D8B9B0D54CBA3C1DBCC0866EE22E87">4</mods:number>
</mods:detail>
<mods:extent id="BB8065AE403B44CADB04C16557F16A8E" unit="page">
<mods:start id="64BBBAC372EE0AD7563432FE1AD67C67">451</mods:start>
<mods:end id="2FD7C885DED103E30C02D26BF6C99FD1">507</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="CD29C45CC3011FF44A81075E4C97C519">journal article</mods:classification>
<mods:identifier id="D81C7C33CA7AEC0D2696159048A91482" type="CLB-Dataset">25263</mods:identifier>
<mods:identifier id="FFD90A99A8B48D73700419CE0180B9F4" type="DOI">10.11646/zootaxa.4683.4.1</mods:identifier>
<mods:identifier id="1BDDC1AAD478E667DF168B03FF1C13D5" type="GBIF-Dataset">48496621-de1a-4490-b866-1ae69d81ecbc</mods:identifier>
<mods:identifier id="DD16DA53D431A9375C08D403672D35F8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="8568D5B90A1818C14DC6BEF7F9000FF5" type="Zenodo-Dep">3479609</mods:identifier>
<mods:identifier id="B0ABA6DD91E037D852A8D7AC9B876C5B" type="ZooBank">982FE3FB-5610-44A1-AAE3-659B28F45307</mods:identifier>
</mods:mods>
<treatment id="03FF87D6FF8E4451FF6BAD5AFCD9F836" ID-DOI="http://doi.org/10.5281/zenodo.4505256" ID-GBIF-Taxon="159690586" ID-Zenodo-Dep="4505256" LSID="urn:lsid:plazi:treatment:03FF87D6FF8E4451FF6BAD5AFCD9F836" httpUri="http://treatment.plazi.org/id/03FF87D6FF8E4451FF6BAD5AFCD9F836" lastPageNumber="476" pageId="25" pageNumber="476">
<subSubSection id="C34C654BFF8E4451FF6BAD5AFE5EFDC2" box="[151,496,584,611]" pageId="25" pageNumber="476" type="nomenclature">
<paragraph id="8BE936C0FF8E4451FF6BAD5AFE5EFDC2" blockId="25.[151,496,584,647]" box="[151,496,584,611]" pageId="25" pageNumber="476">
<heading id="D0A181ACFF8E4451FF6BAD5AFE5EFDC2" bold="true" box="[151,496,584,611]" fontSize="11" level="1" pageId="25" pageNumber="476" reason="1">
<emphasis id="B922EAD2FF8E4451FF6BAD5AFE5EFDC2" bold="true" box="[151,496,584,611]" pageId="25" pageNumber="476">
<taxonomicName id="4C564D43FF8E4451FF6BAD5AFE25FDC2" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[151,395,584,611]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="sabahensis" status="sp. nov.">
<emphasis id="B922EAD2FF8E4451FF6BAD5AFE25FDC2" bold="true" box="[151,395,584,611]" italics="true" pageId="25" pageNumber="476">Theopea sabahensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21157A9FF8E4451FE6EAD5BFE5EFDC2" box="[402,496,585,611]" pageId="25" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C654BFF8E4451FF6BAD7EFB2AF9D6" pageId="25" pageNumber="476" type="description">
<paragraph id="8BE936C0FF8E4451FF6BAD7EFEDFFD26" blockId="25.[151,496,584,647]" box="[151,369,620,647]" pageId="25" pageNumber="476">
(
<figureCitation id="136D2A45FF8E4451FF63AD7EFF5FFD26" box="[159,241,620,647]" captionStart="FIGURE 11" captionStartId="23.[151,250,1939,1964]" captionTargetBox="[275,1333,594,1918]" captionTargetId="figure@23.[228,1359,589,1920]" captionTargetPageId="23" captionText="FIGURE 11. Habitus of Theopea lui sp. nov. and T. sabahensis sp. nov. 11A. T. lui sp. nov., male, dorsal view; 11B. Same, ventral view; 11C. T. lui sp. nov., female, dorsal view; 11D. T. sabahensis sp. nov., male, dorsal view; 11E. Same, ventral view; 11F. T. sabahensis sp. nov., female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3479631" httpUri="https://zenodo.org/record/3479631/files/figure.png" pageId="25" pageNumber="476">Figs 11</figureCitation>
D–11F, 13)
</paragraph>
<paragraph id="8BE936C0FF8E4451FF6BADA7FBB6FC23" blockId="25.[151,1437,692,1943]" pageId="25" pageNumber="476">
<emphasis id="B922EAD2FF8E4451FF6BADA7FEE5FD6E" bold="true" box="[151,331,692,719]" pageId="25" pageNumber="476">Types (n= 74).</emphasis>
<materialsCitation id="3B3E3C9DFF8E4451FEABADA6FDE6FD53" ID-GBIF-Occurrence="2425473755" collectingDate="2005-06" collectionCode="BMNH" collectorName="H. Takano &amp; T. Owen-Edmunds." country="Malaysia" location="Lower Kinabatangan" municipality="Near Gum Gum" pageId="25" pageNumber="476" specimenCount="1" specimenCount-male="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="54ED8862FF8E4451FEABADA6FE11FD6F" box="[343,447,692,718]" pageId="25" pageNumber="476" type="holotype">Holotype</typeStatus>
<specimenCount id="9D50FD49FF8E4451FE36ADA7FE71FD6E" box="[458,479,693,719]" pageId="25" pageNumber="476" type="male">♂</specimenCount>
(
<collectionCode id="ED47AE05FF8E4451FE0FADA7FDE3FD6E" box="[499,589,693,719]" pageId="25" pageNumber="476">BMNH</collectionCode>
),
<emphasis id="B922EAD2FF8E4451FD98ADA6FD52FD6E" bold="true" box="[612,764,692,719]" pageId="25" pageNumber="476">
<collectingCountry id="F3417650FF8E4451FD98ADA6FD52FD6E" box="[612,764,692,719]" name="Malaysia" pageId="25" pageNumber="476">MALAYSIA</collectingCountry>
</emphasis>
.
<collectingRegion id="4992F822FF8E4451FCF3ADA6FCF9FD6F" box="[783,855,692,718]" country="Malaysia" name="Sabah" pageId="25" pageNumber="476">Sabah</collectingRegion>
.
<collectingMunicipality id="6B8DACBAFF8E4451FC9AADA7FB88FD6E" box="[870,1062,693,719]" pageId="25" pageNumber="476">Near Gum Gum</collectingMunicipality>
,
<location id="8E89601BFF8E4451FBCAADA7FA87FD6F" LSID="urn:lsid:plazi:treatment:03FF87D6FF8E4451FF6BAD5AFCD9F836:8E89601BFF8E4451FBCAADA7FA87FD6F" box="[1078,1321,692,719]" country="Malaysia" municipality="Near Gum Gum" name="Lower Kinabatangan" pageId="25" pageNumber="476" stateProvince="Sabah">Lower Kinabatangan</location>
,
<date id="FFE81000FF8E4451FAC5ADA6FA36FD6E" box="[1337,1432,692,719]" pageId="25" pageNumber="476" value="2005-06">
<collectingDate id="EFACE9E8FF8E4451FAC5ADA6FA36FD6E" box="[1337,1432,692,719]" pageId="25" pageNumber="476" value="2005-06">VI.2005</collectingDate>
</date>
, leg.
<collectorName id="26A35316FF8E4451FF35ADCBFEECFD53" box="[201,322,728,755]" pageId="25" pageNumber="476">H. Takano</collectorName>
&amp;
<collectorName id="26A35316FF8E4451FE95ADCBFDE6FD53" box="[361,584,728,755]" pageId="25" pageNumber="476">T. Owen-Edmunds.</collectorName>
</materialsCitation>
<materialsCitation id="3B3E3C9DFF8E4451FDADADCBFEC4FCB7" ID-GBIF-Occurrence="2425473733" collectingDate="2005-06" collectionCode="BMNH" collectorName="H. Takano &amp; T. Owen-Edmunds." country="Indonesia" location="Lower Kinabatangan" municipality="Near Gum Gum" pageId="25" pageNumber="476" specimenCount="41" specimenCount-female="23" specimenCount-male="18" stateProvince="Kalimantan Selatan" typeStatus="paratype">
<typeStatus id="54ED8862FF8E4451FDADADCBFD6CFD52" box="[593,706,729,755]" pageId="25" pageNumber="476" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="9D50FD49FF8E4451FD31ADCBFCB8FD52" box="[717,790,729,755]" pageId="25" pageNumber="476" type="male">18♂♂</specimenCount>
,
<specimenCount id="9D50FD49FF8E4451FCDFADCBFCC7FD52" box="[803,873,729,755]" pageId="25" pageNumber="476" type="female">23♀♀</specimenCount>
(
<collectionCode id="ED47AE05FF8E4451FC86ADCBFC7AFD52" box="[890,980,729,755]" pageId="25" pageNumber="476">BMNH</collectionCode>
), same data as holotype;
<emphasis id="B922EAD2FF8E4451FB0BADCAFA38FD52" bold="true" box="[1271,1430,728,755]" pageId="25" pageNumber="476">
<collectingCountry id="F3417650FF8E4451FB0BADCAFA38FD52" box="[1271,1430,728,755]" name="Indonesia" pageId="25" pageNumber="476">INDONESIA</collectingCountry>
</emphasis>
.
<collectingRegion id="4992F822FF8E4451FF6BADEEFEC4FCB7" box="[151,362,764,790]" country="Indonesia" name="Kalimantan Selatan" pageId="25" pageNumber="476">South Kalimantan</collectingRegion>
</materialsCitation>
:
<materialsCitation id="3B3E3C9DFF8E4451FE8BADEFFA36FCB7" ID-GBIF-Occurrence="2425473743" box="[375,1432,764,791]" collectingDate="1997-09-03" collectingDateMax="1997-09-22" collectingDateMin="1997-09-03" collectionCode="JBCB" collectorName="S. Jakl" country="Indonesia" location="Kandagan distr." pageId="25" pageNumber="476" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D50FD49FF8E4451FE8BADEFFE32FCB6" box="[375,412,765,791]" pageId="25" pageNumber="476" type="male">1♂</specimenCount>
,
<specimenCount id="9D50FD49FF8E4451FE57ADEFFE4DFCB6" box="[427,483,765,791]" pageId="25" pageNumber="476" type="female">2♀♀</specimenCount>
(
<collectionCode id="ED47AE05FF8E4451FE0AADEFFD90FCB6" box="[502,574,765,791]" pageId="25" pageNumber="476">JBCB</collectionCode>
),
<location id="8E89601BFF8E4451FDA8ADEEFCA2FCB7" LSID="urn:lsid:plazi:treatment:03FF87D6FF8E4451FF6BAD5AFCD9F836:8E89601BFF8E4451FDA8ADEEFCA2FCB7" box="[596,780,764,790]" country="Indonesia" name="Kandagan distr." pageId="25" pageNumber="476">Kandagan distr.</location>
,
<locationDeviation id="9A5B2FE2FF8E4451FCE6ADEFFBB1FCB7" box="[794,1055,764,791]" location="Loksado" pageId="25" pageNumber="476">
<quantity id="4CAE9B25FF8E4451FCE6ADEFFCCAFCB7" box="[794,868,764,791]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="25" pageNumber="476" unit="km" value="17.0">17 km</quantity>
NE of Loksado
</locationDeviation>
vill.,
<date id="FFE81000FF8E4451FB9BADEFFAABFCB6" box="[1127,1285,765,791]" pageId="25" pageNumber="476" value="1997-09-03" valueMax="1997-09-22" valueMin="1997-09-03">
<collectingDate id="EFACE9E8FF8E4451FB9BADEFFAABFCB6" box="[1127,1285,765,791]" pageId="25" pageNumber="476" value="1997-09-03" valueMax="1997-09-22" valueMin="1997-09-03">3–22.IX.1997</collectingDate>
</date>
, leg.
<collectorName id="26A35316FF8E4451FABBADEFFA36FCB7" box="[1351,1432,764,791]" pageId="25" pageNumber="476">S. Jákl</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3E3C9DFF8E4451FF6BAC32FF07FCFE" ID-GBIF-Occurrence="2425473758" collectingDate="1927-07-22" collectingDateMax="1927-08-09" collectingDateMin="1927-07-22" collectionCode="BMNH" country="Malaysia" location="Nr. Sandakan" municipality="Bettotan" pageId="25" pageNumber="476" specimenCount="8" specimenCount-female="4" specimenCount-male="4" stateProvince="Sabah" typeStatus="paratype">
<emphasis id="B922EAD2FF8E4451FF6BAC32FE81FC9A" bold="true" box="[151,303,800,827]" pageId="25" pageNumber="476">
<collectingCountry id="F3417650FF8E4451FF6BAC32FE81FC9A" box="[151,303,800,827]" name="Malaysia" pageId="25" pageNumber="476">MALAYSIA</collectingCountry>
</emphasis>
.
<collectingRegion id="4992F822FF8E4451FEC3AC32FE26FC9B" box="[319,392,800,826]" country="Malaysia" name="Sabah" pageId="25" pageNumber="476">Sabah</collectingRegion>
:
<specimenCount id="9D50FD49FF8E4451FE68AC33FE61FC9A" box="[404,463,801,827]" pageId="25" pageNumber="476" type="male">4♂♂</specimenCount>
,
<specimenCount id="9D50FD49FF8E4451FE20AC33FDBAFC9A" box="[476,532,801,827]" pageId="25" pageNumber="476" type="female">4♀♀</specimenCount>
(
<collectionCode id="ED47AE05FF8E4451FDD9AC33FDD1FC9A" box="[549,639,801,827]" pageId="25" pageNumber="476">BMNH</collectionCode>
),
<collectingMunicipality id="6B8DACBAFF8E4451FD6FAC33FD5BFC9A" box="[659,757,801,827]" pageId="25" pageNumber="476">Bettotan</collectingMunicipality>
,
<location id="8E89601BFF8E4451FCFEAC33FC31FC9B" LSID="urn:lsid:plazi:treatment:03FF87D6FF8E4451FF6BAD5AFCD9F836:8E89601BFF8E4451FCFEAC33FC31FC9B" box="[770,927,800,827]" country="Malaysia" municipality="Bettotan" name="Nr. Sandakan" pageId="25" pageNumber="476" stateProvince="Sabah">Nr. Sandakan</location>
,
<date id="FFE81000FF8E4451FC50AC33FB3CFC9A" box="[940,1170,801,827]" pageId="25" pageNumber="476" value="1927-07-22" valueMax="1927-08-09" valueMin="1927-07-22">
<collectingDate id="EFACE9E8FF8E4451FC50AC33FB3CFC9A" box="[940,1170,801,827]" pageId="25" pageNumber="476" value="1927-07-22" valueMax="1927-08-09" valueMin="1927-07-22">22.VII.–9.VIII.1927</collectingDate>
</date>
, leg. C. B. K. &amp; H. M. P
</materialsCitation>
.;
<materialsCitation id="3B3E3C9DFF8E4451FF47AC57FB6FFCFE" ID-GBIF-Occurrence="2425473718" box="[187,1217,836,863]" collectingDate="1927-07-09" collectingDateMax="1927-07-17" collectingDateMin="1927-07-09" collectionCode="BMNH" country="Malaysia" location="Sandakan" municipality="Samawang" pageId="25" pageNumber="476" specimenCount="18" specimenCount-female="7" specimenCount-male="11" typeStatus="paratype">
<specimenCount id="9D50FD49FF8E4451FF47AC57FEADFCFE" box="[187,259,837,863]" pageId="25" pageNumber="476" type="male">11♂♂</specimenCount>
,
<specimenCount id="9D50FD49FF8E4451FEECAC57FEE6FCFE" box="[272,328,837,863]" pageId="25" pageNumber="476" type="female">7♀♀</specimenCount>
(
<collectionCode id="ED47AE05FF8E4451FEA5AC57FE1DFCFE" box="[345,435,837,863]" pageId="25" pageNumber="476">BMNH</collectionCode>
),
<collectingMunicipality id="6B8DACBAFF8E4451FE3BAC57FDEBFCFE" box="[455,581,837,863]" pageId="25" pageNumber="476">Samawang</collectingMunicipality>
, nr.
<location id="8E89601BFF8E4451FD8BAC56FD49FCFF" LSID="urn:lsid:plazi:treatment:03FF87D6FF8E4451FF6BAD5AFCD9F836:8E89601BFF8E4451FD8BAC56FD49FCFF" box="[631,743,836,862]" country="Malaysia" municipality="Samawang" name="Sandakan" pageId="25" pageNumber="476">Sandakan</location>
,
<date id="FFE81000FF8E4451FD08AC57FC32FCFE" box="[756,924,837,863]" pageId="25" pageNumber="476" value="1927-07-09" valueMax="1927-07-17" valueMin="1927-07-09">
<collectingDate id="EFACE9E8FF8E4451FD08AC57FC32FCFE" box="[756,924,837,863]" pageId="25" pageNumber="476" value="1927-07-09" valueMax="1927-07-17" valueMin="1927-07-09">9–17.VII.1927</collectingDate>
</date>
, leg. C. B. K. &amp; H. M. P
</materialsCitation>
.;
<materialsCitation id="3B3E3C9DFF8E4451FB2FAC57FE26FC23" ID-GBIF-Occurrence="2425473807" collectionCode="USNM" collectorName="Baker" country="Malaysia" location="Sandakan" pageId="25" pageNumber="476" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D50FD49FF8E4451FB2FAC57FB59FCFE" box="[1235,1271,837,863]" pageId="25" pageNumber="476" type="male">1♂</specimenCount>
,
<specimenCount id="9D50FD49FF8E4451FAFAAC57FA87FCFE" box="[1286,1321,837,863]" pageId="25" pageNumber="476" type="female">1♀</specimenCount>
(
<collectionCode id="ED47AE05FF8E4451FAC5AC57FA3EFCFE" box="[1337,1424,837,863]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="25" pageNumber="476" type="Museum">USNM</collectionCode>
),
<location id="8E89601BFF8E4451FF6BAC7AFEA9FC23" LSID="urn:lsid:plazi:treatment:03FF87D6FF8E4451FF6BAD5AFCD9F836:8E89601BFF8E4451FF6BAC7AFEA9FC23" box="[151,263,872,898]" country="Malaysia" name="Sandakan" pageId="25" pageNumber="476">Sandakan</location>
, leg.
<collectorName id="26A35316FF8E4451FEBEAC7AFE26FC23" box="[322,392,872,898]" pageId="25" pageNumber="476">Baker</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3E3C9DFF8E4451FE68AC7BFBB6FC23" ID-GBIF-Occurrence="2425473753" box="[404,1048,872,899]" collectingDate="1983-08-05" collectingDateMax="1983-08-11" collectingDateMin="1983-08-05" collectionCode="EUMJ" collectorName="N. Ohbayashi." country="Malaysia" location="Sepilok" pageId="25" pageNumber="476" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D50FD49FF8E4451FE68AC7BFE19FC22" box="[404,439,873,899]" pageId="25" pageNumber="476" type="male">1♂</specimenCount>
(
<collectionCode id="ED47AE05FF8E4451FE3AAC7BFDBBFC22" box="[454,533,873,899]" country="Japan" httpUri="http://grbio.org/cool/0wyw-sfpi" name="Ehime University" pageId="25" pageNumber="476">EUMJ</collectionCode>
),
<location id="8E89601BFF8E4451FDD4AC7AFD2EFC23" LSID="urn:lsid:plazi:treatment:03FF87D6FF8E4451FF6BAD5AFCD9F836:8E89601BFF8E4451FDD4AC7AFD2EFC23" box="[552,640,872,898]" country="Malaysia" name="Sepilok" pageId="25" pageNumber="476">Sepilok</location>
,
<date id="FFE81000FF8E4451FD77AC7AFC94FC22" box="[651,826,872,899]" pageId="25" pageNumber="476" value="1983-08-05" valueMax="1983-08-11" valueMin="1983-08-05">
<collectingDate id="EFACE9E8FF8E4451FD77AC7AFC94FC22" box="[651,826,872,899]" pageId="25" pageNumber="476" value="1983-08-05" valueMax="1983-08-11" valueMin="1983-08-05">5–11.VIII.1983</collectingDate>
</date>
, leg.
<collectorName id="26A35316FF8E4451FC8BAC7BFBB6FC23" box="[887,1048,872,899]" pageId="25" pageNumber="476">N. Ohbayashi.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BE936C0FF8E4451FF3BAC9EFDC0F9F3" blockId="25.[151,1437,692,1943]" pageId="25" pageNumber="476">
<emphasis id="B922EAD2FF8E4451FF3BAC9EFEF7FC06" bold="true" box="[199,345,908,935]" pageId="25" pageNumber="476">Description.</emphasis>
Length 6.0–
<quantity id="4CAE9B25FF8E4451FE12AC9EFDE7FC06" box="[494,585,908,935]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="25" pageNumber="476" unit="mm" value="6.7">6.7 mm</quantity>
, width
<quantity id="4CAE9B25FF8E4451FD5DAC9FFC80FC06" box="[673,814,909,935]" metricMagnitude="-3" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="25" pageNumber="476" unit="mm" value="2.55" valueMax="2.7" valueMin="2.4">2.4–2.7 mm</quantity>
. General color (
<figureCitation id="136D2A45FF8E4451FC17AC9EFBEEFC06" box="[1003,1088,908,935]" captionStart="FIGURE 11" captionStartId="23.[151,250,1939,1964]" captionTargetBox="[275,1333,594,1918]" captionTargetId="figure@23.[228,1359,589,1920]" captionTargetPageId="23" captionText="FIGURE 11. Habitus of Theopea lui sp. nov. and T. sabahensis sp. nov. 11A. T. lui sp. nov., male, dorsal view; 11B. Same, ventral view; 11C. T. lui sp. nov., female, dorsal view; 11D. T. sabahensis sp. nov., male, dorsal view; 11E. Same, ventral view; 11F. T. sabahensis sp. nov., female, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3479631" httpUri="https://zenodo.org/record/3479631/files/figure.png" pageId="25" pageNumber="476">Figs 11</figureCitation>
D–11F) metallic green; antennomeres I–III greenish black, IV–VIII black, IX–XI white or pale yellow. Antennae filiform (
<figureCitation id="136D2A45FF8E4451FB42ACA2FA8BFC6A" box="[1214,1317,944,971]" captionStart="FIGURE 13" captionStartId="26.[151,250,1929,1954]" captionTargetBox="[158,1428,211,1802]" captionTargetId="figure@26.[151,1436,196,1812]" captionTargetPageId="26" captionText="FIGURE 13. Diagnostic characters of Theopea sabahensis sp. nov. 13A. Antenna, male; 13B. Antenna, female; 13C. Aedeagus, dorsal view; 13D. Aedeagus, lateral view; 13E. Aedeagus, ventral view; 13F. Abdominal ventrite VIII; 13G. Gonocoxae; 13H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479635" httpUri="https://zenodo.org/record/3479635/files/figure.png" pageId="25" pageNumber="476">Fig. 13A</figureCitation>
) in males, length ratios of antennomeres I–XI 1.0: 0.3: 0.8: 0.9: 0.9: 0.9: 0.9: 0.8: 0.8: 0.7: 0.9, length to width ratios of antennomeres I–XI 3.3: 1.3: 3.0: 3.5: 3.1: 3.1: 3.1: 3.0: 3.1: 3.0: 5.6; shorter in females (
<figureCitation id="136D2A45FF8E4451FB66ACEAFB50FBB2" box="[1178,1278,1016,1043]" captionStart="FIGURE 13" captionStartId="26.[151,250,1929,1954]" captionTargetBox="[158,1428,211,1802]" captionTargetId="figure@26.[151,1436,196,1812]" captionTargetPageId="26" captionText="FIGURE 13. Diagnostic characters of Theopea sabahensis sp. nov. 13A. Antenna, male; 13B. Antenna, female; 13C. Aedeagus, dorsal view; 13D. Aedeagus, lateral view; 13E. Aedeagus, ventral view; 13F. Abdominal ventrite VIII; 13G. Gonocoxae; 13H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479635" httpUri="https://zenodo.org/record/3479635/files/figure.png" pageId="25" pageNumber="476">Fig. 13B</figureCitation>
), length ratios of antennomeres I–XI 1.0: 0.3: 0.7: 0.8: 0.8: 0.7: 0.7: 0.7: 0.7: 0.6: 0.8, length to width ratios of antennomeres I–XI 3.3: 1.5: 3.1: 3.4: 3.4: 2.9: 2.7: 2.7: 3.1: 2.8: 3.8. Elytra elongate and parallel-sided, 1.8x longer than wide; disc with dense, coarse punctures, arranged into longitudinal rows, with one weak longitudinal ridge between two longitudinal rows of punctures, three or four ridges near suture basally abbreviated. Tarsomeres I of front legs swollen in males; subparallel in females. Aedeagus (
<figureCitation id="136D2A45FF8E4451FD0AABBEFCE2FB66" box="[758,844,1196,1223]" captionStart="FIGURE 13" captionStartId="26.[151,250,1929,1954]" captionTargetBox="[158,1428,211,1802]" captionTargetId="figure@26.[151,1436,196,1812]" captionTargetPageId="26" captionText="FIGURE 13. Diagnostic characters of Theopea sabahensis sp. nov. 13A. Antenna, male; 13B. Antenna, female; 13C. Aedeagus, dorsal view; 13D. Aedeagus, lateral view; 13E. Aedeagus, ventral view; 13F. Abdominal ventrite VIII; 13G. Gonocoxae; 13H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479635" httpUri="https://zenodo.org/record/3479635/files/figure.png" pageId="25" pageNumber="476">Figs 13</figureCitation>
C–13E) extremely slender, 10.1x longer than wide; sides widest at middle, gradually narrowed towards basal 1/4, abruptly narrowed subapically, apex truncate; tectum well sclerotized, basally broadened; moderately curved in lateral view; apex with short slit from apex to apical 1/10, basal notch closed except for apex; triangular sclerites absent; internal sac with one median, elongate sclerite, parallel-sided, apically tapering, with one longitudinal row of long setae along sides of median sclerite, with one pair of lateral clusters of stout setae located subapically, and one pair of elongate sclerites, apices acute. Gonocoxae (
<figureCitation id="136D2A45FF8E4451FA93AA72FF66FA3E" captionStart="FIGURE 13" captionStartId="26.[151,250,1929,1954]" captionTargetBox="[158,1428,211,1802]" captionTargetId="figure@26.[151,1436,196,1812]" captionTargetPageId="26" captionText="FIGURE 13. Diagnostic characters of Theopea sabahensis sp. nov. 13A. Antenna, male; 13B. Antenna, female; 13C. Aedeagus, dorsal view; 13D. Aedeagus, lateral view; 13E. Aedeagus, ventral view; 13F. Abdominal ventrite VIII; 13G. Gonocoxae; 13H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479635" httpUri="https://zenodo.org/record/3479635/files/figure.png" pageId="25" pageNumber="476">Fig. 13G</figureCitation>
) elongate, widest at apical 1/6, both gonocoxae combined together from basal 1/7 to apical 1/6; apices narrowly rounded, each gonocoxa with eight setae along outer margin from apex to apical 1/6; with one pair of short lateral processes at basal 2/5. Ventrite VIII (
<figureCitation id="136D2A45FF8E4451FDC5AADEFD35FA46" box="[569,667,1484,1511]" captionStart="FIGURE 13" captionStartId="26.[151,250,1929,1954]" captionTargetBox="[158,1428,211,1802]" captionTargetId="figure@26.[151,1436,196,1812]" captionTargetPageId="26" captionText="FIGURE 13. Diagnostic characters of Theopea sabahensis sp. nov. 13A. Antenna, male; 13B. Antenna, female; 13C. Aedeagus, dorsal view; 13D. Aedeagus, lateral view; 13E. Aedeagus, ventral view; 13F. Abdominal ventrite VIII; 13G. Gonocoxae; 13H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479635" httpUri="https://zenodo.org/record/3479635/files/figure.png" pageId="25" pageNumber="476">Fig. 13F</figureCitation>
) elongate and well sclerotized; disc with several long setae laterally and near apical margin, and with dense, short setae along apical margin; spiculum extremely slender. Receptacle of spermatheca (
<figureCitation id="136D2A45FF8E4451FEC8A906FE35F98E" box="[308,411,1556,1583]" captionStart="FIGURE 13" captionStartId="26.[151,250,1929,1954]" captionTargetBox="[158,1428,211,1802]" captionTargetId="figure@26.[151,1436,196,1812]" captionTargetPageId="26" captionText="FIGURE 13. Diagnostic characters of Theopea sabahensis sp. nov. 13A. Antenna, male; 13B. Antenna, female; 13C. Aedeagus, dorsal view; 13D. Aedeagus, lateral view; 13E. Aedeagus, ventral view; 13F. Abdominal ventrite VIII; 13G. Gonocoxae; 13H. Spermatheca." figureDoi="http://doi.org/10.5281/zenodo.3479635" httpUri="https://zenodo.org/record/3479635/files/figure.png" pageId="25" pageNumber="476">Fig. 13H</figureCitation>
) strongly swollen; pump slender and strongly curved; proximal spermathecal duct deeply inserted into receptacle, narrow and short.
</paragraph>
<paragraph id="8BE936C0FF8E4451FF3BA94EFB2AF9D6" blockId="25.[151,1437,692,1943]" box="[199,1156,1628,1655]" pageId="25" pageNumber="476">
<emphasis id="B922EAD2FF8E4451FF3BA94EFEE4F9D6" bold="true" box="[199,330,1628,1655]" pageId="25" pageNumber="476">Variations.</emphasis>
Specimens from
<collectingRegion id="4992F822FF8E4451FDF2A94EFD74F9D7" box="[526,730,1628,1654]" country="Indonesia" name="Kalimantan Selatan" pageId="25" pageNumber="476">South Kalimantan</collectingRegion>
have four apical white antennomeres.
</paragraph>
</subSubSection>
<subSubSection id="C34C654BFF8E4451FF3BA992FA32F8EE" pageId="25" pageNumber="476" type="diagnosis">
<paragraph id="8BE936C0FF8E4451FF3BA992FA32F8EE" blockId="25.[151,1437,692,1943]" pageId="25" pageNumber="476">
<emphasis id="B922EAD2FF8E4451FF3BA992FEECF93A" bold="true" box="[199,322,1664,1691]" pageId="25" pageNumber="476">Diagnosis.</emphasis>
Members of this new species can be separated from others by the metallic green bodies (metallic blue in others), three apical white apical antennomeres (four apical white apical antennomeres in
<emphasis id="B922EAD2FF8E4451FB09A9B7FA9FF91F" box="[1269,1329,1700,1726]" italics="true" pageId="25" pageNumber="476">T. lui</emphasis>
<emphasis id="B922EAD2FF8E4451FAC6A9B7FA3BF91E" bold="true" box="[1338,1429,1701,1727]" pageId="25" pageNumber="476">sp. nov.</emphasis>
, white antennomeres IV–XI in
<taxonomicName id="4C564D43FF8E4451FE1BA9DBFD9DF943" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[487,563,1737,1762]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="guoi" status="sp. nov.">
<emphasis id="B922EAD2FF8E4451FE1BA9DBFD9DF943" box="[487,563,1737,1762]" italics="true" pageId="25" pageNumber="476">T. guoi</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FF8E4451FDC5A9DBFD3FF942" bold="true" box="[569,657,1737,1763]" pageId="25" pageNumber="476">
<taxonomicNameLabel id="A21157A9FF8E4451FDC5A9DBFD3FF942" box="[569,657,1737,1763]" pageId="25" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). The aedeagi of males of
<taxonomicName id="4C564D43FF8E4451FC48A9DBFBE7F943" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[948,1097,1736,1762]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="sabahensis" status="sp. nov.">
<emphasis id="B922EAD2FF8E4451FC48A9DBFBE7F943" box="[948,1097,1736,1762]" italics="true" pageId="25" pageNumber="476">T. sabahensis</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FF8E4451FBB3A9DBFB09F942" bold="true" box="[1103,1191,1737,1763]" pageId="25" pageNumber="476">
<taxonomicNameLabel id="A21157A9FF8E4451FBB3A9DBFB09F942" box="[1103,1191,1737,1763]" pageId="25" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are similar to those of
<emphasis id="B922EAD2FF8E4451FF6BA9FFFF7FF8A7" box="[151,209,1772,1798]" italics="true" pageId="25" pageNumber="476">T. lui</emphasis>
<emphasis id="B922EAD2FF8E4451FF2BA9FFFE9EF8A6" bold="true" box="[215,304,1773,1799]" pageId="25" pageNumber="476">sp. nov.</emphasis>
in possessing apically narrowed and truncate apices (wide apices in
<taxonomicName id="4C564D43FF8E4451FBCCA9FFFB6CF8A7" box="[1072,1218,1772,1798]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="flavipalpis">
<emphasis id="B922EAD2FF8E4451FBCCA9FFFB6CF8A7" box="[1072,1218,1772,1798]" italics="true" pageId="25" pageNumber="476">T. flavipalpis</emphasis>
</taxonomicName>
, rounded in
<taxonomicName id="4C564D43FF8E4451FAB3A9FFFA32F8A7" authority="Lee &amp; Bezdĕk, 2019" authorityName="Lee &amp; Bezdĕk" authorityYear="2019" box="[1359,1436,1773,1798]" class="Insecta" family="Chrysomelidae" genus="Theopea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="476" phylum="Arthropoda" rank="species" species="guoi" status="sp. nov.">
<emphasis id="B922EAD2FF8E4451FAB3A9FFFA32F8A7" box="[1359,1436,1773,1798]" italics="true" pageId="25" pageNumber="476">T. guoi</emphasis>
</taxonomicName>
<emphasis id="B922EAD2FF8E4451FF6BA803FF5FF88A" bold="true" box="[151,241,1809,1835]" pageId="25" pageNumber="476">
<taxonomicNameLabel id="A21157A9FF8E4451FF6BA803FF5FF88A" box="[151,241,1809,1835]" pageId="25" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and medially widened tectum (apically tapering tectum in others), but differing by the absence of a longitudinal row of hook-like setae near the apex of the internal sac (presence of such hook-like setae in
<emphasis id="B922EAD2FF8E4451FB0EA827FA82F8EF" box="[1266,1324,1844,1870]" italics="true" pageId="25" pageNumber="476">T. lui</emphasis>
<emphasis id="B922EAD2FF8E4451FACEA827FA22F8EE" bold="true" box="[1330,1420,1845,1871]" pageId="25" pageNumber="476">sp. nov.</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C34C654BFF8E4451FF3BA84BFC2DF8D3" box="[199,899,1880,1907]" pageId="25" pageNumber="476" type="etymology">
<paragraph id="8BE936C0FF8E4451FF3BA84BFC2DF8D3" blockId="25.[151,1437,692,1943]" box="[199,899,1880,1907]" pageId="25" pageNumber="476">
<emphasis id="B922EAD2FF8E4451FF3BA84BFEE0F8D2" bold="true" box="[199,334,1881,1907]" pageId="25" pageNumber="476">Etymology.</emphasis>
The species name is named after the
<typeStatus id="54ED8862FF8E4451FD0EA84BFC8CF8D2" box="[754,802,1881,1907]" pageId="25" pageNumber="476">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C34C654BFF8E4451FF3BA86EFCD9F836" box="[199,887,1916,1943]" pageId="25" pageNumber="476" type="distribution">
<paragraph id="8BE936C0FF8E4451FF3BA86EFCD9F836" blockId="25.[151,1437,692,1943]" box="[199,887,1916,1943]" pageId="25" pageNumber="476">
<emphasis id="B922EAD2FF8E4451FF3BA86EFECFF836" bold="true" box="[199,353,1916,1943]" pageId="25" pageNumber="476">Distribution.</emphasis>
<collectingCountry id="F3417650FF8E4451FE9BA86EFE76F837" box="[359,472,1916,1942]" name="Indonesia" pageId="25" pageNumber="476">Indonesia</collectingCountry>
:
<collectingRegion id="4992F822FF8E4451FE1FA86EFDC7F837" box="[483,617,1916,1942]" country="Indonesia" name="Kalimantan Utara" pageId="25" pageNumber="476">Kalimantan</collectingRegion>
, East
<collectingCountry id="F3417650FF8E4451FD57A86EFCBCF837" box="[683,786,1916,1942]" name="Malaysia" pageId="25" pageNumber="476">Malaysia</collectingCountry>
(
<collectingRegion id="4992F822FF8E4451FCDDA86EFCC5F836" box="[801,875,1916,1943]" country="Malaysia" name="Sabah" pageId="25" pageNumber="476">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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