139 lines
18 KiB
XML
139 lines
18 KiB
XML
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<mods:title id="F9A88323806864BD01C0E0DDB0EADC24">The centipede family Anopsobiidae new to North America, with the description of a new genus and species and notes on the Henicopidae of North America and the Anopsobiidae of the Northern Hemisphere (Chilopoda, Lithobiomorpha)</mods:title>
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<mods:namePart id="BD33875DBC6185EFFA864D0B77F65090">Shear, William A.</mods:namePart>
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<mods:title id="FF8AA69990BB9C1E13D5D5F894736DFD">Zootaxa</mods:title>
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<mods:date id="B6731E684166D0A518F11191F9256709">2018</mods:date>
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<mods:number id="294CFF391D924CC2BB50A8FA4EDDBF62">4422</mods:number>
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<treatment id="03FF87B7FFDDFF9941EAFEEB1BEDFAD6" ID-DOI="http://doi.org/10.5281/zenodo.5969827" ID-GBIF-Taxon="148671727" ID-Zenodo-Dep="5969827" LSID="urn:lsid:plazi:treatment:03FF87B7FFDDFF9941EAFEEB1BEDFAD6" httpUri="http://treatment.plazi.org/id/03FF87B7FFDDFF9941EAFEEB1BEDFAD6" lastPageNumber="270" pageId="11" pageNumber="270">
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<subSubSection id="C34C652AFFDDFF9941EAFEEB19F3FE8C" box="[151,718,321,347]" pageId="11" pageNumber="270" type="nomenclature">
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<paragraph id="8BE936A1FFDDFF9941EAFEEB19F3FE8C" blockId="11.[151,718,321,347]" box="[151,718,321,347]" pageId="11" pageNumber="270">
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<heading id="D0A181CDFFDDFF9941EAFEEB19F3FE8C" bold="true" box="[151,718,321,347]" fontSize="11" level="1" pageId="11" pageNumber="270" reason="1">
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<taxonomicName id="4C564D22FFDDFF9941EAFEEB19F3FE8C" authority="Williams & Hefner, 1928" authorityName="Williams & Hefner" authorityYear="1928" box="[151,718,321,347]" class="Chilopoda" family="Henicopidae" genus="Buethobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="11" pageNumber="270" phylum="Arthropoda" rank="species" species="huestoni">
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<emphasis id="B922EAB3FFDDFF9941EAFEEB19F3FE8C" bold="true" box="[151,718,321,347]" pageId="11" pageNumber="270">
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<emphasis id="B922EAB3FFDDFF9941EAFEEB1AACFE8C" bold="true" box="[151,401,321,347]" italics="true" pageId="11" pageNumber="270">Buethobius huestoni</emphasis>
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<bibRefCitation id="EFC74B50FFDDFF9940E5FEEB19F3FE8C" box="[408,718,321,347]" pageId="11" pageNumber="270" refId="ref17312" refString="Williams, S. R. & Hefner, R. A. (1928) The millipedes and centipedes of Ohio. Ohio Biological Survey Bulletin, 4, 91 - 146." type="journal article">Williams & Hefner, 1928</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C34C652AFFDDFF9941EAFE221BEDFAD6" pageId="11" pageNumber="270" type="reference_group">
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<paragraph id="8BE936A1FFDDFF9941EAFE221F7AFE49" blockId="11.[151,1095,392,414]" box="[151,1095,392,414]" pageId="11" pageNumber="270">
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<treatmentCitationGroup id="AB46118FFFDDFF9941EAFE221F7AFE49" box="[151,1095,392,414]" pageId="11" pageNumber="270">
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<taxonomicName id="4C564D22FFDDFF9941EAFE2219A0FE49" authority="Williams & Hefner 1928: 139" authorityName="Williams & Hefner" authorityPageNumber="139" authorityYear="1928" box="[151,669,392,414]" class="Chilopoda" family="Henicopidae" genus="Buethobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="11" pageNumber="270" phylum="Arthropoda" rank="species" species="huestoni">
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<emphasis id="B922EAB3FFDDFF9941EAFE221A58FE49" box="[151,357,392,414]" italics="true" pageId="11" pageNumber="270">Buethobius huestoni</emphasis>
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<treatmentCitation id="0AF710B0FFDDFF994016FE2219A0FE49" author="" box="[363,669,392,414]" page="139" pageId="11" pageNumber="270">
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<bibRefCitation id="EFC74B50FFDDFF994016FE221957FE49" box="[363,618,392,414]" pageId="11" pageNumber="270" refId="ref17312" refString="Williams, S. R. & Hefner, R. A. (1928) The millipedes and centipedes of Ohio. Ohio Biological Survey Bulletin, 4, 91 - 146." type="journal article">Williams & Hefner 1928</bibRefCitation>
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: 139
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</taxonomicName>
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;
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<treatmentCitation id="0AF710B0FFDDFF9943DAFE221858FE49" author="" box="[679,869,392,414]" page="44" pageId="11" pageNumber="270">
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<bibRefCitation id="EFC74B50FFDDFF9943DAFE22187FFE49" box="[679,834,392,414]" pageId="11" pageNumber="270" refId="ref16977" refString="Mercurio, R. J. (2010) An annotated catalog of the centipedes (Chilopoda) from the United States of America, Canada and Greenland (1758 - 2008). XLibris Corporation, Bloomington, IN, 559 pp." type="book chapter">Mercurio 2010</bibRefCitation>
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: 44
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</treatmentCitation>
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(complete references)
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</paragraph>
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<paragraph id="8BE936A1FFDDFF9941EAFE601BEDFAD6" blockId="11.[151,1437,458,1281]" pageId="11" pageNumber="270">
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The description is brief, but contains more detail than that of the following species. Nevertheless, key characters go unmentioned. Deep red in color,
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<quantity id="4CAE9B44FFDDFF994378FE45196EFDD1" box="[517,595,495,519]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="270" unit="mm" value="10.0">10 mm</quantity>
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long, “antennae long, more than 30 segments,” coxal pores “2, 3, 3, 2–4, 5, 5, 5.”
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<bibRefCitation id="EFC74B50FFDDFF9941A1FDB81925FDFC" box="[220,536,530,555]" pageId="11" pageNumber="270" refId="ref17312" refString="Williams, S. R. & Hefner, R. A. (1928) The millipedes and centipedes of Ohio. Ohio Biological Survey Bulletin, 4, 91 - 146." type="journal article">Williams and Hefner (1928)</bibRefCitation>
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also state: “No pigmented ocelli; a light spot which resembles a non-pigmented area.” Interestingly, the margins of the coxosternum are said to have no teeth. The descriptions by Williams and Hefner of their
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<taxonomicName id="4C564D22FFDDFF994028FDF11AEEFDA4" ID-CoL="62GTT" authorityName="Chamberlin" authorityYear="1911" box="[341,467,603,627]" class="Chilopoda" family="Henicopidae" genus="Buethobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="11" pageNumber="270" phylum="Arthropoda" rank="genus">
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<emphasis id="B922EAB3FFDDFF994028FDF11AEEFDA4" box="[341,467,603,627]" italics="true" pageId="11" pageNumber="270">Buethobius</emphasis>
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</taxonomicName>
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species are accompanied by two crude drawings, labelled only as “
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<emphasis id="B922EAB3FFDDFF99447AFDF11EA1FDA4" box="[1287,1436,603,627]" italics="true" pageId="11" pageNumber="270">
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<taxonomicName id="4C564D22FFDDFF99447AFDF11EB2FDA4" ID-CoL="62GTT" authorityName="Chamberlin" authorityYear="1911" box="[1287,1423,603,627]" class="Chilopoda" family="Henicopidae" genus="Buethobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="11" pageNumber="270" phylum="Arthropoda" rank="genus">Buethobius</taxonomicName>
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,”
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</emphasis>
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without reference to either species. However,
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<figureCitation id="136D2A24FFDDFF9943E8FDD419C5FD40" box="[661,760,638,663]" captionStart="FIGURE 26" captionStartId="19.[151,250,1114,1136]" captionTargetBox="[151,1436,195,1093]" captionTargetId="figure@19.[151,1436,195,1093]" captionTargetPageId="19" captionText="FIGURE 26. An unidentified troglomorphic anopsobiid centipede from Lost Soldier Cave, Sequoia Kings Canyon National Park, California. Photograph courtesy of Jean Krejca." httpUri="https://zenodo.org/record/1455633/files/figure.png" pageId="11" pageNumber="270">Fig. 26A</figureCitation>
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shows the posterior segments of a female with 5 coxal pores on the 15th coxa, and
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<figureCitation id="136D2A24FFDDFF9940FBFD0F1AD4FD69" box="[390,489,677,702]" captionStart="FIGURE 26" captionStartId="19.[151,250,1114,1136]" captionTargetBox="[151,1436,195,1093]" captionTargetId="figure@19.[151,1436,195,1093]" captionTargetPageId="19" captionText="FIGURE 26. An unidentified troglomorphic anopsobiid centipede from Lost Soldier Cave, Sequoia Kings Canyon National Park, California. Photograph courtesy of Jean Krejca." httpUri="https://zenodo.org/record/1455633/files/figure.png" pageId="11" pageNumber="270">Fig. 26B</figureCitation>
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shows a male with a coxal pore formula of 2, 4, 4, 5. Since
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<taxonomicName id="4C564D22FFDDFF9945F2FD0C1E0DFD6A" ID-CoL="NNVL" authorityName="Williams & Hefner" authorityYear="1928" box="[1167,1328,677,701]" class="Chilopoda" family="Henicopidae" genus="Buethobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="11" pageNumber="270" phylum="Arthropoda" rank="species" species="translucens">
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<emphasis id="B922EAB3FFDDFF9945F2FD0C1E0DFD6A" box="[1167,1328,677,701]" italics="true" pageId="11" pageNumber="270">B. translucens</emphasis>
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</taxonomicName>
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is said to have coxal pores 2, 2, 2, 2, it seems reasonable to assume that the illustrations are of
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<emphasis id="B922EAB3FFDDFF994533FD601FE9FD36" box="[1102,1236,713,737]" italics="true" pageId="11" pageNumber="270">
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<taxonomicName id="4C564D22FFDDFF994533FD601FEDFD36" box="[1102,1232,713,737]" class="Chilopoda" family="Henicopidae" genus="Buethobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="11" pageNumber="270" phylum="Arthropoda" rank="species" species="huestoni">B. huestoni</taxonomicName>
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.
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</emphasis>
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However, neither illustration shows the prominent process of the 15th coxa seen in other
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<taxonomicName id="4C564D22FFDDFF99428CFD5A1F52FCDF" ID-CoL="62GTT" authorityName="Chamberlin" authorityYear="1911" box="[1009,1135,752,776]" class="Chilopoda" family="Henicopidae" genus="Buethobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="11" pageNumber="270" phylum="Arthropoda" rank="genus">
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<emphasis id="B922EAB3FFDDFF99428CFD5A1F52FCDF" box="[1009,1135,752,776]" italics="true" pageId="11" pageNumber="270">Buethobius</emphasis>
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</taxonomicName>
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species. Andrew Weaver collected intensively for many years in
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<collectingRegion id="4992F843FFDDFF99432EFCBE19B6FCFA" box="[595,651,788,813]" country="United States of America" name="Ohio" pageId="11" pageNumber="270">Ohio</collectingRegion>
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and never encountered any species of
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<emphasis id="B922EAB3FFDDFF99453DFCBE1FF8FCFB" box="[1088,1221,788,812]" italics="true" pageId="11" pageNumber="270">
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<taxonomicName id="4C564D22FFDDFF99453DFCBE1FFCFCFB" ID-CoL="62GTT" authorityName="Chamberlin" authorityYear="1911" box="[1088,1217,788,812]" class="Chilopoda" family="Henicopidae" genus="Buethobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="11" pageNumber="270" phylum="Arthropoda" rank="genus">Buethobius</taxonomicName>
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,
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</emphasis>
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though he claimed to have seen one specimen from
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<collectingRegion id="4992F843FFDDFF994377FC921964FC86" box="[522,601,824,849]" country="United States of America" name="Illinois" pageId="11" pageNumber="270">Illinois</collectingRegion>
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matching the description of
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<taxonomicName id="4C564D22FFDDFF9942E0FC931F21FC87" box="[925,1052,824,848]" class="Chilopoda" family="Henicopidae" genus="Buethobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="11" pageNumber="270" phylum="Arthropoda" rank="species" species="huestoni">
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<emphasis id="B922EAB3FFDDFF9942E0FC931F21FC87" box="[925,1052,824,848]" italics="true" pageId="11" pageNumber="270">B. huestoni</emphasis>
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</taxonomicName>
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(A. A. Weaver, unpublished data; this is may be the source of the
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<collectingRegion id="4992F843FFDDFF994084FCF61975FCA2" box="[505,584,860,885]" country="United States of America" name="Illinois" pageId="11" pageNumber="270">Illinois</collectingRegion>
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record mentioned by
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<bibRefCitation id="EFC74B50FFDDFF994243FCF618C4FCA2" box="[830,1017,860,885]" pageId="11" pageNumber="270" refId="ref16977" refString="Mercurio, R. J. (2010) An annotated catalog of the centipedes (Chilopoda) from the United States of America, Canada and Greenland (1758 - 2008). XLibris Corporation, Bloomington, IN, 559 pp." type="book chapter">Mercurio [2010]</bibRefCitation>
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since Weaver shared his unpublished notes widely), based on the inference that the species lacks coxosternal teeth. However, I found two specimens in the USNM collection labelled “
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<taxonomicName id="4C564D22FFDDFF99437AFC0E18A8FC6A" authority="Oxford 1928" authorityName="Oxford" authorityYear="1928" box="[519,917,932,957]" class="Chilopoda" family="Henicopidae" genus="Buethobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="11" pageNumber="270" phylum="Arthropoda" rank="species" species="huestoni">Buethobius huestoni Oxford 1928</taxonomicName>
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.” On the reverse, the handwritten label says “This is what we called
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<taxonomicName id="4C564D22FFDDFF9940CCFC6219A4FC36" authority="Shear, 2018" authorityName="Shear" authorityYear="2018" box="[433,665,968,993]" class="Chilopoda" family="Henicopidae" genus="Buethobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="11" pageNumber="270" phylum="Arthropoda" rank="species" species="huestoni" status="sp. nov.">Buethobius huestoni</taxonomicName>
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<taxonomicNameLabel id="A21157C8FFDDFF9943DEFC6319DCFC37" box="[675,737,969,992]" pageId="11" pageNumber="270" rank="species">n. sp.</taxonomicNameLabel>
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we have only
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<specimenCount id="9D50FD28FFDDFF9942F2FC6218DAFC36" box="[911,999,968,993]" pageId="11" pageNumber="270" type="male">3 males</specimenCount>
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so far.” The specimens, a male and a female, are in poor condition; one complete antenna has 38 segments and the pore counts conform to the range given by
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<bibRefCitation id="EFC74B50FFDDFF994182FBBA1916FBFF" box="[255,555,1040,1065]" pageId="11" pageNumber="270" refId="ref17312" refString="Williams, S. R. & Hefner, R. A. (1928) The millipedes and centipedes of Ohio. Ohio Biological Survey Bulletin, 4, 91 - 146." type="journal article">Williams & Hefner (1928)</bibRefCitation>
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. I could not determine with certainty if an ocellus is present, but it appears it is not. As described in 1928, the forcipular coxosterna of both specimens have a straight, even margin lacking teeth. Leg-pairs 1–12 have single-segmented tarsi; the female retains leg 13, which has an incomplete suture dividing the tarsus. Neither specimen has legpairs 14 and 15. The coxae of legpair 15 do not have a distomesal spinous process. This combination of characters is unique in both anopsobiids and henicopids, so determining the status of
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<taxonomicName id="4C564D22FFDDFF9944F9FB0B1BCBFB0B" authorityName="Shear" authorityYear="2018" class="Chilopoda" family="Henicopidae" genus="Buethobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lithobiomorpha" pageId="11" pageNumber="270" phylum="Arthropoda" rank="species" species="huestoni">
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<emphasis id="B922EAB3FFDDFF9944F9FB0B1BCBFB0B" italics="true" pageId="11" pageNumber="270">B. huestoni</emphasis>
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</taxonomicName>
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must wait for fresh collections. Heuston Woods, Oxford,
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<collectingRegion id="4992F843FFDDFF9942FBFB6E18FCFB0A" box="[902,961,1220,1245]" country="United States of America" name="Ohio" pageId="11" pageNumber="270">Ohio</collectingRegion>
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, the
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<typeStatus id="54ED8803FFDDFF994285FB6F1F15FB0A" box="[1016,1064,1221,1245]" pageId="11" pageNumber="270">type</typeStatus>
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locality, is now a protected state park.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |