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<document id="05E3213AA7E76077A26F1AC4EE81FF91" ID="10.11646/zootaxa.4121.3.1" ID-CLB-Dataset="38912" ID-DOI="10.11646/zootaxa.4121.3.1" ID-GBIF-Dataset="6cb37831-da16-4763-a32c-aef636512990" ID-ISSN="1175-5326" ID-Zenodo-Dep="265007" ID-ZooBank="172021D4-55C8-483F-A6B2-F823C3022F90" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465452580540" checkinUser="plazi" docAuthor="Aston, Paul" docDate="2016" docId="03FF87A85E2B537357B4F940FBA8928E" docLanguage="en" docName="zootaxa.4121.3.1.pdf" docOrigin="Zootaxa 4121 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudognathaphanus Schauberger 1932" docType="treatment" docVersion="10" lastPageNumber="228" masterDocId="FFC6FFD05E30536F5723FFDEFFB59265" masterDocTitle="Catalogue and Bibliography of the Hong Kong Carabidae Latreille, 1802 (Coleoptera: Adephaga), with notes on the historic boundaries of Hong Kong as related to zoological collections" masterLastPageNumber="257" masterPageNumber="201" pageNumber="228" updateTime="1698662791348" updateUser="ExternalLinkService">
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<mods:title id="528D616B9DCF66802B6CA10B97C44188">Catalogue and Bibliography of the Hong Kong Carabidae Latreille, 1802 (Coleoptera: Adephaga), with notes on the historic boundaries of Hong Kong as related to zoological collections</mods:title>
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<mods:namePart id="006325AAA176328632A6BC0F82CAD250">Aston, Paul</mods:namePart>
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<treatment id="03FF87A85E2B537357B4F940FBA8928E" ID-DOI="http://doi.org/10.5281/zenodo.6077354" ID-GBIF-Taxon="120826484" ID-Zenodo-Dep="6077354" LSID="urn:lsid:plazi:treatment:03FF87A85E2B537357B4F940FBA8928E" httpUri="http://treatment.plazi.org/id/03FF87A85E2B537357B4F940FBA8928E" lastPageId="28" lastPageNumber="228" pageId="27" pageNumber="228">
<subSubSection id="C34C65355E2B537457B4F940FD1994D0" box="[151,684,1694,1717]" pageId="27" pageNumber="228" type="nomenclature">
<paragraph id="8BE936BE5E2B537457B4F940FD1994D0" blockId="27.[151,684,1694,1717]" box="[151,684,1694,1717]" pageId="27" pageNumber="228">
<heading id="D0A181D25E2B537457B4F940FD1994D0" bold="true" box="[151,684,1694,1717]" fontSize="10" level="2" pageId="27" pageNumber="228" reason="2">
<emphasis id="B922EAAC5E2B537457B4F940FD1994D0" bold="true" box="[151,684,1694,1717]" pageId="27" pageNumber="228">
Genus
<taxonomicName id="4C564D3D5E2B537457C5F940FD1994D0" ID-CoL="8KYLW" authority="Schauberger, 1932" authorityName="Schauberger" authorityYear="1932" box="[230,684,1694,1717]" class="Insecta" family="Carabidae" genus="Pseudognathaphanus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAAC5E2B537457C5F940FE6194D0" bold="true" box="[230,468,1694,1717]" italics="true" pageId="27" pageNumber="228">Pseudognathaphanus</emphasis>
Schauberger, 1932
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34C65355E2B537357B4F903FBA8928E" lastPageId="28" lastPageNumber="229" pageId="27" pageNumber="228" type="reference_group">
<paragraph id="8BE936BE5E2B537457B4F903FD149496" blockId="27.[151,1436,1757,2024]" box="[151,673,1757,1779]" pageId="27" pageNumber="228">
<taxonomicName id="4C564D3D5E2B537457B4F903FD149496" ID-CoL="4NXHK" authority="MacLeay 1825" authorityName="MacLeay" authorityYear="1825" box="[151,673,1757,1779]" class="Insecta" family="Carabidae" genus="Pseudognathaphanus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="punctilabris">
<emphasis id="B922EAAC5E2B537457B4F903FE449496" box="[151,497,1757,1779]" italics="true" pageId="27" pageNumber="228">Pseudognathaphanus punctilabris</emphasis>
(
<bibRefCitation id="EFC74B4F5E2B537456DDF903FD2F9496" author="MacLeay" box="[510,666,1757,1779]" pageId="27" pageNumber="228" refString="MacLeay, W. S. (1825) Annulosa Javanica, or, An attempt to Illustrate the natural affinities and analogies of the insects collected in Java by Thomas Horsfield, M. D. F. L. &amp; G. S. and deposited by him in the museum of the honourable East - india Company. Number 1. Parbury and Allen, Kingsbury, London, 150 pp." type="book" year="1825">MacLeay 1825</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="8BE936BE5E2B537457E4F923FE41952A" blockId="27.[151,1436,1757,2024]" pageId="27" pageNumber="228">
Bates (1886 &amp; 1889) list
<taxonomicName id="4C564D3D5E2B537456F9F922FC7C9577" ID-CoL="4NXHB" authority="Walker, 1859" authorityName="Walker" authorityYear="1859" box="[474,969,1788,1810]" class="Insecta" family="Carabidae" genus="Pseudognathaphanus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="dispellens">
<emphasis id="B922EAAC5E2B537456F9F922FC969577" box="[474,803,1788,1810]" italics="true" pageId="27" pageNumber="228">Pseudognathaphanus dispellens</emphasis>
(
<bibRefCitation id="EFC74B4F5E2B53745417F922FC779577" author="Walker" box="[820,962,1788,1810]" pageId="27" pageNumber="228" refString="Walker, F. (1859) Characters of some apparently undescribed Ceylon Insects. The Annals and Magazine of Natural History, Series 3, 3, 50 - 56." type="journal article" year="1859">Walker, 1859</bibRefCitation>
)
</taxonomicName>
as occurring in
<collectingCountry id="F341762E5E2B53745358F923FB4F9577" box="[1147,1274,1789,1810]" name="Hong Kong" pageId="27" pageNumber="228">Hong Kong</collectingCountry>
.
<bibRefCitation id="EFC74B4F5E2B53745224F923FA299577" author="Kataev" box="[1287,1436,1788,1810]" pageId="27" pageNumber="228" refString="Kataev, B. M. (2002) On some new and little - known species of the Anisodactylina and Harpalina (the Selenophori group) from East Asia and Oriental region (Coleoptera: Carabidae: Harpalini). Russian Entomological Journal, 11 (3), 241 - 252." type="journal article" year="2002">Kataev (2002)</bibRefCitation>
pointed out that
<taxonomicName id="4C564D3D5E2B5374566AF8C5FE7E9555" ID-CoL="4NXHB" box="[329,459,1818,1840]" class="Insecta" family="Carabidae" genus="Pseudognathaphanus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="dispellens">
<emphasis id="B922EAAC5E2B5374566AF8C5FE7E9555" box="[329,459,1818,1840]" italics="true" pageId="27" pageNumber="228">P. dispellens</emphasis>
</taxonomicName>
is confined to
<collectingCountry id="F341762E5E2B53745550F8C4FD6F9555" box="[627,730,1818,1840]" name="Sri Lanka" pageId="27" pageNumber="228">Sri Lanka</collectingCountry>
and the very similar
<taxonomicName id="4C564D3D5E2B537454E6F8C5FBEB9555" ID-CoL="4NXHK" box="[965,1118,1818,1840]" class="Insecta" family="Carabidae" genus="Pseudognathaphanus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="punctilabris">
<emphasis id="B922EAAC5E2B537454E6F8C5FBEB9555" box="[965,1118,1818,1840]" italics="true" pageId="27" pageNumber="228">P. punctilabris</emphasis>
</taxonomicName>
is the species with the larger distribution covering eastern Asia.
</paragraph>
<paragraph id="8BE936BE5E2B537357E4F886FBA8928E" blockId="27.[151,1436,1757,2024]" lastBlockId="28.[151,1435,152,235]" lastPageId="28" lastPageNumber="229" pageId="27" pageNumber="228">
Specimens collected in
<collectingCountry id="F341762E5E2B537456E0F887FD88950B" box="[451,573,1881,1902]" name="Hong Kong" pageId="27" pageNumber="228">Hong Kong</collectingCountry>
were determined using Kataevs (2002) key. Frontal impressions were present, a character typical of
<taxonomicName id="4C564D3D5E2B53745641F8A8FD8995E9" ID-CoL="8KYLW" box="[354,572,1910,1932]" class="Insecta" family="Carabidae" genus="Pseudognathaphanus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="B922EAAC5E2B53745641F8A8FD8995E9" box="[354,572,1910,1932]" italics="true" pageId="27" pageNumber="228">Pseudognathaphanus</emphasis>
</taxonomicName>
, but these beetles were possibly distinct from
<taxonomicName id="4C564D3D5E2B53745335F8A6FAEE95E9" ID-CoL="4NXHK" authority="MacLeay 1825" authorityName="MacLeay" authorityYear="1825" box="[1046,1371,1910,1933]" class="Insecta" family="Carabidae" genus="Pseudognathaphanus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="punctilabris">
<emphasis id="B922EAAC5E2B53745335F8A6FB1F95E9" box="[1046,1194,1910,1933]" italics="true" pageId="27" pageNumber="228">P. punctilabris</emphasis>
(
<bibRefCitation id="EFC74B4F5E2B5374539BF8A8FAE195E9" author="MacLeay" box="[1208,1364,1910,1932]" pageId="27" pageNumber="228" refString="MacLeay, W. S. (1825) Annulosa Javanica, or, An attempt to Illustrate the natural affinities and analogies of the insects collected in Java by Thomas Horsfield, M. D. F. L. &amp; G. S. and deposited by him in the museum of the honourable East - india Company. Number 1. Parbury and Allen, Kingsbury, London, 150 pp." type="book" year="1825">MacLeay 1825</bibRefCitation>
)
</taxonomicName>
in the male genitalia, as described by
<bibRefCitation id="EFC74B4F5E2B537456F4F848FDD995CE" author="Kataev" box="[471,620,1941,1963]" pageId="27" pageNumber="228" refString="Kataev, B. M. (2002) On some new and little - known species of the Anisodactylina and Harpalina (the Selenophori group) from East Asia and Oriental region (Coleoptera: Carabidae: Harpalini). Russian Entomological Journal, 11 (3), 241 - 252." type="journal article" year="2002">Kataev (2002)</bibRefCitation>
. Using
<bibRefCitation id="EFC74B4F5E2B5374559AF84BFCC295CE" author="Andrewes" box="[697,887,1941,1963]" pageId="27" pageNumber="228" refString="Andrewes, H. E. (1920 b) Notes sur les Carabiques orientaux II. Annales de la Societe Entomologique de Belgique, 60 (2), 106 - 111." type="journal article" year="1920" yearSuffix="b">Andrewes (1920b)</bibRefCitation>
the specimens key to
<taxonomicName id="4C564D3D5E2B53745379F84BFE9695AF" ID-CoL="4NXHL" authority="Bates, 1892" authorityName="Bates" authorityYear="1892" class="Insecta" family="Carabidae" genus="Pseudognathaphanus" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="228" phylum="Arthropoda" rank="species" species="rufitactor">
<emphasis id="B922EAAC5E2B53745379F84BFA2995CE" box="[1114,1436,1941,1963]" italics="true" pageId="27" pageNumber="228">Pseudognathaphanus rufitactor</emphasis>
(
<bibRefCitation id="EFC74B4F5E2B537457BDF86AFEA995AF" author="Bates" box="[158,284,1972,1994]" pageId="27" pageNumber="228" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of Carabidae. Annali del Museo Civico di Storia Natural di Genova, 32, 267 - 428." type="journal article" year="1892">Bates, 1892</bibRefCitation>
)
</taxonomicName>
, but they seem to differ from both
<bibRefCitation id="EFC74B4F5E2B537455B3F86AFCFB95AF" author="Andrewes" box="[656,846,1972,1994]" pageId="27" pageNumber="228" refString="Andrewes, H. E. (1920 b) Notes sur les Carabiques orientaux II. Annales de la Societe Entomologique de Belgique, 60 (2), 106 - 111." type="journal article" year="1920" yearSuffix="b">Andrewes (1920b)</bibRefCitation>
and
<bibRefCitation id="EFC74B4F5E2B537454A1F86BFBB395AF" author="Bates" box="[898,1030,1972,1994]" pageId="27" pageNumber="228" refString="Bates, H. W. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine. XLIV. List of Carabidae. Annali del Museo Civico di Storia Natural di Genova, 32, 267 - 428." type="journal article" year="1892">Bates (1892)</bibRefCitation>
in having antennomere 3 black; elytral stria very weakly punctate and being slightly larger in size.
<emphasis id="B922EAAC5E2B537455DCF80CFCE8958D" bold="true" box="[767,861,2002,2024]" pageId="27" pageNumber="228">Material</emphasis>
: ♀ W.T.L.,
<date id="FFE8107E5E2B537454F9F80CFB94958D" box="[986,1057,2002,2024]" pageId="27" pageNumber="228" value="2012-04-09">9.iv.12</date>
; 1 ex. W.T.L.,
<date id="FFE8107E5E2B53745398F80CFABF958D" box="[1211,1290,2002,2024]" pageId="27" pageNumber="228" value="2015-10-28">28.x.15</date>
; 1 ex. W.T.L.,
<date id="FFE8107E5E2C537357B4FF46FF5592CB" box="[151,224,152,174]" pageId="28" pageNumber="229" value="2012-11-04">4.xi.12</date>
; 1 ex. W.T.L.,
<date id="FFE8107E5E2C537356A4FF46FE6592CB" box="[391,464,152,174]" pageId="28" pageNumber="229" value="2013-11-06">6.xi.13</date>
; 1 ex. W.T.L.
<date id="FFE8107E5E2C53735551FF46FD7292CB" box="[626,711,152,174]" pageId="28" pageNumber="229" value="2015-11-12">12.xi.15</date>
; 1 ex. W.T.L.,
<date id="FFE8107E5E2C5373544CFF46FC7092CB" box="[879,965,152,174]" pageId="28" pageNumber="229" value="2014-11-18">18.xi.14</date>
; ♀ W.T.L.,
<date id="FFE8107E5E2C5373536AFF46FB2A92CB" box="[1097,1183,152,174]" pageId="28" pageNumber="229" value="2014-11-23">23.xi.14</date>
; 1 ex. W.T.,
<date id="FFE8107E5E2C53735213FF46FF7F92A9" pageId="28" pageNumber="229" value="2011-11">November 2011</date>
;sight record W.T.L.
<date id="FFE8107E5E2C5373568AFF68FE4C92A9" box="[425,505,182,204]" pageId="28" pageNumber="229" value="2015-12-02">2.xii.15</date>
; 1 ex. W.T.L.,
<date id="FFE8107E5E2C53735581FF68FD4592A9" box="[674,752,182,204]" pageId="28" pageNumber="229" value="2011-12-07">7.xii.11</date>
; sight record W.T.L.,
<date id="FFE8107E5E2C537354C2FF68FB8492A9" box="[993,1073,182,204]" pageId="28" pageNumber="229" value="2013-12-08">8.xii.13</date>
.
<emphasis id="B922EAAC5E2C53735363FF68FB2792A9" bold="true" box="[1088,1170,182,204]" pageId="28" pageNumber="229">Biology</emphasis>
: Imago found April and December to December. Can be attracted to light, it is fully winged and capable of flight.
</paragraph>
</subSubSection>
</treatment>
</document>