380 lines
57 KiB
XML
380 lines
57 KiB
XML
<document id="210934480B6192C9E3499191057E5B4C" ID-CLB-Dataset="172604" ID-DOI="10.5852/ejt.2022.845.1953" ID-GBIF-Dataset="1d14db17-e238-4cb1-b9ce-c453ad191baf" ID-ISSN="2118-9773" ID-Zenodo-Dep="7234830" ID-ZooBank="B8502709-68B1-4631-995C-5249DC3A7C16" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666360519903" checkinUser="felipe" docAuthor="Szawaryn, Karol & Czerwiński, Tomasz" docDate="2022" docId="03FF1C05FFC0B31A275DE5BDFC13FB8B" docLanguage="en" docName="ejt-845_30-65.pdf" docOrigin="European Journal of Taxonomy 845" docSource="http://dx.doi.org/10.5852/ejt.2022.845.1953" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chnoodes yanayacu Szawaryn & Czerwiński 2022, sp. nov." docType="treatment" docUuid="8FCC7088-DF9A-4416-A348-F99E37EAAE09" docUuidSource="ZooBank" docVersion="5" lastPageNumber="48" masterDocId="FFC6647DFFCFB3082533E15AFF9FFFF4" masterDocTitle="New Coccinellidae (Coleoptera, Coccinelloidea) from Napo Province in Ecuador" masterLastPageNumber="65" masterPageNumber="30" pageNumber="45" updateTime="1699408078409" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="7619860E554E67B304BF12C0EA773029">
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<mods:title id="0DF5073AB2E06E5AB64DB1438BDA8F2A">New Coccinellidae (Coleoptera, Coccinelloidea) from Napo Province in Ecuador</mods:title>
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</mods:titleInfo>
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<mods:name id="E3BD9FC81DC2D5EE512BDD7E52732B29" type="personal">
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<mods:roleTerm id="462B825BFAD82E9F13DEF363F4FCDC9C">Author</mods:roleTerm>
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<mods:namePart id="45F49D7C8A1412C8E326C0D096D7C99E">Szawaryn, Karol</mods:namePart>
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<mods:affiliation id="DFEEB8E7D875FEDAD77F8774D87AEF64">Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679, Warsaw, Poland.</mods:affiliation>
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<mods:nameIdentifier id="22A035CC301BE18BC7E921F0DE143CE2" type="email">k.szawaryn@gmail.com</mods:nameIdentifier>
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<mods:namePart id="754ED6C91CFE42C884BA10707FDC4723">Czerwiński, Tomasz</mods:namePart>
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<mods:affiliation id="6FBD74B0157127CDA94FC1A726B984FA">Museum and Institute of Zoology, Polish Academy of Sciences, Wilcza 64, 00 - 679, Warsaw, Poland.</mods:affiliation>
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<mods:nameIdentifier id="D7F9D4455FFFA058ADC9D8AF517C5232" type="email">tczerwinski@miiz.waw.pl</mods:nameIdentifier>
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<mods:title id="DB4793456E4AD034A057D4AB9DC5B236">European Journal of Taxonomy</mods:title>
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<mods:part id="9E4668EBA92DA3F48784976893258EC1">
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<mods:date id="DC2DC856A5B33ACAE58370275A2DB00A">2022</mods:date>
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<mods:number id="6232D9E2405EF3B292B42ACEEE17B938">2022-10-21</mods:number>
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<mods:number id="D78E859C006F7B30A57727F832C23397">845</mods:number>
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<mods:url id="94268F571A3C4277AC41B6F10DA2C500">http://dx.doi.org/10.5852/ejt.2022.845.1953</mods:url>
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<treatment id="03FF1C05FFC0B31A275DE5BDFC13FB8B" ID-DOI="http://doi.org/10.5281/zenodo.7234906" ID-GBIF-Taxon="203373013" ID-Zenodo-Dep="7234906" LSID="urn:lsid:zoobank.org:act:8FCC7088-DF9A-4416-A348-F99E37EAAE09" httpUri="http://treatment.plazi.org/id/03FF1C05FFC0B31A275DE5BDFC13FB8B" lastPageId="18" lastPageNumber="48" pageId="15" pageNumber="45">
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<subSubSection id="C34CFE98FFC0B307275DE5BDFC59FAF6" box="[622,966,1255,1282]" pageId="15" pageNumber="45" type="nomenclature">
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<paragraph id="8BE9AD13FFC0B307275DE5BDFC59FAF6" blockId="15.[364,1224,1255,1353]" box="[622,966,1255,1282]" pageId="15" pageNumber="45">
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<heading id="D0A11A7FFFC0B307275DE5BDFC59FAF6" box="[622,966,1255,1282]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="45" reason="2">
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<taxonomicName id="4C56D690FFC0B307275DE5BDFCFBFAF6" authority="Szawaryn & Czerwiński, 2022" authorityName="Szawaryn & Czerwiński" authorityYear="2022" box="[622,868,1255,1282]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="yanayacu" status="sp. nov.">
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<emphasis id="B9227101FFC0B307275DE5BDFCFBFAF6" bold="true" box="[622,868,1255,1282]" italics="true" pageId="15" pageNumber="45">Chnoodes yanayacu</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A211CC7AFFC0B307265FE5B2FC59FAF6" box="[876,966,1256,1282]" pageId="15" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34CFE98FFC0B307245FE451FCFBFABD" pageId="15" pageNumber="45" type="description">
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<paragraph id="8BE9AD13FFC0B307245FE451FB57FAD1" blockId="15.[364,1224,1255,1353]" box="[364,1224,1291,1317]" pageId="15" pageNumber="45">
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<uri id="FFC7A111FFC0B307245FE451FB57FAD1" box="[364,1224,1291,1317]" pageId="15" pageNumber="45">
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urn:lsid:zoobank.org:act:
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<uuid id="FFF097C6FFC0B30727BBE451FB57FAD1" box="[648,1224,1291,1317]" pageId="15" pageNumber="45">8FCC7088-DF9A-4416-A348-F99E37EAAE09</uuid>
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</uri>
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||
</paragraph>
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<paragraph id="8BE9AD13FFC0B30727E3E474FCFBFABD" blockId="15.[364,1224,1255,1353]" box="[720,868,1326,1353]" pageId="15" pageNumber="45">
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<figureCitation id="136DB196FFC0B30727E3E474FCD6FABD" box="[720,841,1326,1353]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[266,1322,287,1790]" captionTargetId="figure-12@10.[266,1322,265,1818]" captionTargetPageId="10" captionText="Fig. 5. Habitus. A–C. Eupalea borowieci sp. nov., ♂, holotype (MNHW). A. Dorsal view. B. Lateral view. C. Frontal view. D–F. Psyllobora marshalli Crotch, 1874, sex (MNHW). D. Dorsal view. E. Lateral view. F. Frontal view. G–I. Chnoodes yanayacu sp. nov., ♂, holotype (MNHW). G. Dorsal view. H. Lateral view. I. Frontal view. J–L. Exoplectra misahualli sp. nov., ♂, holotype (MNHW). J. Dorsal view. K. Lateral view. L. Frontal view." figureDoi="http://doi.org/10.5281/zenodo.7234853" httpUri="https://zenodo.org/record/7234853/files/figure.png" pageId="15" pageNumber="45">Figs 5G–I</figureCitation>
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,
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<figureCitation id="136DB196FFC0B3072666E475FCFBFABD" box="[853,868,1327,1353]" captionStart="Fig" captionStartId="17.[189,232,1787,1813]" captionTargetBox="[245,1342,265,1749]" captionTargetId="figure-13@17.[245,1342,265,1749]" captionTargetPageId="17" captionText="Fig. 7. Chnoodes yanayacu sp. nov. A. Paratype (MNHW), ♀, abdomen. B. Holotype (MNHW), ♂, abdomen. C. Paratype (MNHW), ♀, tergite VIII. D. Paratype (MNHW), ♀, sternite VIII. E. Holotype (MNHW), ♂, tergite VIII. F. Holotype (MNHW), ♂, sternite VIII. G. Paratype (MNHW), spermatheca. H. Paratype (MNHW), ♀, genitalia, ventral. I. Holotype (MNHW), penis, lateral. J. Holotype (MNHW), penis tip, inner view. K. Holotype (MNHW), penis tip, lateral view. L. Holotype (MNHW), ♂, segments IX and X, dorsal view. M. Holotype (MNHW), tegmen, lateral view. N. Holotype (MNHW), tegmen, inner view." figureDoi="http://doi.org/10.5281/zenodo.7234871" httpUri="https://zenodo.org/record/7234871/files/figure.png" pageId="15" pageNumber="45">7</figureCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34CFE98FFC0B307258EE4D0FB3FF8DE" pageId="15" pageNumber="45" type="diagnosis">
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<paragraph id="8BE9AD13FFC0B307258EE4D0FEDDFA53" blockId="15.[189,1401,1418,1834]" box="[189,322,1418,1447]" pageId="15" pageNumber="45">
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<emphasis id="B9227101FFC0B307258EE4D0FEDDFA53" bold="true" box="[189,322,1418,1447]" pageId="15" pageNumber="45">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8BE9AD13FFC0B307258EE4F5FB3FF8DE" blockId="15.[189,1401,1418,1834]" pageId="15" pageNumber="45">
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This species is most similar to
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<taxonomicName id="4C56D690FFC0B3072700E4F5FC4BFA3E" authority="Mader, 1957" authorityName="Mader" authorityYear="1957" box="[563,980,1455,1482]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="rufovittata">
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<emphasis id="B9227101FFC0B3072700E4F5FCAFFA3E" box="[563,816,1455,1482]" italics="true" pageId="15" pageNumber="45">Chnoodes rufovittata</emphasis>
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Mader, 1957
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</taxonomicName>
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; however; its body outline is more elongate and maculae are larger, covering more surface of elytra, whereas in
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<taxonomicName id="4C56D690FFC0B3072151E489FA9BFA19" authorityName="Mader" authorityYear="1957" box="[1122,1284,1491,1517]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="rufovittata">
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<emphasis id="B9227101FFC0B3072151E489FA9BFA19" box="[1122,1284,1491,1517]" italics="true" pageId="15" pageNumber="45">C. rufovittata</emphasis>
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</taxonomicName>
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the body is more rounded with smaller elytral maculae.
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<taxonomicName id="4C56D690FFC0B30727C8E4ACFC71F9E4" authority="Szawaryn & Czerwiński, 2022" authorityName="Szawaryn & Czerwiński" authorityYear="2022" box="[763,1006,1526,1552]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="yanayacu" status="sp. nov.">
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<emphasis id="B9227101FFC0B30727C8E4ACFC71F9E4" box="[763,1006,1526,1552]" italics="true" pageId="15" pageNumber="45">Chnoodes yanayacu</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A211CC7AFFC0B30726C9E4ACFBC7F9E4" box="[1018,1112,1526,1552]" pageId="15" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
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is also quite similar to
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<taxonomicName id="4C56D690FFC0B307258EE740FE76F9C7" authority="Sicard, 1912" authorityName="Sicard" authorityYear="1912" box="[189,489,1561,1588]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="centralis">
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<emphasis id="B9227101FFC0B307258EE740FED6F9C7" box="[189,329,1561,1588]" italics="true" pageId="15" pageNumber="45">C. centralis</emphasis>
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Sicard, 1912
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</taxonomicName>
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and
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<taxonomicName id="4C56D690FFC0B3072710E740FB53F9C7" authority="Kruger, Castro-Guedes & Almeida, 2016" authorityName="Kruger, Castro-Guedes & Almeida" authorityYear="2016" box="[547,1228,1561,1588]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="unimaculata">
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<emphasis id="B9227101FFC0B3072710E740FD43F9C7" box="[547,732,1561,1588]" italics="true" pageId="15" pageNumber="45">C. unimaculata</emphasis>
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<bibRefCitation id="EFC7D0E2FFC0B30727D5E740FB53F9C7" author="Kruger T. C. & Castro-Guedes C. F. & Almeida L. M." box="[742,1228,1561,1588]" pageId="15" pageNumber="45" pagination="269 - 283" refId="ref17758" refString="Kruger T. C., Castro-Guedes C. F. & Almeida L. M. 2016. Two new species of Chnoodes Chevrolat (Coleoptera: Coccinellidae) from Brazil. Zootaxa 4078 (1): 269 - 283. https: // doi. org / 10.11646 / zootaxa. 4078.1.24" type="journal article" year="2016">Krüger, Castro-Guedes & Almeida, 2016</bibRefCitation>
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</taxonomicName>
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(
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<bibRefCitation id="EFC7D0E2FFC0B30721EFE743FF64F9A2" author="Kruger T. C. & Castro-Guedes C. F. & Almeida L. M." pageId="15" pageNumber="45" pagination="269 - 283" refId="ref17758" refString="Kruger T. C., Castro-Guedes C. F. & Almeida L. M. 2016. Two new species of Chnoodes Chevrolat (Coleoptera: Coccinellidae) from Brazil. Zootaxa 4078 (1): 269 - 283. https: // doi. org / 10.11646 / zootaxa. 4078.1.24" type="journal article" year="2016">
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Krüger
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<emphasis id="B9227101FFC0B3072009E740FAE8F9C7" box="[1338,1399,1561,1587]" italics="true" pageId="15" pageNumber="45">et al.</emphasis>
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2016
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</bibRefCitation>
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;
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<bibRefCitation id="EFC7D0E2FFC0B3072438E767FE37F9A2" author="Kruger T. C." box="[267,424,1596,1623]" pageId="15" pageNumber="45" refId="ref17726" refString="Kruger T. C. 2018. Revisao das especies brasileiras de Chnoodes Chevrolat, 1849 (Coleoptera: Coccinellidae). PhD thesis, Universidade Federal de Parana, Curitiba." type="book" year="2018">Krüger 2018</bibRefCitation>
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), by general appearance, but it differs from both species by much larger elytral maculae covering almost the entire surface of the elytra, while in both mentioned species the macula are distinctly smaller or forming a ring pattern. This species also differs from
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<taxonomicName id="4C56D690FFC0B3072158E7D9FB69F969" authorityName="Sicard" authorityYear="1912" box="[1131,1270,1667,1693]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="centralis">
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<emphasis id="B9227101FFC0B3072158E7D9FB69F969" box="[1131,1270,1667,1693]" italics="true" pageId="15" pageNumber="45">C. centralis</emphasis>
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</taxonomicName>
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by having the female genitalia with longer styli and constricted base of the spermatheca, from
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<taxonomicName id="4C56D690FFC0B307218EE7FCFAE9F934" authorityName="Kruger, Castro-Guedes & Almeida" authorityYear="2016" box="[1213,1398,1702,1728]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="unimaculata">
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<emphasis id="B9227101FFC0B307218EE7FCFAE9F934" box="[1213,1398,1702,1728]" italics="true" pageId="15" pageNumber="45">C. unimaculata</emphasis>
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</taxonomicName>
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by more elongate coxites and the presence of a mammiliform projection on spermathecal apex. Males of
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<taxonomicName id="4C56D690FFC0B30725D2E7B7FEEFF8F2" authorityName="Sicard" authorityYear="1912" box="[225,368,1772,1799]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="centralis">
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<emphasis id="B9227101FFC0B30725D2E7B7FF6AF8F3" box="[225,245,1773,1799]" italics="true" pageId="15" pageNumber="45">C</emphasis>
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.
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<emphasis id="B9227101FFC0B307243BE7B6FEEFF8F2" box="[264,368,1772,1798]" italics="true" pageId="15" pageNumber="45">centralis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C56D690FFC0B3072481E7B7FDF2F8F2" authorityName="Kruger, Castro-Guedes & Almeida" authorityYear="2016" box="[434,621,1772,1799]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="unimaculata">
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<emphasis id="B9227101FFC0B3072481E7B7FE59F8F3" box="[434,454,1773,1799]" italics="true" pageId="15" pageNumber="45">C</emphasis>
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.
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<emphasis id="B9227101FFC0B30724EAE7B6FDF2F8F2" box="[473,621,1772,1798]" italics="true" pageId="15" pageNumber="45">unimaculata</emphasis>
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</taxonomicName>
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are unknown.
|
||
<taxonomicName id="4C56D690FFC0B307261AE7B7FC52F8F2" authorityName="Mader" authorityYear="1957" box="[809,973,1772,1799]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="rufovittata">
|
||
<emphasis id="B9227101FFC0B307261AE7B7FC52F8F2" box="[809,973,1772,1799]" italics="true" pageId="15" pageNumber="45">C. rufovittata</emphasis>
|
||
</taxonomicName>
|
||
was described from
|
||
<collectingCountry id="F341ED83FFC0B30721E0E7B6FAB6F8F2" box="[1235,1321,1772,1798]" name="Bolivia" pageId="15" pageNumber="45">Bolivia</collectingCountry>
|
||
while
|
||
<taxonomicName id="4C56D690FFC0B307258EE64AFED7F8DD" authorityName="Sicard" authorityYear="1912" box="[189,328,1807,1834]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="centralis">
|
||
<emphasis id="B9227101FFC0B307258EE64AFED7F8DD" box="[189,328,1807,1834]" italics="true" pageId="15" pageNumber="45">C. centralis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F341ED83FFC0B30724BCE655FDDFF8DD" box="[399,576,1807,1833]" name="French Guiana" pageId="15" pageNumber="45">French Guiana</collectingCountry>
|
||
, and
|
||
<taxonomicName id="4C56D690FFC0B307274EE64AFCABF8DD" authorityName="Kruger, Castro-Guedes & Almeida" authorityYear="2016" box="[637,820,1807,1834]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="45" phylum="Arthropoda" rank="species" species="unimaculata">
|
||
<emphasis id="B9227101FFC0B307274EE64AFCABF8DD" box="[637,820,1807,1834]" italics="true" pageId="15" pageNumber="45">C. unimaculata</emphasis>
|
||
</taxonomicName>
|
||
occur in Eastern
|
||
<collectingCountry id="F341ED83FFC0B3072132E655FBD6F8DD" box="[1025,1097,1807,1833]" name="Brazil" pageId="15" pageNumber="45">Brazil</collectingCountry>
|
||
(
|
||
<collectingRegion id="499263F1FFC0B307216AE64AFB0CF8DE" box="[1113,1171,1808,1834]" country="Brazil" name="Para" pageId="15" pageNumber="45">Pará</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34CFE98FFC0B307258EE60BFEB0F849" pageId="15" pageNumber="45" type="etymology">
|
||
<paragraph id="8BE9AD13FFC0B307258EE60BFEDAF89F" blockId="15.[189,325,1873,1899]" box="[189,325,1873,1899]" pageId="15" pageNumber="45">
|
||
<heading id="D0A11A7FFFC0B307258EE60BFEDAF89F" bold="true" box="[189,325,1873,1899]" fontSize="11" level="3" pageId="15" pageNumber="45" reason="3">
|
||
<emphasis id="B9227101FFC0B307258EE60BFEDAF89F" bold="true" box="[189,325,1873,1899]" pageId="15" pageNumber="45">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE9AD13FFC0B307258EE625FEB0F849" blockId="15.[189,1399,1919,1981]" pageId="15" pageNumber="45">
|
||
This species is named after the Yanayacu biological station in
|
||
<collectingCountry id="F341ED83FFC0B30726FBE625FBB5F86D" box="[968,1066,1919,1945]" name="Ecuador" pageId="15" pageNumber="45">Ecuador</collectingCountry>
|
||
where the
|
||
<typeStatus id="54ED13B1FFC0B3072188E6DAFB71F86E" box="[1211,1262,1920,1946]" pageId="15" pageNumber="45">type</typeStatus>
|
||
series was collected.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34CFE98FFDFB318258EE052FEE7FDB7" pageId="16" pageNumber="46" type="materials_examined">
|
||
<paragraph id="8BE9AD13FFDFB318258EE052FE32FED6" blockId="16.[189,429,264,290]" box="[189,429,264,290]" pageId="16" pageNumber="46">
|
||
<heading id="D0A11A7FFFDFB318258EE052FE32FED6" bold="true" box="[189,429,264,290]" fontSize="11" level="3" pageId="16" pageNumber="46" reason="3">
|
||
<emphasis id="B9227101FFDFB318258EE052FE32FED6" bold="true" box="[189,429,264,290]" pageId="16" pageNumber="46">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE9AD13FFDFB31825D9E06DFEBDFE6C" blockId="16.[189,1399,311,408]" pageId="16" pageNumber="46">
|
||
<materialsCitation id="3B3EA74EFFDFB31825D9E06DFEBDFE6C" ID-GBIF-Occurrence="3946725377" collectingDate="2009-11-25" collectionCode="R, MNHW" country="Ecuador" location="Yanayacu" municipality="Cosanga" pageId="16" pageNumber="46" specimenCount="1" specimenCount-male="1" stateProvince="Napo" typeStatus="holotype">
|
||
<emphasis id="B9227101FFDFB31825D9E06DFEC2FEA5" bold="true" box="[234,349,311,337]" pageId="16" pageNumber="46">
|
||
<typeStatus id="54ED13B1FFDFB31825D9E06DFEC2FEA5" box="[234,349,311,337]" pageId="16" pageNumber="46" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F341ED83FFDFB318258EE000FED4FE80" box="[189,331,346,372]" name="Ecuador" pageId="16" pageNumber="46">ECUADOR</collectingCountry>
|
||
•
|
||
<specimenCount id="9D50669AFFDFB3182451E001FEE4FE81" box="[354,379,347,373]" pageId="16" pageNumber="46" type="male">♂</specimenCount>
|
||
; “
|
||
<collectingRegion id="499263F1FFDFB31824A1E000FD86FE81" box="[402,537,346,373]" country="Ecuador" name="Napo" pageId="16" pageNumber="46">Napo prov.</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B8D3769FFDFB3182710E000FD14FE80" box="[547,651,346,372]" pageId="16" pageNumber="46">Cosanga</collectingMunicipality>
|
||
,
|
||
<location id="8E89FBC8FFDFB31827A6E001FC95FE81" LSID="urn:lsid:plazi:treatment:03FF1C05FFC0B31A275DE5BDFC13FB8B:8E89FBC8FFDFB31827A6E001FC95FE81" box="[661,778,347,373]" country="Ecuador" municipality="Cosanga" name="Yanayacu" pageId="16" pageNumber="46" stateProvince="Napo">Yanayacu</location>
|
||
, collecting along the road,
|
||
<date id="FFE88BD3FFDFB3182177E000FB53FE80" box="[1092,1228,346,373]" pageId="16" pageNumber="46" value="2009-11-25">
|
||
<collectingDate id="EFAC723BFFDFB3182177E000FB53FE80" box="[1092,1228,346,373]" pageId="16" pageNumber="46" value="2009-11-25">25 XI 2009</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectionCode id="ED4735D6FFDFB318203BE001FA80FE81" box="[1288,1311,347,373]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="46">R</collectionCode>
|
||
. Ruta”;
|
||
<collectionCode id="ED4735D6FFDFB318258EE024FE81FE6C" box="[189,286,382,408]" pageId="16" pageNumber="46">MNHW</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BE9AD13FFDFB31825D9E0E5FEE7FDB7" blockId="16.[189,1399,447,579]" pageId="16" pageNumber="46">
|
||
<materialsCitation id="3B3EA74EFFDFB31825D9E0E5FBBBFE09" ID-GBIF-Occurrence="3946725412" collectingDate="2009-11-25" collectionCode="MNHW" country="Ecuador" location="Yanayacu" municipality="Cosanga" pageId="16" pageNumber="46" specimenCount="2" specimenCount-female="1" stateProvince="Napo" typeStatus="paratype">
|
||
<emphasis id="B9227101FFDFB31825D9E0E5FEF7FE2D" bold="true" box="[234,360,447,473]" pageId="16" pageNumber="46">
|
||
<typeStatus id="54ED13B1FFDFB31825D9E0E5FEF7FE2D" box="[234,360,447,473]" pageId="16" pageNumber="46" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F341ED83FFDFB318258EE0B8FED4FE08" box="[189,331,482,508]" name="Ecuador" pageId="16" pageNumber="46">ECUADOR</collectingCountry>
|
||
•
|
||
<specimenCount id="9D50669AFFDFB3182459E0B8FE06FE09" box="[362,409,482,509]" pageId="16" pageNumber="46" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount id="9D50669AFFDFB3182498E0B8FE75FE09" box="[427,490,482,509]" pageId="16" pageNumber="46" type="generic">1 ex.</specimenCount>
|
||
; same collection data as for holotype;
|
||
<collectionCode id="ED4735D6FFDFB31826F7E0B9FBBBFE09" box="[964,1060,483,509]" pageId="16" pageNumber="46">MNHW</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B3EA74EFFDFB3182170E0B8FC8CFDD4" ID-GBIF-Occurrence="3946725304" collectingDate="2009-12-02" collectionCode="R, MNHW" country="Ecuador" location="Cosanga" municipality="Las Palmas Island" pageId="16" pageNumber="46" specimenCount="2" typeStatus="paratype">
|
||
<specimenCount id="9D50669AFFDFB3182170E0B8FB1DFE09" box="[1091,1154,482,509]" pageId="16" pageNumber="46" type="generic">2 ex.</specimenCount>
|
||
; “
|
||
<location id="8E89FBC8FFDFB31821AEE0B8FA9AFE08" LSID="urn:lsid:plazi:treatment:03FF1C05FFC0B31A275DE5BDFC13FB8B:8E89FBC8FFDFB31821AEE0B8FA9AFE08" box="[1181,1285,482,508]" country="Ecuador" municipality="Las Palmas Island" name="Cosanga" pageId="16" pageNumber="46">Cosanga</location>
|
||
vic.,
|
||
<collectingMunicipality id="6B8D3769FFDFB318207FE0B9FEFBFDEB" pageId="16" pageNumber="46">Las Palmas Island</collectingMunicipality>
|
||
,
|
||
<date id="FFE88BD3FFDFB3182443E35FFE6AFDEB" box="[368,501,517,544]" pageId="16" pageNumber="46" value="2009-12-02">
|
||
<collectingDate id="EFAC723BFFDFB3182443E35FFE6AFDEB" box="[368,501,517,544]" pageId="16" pageNumber="46" value="2009-12-02">2 XII 2009</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectionCode id="ED4735D6FFDFB3182707E35CFDD4FDD4" box="[564,587,518,544]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="46">R</collectionCode>
|
||
. Ruta ”;
|
||
<collectionCode id="ED4735D6FFDFB3182780E35CFC8CFDD4" box="[691,787,518,544]" pageId="16" pageNumber="46">MNHW</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B3EA74EFFDFB318261FE35FFEEBFDB7" ID-GBIF-Occurrence="3946725403" collectingDate="2009-12-02" collectionCode="R" country="Ecuador" location="Cosanga" municipality="Yanayacu" pageId="16" pageNumber="46" specimenCount="1" typeStatus="paratype">
|
||
<specimenCount id="9D50669AFFDFB318261FE35FFCF6FDD4" box="[812,873,517,544]" pageId="16" pageNumber="46" type="generic">1 ex.</specimenCount>
|
||
; “
|
||
<location id="8E89FBC8FFDFB31826B3E35FFC75FDEB" LSID="urn:lsid:plazi:treatment:03FF1C05FFC0B31A275DE5BDFC13FB8B:8E89FBC8FFDFB31826B3E35FFC75FDEB" box="[896,1002,517,543]" country="Ecuador" municipality="Yanayacu" name="Cosanga" pageId="16" pageNumber="46">Cosanga</location>
|
||
,
|
||
<collectingMunicipality id="6B8D3769FFDFB31826C5E35CFBF4FDD4" box="[1014,1131,518,544]" pageId="16" pageNumber="46">Yanayacu</collectingMunicipality>
|
||
, road,
|
||
<date id="FFE88BD3FFDFB3182188E35FFAA0FDEB" box="[1211,1343,517,544]" pageId="16" pageNumber="46" value="2009-12-02">
|
||
<collectingDate id="EFAC723BFFDFB3182188E35FFAA0FDEB" box="[1211,1343,517,544]" pageId="16" pageNumber="46" value="2009-12-02">2 XII 2009</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectionCode id="ED4735D6FFDFB318258EE373FF4BFDB7" box="[189,212,553,579]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="46">R</collectionCode>
|
||
. Ruta ”; MIZ
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34CFE98FFDFB318258EE330FED5FCCB" pageId="16" pageNumber="46" type="discussion">
|
||
<paragraph id="8BE9AD13FFDFB318258EE330FEAFFD70" blockId="16.[189,304,618,644]" box="[189,304,618,644]" pageId="16" pageNumber="46">
|
||
<heading id="D0A11A7FFFDFB318258EE330FEAFFD70" bold="true" box="[189,304,618,644]" fontSize="11" level="3" pageId="16" pageNumber="46" reason="3">
|
||
<emphasis id="B9227101FFDFB318258EE330FEAFFD70" bold="true" box="[189,304,618,644]" pageId="16" pageNumber="46">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE9AD13FFDFB318258EE3C2FED5FCCB" blockId="16.[189,1399,664,831]" pageId="16" pageNumber="46">
|
||
Guillermo González illustrated the
|
||
<typeStatus id="54ED13B1FFDFB3182755E3C2FD52FD46" box="[614,717,664,690]" pageId="16" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C56D690FFDFB31827C9E3C3FC02FD47" authorityName="Mader" authorityYear="1957" box="[762,925,665,691]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="46" phylum="Arthropoda" rank="species" species="rufovittata">
|
||
<emphasis id="B9227101FFDFB31827C9E3C3FC02FD47" box="[762,925,665,691]" italics="true" pageId="16" pageNumber="46">C. rufovittata</emphasis>
|
||
</taxonomicName>
|
||
on his website (www.coccinellidae.cl), which is deposited in the Museo Nacional de Historia Natural de
|
||
<collectingRegion id="499263F1FFDFB31826E9E3E6FBDBFD22" box="[986,1092,700,726]" country="Chile" name="Region Metropolitana" pageId="16" pageNumber="46">Santiago</collectingRegion>
|
||
,
|
||
<collectingCountry id="F341ED83FFDFB3182161E3E6FB0DFD22" box="[1106,1170,700,726]" name="Chile" pageId="16" pageNumber="46">Chile</collectingCountry>
|
||
(MNHNS), but he was not able to investigate genitalia of that specimen. We provide here a detailed description of male and female genitalia that will allow to verify the status of
|
||
<taxonomicName id="4C56D690FFDFB318265FE258FB9EFCE8" authority="Szawaryn & Czerwiński, 2022" authorityName="Szawaryn & Czerwiński" authorityYear="2022" box="[876,1025,770,796]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="46" phylum="Arthropoda" rank="species" species="yanayacu" status="sp. nov.">
|
||
<emphasis id="B9227101FFDFB318265FE258FB9EFCE8" box="[876,1025,770,796]" italics="true" pageId="16" pageNumber="46">C. yanayacu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A211CC7AFFDFB318213BE259FBFBFCE9" box="[1032,1124,771,797]" pageId="16" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
|
||
when new information is available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34CFE98FFDFB31A258EE23CFC13FB8B" lastPageId="18" lastPageNumber="48" pageId="16" pageNumber="46" type="description">
|
||
<paragraph id="8BE9AD13FFDFB318258EE23CFED0FC74" blockId="16.[189,335,870,896]" box="[189,335,870,896]" pageId="16" pageNumber="46">
|
||
<heading id="D0A11A7FFFDFB318258EE23CFED0FC74" bold="true" box="[189,335,870,896]" fontSize="11" level="3" pageId="16" pageNumber="46" reason="3">
|
||
<emphasis id="B9227101FFDFB318258EE23CFED0FC74" bold="true" box="[189,335,870,896]" pageId="16" pageNumber="46">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BE9AD13FFDFB318258EE2CCFD49FBC9" blockId="16.[189,1398,917,1085]" pageId="16" pageNumber="46">
|
||
BODY. Length
|
||
<quantity id="4CAE00F6FFDFB3182456E2CFFE4FFC44" box="[357,464,917,944]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="16" pageNumber="46" unit="mm" value="3.2">3.20 mm</quantity>
|
||
; TL/EW = 1.15–1.30; PL/PW = 0.52; EL/EW = 0.95–1.00. Body oval (
|
||
<figureCitation id="136DB196FFDFB318203FE2CFFAF7FC44" box="[1292,1384,917,944]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[266,1322,287,1790]" captionTargetId="figure-12@10.[266,1322,265,1818]" captionTargetPageId="10" captionText="Fig. 5. Habitus. A–C. Eupalea borowieci sp. nov., ♂, holotype (MNHW). A. Dorsal view. B. Lateral view. C. Frontal view. D–F. Psyllobora marshalli Crotch, 1874, sex (MNHW). D. Dorsal view. E. Lateral view. F. Frontal view. G–I. Chnoodes yanayacu sp. nov., ♂, holotype (MNHW). G. Dorsal view. H. Lateral view. I. Frontal view. J–L. Exoplectra misahualli sp. nov., ♂, holotype (MNHW). J. Dorsal view. K. Lateral view. L. Frontal view." figureDoi="http://doi.org/10.5281/zenodo.7234853" httpUri="https://zenodo.org/record/7234853/files/figure.png" pageId="16" pageNumber="46">Fig. 5G</figureCitation>
|
||
), moderately convex (
|
||
<figureCitation id="136DB196FFDFB318249EE2E2FD95FC27" box="[429,522,952,979]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[266,1322,287,1790]" captionTargetId="figure-12@10.[266,1322,265,1818]" captionTargetPageId="10" captionText="Fig. 5. Habitus. A–C. Eupalea borowieci sp. nov., ♂, holotype (MNHW). A. Dorsal view. B. Lateral view. C. Frontal view. D–F. Psyllobora marshalli Crotch, 1874, sex (MNHW). D. Dorsal view. E. Lateral view. F. Frontal view. G–I. Chnoodes yanayacu sp. nov., ♂, holotype (MNHW). G. Dorsal view. H. Lateral view. I. Frontal view. J–L. Exoplectra misahualli sp. nov., ♂, holotype (MNHW). J. Dorsal view. K. Lateral view. L. Frontal view." figureDoi="http://doi.org/10.5281/zenodo.7234853" httpUri="https://zenodo.org/record/7234853/files/figure.png" pageId="16" pageNumber="46">Fig. 5H</figureCitation>
|
||
), winged; dorsum black with one big, chestnut brown spot on each elytron and light brown anterolateral edges of pronotum (
|
||
<figureCitation id="136DB196FFDFB318263AE286FCCAFC02" box="[777,853,987,1014]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[266,1322,287,1790]" captionTargetId="figure-12@10.[266,1322,265,1818]" captionTargetPageId="10" captionText="Fig. 5. Habitus. A–C. Eupalea borowieci sp. nov., ♂, holotype (MNHW). A. Dorsal view. B. Lateral view. C. Frontal view. D–F. Psyllobora marshalli Crotch, 1874, sex (MNHW). D. Dorsal view. E. Lateral view. F. Frontal view. G–I. Chnoodes yanayacu sp. nov., ♂, holotype (MNHW). G. Dorsal view. H. Lateral view. I. Frontal view. J–L. Exoplectra misahualli sp. nov., ♂, holotype (MNHW). J. Dorsal view. K. Lateral view. L. Frontal view." figureDoi="http://doi.org/10.5281/zenodo.7234853" httpUri="https://zenodo.org/record/7234853/files/figure.png" pageId="16" pageNumber="46">Fig 5I</figureCitation>
|
||
); ventral side infuscate. Dorsum covered with short hairs, forming weak, wavy pattern on elytra. Dorsum with single size punctures. Punctures on elytra less coarse than on head and pronotum.
|
||
</paragraph>
|
||
<paragraph id="8BE9AD13FFDFB318258EE53EFB55F9F5" blockId="16.[189,1399,1123,1537]" pageId="16" pageNumber="46">
|
||
HEAD. Head partially covered by pronotum (
|
||
<figureCitation id="136DB196FFDFB31827D3E539FCA8FB8A" box="[736,823,1123,1150]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[266,1322,287,1790]" captionTargetId="figure-12@10.[266,1322,265,1818]" captionTargetPageId="10" captionText="Fig. 5. Habitus. A–C. Eupalea borowieci sp. nov., ♂, holotype (MNHW). A. Dorsal view. B. Lateral view. C. Frontal view. D–F. Psyllobora marshalli Crotch, 1874, sex (MNHW). D. Dorsal view. E. Lateral view. F. Frontal view. G–I. Chnoodes yanayacu sp. nov., ♂, holotype (MNHW). G. Dorsal view. H. Lateral view. I. Frontal view. J–L. Exoplectra misahualli sp. nov., ♂, holotype (MNHW). J. Dorsal view. K. Lateral view. L. Frontal view." figureDoi="http://doi.org/10.5281/zenodo.7234853" httpUri="https://zenodo.org/record/7234853/files/figure.png" pageId="16" pageNumber="46">Fig. 5I</figureCitation>
|
||
); ventral antennal grooves deep, straight. Eyes moderately convex, dorsally with inner orbits arcuate; ocular canthus present, protruding, reaching beyond half width of an eye; interocular distance about 0.50 × as wide as head across eyes; interfacetal setae absent. Antenna short, about 0.50 × as long as head capsule width, composed of 10 antennomeres; scape enlarged, distinctly roundly projected anteriorly; pedicel narrower than scape, barrel-shaped, about 1.20 × as long as wide; antennomere 3 distinctly shorter than pedicel. Antennal club consisting of four antennomeres; antennomeres 7–9 transverse, antennomere 10 elongate, truncate apically. Anterior clypeal margin margined with anterolateral lobes roundly projected. Labrum truncate at apex. Mandible bidentate at apex. Maxillary cardo slightly transverse; terminal palpomere securiform, truncate obliquely. Submentum transverse; mentum sub-trapezoidal, covered with sparse long hairs, with lateral longitudinal weak impressions on each side; prementum transverse; labial palps separated by distance about equal to width of palpiger; apical palpomere slightly shorter than penultimate one.
|
||
</paragraph>
|
||
<paragraph id="8BE9AD13FFDFB318258EE773FB19F907" blockId="16.[189,1399,1576,1779]" pageId="16" pageNumber="46">
|
||
PROTHORAX. Pronotum (
|
||
<figureCitation id="136DB196FFDFB31824E0E772FDBCF9B7" box="[467,547,1576,1603]" captionStart="Fig" captionStartId="10.[189,232,1857,1883]" captionTargetBox="[266,1322,287,1790]" captionTargetId="figure-12@10.[266,1322,265,1818]" captionTargetPageId="10" captionText="Fig. 5. Habitus. A–C. Eupalea borowieci sp. nov., ♂, holotype (MNHW). A. Dorsal view. B. Lateral view. C. Frontal view. D–F. Psyllobora marshalli Crotch, 1874, sex (MNHW). D. Dorsal view. E. Lateral view. F. Frontal view. G–I. Chnoodes yanayacu sp. nov., ♂, holotype (MNHW). G. Dorsal view. H. Lateral view. I. Frontal view. J–L. Exoplectra misahualli sp. nov., ♂, holotype (MNHW). J. Dorsal view. K. Lateral view. L. Frontal view." figureDoi="http://doi.org/10.5281/zenodo.7234853" httpUri="https://zenodo.org/record/7234853/files/figure.png" pageId="16" pageNumber="46">Fig. 5I</figureCitation>
|
||
) with anterior corners rounded, slightly produced anteriorly, not swollen with regular border; anterior and lateral margin with entire border; posterior margin without border. Prothoracic hypomeron smooth; prosternal process with sides parallel, with apex rounded, its surface with lateral carinae short, joined with procoxal bordering line. Prosternum in front of coxa about 0.60 × as long as coxal longitudinal diameter at the same position, anterior margin with complete bordering line; procoxal cavity transverse oval; procoxal bordering line incomplete medially.
|
||
</paragraph>
|
||
<paragraph id="8BE9AD13FFDFB31A258EE640FC21FEB3" blockId="16.[189,1399,1817,2019]" lastBlockId="18.[189,1398,265,327]" lastPageId="18" lastPageNumber="48" pageId="16" pageNumber="46">PTEROTHORAX. Mesoventrite with anterior edge straight with complete raised border; mesoventral process at median width of coxa about equal as corresponding coxal diameter; meso-metaventral articulation with suture visible; junction straight. Scutellar shield triangular, bordered; surface punctate and haired. Elytra with sides rounded; lateral margins slender, entirely visible from above; humeral angles present; elytral epipleuron wide, about 3.0 × as wide as corresponding metaepisternum, incomplete apically, with shallow foveae for reception of femora; inner margin with border line fading before base of elytron. Metaventrite with discrimen long but incomplete anteriorly; metaventral postcoxal lines joined at middle, laterally complete, rounded; metaventrite finely punctate.</paragraph>
|
||
<caption id="DF29FD9BFFDEB319258EE7A1FEDFF81C" ID-DOI="http://doi.org/10.5281/zenodo.7234871" ID-Zenodo-Dep="7234871" httpUri="https://zenodo.org/record/7234871/files/figure.png" pageId="17" pageNumber="47" startId="17.[189,232,1787,1813]" targetBox="[245,1342,265,1749]" targetPageId="17">
|
||
<paragraph id="8BE9AD13FFDEB319258EE7A1FEDFF81C" blockId="17.[189,1399,1787,2024]" pageId="17" pageNumber="47">
|
||
<emphasis id="B9227101FFDEB319258EE7A1FE91F8E1" bold="true" box="[189,270,1787,1813]" pageId="17" pageNumber="47">Fig. 7.</emphasis>
|
||
<taxonomicName id="4C56D690FFDEB319242BE7A1FD96F8E1" authority="Szawaryn & Czerwiński, 2022" authorityName="Szawaryn & Czerwiński" authorityYear="2022" box="[280,521,1787,1813]" class="Insecta" family="Coccinellidae" genus="Chnoodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="47" phylum="Arthropoda" rank="species" species="yanayacu" status="sp. nov.">
|
||
<emphasis id="B9227101FFDEB319242BE7A1FD96F8E1" box="[280,521,1787,1813]" italics="true" pageId="17" pageNumber="47">Chnoodes yanayacu</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A211CC7AFFDEB3192720E7A6FDEFF8E2" box="[531,624,1788,1814]" pageId="17" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="B9227101FFDEB319274FE7A1FD10F8E1" bold="true" box="[636,655,1787,1813]" pageId="17" pageNumber="47">A</emphasis>
|
||
. Paratype (MNHW), ♀, abdomen.
|
||
<emphasis id="B9227101FFDEB319210AE7A1FBD2F8E1" bold="true" box="[1081,1101,1787,1813]" pageId="17" pageNumber="47">B</emphasis>
|
||
. Holotype (MNHW), ♂, abdomen.
|
||
<emphasis id="B9227101FFDEB319240BE644FED2F8CC" bold="true" box="[312,333,1822,1848]" pageId="17" pageNumber="47">C</emphasis>
|
||
. Paratype (MNHW), ♀, tergite VIII.
|
||
<emphasis id="B9227101FFDEB319263AE644FC81F8CC" bold="true" box="[777,798,1822,1848]" pageId="17" pageNumber="47">D</emphasis>
|
||
. Paratype (MNHW), ♀, sternite VIII.
|
||
<emphasis id="B9227101FFDEB31921D5E644FB65F8CC" bold="true" box="[1254,1274,1822,1848]" pageId="17" pageNumber="47">E</emphasis>
|
||
. Holotype (MNHW), ♂, tergite VIII.
|
||
<emphasis id="B9227101FFDEB31924C0E618FD9AF8A8" bold="true" box="[499,517,1858,1884]" pageId="17" pageNumber="47">F</emphasis>
|
||
. Holotype (MNHW), ♂, sternite VIII.
|
||
<emphasis id="B9227101FFDEB31926FAE61BFC7FF8AF" bold="true" box="[969,992,1857,1883]" pageId="17" pageNumber="47">G</emphasis>
|
||
. Paratype (MNHW), spermatheca.
|
||
<emphasis id="B9227101FFDEB319258EE63FFF4BF88B" bold="true" box="[189,212,1893,1919]" pageId="17" pageNumber="47">H</emphasis>
|
||
. Paratype (MNHW), ♀, genitalia, ventral.
|
||
<emphasis id="B9227101FFDEB31927F3E63FFD54F88B" bold="true" box="[704,715,1893,1919]" pageId="17" pageNumber="47">I</emphasis>
|
||
. Holotype (MNHW), penis, lateral.
|
||
<emphasis id="B9227101FFDEB319215EE63FFBE3F88B" bold="true" box="[1133,1148,1893,1919]" pageId="17" pageNumber="47">J</emphasis>
|
||
. Holotype (MNHW), penis tip, inner view.
|
||
<emphasis id="B9227101FFDEB3192480E6D2FE55F856" bold="true" box="[435,458,1928,1954]" pageId="17" pageNumber="47">K</emphasis>
|
||
. Holotype (MNHW), penis tip, lateral view.
|
||
<emphasis id="B9227101FFDEB31926E2E6D2FC7AF856" bold="true" box="[977,997,1928,1954]" pageId="17" pageNumber="47">L</emphasis>
|
||
. Holotype (MNHW), ♂, segments IX and X, dorsal view.
|
||
<emphasis id="B9227101FFDEB31924E6E6F1FE6EF831" bold="true" box="[469,497,1963,1989]" pageId="17" pageNumber="47">M</emphasis>
|
||
. Holotype (MNHW), tegmen, lateral view.
|
||
<emphasis id="B9227101FFDEB31926C9E6F1FB90F831" bold="true" box="[1018,1039,1963,1989]" pageId="17" pageNumber="47">N</emphasis>
|
||
. Holotype (MNHW), tegmen, inner view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE9AD13FFDDB31A258EE034FD38FE3A" blockId="18.[189,1399,366,463]" pageId="18" pageNumber="48">LEGS. Legs with trochanters angulate; femora relatively large, excavated to receive tibiae, tibiae shallowly excavated to receive the tarsi, tibial apices without spurs; tarsi with four tarsomeres, third very small; tarsal claws of all legs in both sexes bifid.</paragraph>
|
||
<paragraph id="8BE9AD13FFDDB31A258EE0ACFE27FD17" blockId="18.[189,1399,501,739]" pageId="18" pageNumber="48">
|
||
ABDOMEN. Abdomen with 5 ventrites in both sexes (
|
||
<figureCitation id="136DB196FFDDB31A262FE0AFFC01FDE4" box="[796,926,501,528]" captionStart="Fig" captionStartId="17.[189,232,1787,1813]" captionTargetBox="[245,1342,265,1749]" captionTargetId="figure-13@17.[245,1342,265,1749]" captionTargetPageId="17" captionText="Fig. 7. Chnoodes yanayacu sp. nov. A. Paratype (MNHW), ♀, abdomen. B. Holotype (MNHW), ♂, abdomen. C. Paratype (MNHW), ♀, tergite VIII. D. Paratype (MNHW), ♀, sternite VIII. E. Holotype (MNHW), ♂, tergite VIII. F. Holotype (MNHW), ♂, sternite VIII. G. Paratype (MNHW), spermatheca. H. Paratype (MNHW), ♀, genitalia, ventral. I. Holotype (MNHW), penis, lateral. J. Holotype (MNHW), penis tip, inner view. K. Holotype (MNHW), penis tip, lateral view. L. Holotype (MNHW), ♂, segments IX and X, dorsal view. M. Holotype (MNHW), tegmen, lateral view. N. Holotype (MNHW), tegmen, inner view." figureDoi="http://doi.org/10.5281/zenodo.7234871" httpUri="https://zenodo.org/record/7234871/files/figure.png" pageId="18" pageNumber="48">Fig. 7A–B</figureCitation>
|
||
); ventrite 1, 1.20 × as long as ventrite 2, ventrite 2, 1.20 × as long as ventrites 3 and 4; abdominal postcoxal lines separate medially, incomplete, posteriorly reaching hind margin of ventrite 1; additional oblique line on ventrite 1 present; hind margin of ventrite
|
||
<quantity id="4CAE00F6FFDDB31A2472E305FEEFFD8D" box="[321,368,607,634]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="18" pageNumber="48" unit="in" value="5.0">5 in</quantity>
|
||
both sexes rounded; in male tergite VIII (
|
||
<figureCitation id="136DB196FFDDB31A2659E305FC5BFD8D" box="[874,964,607,633]" captionStart="Fig" captionStartId="17.[189,232,1787,1813]" captionTargetBox="[245,1342,265,1749]" captionTargetId="figure-13@17.[245,1342,265,1749]" captionTargetPageId="17" captionText="Fig. 7. Chnoodes yanayacu sp. nov. A. Paratype (MNHW), ♀, abdomen. B. Holotype (MNHW), ♂, abdomen. C. Paratype (MNHW), ♀, tergite VIII. D. Paratype (MNHW), ♀, sternite VIII. E. Holotype (MNHW), ♂, tergite VIII. F. Holotype (MNHW), ♂, sternite VIII. G. Paratype (MNHW), spermatheca. H. Paratype (MNHW), ♀, genitalia, ventral. I. Holotype (MNHW), penis, lateral. J. Holotype (MNHW), penis tip, inner view. K. Holotype (MNHW), penis tip, lateral view. L. Holotype (MNHW), ♂, segments IX and X, dorsal view. M. Holotype (MNHW), tegmen, lateral view. N. Holotype (MNHW), tegmen, inner view." figureDoi="http://doi.org/10.5281/zenodo.7234871" httpUri="https://zenodo.org/record/7234871/files/figure.png" pageId="18" pageNumber="48">Fig. 7E</figureCitation>
|
||
) and sternite VIII (
|
||
<figureCitation id="136DB196FFDDB31A219EE305FA99FD8D" box="[1197,1286,607,633]" captionStart="Fig" captionStartId="17.[189,232,1787,1813]" captionTargetBox="[245,1342,265,1749]" captionTargetId="figure-13@17.[245,1342,265,1749]" captionTargetPageId="17" captionText="Fig. 7. Chnoodes yanayacu sp. nov. A. Paratype (MNHW), ♀, abdomen. B. Holotype (MNHW), ♂, abdomen. C. Paratype (MNHW), ♀, tergite VIII. D. Paratype (MNHW), ♀, sternite VIII. E. Holotype (MNHW), ♂, tergite VIII. F. Holotype (MNHW), ♂, sternite VIII. G. Paratype (MNHW), spermatheca. H. Paratype (MNHW), ♀, genitalia, ventral. I. Holotype (MNHW), penis, lateral. J. Holotype (MNHW), penis tip, inner view. K. Holotype (MNHW), penis tip, lateral view. L. Holotype (MNHW), ♂, segments IX and X, dorsal view. M. Holotype (MNHW), tegmen, lateral view. N. Holotype (MNHW), tegmen, inner view." figureDoi="http://doi.org/10.5281/zenodo.7234871" httpUri="https://zenodo.org/record/7234871/files/figure.png" pageId="18" pageNumber="48">Fig. 7F</figureCitation>
|
||
) truncate apically; sternite VIII with shallow, but clearly visible central depression; in female tergite VIII (
|
||
<figureCitation id="136DB196FFDDB31A2026E3D8FAF1FD68" box="[1301,1390,642,668]" captionStart="Fig" captionStartId="17.[189,232,1787,1813]" captionTargetBox="[245,1342,265,1749]" captionTargetId="figure-13@17.[245,1342,265,1749]" captionTargetPageId="17" captionText="Fig. 7. Chnoodes yanayacu sp. nov. A. Paratype (MNHW), ♀, abdomen. B. Holotype (MNHW), ♂, abdomen. C. Paratype (MNHW), ♀, tergite VIII. D. Paratype (MNHW), ♀, sternite VIII. E. Holotype (MNHW), ♂, tergite VIII. F. Holotype (MNHW), ♂, sternite VIII. G. Paratype (MNHW), spermatheca. H. Paratype (MNHW), ♀, genitalia, ventral. I. Holotype (MNHW), penis, lateral. J. Holotype (MNHW), penis tip, inner view. K. Holotype (MNHW), penis tip, lateral view. L. Holotype (MNHW), ♂, segments IX and X, dorsal view. M. Holotype (MNHW), tegmen, lateral view. N. Holotype (MNHW), tegmen, inner view." figureDoi="http://doi.org/10.5281/zenodo.7234871" httpUri="https://zenodo.org/record/7234871/files/figure.png" pageId="18" pageNumber="48">Fig. 7C</figureCitation>
|
||
) and sternite VIII (
|
||
<figureCitation id="136DB196FFDDB31A24A7E3FFFE6DFD34" box="[404,498,677,704]" captionStart="Fig" captionStartId="17.[189,232,1787,1813]" captionTargetBox="[245,1342,265,1749]" captionTargetId="figure-13@17.[245,1342,265,1749]" captionTargetPageId="17" captionText="Fig. 7. Chnoodes yanayacu sp. nov. A. Paratype (MNHW), ♀, abdomen. B. Holotype (MNHW), ♂, abdomen. C. Paratype (MNHW), ♀, tergite VIII. D. Paratype (MNHW), ♀, sternite VIII. E. Holotype (MNHW), ♂, tergite VIII. F. Holotype (MNHW), ♂, sternite VIII. G. Paratype (MNHW), spermatheca. H. Paratype (MNHW), ♀, genitalia, ventral. I. Holotype (MNHW), penis, lateral. J. Holotype (MNHW), penis tip, inner view. K. Holotype (MNHW), penis tip, lateral view. L. Holotype (MNHW), ♂, segments IX and X, dorsal view. M. Holotype (MNHW), tegmen, lateral view. N. Holotype (MNHW), tegmen, inner view." figureDoi="http://doi.org/10.5281/zenodo.7234871" httpUri="https://zenodo.org/record/7234871/files/figure.png" pageId="18" pageNumber="48">Fig. 7D</figureCitation>
|
||
) rounded apically. Sternites VIII and ventrites VIII in both sexes with long hairs on hind margin.
|
||
</paragraph>
|
||
<paragraph id="8BE9AD13FFDDB31A258EE251FB00FC20" blockId="18.[189,1399,778,981]" pageId="18" pageNumber="48">
|
||
MALE TERMINALIA AND GENITALIA. Sternite IX (
|
||
<figureCitation id="136DB196FFDDB31A27D2E250FCA3FCD0" box="[737,828,778,804]" captionStart="Fig" captionStartId="17.[189,232,1787,1813]" captionTargetBox="[245,1342,265,1749]" captionTargetId="figure-13@17.[245,1342,265,1749]" captionTargetPageId="17" captionText="Fig. 7. Chnoodes yanayacu sp. nov. A. Paratype (MNHW), ♀, abdomen. B. Holotype (MNHW), ♂, abdomen. C. Paratype (MNHW), ♀, tergite VIII. D. Paratype (MNHW), ♀, sternite VIII. E. Holotype (MNHW), ♂, tergite VIII. F. Holotype (MNHW), ♂, sternite VIII. G. Paratype (MNHW), spermatheca. H. Paratype (MNHW), ♀, genitalia, ventral. I. Holotype (MNHW), penis, lateral. J. Holotype (MNHW), penis tip, inner view. K. Holotype (MNHW), penis tip, lateral view. L. Holotype (MNHW), ♂, segments IX and X, dorsal view. M. Holotype (MNHW), tegmen, lateral view. N. Holotype (MNHW), tegmen, inner view." figureDoi="http://doi.org/10.5281/zenodo.7234871" httpUri="https://zenodo.org/record/7234871/files/figure.png" pageId="18" pageNumber="48">Fig. 7L</figureCitation>
|
||
) with bilobed, partially membranous additional sclerite at base of apodeme; apodeme rod-like, straight; tergite X very short, transverse. Tegmen in inner view (
|
||
<figureCitation id="136DB196FFDDB31A2436E20AFEFFFC9F" box="[261,352,848,875]" captionStart="Fig" captionStartId="17.[189,232,1787,1813]" captionTargetBox="[245,1342,265,1749]" captionTargetId="figure-13@17.[245,1342,265,1749]" captionTargetPageId="17" captionText="Fig. 7. Chnoodes yanayacu sp. nov. A. Paratype (MNHW), ♀, abdomen. B. Holotype (MNHW), ♂, abdomen. C. Paratype (MNHW), ♀, tergite VIII. D. Paratype (MNHW), ♀, sternite VIII. E. Holotype (MNHW), ♂, tergite VIII. F. Holotype (MNHW), ♂, sternite VIII. G. Paratype (MNHW), spermatheca. H. Paratype (MNHW), ♀, genitalia, ventral. I. Holotype (MNHW), penis, lateral. J. Holotype (MNHW), penis tip, inner view. K. Holotype (MNHW), penis tip, lateral view. L. Holotype (MNHW), ♂, segments IX and X, dorsal view. M. Holotype (MNHW), tegmen, lateral view. N. Holotype (MNHW), tegmen, inner view." figureDoi="http://doi.org/10.5281/zenodo.7234871" httpUri="https://zenodo.org/record/7234871/files/figure.png" pageId="18" pageNumber="48">Fig. 7N</figureCitation>
|
||
) with penis guide parallel, apex acute; parameres (
|
||
<figureCitation id="136DB196FFDDB31A2699E20AFB93FC9F" box="[938,1036,848,875]" captionStart="Fig" captionStartId="17.[189,232,1787,1813]" captionTargetBox="[245,1342,265,1749]" captionTargetId="figure-13@17.[245,1342,265,1749]" captionTargetPageId="17" captionText="Fig. 7. Chnoodes yanayacu sp. nov. A. Paratype (MNHW), ♀, abdomen. B. Holotype (MNHW), ♂, abdomen. C. Paratype (MNHW), ♀, tergite VIII. D. Paratype (MNHW), ♀, sternite VIII. E. Holotype (MNHW), ♂, tergite VIII. F. Holotype (MNHW), ♂, sternite VIII. G. Paratype (MNHW), spermatheca. H. Paratype (MNHW), ♀, genitalia, ventral. I. Holotype (MNHW), penis, lateral. J. Holotype (MNHW), penis tip, inner view. K. Holotype (MNHW), penis tip, lateral view. L. Holotype (MNHW), ♂, segments IX and X, dorsal view. M. Holotype (MNHW), tegmen, lateral view. N. Holotype (MNHW), tegmen, inner view." figureDoi="http://doi.org/10.5281/zenodo.7234871" httpUri="https://zenodo.org/record/7234871/files/figure.png" pageId="18" pageNumber="48">Fig. 7M</figureCitation>
|
||
), well developed, simple, more than 2 × as long as penis guide, with fringe of dense, long setae on apices and edges; tegminal strut wide. Penis capsule (
|
||
<figureCitation id="136DB196FFDDB31A245DE2CDFE20FC45" box="[366,447,919,945]" captionStart="Fig" captionStartId="7.[189,232,1892,1918]" captionTargetBox="[189,1398,557,1852]" captionTargetId="figure-13@7.[189,1398,557,1852]" captionTargetPageId="7" captionText="Fig. 3. SEM images. A–D. Eupalea borowieci sp. nov., paratype (MIZ). E. Hyperaspis rutai sp. nov., paratype (MIZ). F. Hinda ecuadorica Gordon & Canepari, 2013, ♂, (MIZ). A. Head and pronotum. B. Head, ventral. C. Prosternum and mesoventrite. D. Habitus, ventral view. E. Head, dorsal view. F. Distal part of abdomen, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7234845" httpUri="https://zenodo.org/record/7234845/files/figure.png" pageId="18" pageNumber="48">Fig. 3I</figureCitation>
|
||
) with inner arm narrow, outer arm broadened but short; penis of equal diameter along entire length, widened on tip (
|
||
<figureCitation id="136DB196FFDDB31A2755E2E0FD5BFC20" box="[614,708,954,980]" captionStart="Fig" captionStartId="17.[189,232,1787,1813]" captionTargetBox="[245,1342,265,1749]" captionTargetId="figure-13@17.[245,1342,265,1749]" captionTargetPageId="17" captionText="Fig. 7. Chnoodes yanayacu sp. nov. A. Paratype (MNHW), ♀, abdomen. B. Holotype (MNHW), ♂, abdomen. C. Paratype (MNHW), ♀, tergite VIII. D. Paratype (MNHW), ♀, sternite VIII. E. Holotype (MNHW), ♂, tergite VIII. F. Holotype (MNHW), ♂, sternite VIII. G. Paratype (MNHW), spermatheca. H. Paratype (MNHW), ♀, genitalia, ventral. I. Holotype (MNHW), penis, lateral. J. Holotype (MNHW), penis tip, inner view. K. Holotype (MNHW), penis tip, lateral view. L. Holotype (MNHW), ♂, segments IX and X, dorsal view. M. Holotype (MNHW), tegmen, lateral view. N. Holotype (MNHW), tegmen, inner view." figureDoi="http://doi.org/10.5281/zenodo.7234871" httpUri="https://zenodo.org/record/7234871/files/figure.png" pageId="18" pageNumber="48">Fig. 7K</figureCitation>
|
||
), tip not partially membranous (
|
||
<figureCitation id="136DB196FFDDB31A210EE2E0FB0EFC20" box="[1085,1169,954,980]" captionStart="Fig" captionStartId="17.[189,232,1787,1813]" captionTargetBox="[245,1342,265,1749]" captionTargetId="figure-13@17.[245,1342,265,1749]" captionTargetPageId="17" captionText="Fig. 7. Chnoodes yanayacu sp. nov. A. Paratype (MNHW), ♀, abdomen. B. Holotype (MNHW), ♂, abdomen. C. Paratype (MNHW), ♀, tergite VIII. D. Paratype (MNHW), ♀, sternite VIII. E. Holotype (MNHW), ♂, tergite VIII. F. Holotype (MNHW), ♂, sternite VIII. G. Paratype (MNHW), spermatheca. H. Paratype (MNHW), ♀, genitalia, ventral. I. Holotype (MNHW), penis, lateral. J. Holotype (MNHW), penis tip, inner view. K. Holotype (MNHW), penis tip, lateral view. L. Holotype (MNHW), ♂, segments IX and X, dorsal view. M. Holotype (MNHW), tegmen, lateral view. N. Holotype (MNHW), tegmen, inner view." figureDoi="http://doi.org/10.5281/zenodo.7234871" httpUri="https://zenodo.org/record/7234871/files/figure.png" pageId="18" pageNumber="48">Fig. 7J</figureCitation>
|
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).
|
||
</paragraph>
|
||
<paragraph id="8BE9AD13FFDDB31A258EE2A6FC13FB8B" blockId="18.[189,1398,1019,1151]" pageId="18" pageNumber="48">
|
||
FEMALE GENITALIA. Sperm duct uniform in diameter, shorter than spermatheca; infundibulum absent; spermatheca vermiform, without clear nodulus and ramus, but with mammiliform projection on apex (
|
||
<figureCitation id="136DB196FFDDB31A25F5E518FEBAFBA8" box="[198,293,1090,1116]" captionStart="Fig" captionStartId="17.[189,232,1787,1813]" captionTargetBox="[245,1342,265,1749]" captionTargetId="figure-13@17.[245,1342,265,1749]" captionTargetPageId="17" captionText="Fig. 7. Chnoodes yanayacu sp. nov. A. Paratype (MNHW), ♀, abdomen. B. Holotype (MNHW), ♂, abdomen. C. Paratype (MNHW), ♀, tergite VIII. D. Paratype (MNHW), ♀, sternite VIII. E. Holotype (MNHW), ♂, tergite VIII. F. Holotype (MNHW), ♂, sternite VIII. G. Paratype (MNHW), spermatheca. H. Paratype (MNHW), ♀, genitalia, ventral. I. Holotype (MNHW), penis, lateral. J. Holotype (MNHW), penis tip, inner view. K. Holotype (MNHW), penis tip, lateral view. L. Holotype (MNHW), ♂, segments IX and X, dorsal view. M. Holotype (MNHW), tegmen, lateral view. N. Holotype (MNHW), tegmen, inner view." figureDoi="http://doi.org/10.5281/zenodo.7234871" httpUri="https://zenodo.org/record/7234871/files/figure.png" pageId="18" pageNumber="48">Fig. 7G</figureCitation>
|
||
); spermathecal accessory gland adjacent to sperm duct. Coxites elongate, rounded on apices, inner surface covered with long hairs; styli distinct (
|
||
<figureCitation id="136DB196FFDDB31A2612E53FFCE0FB8B" box="[801,895,1125,1151]" captionStart="Fig" captionStartId="17.[189,232,1787,1813]" captionTargetBox="[245,1342,265,1749]" captionTargetId="figure-13@17.[245,1342,265,1749]" captionTargetPageId="17" captionText="Fig. 7. Chnoodes yanayacu sp. nov. A. Paratype (MNHW), ♀, abdomen. B. Holotype (MNHW), ♂, abdomen. C. Paratype (MNHW), ♀, tergite VIII. D. Paratype (MNHW), ♀, sternite VIII. E. Holotype (MNHW), ♂, tergite VIII. F. Holotype (MNHW), ♂, sternite VIII. G. Paratype (MNHW), spermatheca. H. Paratype (MNHW), ♀, genitalia, ventral. I. Holotype (MNHW), penis, lateral. J. Holotype (MNHW), penis tip, inner view. K. Holotype (MNHW), penis tip, lateral view. L. Holotype (MNHW), ♂, segments IX and X, dorsal view. M. Holotype (MNHW), tegmen, lateral view. N. Holotype (MNHW), tegmen, inner view." figureDoi="http://doi.org/10.5281/zenodo.7234871" httpUri="https://zenodo.org/record/7234871/files/figure.png" pageId="18" pageNumber="48">Fig. 7H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |