536 lines
73 KiB
XML
536 lines
73 KiB
XML
<document id="72AAD2EC6E701A2337C701D735F66427" ID-CLB-Dataset="112029" ID-DOI="10.11646/zootaxa.5169.3.2" ID-GBIF-Dataset="6ffbcac4-29d1-4be5-9492-5f0d1432eea7" ID-ISSN="1175-5326" ID-Zenodo-Dep="6952146" ID-ZooBank="8419DE17-AF1B-4D58-A486-B9ECD5AC5EA5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1659438232014" checkinUser="plazi" docAuthor="Barjadze, Shalva, Parimuchová, Andrea, Raschmanová, Natália, Maghradze, Eter & Kováč, Ľubomír" docDate="2022" docId="03FF1701192BFF8595AAFA539D2821FD" docLanguage="en" docName="zootaxa.5169.3.2.pdf" docOrigin="Zootaxa 5169 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Plutomurus kharagauliensis Barjadze, Kovac & Parimuchova 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="258" masterDocId="FFC66F791929FF83953DFFB59E082210" masterDocTitle="Two new species of Plutomurus Yosii (Collembola: Tomoceridae) from the Caucasus and central Europe" masterLastPageNumber="266" masterPageNumber="252" pageNumber="254" updateTime="1699362624138" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="F17DC99B805C2A5D389D6BF6B09DE8A2">
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<mods:title id="7E95FB8F518BEED5E932EE214C4D9C82">Two new species of Plutomurus Yosii (Collembola: Tomoceridae) from the Caucasus and central Europe</mods:title>
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<mods:name id="BDE6DF99D33A1F7FAF510F18EAA9D6DC" type="personal">
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<mods:roleTerm id="CC26AB473EBA4782DB9B6D4D1FFD4F9B">Author</mods:roleTerm>
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<mods:namePart id="8B26DFA18227720DFDEDF2BB76FFE2B0">Barjadze, Shalva</mods:namePart>
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<mods:affiliation id="C4BA992A2DA8B9AD7563EFA87DBD576C">Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Republic of Georgia</mods:affiliation>
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<mods:roleTerm id="078834F92DA9556172629616C3C581D5">Author</mods:roleTerm>
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<mods:namePart id="D84120DC013DB6837EDE8F5B95FF0D0E">Parimuchová, Andrea</mods:namePart>
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<mods:nameIdentifier id="359D5283A975C1BA4AA39CC6898FF040" type="ORCID">0000-0002-8977-9991</mods:nameIdentifier>
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<mods:affiliation id="C70B8E90CC8226A20FD6726F4F755EA3">Institute of Biology and Ecology, Faculty of Science, Pavol Jozef Šafárik University in Košice, Košice, Šrobárova 2, 04154 Košice, Slovakia & andrea. parimuchova @ upjs. sk; https: // orcid. org / 0000 - 0002 - 8977 - 9991</mods:affiliation>
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<mods:nameIdentifier id="24FA1A15520C44093FA1F3EF872508D8" type="email">andrea.parimuchova@upjs.sk</mods:nameIdentifier>
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<mods:roleTerm id="4A82ED7417BD6194700ACA9E6534D7DF">Author</mods:roleTerm>
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<mods:namePart id="96590213B78B93BC62C3796683FBBF3D">Raschmanová, Natália</mods:namePart>
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<mods:nameIdentifier id="E9F905BB8DC2F3109A79CECCF46E6B58" type="ORCID">0000-0002-9476-1753</mods:nameIdentifier>
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<mods:affiliation id="35148773128621F8AC0961602109AFC3">Institute of Biology and Ecology, Faculty of Science, Pavol Jozef Šafárik University in Košice, Košice, Šrobárova 2, 04154 Košice, Slovakia & natalia. raschmanova @ upjs. sk; https: // orcid. org / 0000 - 0002 - 9476 - 1753</mods:affiliation>
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<mods:nameIdentifier id="F7C075628F7AEFC185F93E019D95A81C" type="email">natalia.raschmanova@upjs.sk</mods:nameIdentifier>
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<mods:namePart id="B7CA354B85725E4553DF104DCD9B129B">Maghradze, Eter</mods:namePart>
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<mods:nameIdentifier id="EAE9CC11FD7E9E83F27E81F54B596407" type="ORCID">0000-0002-4796-9439</mods:nameIdentifier>
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<mods:affiliation id="F08AC87FFBD159CC6E8624067C4ABFB7">Institute of Zoology, Ilia State University, Giorgi Tsereteli 3, 0162, Tbilisi, Republic of Georgia & eter. magradze. 1 @ iliauni. edu. ge; https: // orcid. org / 0000 - 0002 - 4796 - 9439</mods:affiliation>
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<mods:nameIdentifier id="A5BAEEAACC9B05D793CE26B9EB0DAFB0" type="email">eter.magradze.1@iliauni.edu.ge</mods:nameIdentifier>
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<mods:namePart id="F3FA4CB593F46111B4B0E6049FFFCB06">Kováč, Ľubomír</mods:namePart>
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<mods:affiliation id="3F27C1B9AC271F66BE709F0B1BE0AA86">Institute of Biology and Ecology, Faculty of Science, Pavol Jozef Šafárik University in Košice, Košice, Šrobárova 2, 04154 Košice, Slovakia & lubomir. kovac @ upjs. sk; https: // orcid. org / 0000 - 0001 - 8194 - 2128</mods:affiliation>
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<mods:nameIdentifier id="BF0A91EC4793FB985822BA73A6CB24D6" type="email">lubomir.kovac@upjs.sk</mods:nameIdentifier>
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<mods:title id="45C12F189D0943B5A84E6F6E9AD7D8D9">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="23D20E1F22ECF015320E21580E53E382">
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<mods:date id="F1F1B87D41BA567912366305A1ADF9F4">2022</mods:date>
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<mods:number id="1E64B47EA09142D0896641870E4E634E">2022-07-29</mods:number>
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<mods:detail id="DBE15E47314CC738383C692F5FB4137B" type="volume">
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<mods:number id="0CEEE93D1BC0DFAEF5C7F46AF190FEC6">5169</mods:number>
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<mods:detail id="0DD63DE9D2C8ADC37A18C5B34DD127DD" type="issue">
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<mods:number id="6844BB290B34785B8CE786926F1ECAEA">3</mods:number>
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<mods:start id="48B64E0344D35DE4E6CB629586A6CC72">252</mods:start>
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<mods:classification id="2D9927AB63F5008A2DA6DC07CC9292EA">journal article</mods:classification>
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<mods:identifier id="50344629FECCA771D7111976ACA67D6D" type="CLB-Dataset">112029</mods:identifier>
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<mods:identifier id="5C5271D329267617E971F0E3FB40859A" type="DOI">10.11646/zootaxa.5169.3.2</mods:identifier>
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<mods:identifier id="560037728B2A9239286F700385C06D9A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="C9BEAD2AB4572A6FC0B5E6A0CA59EF84" type="Zenodo-Dep">6952146</mods:identifier>
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<mods:identifier id="54D43B1C5245F5C0058EF1B09515F219" type="ZooBank">8419DE17-AF1B-4D58-A486-B9ECD5AC5EA5</mods:identifier>
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<treatment id="03FF1701192BFF8595AAFA539D2821FD" ID-DOI="http://doi.org/10.5281/zenodo.6957988" ID-GBIF-Taxon="198574462" ID-Zenodo-Dep="6957988" LSID="urn:lsid:plazi:treatment:03FF1701192BFF8595AAFA539D2821FD" httpUri="http://treatment.plazi.org/id/03FF1701192BFF8595AAFA539D2821FD" lastPageId="6" lastPageNumber="258" pageId="2" pageNumber="254">
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<subSubSection id="C34CF59C192BFF8195AAFA539DF42410" box="[151,1020,1509,1537]" pageId="2" pageNumber="254" type="nomenclature">
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<paragraph id="8BE9A617192BFF8195AAFA539DF42410" blockId="2.[151,1020,1509,1573]" box="[151,1020,1509,1537]" pageId="2" pageNumber="254">
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<heading id="D0A1117B192BFF8195AAFA539DF42410" bold="true" box="[151,1020,1509,1537]" fontSize="11" level="1" pageId="2" pageNumber="254" reason="1">
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<emphasis id="B9227A05192BFF8195AAFA539DF42410" bold="true" box="[151,1020,1509,1537]" pageId="2" pageNumber="254">
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<taxonomicName id="4C56DD94192BFF8195AAFA539D9F2410" ID-CoL="B8NWK" authority="Barjadze, Kovac & Parimuchova" authorityName="Barjadze, Kovac & Parimuchova" authorityYear="2022" box="[151,919,1509,1537]" class="Collembola" family="Tomoceridae" genus="Plutomurus" kingdom="Animalia" order="Entomobryomorpha" pageId="2" pageNumber="254" phylum="Arthropoda" rank="species" species="kharagauliensis" status="sp. nov.">
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<emphasis id="B9227A05192BFF8195AAFA539FF82411" bold="true" box="[151,496,1510,1537]" italics="true" pageId="2" pageNumber="254">Plutomurus kharagauliensis</emphasis>
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Barjadze, Kováč & Parimuchová
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</taxonomicName>
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<taxonomicNameLabel id="A211C77E192BFF8196A3FA539DF42410" box="[926,1020,1510,1536]" pageId="2" pageNumber="254" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34CF59C192BFF8195AAF9BF9F0E2435" box="[151,262,1546,1573]" pageId="2" pageNumber="254" type="description">
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<paragraph id="8BE9A617192BFF8195AAF9BF9F0E2435" blockId="2.[151,1020,1509,1573]" box="[151,262,1546,1573]" pageId="2" pageNumber="254">
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<figureCitation id="136DBA92192BFF8195AAF9BF9F0E2435" box="[151,262,1546,1573]" captionStart-0="FIGURE 3" captionStart-1="FIGURES 4–11" captionStart-2="FIGURES 12–14" captionStart-3="FIGURE 15" captionStartId-0="3.[151,250,1111,1135]" captionStartId-1="4.[151,264,1860,1884]" captionStartId-2="5.[151,264,1728,1752]" captionStartId-3="6.[151,250,2016,2040]" captionTargetBox-0="[199,1388,181,1087]" captionTargetBox-1="[199,1391,190,1827]" captionTargetBox-2="[178,1403,190,1695]" captionTargetBox-3="[151,1436,1029,1993]" captionTargetId-0="figure-22@3.[199,1388,181,1087]" captionTargetId-1="figure-17@4.[190,1397,181,1836]" captionTargetId-2="figure-22@5.[163,1425,181,1704]" captionTargetId-3="figure-412@6.[151,1436,1029,1993]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 3. Habitus of Plutomurus kharagauliensis sp. nov. (scale bar: 0.5 mm)." captionText-1="FIGURES 4–11. Plutomurus kharagauliensis sp. nov. 4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm)." captionText-2="FIGURES 12–14. Dorsal chaetotaxy in Plutomurus kharagauliensis sp. nov. 12, Head.A: Anterior macrochaetae, B: interocular Mc, C: postocular Mc; 13, Th. II–III; 14, Abd. I–V (big ovals and round circles near Ant. I represent eyes; large black cir- cle—macrochaeta; open circles—mesochaeta; squares with dots—bothriothricha; circles with a slash represent pseudopores)." captionText-3="FIGURE 15. Plutomurus kharagauliensis sp. nov. Macrochaetae complex on Abd IV (bar: 50 μm)." figureDoi-0="http://doi.org/10.5281/zenodo.6952152" figureDoi-1="http://doi.org/10.5281/zenodo.6952154" figureDoi-2="http://doi.org/10.5281/zenodo.6952156" figureDoi-3="http://doi.org/10.5281/zenodo.6952158" httpUri-0="https://zenodo.org/record/6952152/files/figure.png" httpUri-1="https://zenodo.org/record/6952154/files/figure.png" httpUri-2="https://zenodo.org/record/6952156/files/figure.png" httpUri-3="https://zenodo.org/record/6952158/files/figure.png" pageId="2" pageNumber="254">Figs 3–15</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34CF59C192BFF8195AAF9E79B002578" pageId="2" pageNumber="254" type="materials_examined">
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<paragraph id="8BE9A617192BFF8195AAF9E79CA72480" blockId="2.[151,1436,1618,2040]" pageId="2" pageNumber="254">
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<materialsCitation id="3B3EAC4A192BFF8195AAF9E79CA22480" ID-GBIF-Occurrence="3878837325" country="Georgia" county="Kharagauli" elevation="644" location="Zemo" municipality="Amashuketi" pageId="2" pageNumber="254" specimenCount="1" stateProvince="Imereti" typeStatus="holotype">
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<emphasis id="B9227A05192BFF8195AAF9E79F3E247C" bold="true" box="[151,310,1618,1644]" pageId="2" pageNumber="254">
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<typeStatus id="54ED18B5192BFF8195AAF9E79EDA247C" box="[151,210,1618,1644]" pageId="2" pageNumber="254">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F341E687192BFF819400F9E79FB6247C" box="[317,446,1618,1644]" name="Georgia" pageId="2" pageNumber="254">GEORGIA</collectingCountry>
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,
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<collectingRegion id="499268F5192BFF8194F4F9E79C63247C" box="[457,619,1618,1644]" country="Georgia" name="Imereti" pageId="2" pageNumber="254">Imereti region</collectingRegion>
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,
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<collectingCounty id="6288DE9B192BFF81974AF9E79CFB247C" box="[631,755,1618,1644]" pageId="2" pageNumber="254">Kharagauli</collectingCounty>
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municipality, close to the village
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<collectingMunicipality id="6B8D3C6D192BFF819150F9E79AFF247C" box="[1133,1271,1618,1644]" pageId="2" pageNumber="254">Amashuketi</collectingMunicipality>
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,
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<location id="8E89F0CC192BFF81903EF9E69B4C247D" LSID="urn:lsid:plazi:treatment:03FF1701192BFF8595AAFA539D2821FD:8E89F0CC192BFF81903EF9E69B4C247D" box="[1283,1348,1619,1645]" country="Georgia" county="Kharagauli" municipality="Amashuketi" name="Zemo" pageId="2" pageNumber="254" stateProvince="Imereti">Zemo</location>
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<collectingRegion id="499268F5192BFF819076F9E79B94247C" box="[1355,1436,1618,1644]" country="Georgia" name="Imereti" pageId="2" pageNumber="254">Imereti</collectingRegion>
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<location id="8E89F0CC192BFF8195AAF9C39EE12480" LSID="urn:lsid:plazi:treatment:03FF1701192BFF8595AAFA539D2821FD:8E89F0CC192BFF8195AAF9C39EE12480" box="[151,233,1654,1680]" country="Georgia" county="Kharagauli" municipality="Amashuketi" name="Plateau" pageId="2" pageNumber="254" stateProvince="Imereti">Plateau</location>
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karst massif,
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||
<location id="8E89F0CC192BFF8194B8F9C39C272480" LSID="urn:lsid:plazi:treatment:03FF1701192BFF8595AAFA539D2821FD:8E89F0CC192BFF8194B8F9C39C272480" box="[389,559,1654,1680]" country="Georgia" county="Kharagauli" municipality="Amashuketi" name="Kozmani Cave" pageId="2" pageNumber="254" stateProvince="Imereti">Kozmani Cave</location>
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,
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<elevation id="007B4124192BFF819707F9C39CA22480" box="[570,682,1654,1681]" metricMagnitude="2" metricUnit="m" metricValue="6.44" pageId="2" pageNumber="254" unit="m" value="644.0">
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<quantity id="4CAE0BF2192BFF819707F9C39C892481" box="[570,641,1654,1681]" metricMagnitude="2" metricUnit="m" metricValue="6.44" pageId="2" pageNumber="254" unit="m" value="644.0">644 m</quantity>
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asl
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</elevation>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BE9A617192BFF8195FAF92F9B002578" blockId="2.[151,1436,1618,2040]" pageId="2" pageNumber="254">
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<emphasis id="B9227A05192BFF8195FAF92F9F7224A4" bold="true" box="[199,378,1690,1716]" pageId="2" pageNumber="254">Type material.</emphasis>
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<materialsCitation id="3B3EAC4A192BFF8194BBF92F9D9E24C8" ID-GBIF-Occurrence="3878837312" collectingDate="2021-11-27" collectionCode="IZISU" collectorName="Sh. Barjadze, E & E. Maghradze & M. Gogshelidze" pageId="2" pageNumber="254" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54ED18B5192BFF8194BBF92F9FFB24A4" box="[390,499,1690,1716]" pageId="2" pageNumber="254" type="holotype">Holotype</typeStatus>
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:
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<collectorName id="26A3C3C1192BFF81973EF92F9C5D24A4" box="[515,597,1690,1716]" pageId="2" pageNumber="254">Female</collectorName>
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on slide, dark zone, hand collecting,
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||
<date id="FFE880D7192BFF819121F92F9A9E24A4" box="[1052,1174,1690,1716]" pageId="2" pageNumber="254" value="2021-11-27">
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<collectingDate id="EFAC793F192BFF819121F92F9A9E24A4" box="[1052,1174,1690,1716]" pageId="2" pageNumber="254" value="2021-11-27">27.xi.2021</collectingDate>
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</date>
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, leg.
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||
<collectorName id="26A3C3C1192BFF8191E1F92F9B9024A5" box="[1244,1432,1690,1717]" pageId="2" pageNumber="254">Sh. Barjadze, E</collectorName>
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. Maghradze,
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<collectorName id="26A3C3C1192BFF819418F90A9FD624C8" box="[293,478,1726,1753]" pageId="2" pageNumber="254">M. Gogshelidze</collectorName>
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, code GEOKOZ20211127-01 (
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<collectionCode id="ED473ED2192BFF81967CF90B9D8724C8" box="[833,911,1726,1752]" pageId="2" pageNumber="254">IZISU</collectionCode>
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)
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</materialsCitation>
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;
|
||
<materialsCitation id="3B3EAC4A192BFF819699F90A9ADB24EC" ID-GBIF-Occurrence="3878837307" collectingDate="2021-11-27" collectionCode="IZISU" collectorName="Sh. Barjadze, E & E. Maghradze & M. Gogshelidze" pageId="2" pageNumber="254" specimenCount="1" typeStatus="paratype">
|
||
<typeStatus id="54ED18B5192BFF819699F90A9A1924C9" box="[932,1041,1727,1753]" pageId="2" pageNumber="254" type="paratype">Paratypes</typeStatus>
|
||
(n=13):
|
||
<specimenCount id="9D506D9E192BFF819145F90B9B2624C8" box="[1144,1326,1726,1752]" pageId="2" pageNumber="254" type="generic">eight specimens</specimenCount>
|
||
on slides with the same data as holotype, code GEOKOZ20211127-02, 03, 04, 05, 08, 09, 10, 11 (
|
||
<collectionCode id="ED473ED2192BFF819140F9579AC324EC" box="[1149,1227,1762,1788]" pageId="2" pageNumber="254">IZISU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B3EAC4A192BFF8191E2F9569A4C2530" ID-GBIF-Occurrence="3878837301" collectingDate="2021-09-14" collectionCode="IZISU" collectorName="E. Maghradze" pageId="2" pageNumber="254" specimenCount="1" typeStatus="paratype">
|
||
<specimenCount id="9D506D9E192BFF8191E2F9569B7124EC" box="[1247,1401,1762,1789]" pageId="2" pageNumber="254" type="generic">one specimen</specimenCount>
|
||
on slide, twilight zone,
|
||
<date id="FFE880D7192BFF819444F8B39FFC2530" box="[377,500,1798,1824]" pageId="2" pageNumber="254" value="2021-09-14">
|
||
<collectingDate id="EFAC793F192BFF819444F8B39FFC2530" box="[377,500,1798,1824]" pageId="2" pageNumber="254" value="2021-09-14">14.ix.2021</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26A3C3C1192BFF819713F8B29CC62530" box="[558,718,1798,1825]" pageId="2" pageNumber="254">E. Maghradze</collectorName>
|
||
, GEOKOZ20210914-14 (
|
||
<collectionCode id="ED473ED2192BFF8196D2F8B39A352530" box="[1007,1085,1798,1824]" pageId="2" pageNumber="254">IZISU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B3EAC4A192BFF819172F8B29B6D2554" ID-GBIF-Occurrence="3878837314" collectingDate="2021-11-27" collectionCode="UPJ" collectorName="Sh. Barjadze, E. & Maghradze, M. & Gogshelidze" pageId="2" pageNumber="254" specimenCount="1" typeStatus="paratype">
|
||
<specimenCount id="9D506D9E192BFF819172F8B29AFD2530" box="[1103,1269,1798,1825]" pageId="2" pageNumber="254" type="generic">two specimens</specimenCount>
|
||
on slides, dark zone,
|
||
<date id="FFE880D7192BFF8195E1F89F9F5E2554" box="[220,342,1834,1860]" pageId="2" pageNumber="254" value="2021-11-27">
|
||
<collectingDate id="EFAC793F192BFF8195E1F89F9F5E2554" box="[220,342,1834,1860]" pageId="2" pageNumber="254" value="2021-11-27">27.xi.2021</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26A3C3C1192BFF8194A8F89F9D932554" box="[405,923,1834,1861]" pageId="2" pageNumber="254">Sh. Barjadze, E. Maghradze, M. Gogshelidze</collectorName>
|
||
, code GEOKOZ20211127-06, 07 (
|
||
<collectionCode id="ED473ED2192BFF819015F89F9B542554" box="[1320,1372,1834,1860]" pageId="2" pageNumber="254">UPJ</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B3EAC4A192BFF81904FF89E9B0C2578" ID-GBIF-Occurrence="3878837324" collectingDate="2021-09-14" collectionCode="UPJ" collectorName="E. Maghradze" pageId="2" pageNumber="254" specimenCount="1" typeStatus="paratype">
|
||
<specimenCount id="9D506D9E192BFF81904FF89E9F042578" pageId="2" pageNumber="254" type="generic">two specimens</specimenCount>
|
||
on slides, twilight zone,
|
||
<date id="FFE880D7192BFF819718F8FB9C972578" box="[549,671,1870,1896]" pageId="2" pageNumber="254" value="2021-09-14">
|
||
<collectingDate id="EFAC793F192BFF819718F8FB9C972578" box="[549,671,1870,1896]" pageId="2" pageNumber="254" value="2021-09-14">14.ix.2021</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26A3C3C1192BFF8197E6F8FA9D732578" box="[731,891,1870,1897]" pageId="2" pageNumber="254">E. Maghradze</collectorName>
|
||
, GEOKOZ20210914-12, 13 (
|
||
<collectionCode id="ED473ED2192BFF8191FAF8FB9AF42578" box="[1223,1276,1870,1896]" pageId="2" pageNumber="254">UPJ</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34CF59C192BFF8595FAF8C79DFE23BD" lastPageId="6" lastPageNumber="258" pageId="2" pageNumber="254" type="description">
|
||
<paragraph id="8BE9A617192BFF8195FAF8C79CFC259C" blockId="2.[151,1436,1618,2040]" box="[199,756,1906,1933]" pageId="2" pageNumber="254">
|
||
<emphasis id="B9227A05192BFF8195FAF8C79F51259C" bold="true" box="[199,345,1906,1932]" pageId="2" pageNumber="254">Description.</emphasis>
|
||
Body length
|
||
<quantity id="4CAE0BF2192BFF8194CDF8C79C9B259D" box="[496,659,1906,1933]" metricMagnitude="-3" metricUnit="m" metricValue="3.2299999999999995" metricValueMax="4.02" metricValueMin="2.44" pageId="2" pageNumber="254" unit="mm" value="3.2299999999999995" valueMax="4.02" valueMin="2.44">2.44–4.02 mm</quantity>
|
||
(
|
||
<figureCitation id="136DBA92192BFF81979FF8C79CEF259C" box="[674,743,1906,1932]" captionStart="FIGURE 3" captionStartId="3.[151,250,1111,1135]" captionTargetBox="[199,1388,181,1087]" captionTargetId="figure-22@3.[199,1388,181,1087]" captionTargetPageId="3" captionText="FIGURE 3. Habitus of Plutomurus kharagauliensis sp. nov. (scale bar: 0.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.6952152" httpUri="https://zenodo.org/record/6952152/files/figure.png" pageId="2" pageNumber="254">Fig. 3</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE9A617192BFF8195FAF8239FF225A0" blockId="2.[151,1436,1618,2040]" box="[199,506,1942,1969]" pageId="2" pageNumber="254">Colour. Body grey colored.</paragraph>
|
||
<paragraph id="8BE9A617192BFF8195FAF80F9C9025E8" blockId="2.[151,1436,1618,2040]" pageId="2" pageNumber="254">Scale distribution. Scales are present dorsally on Ant. I–II, head, Th. and Abd. segments, all leg segments, both sides of ventral tube and ventral side of furca.</paragraph>
|
||
<caption id="DF29F69F192AFF8095AAFBE29DDA267F" ID-DOI="http://doi.org/10.5281/zenodo.6952152" ID-Zenodo-Dep="6952152" box="[151,978,1111,1135]" httpUri="https://zenodo.org/record/6952152/files/figure.png" pageId="3" pageNumber="255" startId="3.[151,250,1111,1135]" targetBox="[199,1388,181,1087]" targetPageId="3">
|
||
<paragraph id="8BE9A617192AFF8095AAFBE29DDA267F" blockId="3.[151,978,1111,1135]" box="[151,978,1111,1135]" pageId="3" pageNumber="255">
|
||
<emphasis id="B9227A05192AFF8095AAFBE29F1B267F" bold="true" box="[151,275,1111,1135]" pageId="3" pageNumber="255">FIGURE 3.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="4C56DD94192AFF8094B7FBE29CA1267F" authority="Barjadze & Parimuchová & Raschmanová & Maghradze & Kováč, 2022" authorityName="Barjadze & Parimuchová & Raschmanová & Maghradze & Kováč" authorityYear="2022" box="[394,681,1111,1135]" class="Collembola" family="Tomoceridae" genus="Plutomurus" kingdom="Animalia" order="Entomobryomorpha" pageId="3" pageNumber="255" phylum="Arthropoda" rank="species" species="kharagauliensis" status="sp. nov.">
|
||
<emphasis id="B9227A05192AFF8094B7FBE29CA1267F" box="[394,681,1111,1135]" italics="true" pageId="3" pageNumber="255">Plutomurus kharagauliensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9227A05192AFF809792FBE29CF7267F" bold="true" box="[687,767,1111,1135]" pageId="3" pageNumber="255">
|
||
<taxonomicNameLabel id="A211C77E192AFF809792FBE29CF7267F" box="[687,767,1111,1135]" pageId="3" pageNumber="255" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(scale bar: 0.5 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE9A617192AFF8095FAFB289AC3277C" blockId="3.[151,1437,1181,2036]" pageId="3" pageNumber="255">
|
||
Head. Ratio of antennal length: head length as 2.67–3.10. Ant. III–IV annulated. Eye patch typical for genus, with six well developed eyes (
|
||
<figureCitation id="136DBA92192AFF8094DEFB749C3D26CB" box="[483,565,1217,1243]" captionStart="FIGURES 12–14" captionStartId="5.[151,264,1728,1752]" captionTargetBox="[178,1403,190,1695]" captionTargetId="figure-22@5.[163,1425,181,1704]" captionTargetPageId="5" captionText="FIGURES 12–14. Dorsal chaetotaxy in Plutomurus kharagauliensis sp. nov. 12, Head.A: Anterior macrochaetae, B: interocular Mc, C: postocular Mc; 13, Th. II–III; 14, Abd. I–V (big ovals and round circles near Ant. I represent eyes; large black cir- cle—macrochaeta; open circles—mesochaeta; squares with dots—bothriothricha; circles with a slash represent pseudopores)." figureDoi="http://doi.org/10.5281/zenodo.6952156" httpUri="https://zenodo.org/record/6952156/files/figure.png" pageId="3" pageNumber="255">Fig. 12</figureCitation>
|
||
). Head dorsally with one unpaired (A
|
||
<subScript id="17D2A452192AFF8096EEFB619DD326F2" attach="left" box="[979,987,1236,1250]" fontSize="6" pageId="3" pageNumber="255">0</subScript>
|
||
), and seven paired Mc: two anterior (A
|
||
<subScript id="17D2A452192AFF8090B0FB619B9D26F2" attach="left" box="[1421,1429,1236,1250]" fontSize="6" pageId="3" pageNumber="255">2</subScript>
|
||
, A
|
||
<subScript id="17D2A452192AFF809596FB4D9EBB2716" attach="left" box="[171,179,1272,1286]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
), two interocular (S
|
||
<subScript id="17D2A452192AFF8094A4FB4D9FA92716" attach="left" box="[409,417,1272,1286]" fontSize="6" pageId="3" pageNumber="255">2</subScript>
|
||
, S
|
||
<subScript id="17D2A452192AFF8094FDFB4D9FC02716" attach="left" box="[448,456,1272,1286]" fontSize="6" pageId="3" pageNumber="255">4</subScript>
|
||
) and three postocular (Pa
|
||
<subScript id="17D2A452192AFF8097D1FB4D9CFC2716" attach="left" box="[748,756,1272,1286]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
, Pa
|
||
<subScript id="17D2A452192AFF80961DFB4D9D252716" attach="left" box="[800,813,1272,1286]" fontSize="6" pageId="3" pageNumber="255">4,</subScript>
|
||
Pa
|
||
<subScript id="17D2A452192AFF809670FB429D5D2715" attach="left" box="[845,853,1271,1285]" fontSize="6" pageId="3" pageNumber="255">5</subScript>
|
||
) distributed as in
|
||
<figureCitation id="136DBA92192AFF809119FB509A7F26EF" box="[1060,1143,1253,1279]" captionStart="FIGURES 12–14" captionStartId="5.[151,264,1728,1752]" captionTargetBox="[178,1403,190,1695]" captionTargetId="figure-22@5.[163,1425,181,1704]" captionTargetPageId="5" captionText="FIGURES 12–14. Dorsal chaetotaxy in Plutomurus kharagauliensis sp. nov. 12, Head.A: Anterior macrochaetae, B: interocular Mc, C: postocular Mc; 13, Th. II–III; 14, Abd. I–V (big ovals and round circles near Ant. I represent eyes; large black cir- cle—macrochaeta; open circles—mesochaeta; squares with dots—bothriothricha; circles with a slash represent pseudopores)." figureDoi="http://doi.org/10.5281/zenodo.6952156" httpUri="https://zenodo.org/record/6952156/files/figure.png" pageId="3" pageNumber="255">Fig. 12</figureCitation>
|
||
. Posterior margin of head with a row of mesochaetae. Prelabral and labral chaetae smooth: prelabral chaetae four (2+2) (
|
||
<figureCitation id="136DBA92192AFF809184FABC9AF72733" box="[1209,1279,1289,1315]" captionStart="FIGURES 4–11" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[199,1391,190,1827]" captionTargetId="figure-17@4.[190,1397,181,1836]" captionTargetPageId="4" captionText="FIGURES 4–11. Plutomurus kharagauliensis sp. nov. 4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.6952154" httpUri="https://zenodo.org/record/6952154/files/figure.png" pageId="3" pageNumber="255">Fig. 4</figureCitation>
|
||
); labrum with 554 papillate chaetae as typical for genus; distal margin of labrum with four elongate, thin–walled, flexible papillae. OML trifurcate, basal chaetae shorter than apical process; OML with four chaeta-like processes.
|
||
</paragraph>
|
||
<paragraph id="8BE9A617192AFF8095FAFAC09ACB2453" blockId="3.[151,1437,1181,2036]" pageId="3" pageNumber="255">
|
||
Body. Dorsal bothriotrichal formula as 2,1/0,0,1,2,0 (
|
||
<figureCitation id="136DBA92192AFF809614FAC09DA6279F" box="[809,942,1397,1423]" captionStart="FIGURES 12–14" captionStartId="5.[151,264,1728,1752]" captionTargetBox="[178,1403,190,1695]" captionTargetId="figure-22@5.[163,1425,181,1704]" captionTargetPageId="5" captionText="FIGURES 12–14. Dorsal chaetotaxy in Plutomurus kharagauliensis sp. nov. 12, Head.A: Anterior macrochaetae, B: interocular Mc, C: postocular Mc; 13, Th. II–III; 14, Abd. I–V (big ovals and round circles near Ant. I represent eyes; large black cir- cle—macrochaeta; open circles—mesochaeta; squares with dots—bothriothricha; circles with a slash represent pseudopores)." figureDoi="http://doi.org/10.5281/zenodo.6952156" httpUri="https://zenodo.org/record/6952156/files/figure.png" pageId="3" pageNumber="255">Figs 13–14</figureCitation>
|
||
). Dorsal Mc formula: 4,2/3,3,4,6,3–4 (
|
||
<figureCitation id="136DBA92192AFF809056FAC09EE927A3" captionStart="FIGURES 12–14" captionStartId="5.[151,264,1728,1752]" captionTargetBox="[178,1403,190,1695]" captionTargetId="figure-22@5.[163,1425,181,1704]" captionTargetPageId="5" captionText="FIGURES 12–14. Dorsal chaetotaxy in Plutomurus kharagauliensis sp. nov. 12, Head.A: Anterior macrochaetae, B: interocular Mc, C: postocular Mc; 13, Th. II–III; 14, Abd. I–V (big ovals and round circles near Ant. I represent eyes; large black cir- cle—macrochaeta; open circles—mesochaeta; squares with dots—bothriothricha; circles with a slash represent pseudopores)." figureDoi="http://doi.org/10.5281/zenodo.6952156" httpUri="https://zenodo.org/record/6952156/files/figure.png" pageId="3" pageNumber="255">Figs 13–14</figureCitation>
|
||
). Thoracic macrochaetotaxy as in
|
||
<figureCitation id="136DBA92192AFF809754FA2C9CB527A3" box="[617,701,1433,1459]" captionStart="FIGURES 12–14" captionStartId="5.[151,264,1728,1752]" captionTargetBox="[178,1403,190,1695]" captionTargetId="figure-22@5.[163,1425,181,1704]" captionTargetPageId="5" captionText="FIGURES 12–14. Dorsal chaetotaxy in Plutomurus kharagauliensis sp. nov. 12, Head.A: Anterior macrochaetae, B: interocular Mc, C: postocular Mc; 13, Th. II–III; 14, Abd. I–V (big ovals and round circles near Ant. I represent eyes; large black cir- cle—macrochaeta; open circles—mesochaeta; squares with dots—bothriothricha; circles with a slash represent pseudopores)." figureDoi="http://doi.org/10.5281/zenodo.6952156" httpUri="https://zenodo.org/record/6952156/files/figure.png" pageId="3" pageNumber="255">Fig. 13</figureCitation>
|
||
. Th. II with two anterior (m
|
||
<subScript id="17D2A452192AFF8096C2FA199A0F27AA" attach="left" box="[1023,1031,1452,1466]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
, m
|
||
<subScript id="17D2A452192AFF809110FA199A3D27AA" attach="left" box="[1069,1077,1452,1466]" fontSize="6" pageId="3" pageNumber="255">4</subScript>
|
||
) and two posterior (p
|
||
<subScript id="17D2A452192AFF809012FA199B3F27AA" attach="left" box="[1327,1335,1452,1466]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
, p
|
||
<subScript id="17D2A452192AFF809068FA1E9B5527A9" attach="left" box="[1365,1373,1451,1465]" fontSize="6" pageId="3" pageNumber="255">5</subScript>
|
||
) Mc; Th. III with two posterior (p
|
||
<subScript id="17D2A452192AFF8094ECFA659FD127CE" attach="right" box="[465,473,1488,1502]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
, p
|
||
<subScript id="17D2A452192AFF8094C9FA659FF427CE" attach="left" box="[500,508,1488,1502]" fontSize="6" pageId="3" pageNumber="255">4</subScript>
|
||
) Mc. Abdominal macrochaetotaxy as in
|
||
<figureCitation id="136DBA92192AFF8096FFFA089A4B27C8" box="[962,1091,1469,1496]" captionStart-0="FIGURES 12–14" captionStart-1="FIGURE 15" captionStartId-0="5.[151,264,1728,1752]" captionStartId-1="6.[151,250,2016,2040]" captionTargetBox-0="[178,1403,190,1695]" captionTargetBox-1="[151,1436,1029,1993]" captionTargetId-0="figure-22@5.[163,1425,181,1704]" captionTargetId-1="figure-412@6.[151,1436,1029,1993]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 12–14. Dorsal chaetotaxy in Plutomurus kharagauliensis sp. nov. 12, Head.A: Anterior macrochaetae, B: interocular Mc, C: postocular Mc; 13, Th. II–III; 14, Abd. I–V (big ovals and round circles near Ant. I represent eyes; large black cir- cle—macrochaeta; open circles—mesochaeta; squares with dots—bothriothricha; circles with a slash represent pseudopores)." captionText-1="FIGURE 15. Plutomurus kharagauliensis sp. nov. Macrochaetae complex on Abd IV (bar: 50 μm)." figureDoi-0="http://doi.org/10.5281/zenodo.6952156" figureDoi-1="http://doi.org/10.5281/zenodo.6952158" httpUri-0="https://zenodo.org/record/6952156/files/figure.png" httpUri-1="https://zenodo.org/record/6952158/files/figure.png" pageId="3" pageNumber="255">Figs 14–15</figureCitation>
|
||
: Abd. I–II each with three long posterior Mc (m
|
||
<subScript id="17D2A452192AFF809476FA419F5B2412" attach="left" box="[331,339,1524,1538]" fontSize="6" pageId="3" pageNumber="255">2</subScript>
|
||
, m
|
||
<subScript id="17D2A452192AFF80944AFA419F772412" attach="left" box="[375,383,1524,1538]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
, m
|
||
<subScript id="17D2A452192AFF80949FFA419FA22412" attach="left" box="[418,426,1524,1538]" fontSize="6" pageId="3" pageNumber="255">4</subScript>
|
||
); Abd. III with four long Mc: two anterior (m
|
||
<subScript id="17D2A452192AFF809697FA419DBA2412" attach="right" box="[938,946,1524,1538]" fontSize="6" pageId="3" pageNumber="255">2</subScript>
|
||
, m
|
||
<subScript id="17D2A452192AFF8096E8FA469DD52411" attach="left" box="[981,989,1523,1537]" fontSize="6" pageId="3" pageNumber="255">6</subScript>
|
||
) and two posterior (p
|
||
<subScript id="17D2A452192AFF8091F1FA419ADC2412" attach="left" box="[1228,1236,1524,1538]" fontSize="6" pageId="3" pageNumber="255">1</subScript>
|
||
, p
|
||
<subScript id="17D2A452192AFF8091D2FA469AFF2411" attach="left" box="[1263,1271,1523,1537]" fontSize="6" pageId="3" pageNumber="255">6</subScript>
|
||
) and one short Mc (p
|
||
<subScript id="17D2A452192AFF8095E7F9AD9EEA2436" attach="left" box="[218,226,1560,1574]" fontSize="6" pageId="3" pageNumber="255">7</subScript>
|
||
) as typical for genus; Abd. IV with six posterior Mc: two long (C
|
||
<subScript id="17D2A452192AFF80968FF9A29DB22435" attach="left" box="[946,954,1559,1573]" fontSize="6" pageId="3" pageNumber="255">6</subScript>
|
||
, E
|
||
<subScript id="17D2A452192AFF8096E5F9AD9DE82436" attach="left" box="[984,992,1560,1574]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
) and four short Mc (A
|
||
<subScript id="17D2A452192AFF8091E4F9A29AE92435" attach="left" box="[1241,1249,1559,1573]" fontSize="6" pageId="3" pageNumber="255">6</subScript>
|
||
, B
|
||
<subScript id="17D2A452192AFF80903DF9A29B002435" attach="left" box="[1280,1288,1559,1573]" fontSize="6" pageId="3" pageNumber="255">6</subScript>
|
||
, D
|
||
<subScript id="17D2A452192AFF809017F9AD9B3A2436" attach="right" box="[1322,1330,1560,1574]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
, T
|
||
<subScript id="17D2A452192AFF809072F9AD9B5F2436" attach="left" box="[1359,1367,1560,1574]" fontSize="6" pageId="3" pageNumber="255">7</subScript>
|
||
) (
|
||
<figureCitation id="136DBA92192AFF809052F9B09EBE2454" captionStart="FIGURE 15" captionStartId="6.[151,250,2016,2040]" captionTargetBox="[151,1436,1029,1993]" captionTargetId="figure-412@6.[151,1436,1029,1993]" captionTargetPageId="6" captionText="FIGURE 15. Plutomurus kharagauliensis sp. nov. Macrochaetae complex on Abd IV (bar: 50 μm)." figureDoi="http://doi.org/10.5281/zenodo.6952158" httpUri="https://zenodo.org/record/6952158/files/figure.png" pageId="3" pageNumber="255">Fig. 15</figureCitation>
|
||
); Abd. V with 2–3 posterior (p
|
||
<subScript id="17D2A452192AFF809733F9899C1E245A" attach="left" box="[526,534,1596,1610]" fontSize="6" pageId="3" pageNumber="255">2</subScript>
|
||
, p
|
||
<subScript id="17D2A452192AFF80970FF9899C32245A" attach="left" box="[562,570,1596,1610]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
or p
|
||
<subScript id="17D2A452192AFF809750F9899C7D245A" attach="left" box="[621,629,1596,1610]" fontSize="6" pageId="3" pageNumber="255">2</subScript>
|
||
, p
|
||
<subScript id="17D2A452192AFF8097AFF9899C92245A" attach="left" box="[658,666,1596,1610]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
, p
|
||
<subScript id="17D2A452192AFF80978BF9899CB6245A" attach="left" box="[694,702,1596,1610]" fontSize="6" pageId="3" pageNumber="255">4</subScript>
|
||
) long Mc and one long Mc laterally (
|
||
<figureCitation id="136DBA92192AFF80915FF99C9ABE2453" box="[1122,1206,1577,1603]" captionStart="FIGURES 12–14" captionStartId="5.[151,264,1728,1752]" captionTargetBox="[178,1403,190,1695]" captionTargetId="figure-22@5.[163,1425,181,1704]" captionTargetPageId="5" captionText="FIGURES 12–14. Dorsal chaetotaxy in Plutomurus kharagauliensis sp. nov. 12, Head.A: Anterior macrochaetae, B: interocular Mc, C: postocular Mc; 13, Th. II–III; 14, Abd. I–V (big ovals and round circles near Ant. I represent eyes; large black cir- cle—macrochaeta; open circles—mesochaeta; squares with dots—bothriothricha; circles with a slash represent pseudopores)." figureDoi="http://doi.org/10.5281/zenodo.6952156" httpUri="https://zenodo.org/record/6952156/files/figure.png" pageId="3" pageNumber="255">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE9A617192AFF8095FAF9F89B2824E8" blockId="3.[151,1437,1181,2036]" pageId="3" pageNumber="255">
|
||
<emphasis id="B9227A05192AFF8095FAF9F89F492477" bold="true" box="[199,321,1613,1639]" pageId="3" pageNumber="255">Mc length</emphasis>
|
||
: Mc on Th. II (m
|
||
<subScript id="17D2A452192AFF809734F9D59C19247E" attach="left" box="[521,529,1632,1646]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
, m
|
||
<subScript id="17D2A452192AFF80970BF9D59C4C247E" attach="left" box="[566,580,1632,1646]" fontSize="6" pageId="3" pageNumber="255">4,</subScript>
|
||
p
|
||
<subScript id="17D2A452192AFF809768F9D59C55247E" attach="left" box="[597,605,1632,1646]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
, p
|
||
<subScript id="17D2A452192AFF809746F9EA9C8B247D" attach="left" box="[635,643,1631,1645]" fontSize="6" pageId="3" pageNumber="255">5</subScript>
|
||
) and Th. III (p
|
||
<subScript id="17D2A452192AFF809613F9D59D3E247E" attach="right" box="[814,822,1632,1646]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
, p
|
||
<subScript id="17D2A452192AFF80966EF9D59D53247E" attach="left" box="[851,859,1632,1646]" fontSize="6" pageId="3" pageNumber="255">4</subScript>
|
||
)—120–167 µm; Mc on Abd. I–II (m
|
||
<subScript id="17D2A452192AFF80903DF9D59B00247E" attach="left" box="[1280,1288,1632,1646]" fontSize="6" pageId="3" pageNumber="255">2</subScript>
|
||
, m
|
||
<subScript id="17D2A452192AFF809010F9D59B3D247E" attach="left" box="[1325,1333,1632,1646]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
, m
|
||
<subScript id="17D2A452192AFF809067F9D59B6A247E" attach="left" box="[1370,1378,1632,1646]" fontSize="6" pageId="3" pageNumber="255">4</subScript>
|
||
) and Abd. III (m
|
||
<subScript id="17D2A452192AFF809424F9319F292482" attach="left" box="[281,289,1668,1682]" fontSize="6" pageId="3" pageNumber="255">2</subScript>
|
||
, m
|
||
<subScript id="17D2A452192AFF80947BF9369F5B2481" attach="left" box="[326,339,1667,1681]" fontSize="6" pageId="3" pageNumber="255">6,</subScript>
|
||
p
|
||
<subScript id="17D2A452192AFF809458F9319F652482" attach="left" box="[357,365,1668,1682]" fontSize="6" pageId="3" pageNumber="255">1</subScript>
|
||
, p
|
||
<subScript id="17D2A452192AFF8094B7F9369F9A2481" attach="left" box="[394,402,1667,1681]" fontSize="6" pageId="3" pageNumber="255">6</subScript>
|
||
)—178–220 µm; Mc on Abd. IV-E
|
||
<subScript id="17D2A452192AFF809626F9319D2B2482" attach="left" box="[795,803,1668,1682]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
238 µm, C
|
||
<subScript id="17D2A452192AFF80969CF9369DA12481" attach="left" box="[929,937,1667,1681]" fontSize="6" pageId="3" pageNumber="255">6</subScript>
|
||
: 168 µm (found only in a single specimen), A
|
||
<subScript id="17D2A452192AFF809596F9129EBB24A5" attach="left" box="[171,179,1703,1717]" fontSize="6" pageId="3" pageNumber="255">6</subScript>
|
||
: 93–135 µm, B
|
||
<subScript id="17D2A452192AFF80945EF9129F6324A5" attach="left" box="[355,363,1703,1717]" fontSize="6" pageId="3" pageNumber="255">6</subScript>
|
||
: 110–140 µm, D
|
||
<subScript id="17D2A452192AFF809714F91D9C3924A6" attach="left" box="[553,561,1704,1718]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
: 100–155 µm and in T
|
||
<subScript id="17D2A452192AFF80960CF91D9D3124A6" attach="left" box="[817,825,1704,1718]" fontSize="6" pageId="3" pageNumber="255">7</subScript>
|
||
Mc length is unknown (missing in all individuals). Diameter of Mc sockets on Abd IV-E
|
||
<subScript id="17D2A452192AFF809724F9799C2924CA" attach="left" box="[537,545,1740,1754]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
and C
|
||
<subScript id="17D2A452192AFF809755F97E9C7824C9" attach="left" box="[616,624,1739,1753]" fontSize="6" pageId="3" pageNumber="255">6</subScript>
|
||
: 8–10 µm, 4 short Mc (A
|
||
<subScript id="17D2A452192AFF8096B1F97E9D9C24C9" attach="left" box="[908,916,1739,1753]" fontSize="6" pageId="3" pageNumber="255">6</subScript>
|
||
, B
|
||
<subScript id="17D2A452192AFF809689F97E9DB424C9" attach="left" box="[948,956,1739,1753]" fontSize="6" pageId="3" pageNumber="255">6</subScript>
|
||
, D
|
||
<subScript id="17D2A452192AFF8096E3F9799DEE24CA" attach="left" box="[990,998,1740,1754]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
, T
|
||
<subScript id="17D2A452192AFF809139F9799A0424CA" attach="left" box="[1028,1036,1740,1754]" fontSize="6" pageId="3" pageNumber="255">7</subScript>
|
||
) subequalor in diameter.A
|
||
<subScript id="17D2A452192AFF809007F97E9B4A24C9" attach="left" box="[1338,1346,1739,1753]" fontSize="6" pageId="3" pageNumber="255">6</subScript>
|
||
slightly longer (5.5–7.5 µm) than in remaining 3 short Mc: B
|
||
<subScript id="17D2A452192AFF8097D5F95A9CF824ED" attach="left" box="[744,752,1775,1789]" fontSize="6" pageId="3" pageNumber="255">6</subScript>
|
||
: 5.5–6.5 µm; D
|
||
<subScript id="17D2A452192AFF80969FF9459DA224EE" attach="left" box="[930,938,1776,1790]" fontSize="6" pageId="3" pageNumber="255">3</subScript>
|
||
: 5.0–6.5 µm and T
|
||
<subScript id="17D2A452192AFF8091BCF9459A8124EE" attach="left" box="[1153,1161,1776,1790]" fontSize="6" pageId="3" pageNumber="255">7</subScript>
|
||
: 5.5–6.0 µm.
|
||
</paragraph>
|
||
<paragraph id="8BE9A617192AFF8095FAF8B79D2425E4" blockId="3.[151,1437,1181,2036]" pageId="3" pageNumber="255">
|
||
Legs. Hind legs with well–developed trochanteral (17–36 chaetae) and femoral (20–35 chaetae) organs (
|
||
<figureCitation id="136DBA92192AFF809052F8B49EAF252F" captionStart="FIGURES 4–11" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[199,1391,190,1827]" captionTargetId="figure-17@4.[190,1397,181,1836]" captionTargetPageId="4" captionText="FIGURES 4–11. Plutomurus kharagauliensis sp. nov. 4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.6952154" httpUri="https://zenodo.org/record/6952154/files/figure.png" pageId="3" pageNumber="255">Fig. 9</figureCitation>
|
||
), with several elongated chaetae; tibiotarsal spine formula as 002, posterior side of tibiotarsus III with two outstanding inner, Spine-like chaetae, (arrows in
|
||
<figureCitation id="136DBA92192AFF80979FF8FC9CFE2573" box="[674,758,1865,1891]" captionStart="FIGURES 4–11" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[199,1391,190,1827]" captionTargetId="figure-17@4.[190,1397,181,1836]" captionTargetPageId="4" captionText="FIGURES 4–11. Plutomurus kharagauliensis sp. nov. 4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.6952154" httpUri="https://zenodo.org/record/6952154/files/figure.png" pageId="3" pageNumber="255">Fig. 10</figureCitation>
|
||
). Tenent hair capitate (
|
||
<figureCitation id="136DBA92192AFF8096C1F8FC9A7B2573" box="[1020,1139,1865,1892]" captionStart="FIGURES 4–11" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[199,1391,190,1827]" captionTargetId="figure-17@4.[190,1397,181,1836]" captionTargetPageId="4" captionText="FIGURES 4–11. Plutomurus kharagauliensis sp. nov. 4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.6952154" httpUri="https://zenodo.org/record/6952154/files/figure.png" pageId="3" pageNumber="255">Figs 5, 10</figureCitation>
|
||
). Ratio of claw III: Emp. III: tenent hair III as 1.66–3.14: 0.91–1.71: 1. Inner edge of claw on all legs with one characteristically minute proximal unpaired tooth (as in
|
||
<figureCitation id="136DBA92192AFF8094C9F8249C3D25BC" box="[500,565,1937,1964]" captionStart="FIGURES 4–11" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[199,1391,190,1827]" captionTargetId="figure-17@4.[190,1397,181,1836]" captionTargetPageId="4" captionText="FIGURES 4–11. Plutomurus kharagauliensis sp. nov. 4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.6952154" httpUri="https://zenodo.org/record/6952154/files/figure.png" pageId="3" pageNumber="255">Fig. 5</figureCitation>
|
||
) and 2–4 distal unpaired teeth,
|
||
<quantity id="4CAE0BF2192AFF8096A7F8249DE925BB" box="[922,993,1937,1964]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="3" pageNumber="255" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">3–5 in</quantity>
|
||
total (b, c, d in
|
||
<figureCitation id="136DBA92192AFF8091A8F8249AD125BC" box="[1173,1241,1937,1964]" captionStart="FIGURES 4–11" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[199,1391,190,1827]" captionTargetId="figure-17@4.[190,1397,181,1836]" captionTargetPageId="4" captionText="FIGURES 4–11. Plutomurus kharagauliensis sp. nov. 4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.6952154" httpUri="https://zenodo.org/record/6952154/files/figure.png" pageId="3" pageNumber="255">Fig. 5</figureCitation>
|
||
). Claw with long pseudonychia (
|
||
<figureCitation id="136DBA92192AFF809402F8009FB925DF" box="[319,433,1973,2000]" captionStart="FIGURES 4–11" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[199,1391,190,1827]" captionTargetId="figure-17@4.[190,1397,181,1836]" captionTargetPageId="4" captionText="FIGURES 4–11. Plutomurus kharagauliensis sp. nov. 4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.6952154" httpUri="https://zenodo.org/record/6952154/files/figure.png" pageId="3" pageNumber="255">Figs 5, 10</figureCitation>
|
||
). Ratio of pseudonychium III: claw III inner edge as 0.38–0.62: 1. Empodium lanceolate, tapered, with 2 internal lamellae bearing 0–6 teeth (
|
||
<figureCitation id="136DBA92192AFF8097E4F86C9D1625E4" box="[729,798,2009,2036]" captionStart="FIGURES 4–11" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[199,1391,190,1827]" captionTargetId="figure-17@4.[190,1397,181,1836]" captionTargetPageId="4" captionText="FIGURES 4–11. Plutomurus kharagauliensis sp. nov. 4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.6952154" httpUri="https://zenodo.org/record/6952154/files/figure.png" pageId="3" pageNumber="255">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF29F69F192DFF8795AAF8F19DEC25FC" ID-DOI="http://doi.org/10.5281/zenodo.6952154" ID-Zenodo-Dep="6952154" httpUri="https://zenodo.org/record/6952154/files/figure.png" pageId="4" pageNumber="256" startId="4.[151,264,1860,1884]" targetBox="[199,1391,190,1827]" targetPageId="4">
|
||
<paragraph id="8BE9A617192DFF8795AAF8F19DEC25FC" blockId="4.[151,1437,1860,2028]" pageId="4" pageNumber="256">
|
||
<emphasis id="B9227A05192DFF8795AAF8F19F4E254C" bold="true" box="[151,326,1860,1884]" pageId="4" pageNumber="256">FIGURES 4–11.</emphasis>
|
||
<taxonomicName id="4C56DD94192DFF879471F8F19C62254C" authority="Barjadze & Parimuchová & Raschmanová & Maghradze & Kováč, 2022" authorityName="Barjadze & Parimuchová & Raschmanová & Maghradze & Kováč" authorityYear="2022" box="[332,618,1860,1884]" class="Collembola" family="Tomoceridae" genus="Plutomurus" kingdom="Animalia" order="Entomobryomorpha" pageId="4" pageNumber="256" phylum="Arthropoda" rank="species" species="kharagauliensis" status="sp. nov.">
|
||
<emphasis id="B9227A05192DFF879471F8F19C62254C" box="[332,618,1860,1884]" italics="true" pageId="4" pageNumber="256">Plutomurus kharagauliensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9227A05192DFF87974DF8F19CC8254C" bold="true" box="[624,704,1860,1884]" pageId="4" pageNumber="256">
|
||
<taxonomicNameLabel id="A211C77E192DFF87974DF8F19CC8254C" box="[624,704,1860,1884]" pageId="4" pageNumber="256" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF29F69F192CFF8695AAF9759B982530" ID-DOI="http://doi.org/10.5281/zenodo.6952156" ID-Zenodo-Dep="6952156" httpUri="https://zenodo.org/record/6952156/files/figure.png" pageId="5" pageNumber="257" startId="5.[151,264,1728,1752]" targetBox="[178,1403,190,1695]" targetPageId="5">
|
||
<paragraph id="8BE9A617192CFF8695AAF9759B982530" blockId="5.[151,1436,1728,1824]" pageId="5" pageNumber="257">
|
||
<emphasis id="B9227A05192CFF8695AAF9759F5C24C8" bold="true" box="[151,340,1728,1752]" pageId="5" pageNumber="257">FIGURES 12–14.</emphasis>
|
||
Dorsal chaetotaxy in
|
||
<taxonomicName id="4C56DD94192CFF86970CF9759D4724C8" authority="Barjadze & Parimuchová & Raschmanová & Maghradze & Kováč, 2022" authorityName="Barjadze & Parimuchová & Raschmanová & Maghradze & Kováč" authorityYear="2022" box="[561,847,1728,1752]" class="Collembola" family="Tomoceridae" genus="Plutomurus" kingdom="Animalia" order="Entomobryomorpha" pageId="5" pageNumber="257" phylum="Arthropoda" rank="species" species="kharagauliensis" status="sp. nov.">
|
||
<emphasis id="B9227A05192CFF86970CF9759D4724C8" box="[561,847,1728,1752]" italics="true" pageId="5" pageNumber="257">Plutomurus kharagauliensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9227A05192CFF869668F9749DAD24C9" bold="true" box="[853,933,1729,1753]" pageId="5" pageNumber="257">
|
||
<taxonomicNameLabel id="A211C77E192CFF869668F9749DAD24C9" box="[853,933,1729,1753]" pageId="5" pageNumber="257" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
12, Head. A: Anterior macrochaetae, B: interocular Mc, C: postocular Mc; 13, Th. II–III; 14, Abd. I–V (big ovals and round circles near Ant. I represent eyes; large black circle—macrochaeta; open circles—mesochaeta; squares with dots—bothriothricha; circles with a slash represent pseudopores).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BE9A617192CFF8695FAF8FA9C8F259D" blockId="5.[151,1437,1870,2041]" pageId="5" pageNumber="257">
|
||
Ventral tube. Hirsute, in
|
||
<specimenCount id="9D506D9E192CFF8694DDF8FA9C722579" box="[480,634,1871,1897]" pageId="5" pageNumber="257" type="generic">one specimen</specimenCount>
|
||
anterior basal, posterior basal and apicolateral sides with 36, about 24 and 35 smooth chaetae respectively (
|
||
<figureCitation id="136DBA92192CFF869708F8C69C71259D" box="[565,633,1907,1933]" captionStart="FIGURES 4–11" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[199,1391,190,1827]" captionTargetId="figure-17@4.[190,1397,181,1836]" captionTargetPageId="4" captionText="FIGURES 4–11. Plutomurus kharagauliensis sp. nov. 4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.6952154" httpUri="https://zenodo.org/record/6952154/files/figure.png" pageId="5" pageNumber="257">Fig. 8</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE9A617192CFF8695FAF8229A0525A0" blockId="5.[151,1437,1870,2041]" box="[199,1037,1942,1969]" pageId="5" pageNumber="257">
|
||
Tenaculum. Corpus with one smooth chaeta; rami with 4 + 4 teeth (
|
||
<figureCitation id="136DBA92192CFF869686F8229DF725A0" box="[955,1023,1942,1969]" captionStart="FIGURES 4–11" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[199,1391,190,1827]" captionTargetId="figure-17@4.[190,1397,181,1836]" captionTargetPageId="4" captionText="FIGURES 4–11. Plutomurus kharagauliensis sp. nov. 4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.6952154" httpUri="https://zenodo.org/record/6952154/files/figure.png" pageId="5" pageNumber="257">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE9A617192CFF8595FAF80E9C4F2399" blockId="5.[151,1437,1870,2041]" lastBlockId="6.[151,1437,151,1005]" lastPageId="6" lastPageNumber="258" pageId="5" pageNumber="257">
|
||
Furca. Ratio manubrium: dens: mucro as 5.54–9.71: 8.11–11.43: 1. Outer margin of basal segment of dens with 2–6 apically acuminate macrochaetae (sockets with arrows in
|
||
<figureCitation id="136DBA92192CFF869669F86A9DAD25E9" box="[852,933,2015,2041]" captionStart="FIGURES 4–11" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[199,1391,190,1827]" captionTargetId="figure-17@4.[190,1397,181,1836]" captionTargetPageId="4" captionText="FIGURES 4–11. Plutomurus kharagauliensis sp. nov. 4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.6952154" httpUri="https://zenodo.org/record/6952154/files/figure.png" pageId="5" pageNumber="257">Fig. 11</figureCitation>
|
||
). Inner edge of dens with well differentiated spines (spinal sockets shown in
|
||
<figureCitation id="136DBA92192FFF85973CFF229C5A22A1" box="[513,594,151,177]" captionStart="FIGURES 4–11" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[199,1391,190,1827]" captionTargetId="figure-17@4.[190,1397,181,1836]" captionTargetPageId="4" captionText="FIGURES 4–11. Plutomurus kharagauliensis sp. nov. 4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.6952154" httpUri="https://zenodo.org/record/6952154/files/figure.png" pageId="6" pageNumber="258">Fig. 11</figureCitation>
|
||
); spines on basal segment of dens forming 2–3 short and poorly organized rows with 1–3 large spines apically; spines on distal segment of dens forming a single row extending maximum
|
||
<superScript id="7C230B5F192FFF8590B9FF0F9B8422D8" attach="right" box="[1412,1420,186,200]" fontSize="6" pageId="6" pageNumber="258">1</superScript>
|
||
/
|
||
<subScript id="17D2A452192FFF8590A9FF789B9422CB" attach="none" box="[1428,1436,205,219]" fontSize="6" pageId="6" pageNumber="258">3</subScript>
|
||
or
|
||
<superScript id="7C230B5F192FFF859588FF6B9EB522FC" attach="right" box="[181,189,222,236]" fontSize="6" pageId="6" pageNumber="258">1</superScript>
|
||
/
|
||
<subScript id="17D2A452192FFF8595F8FF449EC522EF" attach="left" box="[197,205,241,255]" fontSize="6" pageId="6" pageNumber="258">2</subScript>
|
||
of length of distal segment of dens; proximal edge of distal segment of dens always with small spines intercalated between long spines, terminal one always largest long spine in row (dental sockets in
|
||
<figureCitation id="136DBA92192FFF8591B3FEB69AD6230D" box="[1166,1246,259,285]" captionStart="FIGURES 4–11" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[199,1391,190,1827]" captionTargetId="figure-17@4.[190,1397,181,1836]" captionTargetPageId="4" captionText="FIGURES 4–11. Plutomurus kharagauliensis sp. nov. 4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.6952154" httpUri="https://zenodo.org/record/6952154/files/figure.png" pageId="6" pageNumber="258">Fig. 11</figureCitation>
|
||
). Dental formula variable, as 4–11
|
||
<emphasis id="B9227A05192FFF859461FE929F9D2351" bold="true" box="[348,405,295,321]" italics="true" pageId="6" pageNumber="258">I–III</emphasis>
|
||
/1–7
|
||
<emphasis id="B9227A05192FFF8594F3FE929FD12351" bold="true" box="[462,473,295,321]" italics="true" pageId="6" pageNumber="258">I</emphasis>
|
||
1–4
|
||
<emphasis id="B9227A05192FFF85972CFE929C4D2351" bold="true" box="[529,581,295,321]" italics="true" pageId="6" pageNumber="258">I(II)</emphasis>
|
||
0–2
|
||
<emphasis id="B9227A05192FFF859741FE929C8F2351" bold="true" box="[636,647,295,321]" italics="true" pageId="6" pageNumber="258">I</emphasis>
|
||
0–2
|
||
<emphasis id="B9227A05192FFF859782FE929CC22351" bold="true" box="[703,714,295,321]" italics="true" pageId="6" pageNumber="258">I</emphasis>
|
||
0–2
|
||
<emphasis id="B9227A05192FFF85963FFE929D052351" bold="true" box="[770,781,295,321]" italics="true" pageId="6" pageNumber="258">I</emphasis>
|
||
(Arabic numbers represent small spines; Roman numerals in bold and Italics represent large spines, on basal/distal segment of dens). Mucro with two basal, 0 intermediate and two distal teeth (202 formula) (
|
||
<figureCitation id="136DBA92192FFF8594C8FEDA9C312399" box="[501,569,367,393]" captionStart="FIGURES 4–11" captionStartId="4.[151,264,1860,1884]" captionTargetBox="[199,1391,190,1827]" captionTargetId="figure-17@4.[190,1397,181,1836]" captionTargetPageId="4" captionText="FIGURES 4–11. Plutomurus kharagauliensis sp. nov. 4, Prelabral chaetae (bar: 0.04 mm); 5, Claw complex III (a, basal small tooth to internal lamellae; b, largest unpaired tooth and c and d additional unpaired teeth) (bar: 0.05 mm); 6, Tenaculum (bar: 0.06 mm); 7, Mucro, chaetae and chaetal sockets omitted (bar: 0.02 mm); 8, Ventral tube (bar: 0.08 mm); 9, Trochanteral and femoral organs (bar: 0.06 mm); 10, Hind tibiotarsus, with spine-like chaetae (arrows on them) (bar: 0.04 mm); 11, Dens with sockets of dental spines and lateral macrochaetae (arrows on them) (bar: 0.05 mm)." figureDoi="http://doi.org/10.5281/zenodo.6952154" httpUri="https://zenodo.org/record/6952154/files/figure.png" pageId="6" pageNumber="258">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BE9A617192FFF8595FAFE279DFE23BD" blockId="6.[151,1437,151,1005]" box="[199,1014,402,429]" pageId="6" pageNumber="258">
|
||
<emphasis id="B9227A05192FFF8595FAFE279F4523BC" bold="true" box="[199,333,402,428]" pageId="6" pageNumber="258">Variability.</emphasis>
|
||
All constant and variable characters are given in
|
||
<tableCitation id="C6D493AC192FFF859648FE269DF923BD" box="[885,1009,403,429]" captionStart-0="TABLE 1" captionStart-1="TABLE 2" captionStartId-0="11.[151,239,151,175]" captionStartId-1="11.[151,239,506,530]" captionTargetPageId-0="11" captionTargetPageId-1="11" captionText-0="TABLE 1. Constant characters of Plutomurus kharagauliensis sp. nov. and P. ruseki sp. nov." captionText-1="TABLE 2. Variable characters of Plutomurus kharagauliensis sp. nov. and P. ruseki sp. nov." httpUri-0="http://table.plazi.org/id/DF29F69F1922FF8895AAFF229A7322BF" httpUri-1="http://table.plazi.org/id/DF29F69F1922FF8895AAFE4F9A7D2002" pageId="6" pageNumber="258" tableUuid-0="DF29F69F1922FF8895AAFF229A7322BF" tableUuid-1="DF29F69F1922FF8895AAFE4F9A7D2002">Tables 1–2</tableCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34CF59C192FFF8595FAFE039A8920E1" pageId="6" pageNumber="258" type="discussion">
|
||
<paragraph id="8BE9A617192FFF8595FAFE039D49202D" blockId="6.[151,1437,151,1005]" pageId="6" pageNumber="258">
|
||
<emphasis id="B9227A05192FFF8595FAFE039F4423C0" bold="true" box="[199,332,438,464]" pageId="6" pageNumber="258">Discussion.</emphasis>
|
||
The new species is most similar to
|
||
<taxonomicName id="4C56DD94192FFF8597DBFE029AB123C1" authority="Djanashvili & Barjadze, 2011" authorityName="Djanashvili & Barjadze" authorityYear="2011" box="[742,1209,439,465]" class="Collembola" family="Tomoceridae" genus="Plutomurus" kingdom="Animalia" order="Entomobryomorpha" pageId="6" pageNumber="258" phylum="Arthropoda" rank="species" species="birsteini">
|
||
<emphasis id="B9227A05192FFF8597DBFE029D6823C1" box="[742,864,439,465]" italics="true" pageId="6" pageNumber="258">P. birsteini</emphasis>
|
||
<bibRefCitation id="EFC7DBE6192FFF859655FE029AB123C1" author="Djanashvili, R. & Barjadze, S." box="[872,1209,439,465]" pageId="6" pageNumber="258" pagination="28 - 30" refId="ref10317" refString="Djanashvili, R. & Barjadze, S. (2011) A new species of the genus Plutomurus Yosii, 1956 (Collembola, Tomoceridae) from Georgian caves. Journal of Cave and Karst Studies, 73, 28 - 30. https: // doi. org / 10.4311 / jcks 2010 lsc 0147" type="journal article" year="2011">Djanashvili & Barjadze, 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
by the combination of the following characters: 1) number of prelabral chaetae (2+2); 2) shape of tenent hair (capitate); 3) formula of tibiotarsal spine-like chaetae (002); 4) number of eyes—6 and 5) number of Mc on Th. II (
|
||
<bibRefCitation id="EFC7DBE6192FFF8591ACFE4A9ED9202D" author="Djanashvili, R. & Barjadze, S." pageId="6" pageNumber="258" pagination="28 - 30" refId="ref10317" refString="Djanashvili, R. & Barjadze, S. (2011) A new species of the genus Plutomurus Yosii, 1956 (Collembola, Tomoceridae) from Georgian caves. Journal of Cave and Karst Studies, 73, 28 - 30. https: // doi. org / 10.4311 / jcks 2010 lsc 0147" type="journal article" year="2011">Djanashvili & Barjadze 2011</bibRefCitation>
|
||
; Barjadze
|
||
<emphasis id="B9227A05192FFF85947AFD969F88202D" box="[327,384,547,573]" italics="true" pageId="6" pageNumber="258">et al.</emphasis>
|
||
2020a). Both species are cave dwellers.
|
||
</paragraph>
|
||
<paragraph id="8BE9A617192FFF8595FAFDF29A8920E1" blockId="6.[151,1437,151,1005]" pageId="6" pageNumber="258">
|
||
The new species differs from
|
||
<taxonomicName id="4C56DD94192FFF859728FDF29C862071" authorityName="Djanashvili & Barjadze" authorityYear="2011" box="[533,654,583,609]" class="Collembola" family="Tomoceridae" genus="Plutomurus" kingdom="Animalia" order="Entomobryomorpha" pageId="6" pageNumber="258" phylum="Arthropoda" rank="species" species="birsteini">
|
||
<emphasis id="B9227A05192FFF859728FDF29C862071" box="[533,654,583,609]" italics="true" pageId="6" pageNumber="258">P. birsteini</emphasis>
|
||
</taxonomicName>
|
||
by the number of Mc on the head and abdominal terga: 1) three postocular Mc on the head dorsum present in the new species, while 2 Mc in
|
||
<taxonomicName id="4C56DD94192FFF8596F6FDDE9A4B2095" authorityName="Djanashvili & Barjadze" authorityYear="2011" box="[971,1091,619,645]" class="Collembola" family="Tomoceridae" genus="Plutomurus" kingdom="Animalia" order="Entomobryomorpha" pageId="6" pageNumber="258" phylum="Arthropoda" rank="species" species="birsteini">
|
||
<emphasis id="B9227A05192FFF8596F6FDDE9A4B2095" box="[971,1091,619,645]" italics="true" pageId="6" pageNumber="258">P. birsteini</emphasis>
|
||
</taxonomicName>
|
||
and 2) Abd. IV with six Mc in the new species, while
|
||
<taxonomicName id="4C56DD94192FFF8594ABFD3A9C0620B9" authorityName="Djanashvili & Barjadze" authorityYear="2011" box="[406,526,655,681]" class="Collembola" family="Tomoceridae" genus="Plutomurus" kingdom="Animalia" order="Entomobryomorpha" pageId="6" pageNumber="258" phylum="Arthropoda" rank="species" species="birsteini">
|
||
<emphasis id="B9227A05192FFF8594ABFD3A9C0620B9" box="[406,526,655,681]" italics="true" pageId="6" pageNumber="258">P. birsteini</emphasis>
|
||
</taxonomicName>
|
||
has three (Barjadze
|
||
<emphasis id="B9227A05192FFF8597CDFD3A9D2F20B9" box="[752,807,655,681]" italics="true" pageId="6" pageNumber="258">et al.</emphasis>
|
||
2020a).
|
||
<taxonomicName id="4C56DD94192FFF8596B5FD3A9A6620B9" authorityName="Djanashvili & Barjadze" authorityYear="2011" box="[904,1134,655,681]" class="Collembola" family="Tomoceridae" genus="Plutomurus" kingdom="Animalia" order="Entomobryomorpha" pageId="6" pageNumber="258" phylum="Arthropoda" rank="species" species="birsteini">
|
||
<emphasis id="B9227A05192FFF8596B5FD3A9A6620B9" box="[904,1134,655,681]" italics="true" pageId="6" pageNumber="258">Plutomurus birsteini</emphasis>
|
||
</taxonomicName>
|
||
was described from eleven caves located in five different karst massifs of western
|
||
<collectingCountry id="F341E687192FFF85962FFD069D6420DD" box="[786,876,691,717]" name="Georgia" pageId="6" pageNumber="258">Georgia</collectingCountry>
|
||
and molecular investigations (COI) show that
|
||
<taxonomicName id="4C56DD94192FFF8590B5FD069EFC20E1" authorityName="Djanashvili & Barjadze" authorityYear="2011" class="Collembola" family="Tomoceridae" genus="Plutomurus" kingdom="Animalia" order="Entomobryomorpha" pageId="6" pageNumber="258" phylum="Arthropoda" rank="species" species="birsteini">
|
||
<emphasis id="B9227A05192FFF8590B5FD069EFC20E1" italics="true" pageId="6" pageNumber="258">P. birsteini</emphasis>
|
||
</taxonomicName>
|
||
is a complex of genetically distinct cryptic species (Barjadze, unpublished data).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34CF59C192FFF8595FAFD4E9F582129" pageId="6" pageNumber="258" type="etymology">
|
||
<paragraph id="8BE9A617192FFF8595FAFD4E9F582129" blockId="6.[151,1437,151,1005]" pageId="6" pageNumber="258">
|
||
<emphasis id="B9227A05192FFF8595FAFD4E9F462105" bold="true" box="[199,334,763,789]" pageId="6" pageNumber="258">Etymology.</emphasis>
|
||
The specific name originates from the
|
||
<typeStatus id="54ED18B5192FFF85962BFD4E9D4E2105" box="[790,838,763,789]" pageId="6" pageNumber="258">type</typeStatus>
|
||
locality––Kharagauli district in which the Kozmani Cave is situated.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34CF59C192FFF8595FAFCF69CE82191" pageId="6" pageNumber="258" type="biology_ecology">
|
||
<paragraph id="8BE9A617192FFF8595FAFCF69CE82191" blockId="6.[151,1437,151,1005]" pageId="6" pageNumber="258">
|
||
<emphasis id="B9227A05192FFF8595FAFCF69F24214D" bold="true" box="[199,300,835,861]" pageId="6" pageNumber="258">Ecology.</emphasis>
|
||
It is a troglophilous species, considering the presence of body pigment and 6+6 well-developed eyes. It occupies both twilight and dark zones of the cave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34CF59C192FFF8595FAFC3F9D2821FD" pageId="6" pageNumber="258" type="distribution">
|
||
<paragraph id="8BE9A617192FFF8595FAFC3F9D2821FD" blockId="6.[151,1437,151,1005]" pageId="6" pageNumber="258">
|
||
<emphasis id="B9227A05192FFF8595FAFC3F9F6921B4" bold="true" box="[199,353,906,932]" pageId="6" pageNumber="258">Distribution.</emphasis>
|
||
This species is known only from a single cave. This cave is the remotest one from Zemo
|
||
<collectingRegion id="499268F5192FFF859076FC3E9B9421B5" box="[1355,1436,907,933]" country="Georgia" name="Imereti" pageId="6" pageNumber="258">Imereti</collectingRegion>
|
||
plateau karst massif and it is isolated from other known caves in the same karst massif (
|
||
<bibRefCitation id="EFC7DBE6192FFF859159FC1A9B5721D9" author="Tatashidze, Z. K. & Tsikarishvili, K. D. & Jishkariani, J. M." box="[1124,1375,943,969]" pageId="6" pageNumber="258" refId="ref11698" refString="Tatashidze, Z. K., Tsikarishvili, K. D. & Jishkariani, J. M. (2009) The Cadastre of the Karst Caves of Georgia. Petiti Publishing House, Tbilisi, 670 pp. [in Georgian]" type="book" year="2009">
|
||
Tatashidze
|
||
<emphasis id="B9227A05192FFF8591D9FC1A9B1421D9" box="[1252,1308,943,969]" italics="true" pageId="6" pageNumber="258">et al.</emphasis>
|
||
2009
|
||
</bibRefCitation>
|
||
). The new species is the first known taxon from Kozmani Cave.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |