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<document id="91B5580646A82E9F0C3806408627B4C2" ID="10.11646/zootaxa.4139.2.1" ID-DOI="10.11646/zootaxa.4139.2.1" ID-GBIF-Dataset="aeecbbb4-4764-41dd-9562-b25f91f9f70e" ID-ISSN="1175-5326" ID-Zenodo-Dep="265016" ID-ZooBank="2EAFF1B9-1FA2-44B1-89A3-476359314C5E" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1468994861279" checkinUser="plazi" docAuthor="Nguyen, Anh D. &amp; Golovatch, Sergei I." docDate="2016" docId="03FECD21FFAFFFF3FF4F3E20FDCBF928" docLanguage="en" docName="zootaxa.4139.2.1.pdf" docOrigin="Zootaxa 4139 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Enghoffosoma fedorenkoi Golovatch 2016, comb. nov." docType="treatment" docVersion="11" lastPageNumber="156" masterDocId="FFC7B559FFACFFF6FFD83D24FFAEFF84" masterDocTitle="The millipede genus Enghoffosoma Golovatch, 1993 recorded in Vietnam for the first time, with descriptions of three new species (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="166" masterPageNumber="151" pageNumber="154" updateTime="1698365308467" updateUser="plazi">
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<mods:title id="E24EE9CD3A55B40C112F6FE8A4D8FFFA">The millipede genus Enghoffosoma Golovatch, 1993 recorded in Vietnam for the first time, with descriptions of three new species (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
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<mods:name id="191113129A4A35100E48B4F72C377C32" type="personal">
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<mods:namePart id="44EDF9518FBB8287FE3572FF0F864555">Nguyen, Anh D.</mods:namePart>
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<mods:namePart id="7E5EF2744F9812FBDA656ED56BA722F8">Golovatch, Sergei I.</mods:namePart>
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<mods:title id="EE6B0D0BA80A980D9C8B895A5DB738F8">Zootaxa</mods:title>
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<mods:date id="6141A50C82C93164305894A394A2B831">2016</mods:date>
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<treatment id="03FECD21FFAFFFF3FF4F3E20FDCBF928" ID-DOI="http://doi.org/10.5281/zenodo.6067529" ID-GBIF-Taxon="123198411" ID-Zenodo-Dep="6067529" LSID="urn:lsid:plazi:treatment:03FECD21FFAFFFF3FF4F3E20FDCBF928" httpUri="http://treatment.plazi.org/id/03FECD21FFAFFFF3FF4F3E20FDCBF928" lastPageId="5" lastPageNumber="156" pageId="3" pageNumber="154">
<subSubSection id="C34D2FBCFFAFFFF5FF4F3E20FEA4FCBB" pageId="3" pageNumber="154" type="nomenclature">
<paragraph id="8BE87C37FFAFFFF5FF4F3E20FCEBFC9A" blockId="3.[151,837,772,831]" box="[151,837,772,798]" pageId="3" pageNumber="154">
<heading id="D0A0CB5BFFAFFFF5FF4F3E20FCEBFC9A" bold="true" box="[151,837,772,798]" fontSize="11" level="1" pageId="3" pageNumber="154" reason="1">
<emphasis id="B923A025FFAFFFF5FF4F3E20FCEBFC9A" bold="true" box="[151,837,772,798]" pageId="3" pageNumber="154">
<taxonomicName id="4C5707B4FFAFFFF5FF4F3E20FD16FC9A" ID-CoL="39T4D" authority="Golovatch, 2016" authorityName="Golovatch" authorityYear="2016" box="[151,696,772,798]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="fedorenkoi" status="comb. nov.">
<emphasis id="B923A025FFAFFFF5FF4F3E20FE7DFC9A" bold="true" box="[151,467,772,798]" italics="true" pageId="3" pageNumber="154">Enghoffosoma fedorenkoi</emphasis>
(
<bibRefCitation id="EFC601C6FFAFFFF5FE3A3E20FD01FC9A" author="Golovatch" box="[482,687,772,798]" pageId="3" pageNumber="154" refString="Golovatch, S. I. (2016) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XIX. Arthropoda Selecta, 25 (2), 131 - 152." type="journal article" year="2016">Golovatch, 2016</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2101D5EFFAFFFF5FD673E20FCEBFC9A" box="[703,837,772,798]" pageId="3" pageNumber="154">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE87C37FFAFFFF5FF4F3E02FEA4FCBB" blockId="3.[151,837,772,831]" box="[151,266,806,831]" pageId="3" pageNumber="154">
(
<figureCitation id="136C60B2FFAFFFF5FF473E02FEACFCBB" box="[159,258,806,831]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="3.[151,250,1910,1932]" captionStartId-1="4.[151,250,1963,1985]" captionStartId-2="5.[151,250,1190,1212]" captionTargetBox-0="[185,1408,941,1888]" captionTargetBox-1="[197,1389,882,1933]" captionTargetBox-2="[151,1436,193,1168]" captionTargetId-0="figure@3.[175,1417,933,1897]" captionTargetId-1="figure@4.[189,1398,877,1941]" captionTargetId-2="figure@5.[151,1436,193,1169]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 3. Enghoffosoma fedorenkoi (Golovatch, 2016), comb. nov., male from Chu Yan Sin National Park, Vietnam. Anterior segments, dorsal view (A), lateral view (B); midbody segments, dorsal view (D), lateral view (E), ventral view (F); posterior segments, dorsal view (G), lateral view (H), ventral view (I); sternum between coxae 4, ventral view (C). Scale = 1 mm." captionText-1="FIGURE 4. Enghoffosoma fedorenkoi (Golovatch, 2016), comb. nov., male from Chu Yan Sin National Park, Vietnam. Right gonopod, lateral view (A); mesal view (B). Scale = 1 mm." captionText-2="FIGURE 5. Enghoffosoma fedorenkoi (Golovatch, 2016), comb. nov., male from Chu Yan Sin National Park; Vietnam. Left gonopod, lateroventral view (A), mesoventral view (B)." httpUri-0="https://zenodo.org/record/265019/files/figure.png" httpUri-1="https://zenodo.org/record/265020/files/figure.png" httpUri-2="https://zenodo.org/record/265021/files/figure.png" pageId="3" pageNumber="154">Figs 35</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34D2FBCFFAFFFF5FF4F3E4DFD79FCFB" box="[151,727,873,895]" pageId="3" pageNumber="154" type="reference_group">
<paragraph id="8BE87C37FFAFFFF5FF4F3E4DFD79FCFB" blockId="3.[151,727,873,895]" box="[151,727,873,895]" pageId="3" pageNumber="154">
<treatmentCitationGroup id="AB475B19FFAFFFF5FF4F3E4DFD79FCFB" box="[151,727,873,895]" pageId="3" pageNumber="154">
<taxonomicName id="4C5707B4FFAFFFF5FF4F3E4DFDF7FCFB" ID-CoL="53G7D" authority="Golovatch, 2016: 149" authorityName="Golovatch" authorityPageNumber="149" authorityYear="2016" box="[151,601,873,895]" class="Diplopoda" family="Paradoxosomatidae" genus="Sundanina" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="fedorenkoi">
<emphasis id="B923A025FFAFFFF5FF4F3E4DFEDBFCFA" box="[151,373,873,894]" italics="true" pageId="3" pageNumber="154">Sundanina fedorenkoi</emphasis>
<treatmentCitation id="0AF65A26FFAFFFF5FEA33E4DFDF7FCFB" author="Golovatch" box="[379,601,873,895]" page="149" pageId="3" pageNumber="154" year="2016">
<bibRefCitation id="EFC601C6FFAFFFF5FEA33E4DFD89FCFB" author="Golovatch" box="[379,551,873,895]" pageId="3" pageNumber="154" refString="Golovatch, S. I. (2016) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XIX. Arthropoda Selecta, 25 (2), 131 - 152." type="journal article" year="2016">Golovatch, 2016</bibRefCitation>
: 149
</treatmentCitation>
</taxonomicName>
, figs 7990.
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</paragraph>
</subSubSection>
<caption id="DF282CBFFFAFFFF5FF4F3A52FF6BF874" httpUri="https://zenodo.org/record/265019/files/figure.png" pageId="3" pageNumber="154" targetBox="[185,1408,941,1888]" targetPageId="3">
<paragraph id="8BE87C37FFAFFFF5FF4F3A52FF6BF874" blockId="3.[151,1437,1910,2032]" pageId="3" pageNumber="154">
<emphasis id="B923A025FFAFFFF5FF4F3A52FEB9F808" bold="true" box="[151,279,1910,1932]" pageId="3" pageNumber="154">FIGURE 3.</emphasis>
<taxonomicName id="4C5707B4FFAFFFF5FEFA3A52FD59F808" authority="Golovatch, 2016" authorityName="Golovatch" authorityYear="2016" box="[290,759,1910,1932]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="fedorenkoi" status="comb. nov.">
<emphasis id="B923A025FFAFFFF5FEFA3A52FD85F80F" box="[290,555,1910,1931]" italics="true" pageId="3" pageNumber="154">Enghoffosoma fedorenkoi</emphasis>
(Golovatch, 2016)
</taxonomicName>
,
<emphasis id="B923A025FFAFFFF5FCDE3A52FCD0F808" bold="true" box="[774,894,1910,1932]" pageId="3" pageNumber="154">
<taxonomicNameLabel id="A2101D5EFFAFFFF5FCDE3A52FCD0F808" box="[774,894,1910,1932]" pageId="3" pageNumber="154">comb. nov.</taxonomicNameLabel>
</emphasis>
, male from Chu Yan Sin National Park, Vietnam. Anterior segments, dorsal view (A), lateral view (B); midbody segments, dorsal view (D), lateral view (E), ventral view (F); posterior segments, dorsal view (G), lateral view (H), ventral view (I); sternum between coxae 4, ventral view (C). Scale = 1 mm.
</paragraph>
</caption>
<subSubSection id="C34D2FBCFFA8FFF2FF4F3DB3FAB1FCBC" pageId="4" pageNumber="155" type="materials_examined">
<paragraph id="8BE87C37FFA8FFF2FF4F3DB3FD3BFE99" blockId="4.[151,1437,151,824]" pageId="4" pageNumber="155">
<emphasis id="B923A025FFA8FFF2FF4F3DB3FE2CFF34" bold="true" box="[151,386,151,176]" pageId="4" pageNumber="155">Material examined.</emphasis>
<specimenCount id="9D51B7BEFFA8FFF2FE533DBCFE76FF34" box="[395,472,151,176]" pageId="4" pageNumber="155" type="male">1 male</specimenCount>
(
<emphasis id="B923A025FFA8FFF2FE333DB3FD0EFF34" bold="true" box="[491,672,151,176]" pageId="4" pageNumber="155">IEBR-Myr 249</emphasis>
),
<collectingCountry id="F3403CA7FFA8FFF2FD623DB3FCB2FF34" box="[698,796,151,176]" name="Vietnam" pageId="4" pageNumber="155">Vietnam</collectingCountry>
,
<collectingRegion id="4993B2D5FFA8FFF2FCF03DB3FC22FF34" box="[808,908,151,176]" country="Vietnam" name="Dak Lak" pageId="4" pageNumber="155">Dak Lak</collectingRegion>
Prov., Chu Yan Sin National Park, coniferous and broadleaved forests,
<quantity id="4CAFD1D2FFA8FFF2FE763D98FDE6FF50" box="[430,584,188,213]" metricMagnitude="3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="4" pageNumber="155" unit="m" value="1250.0" valueMax="1500.0" valueMin="1000.0">10001500 m</quantity>
a.s.l.,
<date id="FFE95AF7FFA8FFF2FD483D98FC04FF50" box="[656,938,188,213]" pageId="4" pageNumber="155" value="2012-03-28" valueMax="2012-04-12" valueMin="2012-03-28">28 March12 April 2012</date>
, leg. Le X. Son;
<specimenCount id="9D51B7BEFFA8FFF2FBBC3D98FB01FF51" box="[1124,1199,188,213]" pageId="4" pageNumber="155" type="male">1 male</specimenCount>
(damaged),
<specimenCount id="9D51B7BEFFA8FFF2FAE43D98FA32FF51" box="[1340,1436,188,213]" pageId="4" pageNumber="155" type="female">1 female</specimenCount>
(
<emphasis id="B923A025FFA8FFF2FF783DFBFEFAFF7C" bold="true" box="[160,340,223,248]" pageId="4" pageNumber="155">IEBR-Myr 119</emphasis>
),
<collectingCountry id="F3403CA7FFA8FFF2FEB53DFBFE61FF7C" box="[365,463,223,248]" name="Vietnam" pageId="4" pageNumber="155">Vietnam</collectingCountry>
,
<collectingRegion id="4993B2D5FFA8FFF2FE043DC4FDF9FF7C" box="[476,599,224,248]" country="Vietnam" name="Lam Dong" pageId="4" pageNumber="155">Lam Dong</collectingRegion>
Prov., Bi Doup-Nui Ba National Park,
<taxonomicName id="4C5707B4FFA8FFF2FBC43DC4FBF2FF73" box="[1052,1116,224,247]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="4" pageNumber="155" phylum="Tracheophyta" rank="genus">
<emphasis id="B923A025FFA8FFF2FBC43DC4FBF2FF73" box="[1052,1116,224,247]" italics="true" pageId="4" pageNumber="155">Pinus</emphasis>
</taxonomicName>
forest,
<quantity id="4CAFD1D2FFA8FFF2FB6D3DC4FAA2FF73" box="[1205,1292,224,248]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="155" unit="m" value="1400.0">1400 m</quantity>
a.s.l., pitfall trapping,
<date id="FFE95AF7FFA8FFF2FEDA3C21FE22FE98" box="[258,396,260,285]" pageId="4" pageNumber="155" value="2009-05-07">7 May 2009</date>
, leg. Anh Duc Nguyen.
</paragraph>
<paragraph id="8BE87C37FFA8FFF2FF1F3C03FBDAFE29" blockId="4.[151,1437,151,824]" pageId="4" pageNumber="155">
<emphasis id="B923A025FFA8FFF2FF1F3C03FE32FEC4" bold="true" box="[199,412,295,320]" pageId="4" pageNumber="155">Descriptive notes.</emphasis>
There are a number of apparently intraspecific variations observed in the new samples as compared to the original description (
<bibRefCitation id="EFC601C6FFA8FFF2FD9F3C68FCAAFEE1" author="Golovatch" box="[583,772,332,357]" pageId="4" pageNumber="155" refString="Golovatch, S. I. (2016) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XIX. Arthropoda Selecta, 25 (2), 131 - 152." type="journal article" year="2016">Golovatch 2016</bibRefCitation>
). In comparison with
<typeStatus id="54ECC295FFA8FFF2FBD83C68FBCBFEE1" box="[1024,1125,332,357]" pageId="4" pageNumber="155" type="holotype">holotype</typeStatus>
, a sympatric material from Chu Yan Sin National Park is considerably larger in size, up to
<quantity id="4CAFD1D2FFA8FFF2FC833C4BFC6EFE03" box="[859,960,367,392]" metricMagnitude="-2" metricUnit="m" metricValue="3.56" pageId="4" pageNumber="155" unit="mm" value="35.6">35.6 mm</quantity>
long (vs ca
<quantity id="4CAFD1D2FFA8FFF2FB903C54FB6BFE0C" box="[1096,1221,368,392]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.7" metricValueMin="2.2" pageId="4" pageNumber="155" unit="mm" value="24.5" valueMax="27.0" valueMin="22.0">2227 mm</quantity>
); width of pro- and metazonae
<quantity id="4CAFD1D2FFA8FFF2FECE3CB0FEC5FE28" box="[278,363,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="4" pageNumber="155" unit="mm" value="3.2">3.2 mm</quantity>
and
<quantity id="4CAFD1D2FFA8FFF2FE7A3CB0FE54FE28" box="[418,506,404,428]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="4" pageNumber="155" unit="mm" value="3.8">3.8 mm</quantity>
, respectively (vs 2.53.3 and 3.04.0 mm, respectively).
</paragraph>
<paragraph id="8BE87C37FFA8FFF2FF1F3C93FB2CFD2C" blockId="4.[151,1437,151,824]" pageId="4" pageNumber="155">
The new material shows a subtriangular hypoproct with two separated distolateral setae borne on small knobs (
<figureCitation id="136C60B2FFA8FFF2FF473CF8FF4DFE70" box="[159,227,476,501]" captionStart="FIGURE 3" captionStartId="3.[151,250,1910,1932]" captionTargetBox="[185,1408,941,1888]" captionTargetId="figure@3.[175,1417,933,1897]" captionTargetPageId="3" captionText="FIGURE 3. Enghoffosoma fedorenkoi (Golovatch, 2016), comb. nov., male from Chu Yan Sin National Park, Vietnam. Anterior segments, dorsal view (A), lateral view (B); midbody segments, dorsal view (D), lateral view (E), ventral view (F); posterior segments, dorsal view (G), lateral view (H), ventral view (I); sternum between coxae 4, ventral view (C). Scale = 1 mm." httpUri="https://zenodo.org/record/265019/files/figure.png" pageId="4" pageNumber="155">
Fig.
<date id="FFE95AF7FFA8FFF2FF0C3CF8FF4DFE70" box="[212,227,476,500]" pageId="4" pageNumber="155">3</date>
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I); male sternum 5 with a densely setose, bifid, trapeziform lobe between coxae 4 (
<figureCitation id="136C60B2FFA8FFF2FB493CF8FB7AFE70" box="[1169,1236,476,501]" captionStart="FIGURE 3" captionStartId="3.[151,250,1910,1932]" captionTargetBox="[185,1408,941,1888]" captionTargetId="figure@3.[175,1417,933,1897]" captionTargetPageId="3" captionText="FIGURE 3. Enghoffosoma fedorenkoi (Golovatch, 2016), comb. nov., male from Chu Yan Sin National Park, Vietnam. Anterior segments, dorsal view (A), lateral view (B); midbody segments, dorsal view (D), lateral view (E), ventral view (F); posterior segments, dorsal view (G), lateral view (H), ventral view (I); sternum between coxae 4, ventral view (C). Scale = 1 mm." httpUri="https://zenodo.org/record/265019/files/figure.png" pageId="4" pageNumber="155">Fig. 3</figureCitation>
C); legs stout and strong, especially three last pairs of legs, 1.21.3 times as long as midbody height; and ventral brushes on tarsi being present until segment 12, thereafter thinning out. In contrast, the
<typeStatus id="54ECC295FFA8FFF2FC683F00FBBCFDB9" box="[944,1042,548,573]" pageId="4" pageNumber="155" type="holotype">holotype</typeStatus>
has a semi-circular hypoproct with two separated distolateral setae which are not borne on knobs; male sternum 5 shows a pair of small, densely setose, basally subcontiguous, rounded lobes between coxae 4; the legs are longer (in the male about 1.41.5 times as long as midbody height); and ventral brushes are present on all tarsi (
<bibRefCitation id="EFC601C6FFA8FFF2FC633FABFBD8FD2C" author="Golovatch" box="[955,1142,655,680]" pageId="4" pageNumber="155" refString="Golovatch, S. I. (2016) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XIX. Arthropoda Selecta, 25 (2), 131 - 152." type="journal article" year="2016">Golovatch 2016</bibRefCitation>
).
</paragraph>
<paragraph id="8BE87C37FFA8FFF2FF1F3F91FAB1FCBC" blockId="4.[151,1437,151,824]" pageId="4" pageNumber="155">
Except for the above differences, the new material totally agrees with the
<typeStatus id="54ECC295FFA8FFF2FBA83F90FB7CFD49" box="[1136,1234,692,717]" pageId="4" pageNumber="155" type="holotype">holotype</typeStatus>
in the gonopod conformation. The gonopod femorite is slender, slightly curved, with neither processes nor other modifications. The postfemoral region is separated from the femorite by a distinct cingulum and consists only of a long parabasal process and a long, thick, gradually attenuated solenomere. The distal part of the solenomere is twisted.
</paragraph>
</subSubSection>
<caption id="DF282CBFFFA8FFF2FF4F3A8FFD4AF85B" httpUri="https://zenodo.org/record/265020/files/figure.png" pageId="4" pageNumber="155" targetBox="[197,1389,882,1933]" targetPageId="4">
<paragraph id="8BE87C37FFA8FFF2FF4F3A8FFD4AF85B" blockId="4.[151,1435,1963,2015]" pageId="4" pageNumber="155">
<emphasis id="B923A025FFA8FFF2FF4F3A8FFEBDF845" bold="true" box="[151,275,1963,1985]" pageId="4" pageNumber="155">FIGURE 4.</emphasis>
<taxonomicName id="4C5707B4FFA8FFF2FEC23A8FFD4CF845" authority="Golovatch, 2016" authorityName="Golovatch" authorityYear="2016" box="[282,738,1963,1985]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="fedorenkoi" status="comb. nov.">
<emphasis id="B923A025FFA8FFF2FEC23A8FFDB0F844" box="[282,542,1963,1984]" italics="true" pageId="4" pageNumber="155">Enghoffosoma fedorenkoi</emphasis>
(Golovatch, 2016)
</taxonomicName>
,
<emphasis id="B923A025FFA8FFF2FD353A8FFCCFF845" bold="true" box="[749,865,1963,1985]" pageId="4" pageNumber="155">
<taxonomicNameLabel id="A2101D5EFFA8FFF2FD353A8FFCCFF845" box="[749,865,1963,1985]" pageId="4" pageNumber="155">comb. nov.</taxonomicNameLabel>
</emphasis>
, male from Chu Yan Sin National Park, Vietnam. Right gonopod, lateral view (A); mesal view (B). Scale = 1 mm.
</paragraph>
</caption>
<caption id="DF282CBFFFA9FFF3FF4F3982FD65FB5F" httpUri="https://zenodo.org/record/265021/files/figure.png" pageId="5" pageNumber="156" targetBox="[151,1436,193,1168]" targetPageId="5">
<paragraph id="8BE87C37FFA9FFF3FF4F3982FD65FB5F" blockId="5.[151,1435,1190,1243]" pageId="5" pageNumber="156">
<emphasis id="B923A025FFA9FFF3FF4F3982FEBAFB38" bold="true" box="[151,276,1190,1212]" pageId="5" pageNumber="156">FIGURE 5.</emphasis>
<taxonomicName id="4C5707B4FFA9FFF3FEC43982FD48FB38" authority="Golovatch, 2016" authorityName="Golovatch" authorityYear="2016" box="[284,742,1190,1212]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="fedorenkoi" status="comb. nov.">
<emphasis id="B923A025FFA9FFF3FEC43982FD8FFB3F" box="[284,545,1190,1211]" italics="true" pageId="5" pageNumber="156">Enghoffosoma fedorenkoi</emphasis>
(Golovatch, 2016)
</taxonomicName>
,
<emphasis id="B923A025FFA9FFF3FD2A3982FCC9FB38" bold="true" box="[754,871,1190,1212]" pageId="5" pageNumber="156">
<taxonomicNameLabel id="A2101D5EFFA9FFF3FD2A3982FCC9FB38" box="[754,871,1190,1212]" pageId="5" pageNumber="156">comb. nov.</taxonomicNameLabel>
</emphasis>
, male from Chu Yan Sin National Park; Vietnam. Left gonopod, lateroventral view (A), mesoventral view (B).
</paragraph>
</caption>
<subSubSection id="C34D2FBCFFA9FFF3FF1F3823FDCBF928" pageId="5" pageNumber="156" type="discussion">
<paragraph id="8BE87C37FFA9FFF3FF1F3823FC6EF99F" blockId="5.[151,1437,1287,1708]" pageId="5" pageNumber="156">
<emphasis id="B923A025FFA9FFF3FF1F3823FE95FAA4" bold="true" box="[199,315,1287,1312]" pageId="5" pageNumber="156">Remarks.</emphasis>
<bibRefCitation id="EFC601C6FFA9FFF3FE9C3823FDA2FAA4" author="Golovatch" box="[324,524,1287,1312]" pageId="5" pageNumber="156" refString="Golovatch, S. I. (2016) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XIX. Arthropoda Selecta, 25 (2), 131 - 152." type="journal article" year="2016">Golovatch (2016)</bibRefCitation>
described this species as
<taxonomicName id="4C5707B4FFA9FFF3FCEB3823FB85FA9B" box="[819,1067,1287,1311]" class="Diplopoda" family="Paradoxosomatidae" genus="Sundanina" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="fedorenkoi">
<emphasis id="B923A025FFA9FFF3FCEB3823FB85FA9B" box="[819,1067,1287,1311]" italics="true" pageId="5" pageNumber="156">Sundanina fedorenkoi</emphasis>
</taxonomicName>
, based on
<specimenCount id="9D51B7BEFFA9FFF3FB70382CFAAEFAA4" box="[1192,1280,1287,1312]" pageId="5" pageNumber="156" type="male">2 males</specimenCount>
coming from the Chu Yan Sin National Park. So some of our material represents a strict topotype. Golovatch also emphasized his strong hesitation in placing that species in the genus
<taxonomicName id="4C5707B4FFA9FFF3FD34386BFBA8FAEC" authority="Attems, 1914" authorityName="Attems" authorityYear="1914" box="[748,1030,1359,1384]" class="Diplopoda" family="Paradoxosomatidae" genus="Sundanina" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="156" phylum="Arthropoda" rank="genus">
<emphasis id="B923A025FFA9FFF3FD34386BFCCAFAE3" box="[748,868,1359,1383]" italics="true" pageId="5" pageNumber="156">Sundanina</emphasis>
Attems, 1914
</taxonomicName>
, a member of the tribe Sundaninini. Indeed, the gonopods in the latter genus and tribe always show a (disto)femoral outgrowth or process and a slender flagelliform solenomere, whereas in
<taxonomicName id="4C5707B4FFA9FFF3FDE038BCFD62FA34" box="[568,716,1432,1456]" class="Diplopoda" family="Paradoxosomatidae" genus="Sundanina" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="fedorenkoi">
<emphasis id="B923A025FFA9FFF3FDE038BCFD62FA34" box="[568,716,1432,1456]" italics="true" pageId="5" pageNumber="156">S. fedorenkoi</emphasis>
</taxonomicName>
the former is absent while the latter is conspicuously thick. As such conditions fully fit in the diagnosis of
<taxonomicName id="4C5707B4FFA9FFF3FD593898FC8DFA50" box="[641,803,1468,1492]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="156" phylum="Arthropoda" rank="genus">
<emphasis id="B923A025FFA9FFF3FD593898FC8DFA50" box="[641,803,1468,1492]" italics="true" pageId="5" pageNumber="156">Enghoffosoma</emphasis>
</taxonomicName>
as given by
<bibRefCitation id="EFC601C6FFA9FFF3FC6A389FFB7EFA50" box="[946,1232,1467,1492]" pageId="5" pageNumber="156" refString="Likhitrakarn, K., Golovatch, S. I. &amp; Panha, S. (2014) Review of the Southeast Asian millipede genus Enghoffosoma Golovatch, 1993 (Diplopoda, Polydesmida, Paradoxosomatidae), with descriptions of new species. Zootaxa, 3811 (4), 491 - 514. http: // dx. doi. org / 10.11646 / zootaxa. 3811.4.4" type="journal article">
Likhitrakarn
<emphasis id="B923A025FFA9FFF3FB9D3898FBD3FA50" box="[1093,1149,1468,1492]" italics="true" pageId="5" pageNumber="156">et al.</emphasis>
(2014)
</bibRefCitation>
and above, we do not hesitate to formally transfer
<taxonomicName id="4C5707B4FFA9FFF3FDD538C4FD0DFA7C" box="[525,675,1504,1528]" class="Diplopoda" family="Paradoxosomatidae" genus="Sundanina" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="fedorenkoi">
<emphasis id="B923A025FFA9FFF3FDD538C4FD0DFA7C" box="[525,675,1504,1528]" italics="true" pageId="5" pageNumber="156">S. fedorenkoi</emphasis>
</taxonomicName>
to
<emphasis id="B923A025FFA9FFF3FD1438C4FCDBFA7C" box="[716,885,1504,1528]" italics="true" pageId="5" pageNumber="156">
<taxonomicName id="4C5707B4FFA9FFF3FD1438C4FCDCFA7C" box="[716,882,1504,1528]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="156" phylum="Arthropoda" rank="genus">Enghoffosoma</taxonomicName>
.
</emphasis>
This results in the following new combination:
<taxonomicName id="4C5707B4FFA9FFF3FF4F3B20FD23F998" authority="Golovatch, 2016" authorityName="Golovatch" authorityYear="2016" box="[151,653,1539,1564]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="fedorenkoi" status="comb. nov.">
<emphasis id="B923A025FFA9FFF3FF4F3B20FE1BF998" box="[151,437,1540,1564]" italics="true" pageId="5" pageNumber="156">Enghoffosoma fedorenkoi</emphasis>
(
<bibRefCitation id="EFC601C6FFA9FFF3FE1C3B27FD2BF998" author="Golovatch" box="[452,645,1539,1564]" pageId="5" pageNumber="156" refString="Golovatch, S. I. (2016) On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), XIX. Arthropoda Selecta, 25 (2), 131 - 152." type="journal article" year="2016">Golovatch, 2016</bibRefCitation>
)
</taxonomicName>
,
<emphasis id="B923A025FFA9FFF3FD403B27FCB6F998" bold="true" box="[664,792,1539,1564]" pageId="5" pageNumber="156">
<taxonomicNameLabel id="A2101D5EFFA9FFF3FD403B27FCB6F998" box="[664,792,1539,1564]" pageId="5" pageNumber="156">comb. nov.</taxonomicNameLabel>
</emphasis>
ex
<taxonomicName id="4C5707B4FFA9FFF3FC993B20FC17F998" box="[833,953,1540,1564]" class="Diplopoda" family="Paradoxosomatidae" genus="Sundanina" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="156" phylum="Arthropoda" rank="genus">
<emphasis id="B923A025FFA9FFF3FC993B20FC17F998" box="[833,953,1540,1564]" italics="true" pageId="5" pageNumber="156">Sundanina</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE87C37FFA9FFF3FF1F3B03FDCBF928" blockId="5.[151,1437,1287,1708]" pageId="5" pageNumber="156">
The species is fairly similar to
<taxonomicName id="4C5707B4FFA9FFF3FDED3B0DFD7BF9C4" box="[565,725,1576,1600]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="triangulare" status="sp. nov.">
<emphasis id="B923A025FFA9FFF3FDED3B0DFD7BF9C4" box="[565,725,1576,1600]" italics="true" pageId="5" pageNumber="156">E. triangulare</emphasis>
</taxonomicName>
<emphasis id="B923A025FFA9FFF3FD383B03FC93F9C4" bold="true" box="[736,829,1575,1600]" pageId="5" pageNumber="156">
<taxonomicNameLabel id="A2101D5EFFA9FFF3FD383B03FC93F9C4" box="[736,829,1575,1600]" pageId="5" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in the paraterga being well-developed, the gonopod femorite slender and devoid of processes, the postfemoral region consisting only of a solenomere and a process
<emphasis id="B923A025FFA9FFF3FA503B6FFA3AF9E0" bold="true" box="[1416,1428,1611,1636]" pageId="5" pageNumber="156">e</emphasis>
, and the solenomere showing a distal tooth terminating the seminal groove. Process
<emphasis id="B923A025FFA9FFF3FBE73B4BFBE5F90C" bold="true" box="[1087,1099,1647,1672]" pageId="5" pageNumber="156">e</emphasis>
is triangular in
<taxonomicName id="4C5707B4FFA9FFF3FB263B55FA35F90C" box="[1278,1435,1648,1672]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="triangulare" status="sp. nov.">
<emphasis id="B923A025FFA9FFF3FB263B55FA35F90C" box="[1278,1435,1648,1672]" italics="true" pageId="5" pageNumber="156">E. triangulare</emphasis>
</taxonomicName>
<emphasis id="B923A025FFA9FFF3FF4F3BB7FF5EF928" bold="true" box="[151,240,1683,1708]" pageId="5" pageNumber="156">
<taxonomicNameLabel id="A2101D5EFFA9FFF3FF4F3BB7FF5EF928" box="[151,240,1683,1708]" pageId="5" pageNumber="156" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, but oar-shaped in
<emphasis id="B923A025FFA9FFF3FE1F3BB0FDCBF928" box="[455,613,1684,1708]" italics="true" pageId="5" pageNumber="156">
<taxonomicName id="4C5707B4FFA9FFF3FE1F3BB0FDCFF928" box="[455,609,1684,1708]" class="Diplopoda" family="Paradoxosomatidae" genus="Enghoffosoma" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="156" phylum="Arthropoda" rank="species" species="fedorenkoi">E. fedorenkoi</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>