treatments-xml/data/03/FE/A5/03FEA553FFF7C873FF0DBACCFC26FE77.xml
2024-06-21 12:22:17 +02:00

591 lines
76 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="3D57A601DFD1A2EADEF5F97A696BB1E0" ID-CLB-Dataset="55574" ID-DOI="10.11646/zootaxa.5134.3.2" ID-GBIF-Dataset="5bbba3c2-785f-4240-b8b2-33c32fb0c5a0" ID-ISSN="1175-5326" ID-Zenodo-Dep="6538197" ID-ZooBank="C28B3BDB-6854-42EB-A09B-15A53246397D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1652257711962" checkinUser="plazi" docAuthor="Pont, Adrian C." docDate="2022" docId="03FEA553FFF7C873FF0DBACCFC26FE77" docLanguage="en" docName="zootaxa.5134.3.2.pdf" docOrigin="Zootaxa 5134 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Drymeia fratercula Pont 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="368" masterDocId="FFC7DD2BFFFEC87DFF9ABA5BFFD1FFDA" masterDocTitle="The genus Drymeia Meigen in the Caucasus Mountains, with a note on the identity of Aspilia glacialis Rondani, 1866 (Diptera: Muscidae)" masterLastPageNumber="382" masterPageNumber="355" pageNumber="364" updateTime="1699330171174" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="35FF54C5B0D3D40ECDAD29971AF2AA59" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="94FCCF0F20B07104E410D3EC586F8A8E">
<mods:title id="45DF2A048DCC9A700BD788520AAD5DA0">The genus Drymeia Meigen in the Caucasus Mountains, with a note on the identity of Aspilia glacialis Rondani, 1866 (Diptera: Muscidae)</mods:title>
</mods:titleInfo>
<mods:name id="9906C5D8BC6E8F347C2D0C8D2532F62A" type="personal">
<mods:role id="E392C18C1BDCA0F9DB535BFB11B533CC">
<mods:roleTerm id="AED53006C32A9A1FE0FE53FBFD9DEB66">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="943E34751F9C9F55248376708EA178EB">Pont, Adrian C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C10C4D0FF699BAA2444A212D6F8D1930">text</mods:typeOfResource>
<mods:relatedItem id="833B1C89CD9488BE1E0CF6415AC8E939" type="host">
<mods:titleInfo id="2F78F3DA0268F4E061DEF64EC42BCFC6">
<mods:title id="26243730DF158A726415FB7C0836A537">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="BB917A2BBBD0A753B467D8AF12BFA508">
<mods:date id="4E14D8AFD2D6C490AAF95E04C3A09372">2022</mods:date>
<mods:detail id="E2A4A2F6C17B1EC2CE2AE8CDF40908FC" type="pubDate">
<mods:number id="36FFECB8D2594272AE4CF284E3660501">2022-05-11</mods:number>
</mods:detail>
<mods:detail id="56A7F07514A8643796A4476EFF203F7F" type="volume">
<mods:number id="66318154F511F7773D512AF06C8BB1B1">5134</mods:number>
</mods:detail>
<mods:detail id="86A5F71A8493A9E56E0CB4C978D37437" type="issue">
<mods:number id="68F6191EDA49C54485CC1FA216BE60EE">3</mods:number>
</mods:detail>
<mods:extent id="67D68E560521FBFB77BF96CCDE8EDDA7" unit="page">
<mods:start id="C45F41CFD2D9D077FE3D4DFF9860D459">355</mods:start>
<mods:end id="F05BFC3049D31A7B8C85537335C6248D">382</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A72D2C211393A8CE907D99BC438EC916">journal article</mods:classification>
<mods:identifier id="F49778B13DF7A0F195C22C2C48E2845D" type="CLB-Dataset">55574</mods:identifier>
<mods:identifier id="2EE5420BA97C09C0E5BC5DAE3D3424C9" type="DOI">10.11646/zootaxa.5134.3.2</mods:identifier>
<mods:identifier id="5839A6283324E130528A075A3AB05A06" type="GBIF-Dataset">5bbba3c2-785f-4240-b8b2-33c32fb0c5a0</mods:identifier>
<mods:identifier id="5102F84428C83890F4ED9936D71CA34D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="C3D9DB3CDEF415F9C799D4E5195B15D6" type="Zenodo-Dep">6538197</mods:identifier>
<mods:identifier id="B94B58284F9344D034878739D60A9A1D" type="ZooBank">C28B3BDB-6854-42EB-A09B-15A53246397D</mods:identifier>
</mods:mods>
<treatment id="03FEA553FFF7C873FF0DBACCFC26FE77" ID-DOI="http://doi.org/10.5281/zenodo.6539281" ID-GBIF-Taxon="195298839" ID-Zenodo-Dep="6539281" LSID="urn:lsid:plazi:treatment:03FEA553FFF7C873FF0DBACCFC26FE77" httpUri="http://treatment.plazi.org/id/03FEA553FFF7C873FF0DBACCFC26FE77" lastPageId="14" lastPageNumber="368" pageId="9" pageNumber="364">
<subSubSection id="C34D47CEFFF7C874FF0DBACCFE35FF6B" box="[151,484,151,178]" pageId="9" pageNumber="364" type="nomenclature">
<paragraph id="8BE81445FFF7C874FF0DBACCFE35FF6B" blockId="9.[151,484,151,213]" box="[151,484,151,178]" pageId="9" pageNumber="364">
<heading id="D0A0A329FFF7C874FF0DBACCFE35FF6B" bold="true" box="[151,484,151,178]" fontSize="11" level="1" pageId="9" pageNumber="364" reason="1">
<emphasis id="B923C857FFF7C874FF0DBACCFE35FF6B" bold="true" box="[151,484,151,178]" pageId="9" pageNumber="364">
<taxonomicName id="4C576FC6FFF7C874FF0DBACCFE51FF68" ID-CoL="9J2PD" authority="Pont, 2022" authorityName="Pont" authorityYear="2022" box="[151,384,151,178]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="fratercula" status="sp. nov.">
<emphasis id="B923C857FFF7C874FF0DBACCFE51FF68" bold="true" box="[151,384,151,178]" italics="true" pageId="9" pageNumber="364">Drymeia fratercula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A210752CFFF7C874FE1DBACCFE35FF6B" box="[391,484,151,177]" pageId="9" pageNumber="364" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34D47CEFFF7C874FF0DBAE0FEC4FF0F" box="[151,277,187,213]" pageId="9" pageNumber="364" type="description">
<paragraph id="8BE81445FFF7C874FF0DBAE0FEC4FF0F" blockId="9.[151,484,151,213]" box="[151,277,187,213]" pageId="9" pageNumber="364">
<figureCitation id="136C08C0FFF7C874FF0DBAE0FEC4FF0F" box="[151,277,187,213]" captionStart-0="FIGURES 1011" captionStart-1="FIGURES 1216" captionStart-2="FIGURES 1718" captionStartId-0="9.[151,262,1964,1988]" captionStartId-1="11.[151,264,1886,1910]" captionStartId-2="12.[151,264,1909,1933]" captionTargetBox-0="[479,1110,237,1933]" captionTargetBox-1="[206,1381,286,1845]" captionTargetBox-2="[220,1383,196,1871]" captionTargetId-0="figure-23@9.[477,1110,237,1940]" captionTargetId-1="figure-57@11.[189,1398,273,1863]" captionTargetId-2="figure-38@12.[204,1383,181,1884]" captionTargetPageId-0="9" captionTargetPageId-1="11" captionTargetPageId-2="12" captionText-0="FIGURES 1011. Drymeia fratercula sp. nov., ♂ habitus. 10. Lateral view. 11. Dorsal view. (holotype; Georgia, Kazbegi, Mt Koltesh, meadow/heath, 2300 m, 8.vii.1983.)" captionText-1="FIGURES 1216. Drymeia fratercula sp.nov. 12. ♂ head, lateral view. 13. ♂ hind tibia and tarsomere 1, posterior view. 14. ♀ fore tarsomeres, anterior view. 15. ♂ sternite 5, dorsal view. 16. ♀ tergite 8. Scale-line 0.25 mm (15), 0.5 mm (16). (Georgia, Ka- zbegi, Mt Koltesh, meadow/heath, 2300 m, 8.vii.1983 (12, 13) and 30.vi.1983 (14, 16); Mt Koltesh, heath, 2500 m, 30.vi.1983 (15).)" captionText-2="FIGURES 1718. Drymeia fratercula sp.nov., ♂ terminalia. 17. Cercal plate, dorsal view. 18. Hypopygium, lateral view. Scale lines 0.25 mm. (Georgia, Kazbegi, Mt Koltesh, heath, 2500 m, 30.vi.1983.)" figureDoi-0="http://doi.org/10.5281/zenodo.6538211" figureDoi-1="http://doi.org/10.5281/zenodo.6538213" figureDoi-2="http://doi.org/10.5281/zenodo.6538215" httpUri-0="https://zenodo.org/record/6538211/files/figure.png" httpUri-1="https://zenodo.org/record/6538213/files/figure.png" httpUri-2="https://zenodo.org/record/6538215/files/figure.png" pageId="9" pageNumber="364">Figs 1018</figureCitation>
</paragraph>
</subSubSection>
<caption id="DF2844CDFFF7C874FF0DBDF7FD8BF833" ID-DOI="http://doi.org/10.5281/zenodo.6538211" ID-Zenodo-Dep="6538211" httpUri="https://zenodo.org/record/6538211/files/figure.png" pageId="9" pageNumber="364" startId="9.[151,262,1964,1988]" targetBox="[479,1110,237,1933]" targetPageId="9">
<paragraph id="8BE81445FFF7C874FF0DBDF7FD8BF833" blockId="9.[151,1436,1964,2025]" pageId="9" pageNumber="364">
<emphasis id="B923C857FFF7C874FF0DBDF7FE83F81F" bold="true" box="[151,338,1964,1989]" pageId="9" pageNumber="364">FIGURES 1011.</emphasis>
<taxonomicName id="4C576FC6FFF7C874FEC3BDF6FDCDF81F" authority="Pont, 2022" authorityName="Pont" authorityYear="2022" box="[345,540,1965,1989]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="364" phylum="Arthropoda" rank="species" species="fratercula" status="sp. nov.">
<emphasis id="B923C857FFF7C874FEC3BDF6FDCDF81F" box="[345,540,1965,1989]" italics="true" pageId="9" pageNumber="364">Drymeia fratercula</emphasis>
</taxonomicName>
<emphasis id="B923C857FFF7C874FDB9BDF6FDA5F81F" bold="true" box="[547,628,1965,1989]" pageId="9" pageNumber="364">
<taxonomicNameLabel id="A210752CFFF7C874FDB9BDF6FDA5F81F" box="[547,628,1965,1989]" pageId="9" pageNumber="364" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♂ habitus.
<emphasis id="B923C857FFF7C874FD69BDF6FCC3F81F" bold="true" box="[755,786,1965,1989]" pageId="9" pageNumber="364">10.</emphasis>
Lateral view.
<emphasis id="B923C857FFF7C874FC3FBDF6FC12F81F" bold="true" box="[933,963,1965,1989]" pageId="9" pageNumber="364">11.</emphasis>
Dorsal view. (holotype; Georgia, Kazbegi, Mt Koltesh, meadow/heath, 2300 m, 8.vii.1983.)
</paragraph>
</caption>
<subSubSection id="C34D47CEFFF4C877FF0DBACDFC15FE9B" pageId="10" pageNumber="365" type="diagnosis">
<paragraph id="8BE81445FFF4C877FF0DBACDFC15FE9B" blockId="10.[151,1437,150,682]" pageId="10" pageNumber="365">
<emphasis id="B923C857FFF4C877FF0DBACDFEC3FF6A" bold="true" box="[151,274,150,176]" pageId="10" pageNumber="365">Diagnosis.</emphasis>
This species is close to the Alpine
<taxonomicName id="4C576FC6FFF4C877FD3EBACCFBFCFF68" authority="(Rondani, 1866)" authorityName="" baseAuthorityName="Rondani" baseAuthorityYear="1866" box="[676,1069,151,178]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="365" phylum="Arthropoda" rank="species" species="glacialis">
<emphasis id="B923C857FFF4C877FD3EBACCFCBDFF6B" box="[676,876,151,177]" italics="true" pageId="10" pageNumber="365">Drymeia glacialis</emphasis>
(
<bibRefCitation id="EFC669B4FFF4C877FCE4BACCFBF7FF68" author="Rondani, C." box="[894,1062,151,178]" pageId="10" pageNumber="365" pagination="68 - 217" refId="ref14028" refString="Rondani, C. (1866) Anthomyinae Italicae collectae distinctae et in ordinem dispositae [a prof. Camillo Rondani]. Atti della Societa italiana di Scienze naturali, 9, 68 - 217." type="journal article" year="1866">Rondani, 1866</bibRefCitation>
)
</taxonomicName>
, e.g. 3 postsutural dorsocentrals, very broad parafacial, and conspicuous facial snout. However,
<taxonomicName id="4C576FC6FFF4C877FCDEBAE0FC15FF0F" authorityName="" baseAuthorityName="Rondani" baseAuthorityYear="1866" box="[836,964,187,213]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="365" phylum="Arthropoda" rank="species" species="glacialis">
<emphasis id="B923C857FFF4C877FCDEBAE0FC15FF0F" box="[836,964,187,213]" italics="true" pageId="10" pageNumber="365">D. glacialis</emphasis>
</taxonomicName>
is a larger, more robust species with longer and stronger setae, and the male has the frontal width equal to diameter of anterior ocellus; hind femur with no or few posteroventrals in apical half; fore tarsus with tarsomere 1 produced posteroventrally at tip into a conspicuous thorn, and tarsomere 1 almost as long as the length of tarsomeres 25 combined.
</paragraph>
</subSubSection>
<subSubSection id="C34D47CEFFF4C877FF5DBB10FD0AFD73" pageId="10" pageNumber="365" type="etymology">
<paragraph id="8BE81445FFF4C877FF5DBB10FD64FE50" blockId="10.[151,1437,150,682]" pageId="10" pageNumber="365">
<emphasis id="B923C857FFF4C877FF5DBB10FE9FFEBF" bold="true" box="[199,334,331,357]" pageId="10" pageNumber="365">Etymology.</emphasis>
The specific name “
<taxonomicName id="4C576FC6FFF4C877FDA8BB10FD70FEBF" authorityName="Pont" authorityYear="2022" box="[562,673,331,357]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="365" phylum="Arthropoda" rank="species" species="fratercula">
<emphasis id="B923C857FFF4C877FDA8BB10FD70FEBF" box="[562,673,331,357]" italics="true" pageId="10" pageNumber="365">fratercula</emphasis>
</taxonomicName>
” is Latin and means “little brother”. This reflects the close similarity between this species and
<taxonomicName id="4C576FC6FFF4C877FE28BB34FD60FE50" authority="(Rondani)" authorityName="" baseAuthorityName="Rondani" baseAuthorityYear="1866" box="[434,689,367,394]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="365" phylum="Arthropoda" rank="species" species="glacialis">
<emphasis id="B923C857FFF4C877FE28BB34FDE5FE53" box="[434,564,367,393]" italics="true" pageId="10" pageNumber="365">D. glacialis</emphasis>
(Rondani)
</taxonomicName>
.
</paragraph>
<paragraph id="8BE81445FFF4C877FF5DBBC9FD0AFD73" blockId="10.[151,1437,150,682]" pageId="10" pageNumber="365">
<emphasis id="B923C857FFF4C877FF5DBBC9FE25FE76" bold="true" box="[199,500,402,428]" pageId="10" pageNumber="365">Type-material Examined.</emphasis>
<materialsCitation id="3B3F1E18FFF4C877FE62BBC8FCDAFE0B" ID-GBIF-Occurrence="3766120303" collectingDate="1983-07-08" collectionCode="BMNH" collectorName="A. C. Pont" country="Georgia" elevation="2300" latitude="42.616665" location="Mt Koltesh" longLatPrecision="1149" longitude="44.75" pageId="10" pageNumber="365" specimenCount="1" specimenCount-male="1" stateProvince="Mtskheta-Mtianeti" typeStatus="holotype">
<typeStatus id="54ECAAE7FFF4C877FE62BBC8FDB1FE77" box="[504,608,403,429]" pageId="10" pageNumber="365" type="holotype">Holotype</typeStatus>
<specimenCount id="9D51DFCCFFF4C877FDFEBBC8FDADFE77" box="[612,636,403,429]" pageId="10" pageNumber="365" type="male"></specimenCount>
.
<collectingCountry id="F34054D5FFF4C877FD19BBC8FCD5FE77" box="[643,772,403,429]" name="Georgia" pageId="10" pageNumber="365">GEORGIA</collectingCountry>
:
<collectingRegion id="4993DAA7FFF4C877FC91BBC8FBFAFE77" box="[779,1067,403,429]" country="Georgia" name="Mtskheta-Mtianeti" pageId="10" pageNumber="365">Mtskheta-Mtianeti region</collectingRegion>
:
<location id="8E88429EFFF4C877FBABBBC8FB7AFE77" LSID="urn:lsid:plazi:treatment:03FEA553FFF7C873FF0DBACCFC26FE77:8E88429EFFF4C877FBABBBC8FB7AFE77" box="[1073,1195,403,429]" country="Georgia" latitude="42.616665" longLatPrecision="1149" longitude="44.75" name="Mt Koltesh" pageId="10" pageNumber="365" stateProvince="Mtskheta-Mtianeti">Mt Koltesh</location>
[~
<geoCoordinate id="EE637282FFF4C877FB50BBC8FAFBFE77" box="[1226,1322,403,429]" degrees="42" direction="north" minutes="37" orientation="latitude" pageId="10" pageNumber="365" precision="925" value="42.616665">42°37N</geoCoordinate>
<geoCoordinate id="EE637282FFF4C877FAB4BBC9FA40FE77" box="[1326,1425,402,429]" degrees="44" direction="east" minutes="45" orientation="longitude" pageId="10" pageNumber="365" precision="925" value="44.75">44°45E</geoCoordinate>
], meadow/heath,
<quantity id="4CAFB9A0FFF4C877FEDDBBECFE71FE0B" box="[327,416,439,465]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="10" pageNumber="365" unit="m" value="2300.0">
<elevation id="007AF376FFF4C877FEDDBBECFE71FE0B" box="[327,416,439,465]" metricMagnitude="3" metricUnit="m" metricValue="2.3" pageId="10" pageNumber="365" unit="m" value="2300.0">2300 m</elevation>
</quantity>
,
<date id="FFE93285FFF4C877FE37BBECFDCFFE0B" box="[429,542,439,465]" pageId="10" pageNumber="365" value="1983-07-08">
<collectingDate id="EFADCB6DFFF4C877FE37BBECFDCFFE0B" box="[429,542,439,465]" pageId="10" pageNumber="365" value="1983-07-08">8.vii.1983</collectingDate>
</date>
,
<collectorName id="26A27193FFF4C877FDB0BBECFD48FE0B" box="[554,665,439,465]" pageId="10" pageNumber="365">A.C. Pont</collectorName>
(
<collectionCode id="ED468C80FFF4C877FD30BBECFCD2FE0B" box="[682,771,439,465]" pageId="10" pageNumber="365">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3F1E18FFF4C877FC82BBECFDF0FE2F" ID-GBIF-Occurrence="3766120320" collectingDate="1983-07-08" collectionCode="BMNH" collectorName="A. C. Pont" country="Georgia" elevation="2300" latitude="42.616665" location="Mt Koltesh" longLatPrecision="1149" longitude="44.75" pageId="10" pageNumber="365" specimenCount="22" specimenCount-female="11" specimenCount-male="11" stateProvince="Mtskheta-Mtianeti" typeStatus="paratype">
<typeStatus id="54ECAAE7FFF4C877FC82BBECFC52FE0B" box="[792,899,439,465]" pageId="10" pageNumber="365" type="paratype">Paratypes</typeStatus>
<specimenCount id="9D51DFCCFFF4C877FC11BBECFC6AFE0B" box="[907,955,439,465]" pageId="10" pageNumber="365" type="male">11♂</specimenCount>
<specimenCount id="9D51DFCCFFF4C877FC5EBBECFC27FE0B" box="[964,1014,439,465]" pageId="10" pageNumber="365" type="female">11♀</specimenCount>
.
<collectingCountry id="F34054D5FFF4C877FB99BBECFB55FE0B" box="[1027,1156,439,465]" name="Georgia" pageId="10" pageNumber="365">GEORGIA</collectingCountry>
:
<collectingRegion id="4993DAA7FFF4C877FB15BBECFA8EFE0B" box="[1167,1375,439,465]" country="Georgia" name="Mtskheta-Mtianeti" pageId="10" pageNumber="365">Mtskheta-Mtianeti</collectingRegion>
(Kazbegi district): data as for holotype
</materialsCitation>
,
<materialsCitation id="3B3F1E18FFF4C877FDB7BB80FA46FE2F" ID-GBIF-Occurrence="3766120308" box="[557,1431,475,501]" collectingDate="1983-06-30" collectionCode="OUMNH, ZMUM, BMNH" collectorName="A. C. Pont" country="Georgia" elevation="2300" latitude="42.616665" location="Mt Koltesh" longLatPrecision="1149" longitude="44.75" pageId="10" pageNumber="365" specimenCount="9" specimenCount-female="3" specimenCount-male="6" stateProvince="Mtskheta-Mtianeti" typeStatus="paratype">
<specimenCount id="9D51DFCCFFF4C877FDB7BB80FD81FE2F" box="[557,592,475,501]" pageId="10" pageNumber="365" type="male">4♂</specimenCount>
<specimenCount id="9D51DFCCFFF4C877FDCDBB80FDABFE2F" box="[599,634,475,501]" pageId="10" pageNumber="365" type="female">3♀</specimenCount>
(
<specimenCount id="9D51DFCCFFF4C877FD10BB80FD7FFE2F" box="[650,686,475,501]" pageId="10" pageNumber="365" type="male">2♂</specimenCount>
<collectionCode id="ED468C80FFF4C877FD2FBB80FCCFFE2F" box="[693,798,475,501]" pageId="10" pageNumber="365">OUMNH</collectionCode>
&amp;
<collectionCode id="ED468C80FFF4C877FCD9BB80FC4DFE2F" box="[835,924,475,501]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="10" pageNumber="365">ZMUM</collectionCode>
, rest
<collectionCode id="ED468C80FFF4C877FC4DBB80FBFEFE2F" box="[983,1071,475,501]" pageId="10" pageNumber="365">BMNH</collectionCode>
); same data but date,
<date id="FFE93285FFF4C877FABABB80FA46FE2F" box="[1312,1431,475,501]" pageId="10" pageNumber="365" value="1983-06-30">
<collectingDate id="EFADCB6DFFF4C877FABABB80FA46FE2F" box="[1312,1431,475,501]" pageId="10" pageNumber="365" value="1983-06-30">30.vi.1983</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B3F1E18FFF4C877FF0DBBA4FBBFFDC3" ID-GBIF-Occurrence="3766120345" box="[151,1134,510,537]" collectingDate="1983-06-30" collectionCode="SZNM, BMNH" collectorName="A. C. Pont" country="Georgia" elevation="2500" location="Mt Koltesh" pageId="10" pageNumber="365" specimenCount="7" specimenCount-female="3" specimenCount-male="4" typeStatus="paratype">
<specimenCount id="9D51DFCCFFF4C877FF0DBBA4FF68FDC3" box="[151,185,511,537]" pageId="10" pageNumber="365" type="male">3♂</specimenCount>
<specimenCount id="9D51DFCCFFF4C877FF5BBBA4FF32FDC3" box="[193,227,511,537]" pageId="10" pageNumber="365" type="female">2♀</specimenCount>
(
<specimenCount id="9D51DFCCFFF4C877FF6EBBA4FEC7FDC3" box="[244,278,511,537]" pageId="10" pageNumber="365" type="male">1♂</specimenCount>
<specimenCount id="9D51DFCCFFF4C877FE87BBA4FE91FDC3" box="[285,320,511,537]" pageId="10" pageNumber="365" type="female">1♀</specimenCount>
<collectionCode id="ED468C80FFF4C877FEDCBBA4FE47FDC3" box="[326,406,511,537]" pageId="10" pageNumber="365">SZNM</collectionCode>
, rest
<collectionCode id="ED468C80FFF4C877FE4ABBA4FDF9FDC3" box="[464,552,511,537]" pageId="10" pageNumber="365">BMNH</collectionCode>
);
<location id="8E88429EFFF4C877FDA0BBA4FD68FDC3" LSID="urn:lsid:plazi:treatment:03FEA553FFF7C873FF0DBACCFC26FE77:8E88429EFFF4C877FDA0BBA4FD68FDC3" box="[570,697,511,537]" country="Georgia" name="Mt Koltesh" pageId="10" pageNumber="365">Mt Koltesh</location>
, heath,
<quantity id="4CAFB9A0FFF4C877FC94BBA5FCB5FDC3" box="[782,868,510,537]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="365" unit="m" value="2500.0">
<elevation id="007AF376FFF4C877FC94BBA5FCB5FDC3" box="[782,868,510,537]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="365" unit="m" value="2500.0">2500 m</elevation>
</quantity>
,
<date id="FFE93285FFF4C877FCF5BBA4FC34FDC3" box="[879,997,511,537]" pageId="10" pageNumber="365" value="1983-06-30">
<collectingDate id="EFADCB6DFFF4C877FCF5BBA4FC34FDC3" box="[879,997,511,537]" pageId="10" pageNumber="365" value="1983-06-30">30.vi.1983</collectingDate>
</date>
(
<collectorName id="26A27193FFF4C877FC6FBBA4FBB6FDC3" box="[1013,1127,511,537]" pageId="10" pageNumber="365">A.C. Pont</collectorName>
)
</materialsCitation>
,
<materialsCitation id="3B3F1E18FFF4C877FBE3BBA4FBE5FDE7" ID-GBIF-Occurrence="3766120312" collectingDate="1983-06-11" collectionCode="OUMNH, BMNH" collectorName="A. C. Pont" country="Georgia" elevation="2650" latitude="42.7" location="Gergeti" longLatPrecision="1148" longitude="44.516666" pageId="10" pageNumber="365" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D51DFCCFFF4C877FBE3BBA4FB4DFDC3" box="[1145,1180,511,537]" pageId="10" pageNumber="365" type="female">2♀</specimenCount>
(
<collectionCode id="ED468C80FFF4C877FB30BBA4FAC5FDC3" box="[1194,1300,511,537]" pageId="10" pageNumber="365">OUMNH</collectionCode>
&amp;
<collectionCode id="ED468C80FFF4C877FAA2BBA4FA41FDC3" box="[1336,1424,511,537]" pageId="10" pageNumber="365">BMNH</collectionCode>
);
<location id="8E88429EFFF4C877FF0DB878FF38FDE7" LSID="urn:lsid:plazi:treatment:03FEA553FFF7C873FF0DBACCFC26FE77:8E88429EFFF4C877FF0DB878FF38FDE7" box="[151,233,547,573]" country="Georgia" latitude="42.7" longLatPrecision="1148" longitude="44.516666" name="Gergeti" pageId="10" pageNumber="365">Gergeti</location>
[~
<geoCoordinate id="EE637282FFF4C877FE8BB878FEA3FDE7" box="[273,370,547,573]" degrees="42" direction="north" minutes="42" orientation="latitude" pageId="10" pageNumber="365" precision="925" value="42.7">42°42N</geoCoordinate>
<geoCoordinate id="EE637282FFF4C877FEE0B879FE0DFDE7" box="[378,476,546,573]" degrees="44" direction="east" minutes="31" orientation="longitude" pageId="10" pageNumber="365" precision="925" value="44.516666">44°31E</geoCoordinate>
], upper heaths,
<quantity id="4CAFB9A0FFF4C877FD11B878FCF6FDE7" box="[651,807,547,573]" metricMagnitude="3" metricUnit="m" metricValue="2.65" metricValueMax="3.0" metricValueMin="2.3" pageId="10" pageNumber="365" unit="m" value="2650.0" valueMax="3000.0" valueMin="2300.0">
<elevation id="007AF376FFF4C877FD11B878FCF6FDE7" box="[651,807,547,573]" metricMagnitude="3" metricUnit="m" metricValue="2.65" metricValueMax="3.0" metricValueMin="2.3" pageId="10" pageNumber="365" unit="m" value="2650.0" valueMax="3000.0" valueMin="2300.0">23003000 m</elevation>
</quantity>
,
<date id="FFE93285FFF4C877FCAEB878FC79FDE7" box="[820,936,547,573]" pageId="10" pageNumber="365" value="1983-06-11">
<collectingDate id="EFADCB6DFFF4C877FCAEB878FC79FDE7" box="[820,936,547,573]" pageId="10" pageNumber="365" value="1983-06-11">11.vi.1983</collectingDate>
</date>
(
<collectorName id="26A27193FFF4C877FC23B878FBFDFDE7" box="[953,1068,547,573]" pageId="10" pageNumber="365">A.C. Pont</collectorName>
)
</materialsCitation>
,
<materialsCitation id="3B3F1E18FFF4C877FBA5B878FB4EFDBB" ID-GBIF-Occurrence="3766120334" collectingDate="2015-07-21" collectionCode="ZMUM, BMNH" collectorName="A. C. Pont" country="Georgia" elevation="3400" latitude="40.483334" location="Mt Aragats" longLatPrecision="1162" longitude="44.183334" municipality="Aragatsotn" pageId="10" pageNumber="365" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCount id="9D51DFCCFFF4C877FBA5B878FBB3FDE7" box="[1087,1122,547,573]" pageId="10" pageNumber="365" type="female">3♀</specimenCount>
(1
<collectionCode id="ED468C80FFF4C877FB13B878FB33FDE7" box="[1161,1250,547,573]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="10" pageNumber="365">ZMUM</collectionCode>
, 2
<collectionCode id="ED468C80FFF4C877FA99B878FA8AFDE7" box="[1283,1371,547,573]" pageId="10" pageNumber="365">BMNH</collectionCode>
). ARMENIA:
<emphasis id="B923C857FFF4C877FE98B81CFEAAFDBB" box="[258,379,583,609]" italics="true" pageId="10" pageNumber="365">
<collectingMunicipality id="6B8C8E3FFFF4C877FE98B81CFEAAFDBB" box="[258,379,583,609]" pageId="10" pageNumber="365">Aragatsotn</collectingMunicipality>
</emphasis>
:
<location id="8E88429EFFF4C877FE16B81CFDDBFDBB" LSID="urn:lsid:plazi:treatment:03FEA553FFF7C873FF0DBACCFC26FE77:8E88429EFFF4C877FE16B81CFDDBFDBB" box="[396,522,583,609]" country="Georgia" latitude="40.483334" longLatPrecision="1162" longitude="44.183334" municipality="Aragatsotn" name="Mt Aragats" pageId="10" pageNumber="365">Mt Aragats</location>
,
<geoCoordinate id="EE637282FFF4C877FD8DB81CFDA7FDBB" box="[535,630,583,609]" degrees="40" direction="north" minutes="29" orientation="latitude" pageId="10" pageNumber="365" precision="925" value="40.483334">40°29N</geoCoordinate>
<geoCoordinate id="EE637282FFF4C877FDE5B81CFD0AFDBB" box="[639,731,583,609]" degrees="44" direction="east" minutes="11" orientation="longitude" pageId="10" pageNumber="365" precision="925" value="44.183334">44°11E</geoCoordinate>
<quantity id="4CAFB9A0FFF4C877FD7EB81CFC52FDBB" box="[740,899,583,609]" metricMagnitude="3" metricUnit="m" metricValue="3.4" metricValueMax="3.42" metricValueMin="3.38" pageId="10" pageNumber="365" unit="m" value="3400.0" valueMax="3420.0" valueMin="3380.0">
<elevation id="007AF376FFF4C877FD7EB81CFC52FDBB" box="[740,899,583,609]" metricMagnitude="3" metricUnit="m" metricValue="3.4" metricValueMax="3.42" metricValueMin="3.38" pageId="10" pageNumber="365" unit="m" value="3400.0" valueMax="3420.0" valueMin="3380.0">33803420 m</elevation>
</quantity>
,
<date id="FFE93285FFF4C877FC0BB81DFBC0FDBB" box="[913,1041,582,609]" pageId="10" pageNumber="365" value="2015-07-21">
<collectingDate id="EFADCB6DFFF4C877FC0BB81DFBC0FDBB" box="[913,1041,582,609]" pageId="10" pageNumber="365" value="2015-07-21">21.vii.2015</collectingDate>
</date>
(
<collectorName id="26A27193FFF4C877FBB9B81CFB46FDBB" box="[1059,1175,583,609]" pageId="10" pageNumber="365">A.C. Pont</collectorName>
)
</materialsCitation>
,
<materialsCitation id="3B3F1E18FFF4C877FB36B81CFCF1FD5F" ID-GBIF-Occurrence="3766120315" collectingDate="2015-07-21" collectionCode="OUMNH" collectorName="A. C. Pont" country="Georgia" elevation="3190" latitude="40.466667" location="Lake Kari" longLatPrecision="1162" longitude="44.183334" pageId="10" pageNumber="365" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D51DFCCFFF4C877FB36B81CFB1EFDBB" box="[1196,1231,583,609]" pageId="10" pageNumber="365" type="male">1♂</specimenCount>
(
<collectionCode id="ED468C80FFF4C877FB7AB81CFA81FDBB" box="[1248,1360,583,609]" pageId="10" pageNumber="365">OUMNH</collectionCode>
);
<location id="8E88429EFFF4C877FAFFB81CFF1BFD5F" LSID="urn:lsid:plazi:treatment:03FEA553FFF7C873FF0DBACCFC26FE77:8E88429EFFF4C877FAFFB81CFF1BFD5F" country="Georgia" latitude="40.466667" longLatPrecision="1162" longitude="44.183334" name="Lake Kari" pageId="10" pageNumber="365">Lake Kari</location>
,
<geoCoordinate id="EE637282FFF4C877FF42B830FEE9FD5F" box="[216,312,619,645]" degrees="40" direction="north" minutes="28" orientation="latitude" pageId="10" pageNumber="365" precision="925" value="40.466667">40°28N</geoCoordinate>
<geoCoordinate id="EE637282FFF4C877FED8B830FE4EFD5F" box="[322,415,619,645]" degrees="44" direction="east" minutes="11" orientation="longitude" pageId="10" pageNumber="365" precision="925" value="44.183334">44°11E</geoCoordinate>
,
<quantity id="4CAFB9A0FFF4C877FE34B830FDD7FD5F" box="[430,518,619,645]" metricMagnitude="3" metricUnit="m" metricValue="3.19" pageId="10" pageNumber="365" unit="m" value="3190.0">
<elevation id="007AF376FFF4C877FE34B830FDD7FD5F" box="[430,518,619,645]" metricMagnitude="3" metricUnit="m" metricValue="3.19" pageId="10" pageNumber="365" unit="m" value="3190.0">3190 m</elevation>
</quantity>
,
<date id="FFE93285FFF4C877FD8EB831FD41FD5F" box="[532,656,618,645]" pageId="10" pageNumber="365" value="2015-07-21">
<collectingDate id="EFADCB6DFFF4C877FD8EB831FD41FD5F" box="[532,656,618,645]" pageId="10" pageNumber="365" value="2015-07-21">21.vii.2015</collectingDate>
</date>
(
<collectorName id="26A27193FFF4C877FD39B830FCC9FD5F" box="[675,792,619,645]" pageId="10" pageNumber="365">A.C. Pont</collectorName>
)
</materialsCitation>
,
<materialsCitation id="3B3F1E18FFF4C877FCB4B830FDE7FD70" ID-GBIF-Occurrence="3766120362" collectingDate="2008-07-11" collectingDateMax="2008-07-12" collectingDateMin="2008-07-11" collectionCode="OUMNH" collectorName="O. Kosterin" country="Russia" location="Lagonaki Plateau" municipality="Caucasus Reserve" pageId="10" pageNumber="365" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D51DFCCFFF4C877FCB4B830FC81FD5F" box="[814,848,619,645]" pageId="10" pageNumber="365" type="female">1♀</specimenCount>
(
<collectionCode id="ED468C80FFF4C877FCF8B830FC01FD5F" box="[866,976,619,645]" pageId="10" pageNumber="365">OUMNH</collectionCode>
).
<collectingCountry id="F34054D5FFF4C877FC7FB830FB9DFD5F" box="[997,1100,619,645]" name="Russia" pageId="10" pageNumber="365">RUSSIA</collectingCountry>
:
<collectingMunicipality id="6B8C8E3FFFF4C877FBC3B830FAF7FD5F" box="[1113,1318,619,645]" pageId="10" pageNumber="365">Caucasus Reserve</collectingMunicipality>
,
<location id="8E88429EFFF4C877FAAEB830FF3AFD73" LSID="urn:lsid:plazi:treatment:03FEA553FFF7C873FF0DBACCFC26FE77:8E88429EFFF4C877FAAEB830FF3AFD73" country="Russia" municipality="Caucasus Reserve" name="Lagonaki Plateau" pageId="10" pageNumber="365">Lagonaki Plateau</location>
,
<date id="FFE93285FFF4C877FF6CB8D4FE4AFD73" box="[246,411,655,681]" pageId="10" pageNumber="365" value="2008-07-11" valueMax="2008-07-12" valueMin="2008-07-11">
<collectingDate id="EFADCB6DFFF4C877FF6CB8D4FE4AFD73" box="[246,411,655,681]" pageId="10" pageNumber="365" value="2008-07-11" valueMax="2008-07-12" valueMin="2008-07-11">1112.vii.2008</collectingDate>
</date>
(
<collectorName id="26A27193FFF4C877FE31B8D4FDFEFD70" box="[427,559,655,682]" pageId="10" pageNumber="365">O. Kosterin</collectorName>
)
</materialsCitation>
,
<materialsCitation id="3B3F1E18FFF4C877FDD8B8D4FD06FD73" ID-GBIF-Occurrence="3766120329" box="[578,727,655,681]" collectionCode="ZMUM" country="Russia" pageId="10" pageNumber="365" specimenCount="3" specimenCount-male="3" typeStatus="paratype">
<specimenCount id="9D51DFCCFFF4C877FDD8B8D4FDB5FD73" box="[578,612,655,681]" pageId="10" pageNumber="365" type="male">3♂</specimenCount>
(
<collectionCode id="ED468C80FFF4C877FDE9B8D4FD01FD73" box="[627,720,655,681]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="10" pageNumber="365">ZMUM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D47CEFFF4C870FF0DB88DFC7DFD2B" lastPageId="13" lastPageNumber="368" pageId="10" pageNumber="365" type="description">
<paragraph id="8BE81445FFF4C877FF0DB88DFE9AFD2B" blockId="10.[151,331,726,753]" box="[151,331,726,753]" pageId="10" pageNumber="365">
<emphasis id="B923C857FFF4C877FF0DB88DFEF8FD2A" bold="true" box="[151,297,726,752]" pageId="10" pageNumber="365">Description.</emphasis>
<specimenCount id="9D51DFCCFFF4C877FEAAB88CFE99FD2B" box="[304,328,727,753]" pageId="10" pageNumber="365" type="male"></specimenCount>
.
</paragraph>
<paragraph id="8BE81445FFF4C877FF0DB944FCC7FB7B" blockId="10.[151,1437,799,2013]" pageId="10" pageNumber="365">
<emphasis id="B923C857FFF4C877FF0DB944FF05FCE3" box="[151,212,799,825]" italics="true" pageId="10" pageNumber="365">Head</emphasis>
(
<figureCitation id="136C08C0FFF4C877FF78B944FEE4FCE3" box="[226,309,799,825]" captionStart="FIGURES 1216" captionStartId="11.[151,264,1886,1910]" captionTargetBox="[206,1381,286,1845]" captionTargetId="figure-57@11.[189,1398,273,1863]" captionTargetPageId="11" captionText="FIGURES 1216. Drymeia fratercula sp.nov. 12. ♂ head, lateral view. 13. ♂ hind tibia and tarsomere 1, posterior view. 14. ♀ fore tarsomeres, anterior view. 15. ♂ sternite 5, dorsal view. 16. ♀ tergite 8. Scale-line 0.25 mm (15), 0.5 mm (16). (Georgia, Ka- zbegi, Mt Koltesh, meadow/heath, 2300 m, 8.vii.1983 (12, 13) and 30.vi.1983 (14, 16); Mt Koltesh, heath, 2500 m, 30.vi.1983 (15).)" figureDoi="http://doi.org/10.5281/zenodo.6538213" httpUri="https://zenodo.org/record/6538213/files/figure.png" pageId="10" pageNumber="365">Fig. 12</figureCitation>
). Ground-colour black. Eye bare. Fronto-orbital plate silvery-white pruinose below, dull above; parafacial silvery-white pruinose; face grey, and gena light grey pruinose; most of occiput subshining. Frons at narrowest point separated by a distance equal to 23 times diameter of anterior ocellus, the margins of each fronto-orbital plate touching or narrowly separated on upper half of frons where the frontal vitta may be obsolete. Ocellar setae long. 12 pairs of frontal setae, including interstitials, and above them, just in front of ocellar tubercle, with 2 pairs of short, fine, reclinate orbitals.Antenna black, postpedicel twice as long as wide. Arista short-pubescent, the individual hairs shorter than its basal diameter. Parafacial at level of insertion of arista twice width of postpedicel, narrowing sharply below. Upper part of face with a conspicuous knob separating antennal bases. In lateral view, facial edge in front of level of profrons. Gena broad, depth below lowest eye-margin equal to length of postpedicel, densely setose and with a group of upcurved setae on anterior part of genal dilation. Occipital setulae long and dense. Palpus black, slender. Proboscis very long, slender, prementum dusted.
</paragraph>
<paragraph id="8BE81445FFF4C877FF5DBEF0FE42FA47" blockId="10.[151,1437,799,2013]" pageId="10" pageNumber="365">
<emphasis id="B923C857FFF4C877FF5DBEF0FEC9FB1F" box="[199,280,1195,1221]" italics="true" pageId="10" pageNumber="365">Thorax</emphasis>
. Ground-colour black. Scutum matt, black, with very indistinct grey vittae before suture, running inside the line of the dorsocentrals, and brownish-grey vittae running between the intra-alar and dorsocentral rows before and after suture (
<figureCitation id="136C08C0FFF4C877FEC5BEA8FE62FAD7" box="[351,435,1267,1293]" captionStart="FIGURES 1011" captionStartId="9.[151,262,1964,1988]" captionTargetBox="[479,1110,237,1933]" captionTargetId="figure-23@9.[477,1110,237,1940]" captionTargetPageId="9" captionText="FIGURES 1011. Drymeia fratercula sp. nov., ♂ habitus. 10. Lateral view. 11. Dorsal view. (holotype; Georgia, Kazbegi, Mt Koltesh, meadow/heath, 2300 m, 8.vii.1983.)" figureDoi="http://doi.org/10.5281/zenodo.6538211" httpUri="https://zenodo.org/record/6538211/files/figure.png" pageId="10" pageNumber="365">Fig. 11</figureCitation>
). Ground-setulae fine, erect. Postpronotal lobe light grey dusted. Pleura thinly dusted. Acrostichals 0+1, the presutural setulae in 34 irregular rows. Dorsocentrals 2+3, exceptionally 2+4. Prealar strong, much longer than 2nd notopleural. Notopleuron densely setulose. Prosternum bare. Katepisternal setae 1+2, with 23 stronger setulae below posterior seta, anterior seta fine. Scutellum black and undusted in posterior view; bare laterally and ventrally.
</paragraph>
<paragraph id="8BE81445FFF4C877FF5DBFFCFE2FF897" blockId="10.[151,1437,799,2013]" pageId="10" pageNumber="365">
<emphasis id="B923C857FFF4C877FF5DBFFCFF2DFA1A" box="[199,252,1447,1472]" italics="true" pageId="10" pageNumber="365">Legs</emphasis>
. Black. Fore tibia with 3 posteroventral setae and 23 stronger anterodorsal setulae just before apex; apical posteroventral seta at most 2/3 as long as length of fore tarsomere 1. Fore tarsomere 1 not as long as the length of tarsomeres 25 combined, not modified, with the posteroventral apical thorn very inconspicuous. Fore claws very short, half as long as length of tarsomere 5, blunt-tipped. Mid femur weakly curved; with complete rows of anteroventral and posteroventral setae, all longer than femoral depth, the posteroventrals rather shorter than the anteroventrals; 0 anterior and 2 posterodorsal to posterior preapical setae. Mid tibia with 3 strong and 2 short anterodorsals, 3 strong and 2 short posterodorsals, 45 short anteroventrals, and 4 posteroventrals. Mid tarsomere 1 with the ventral spinules shorter than tarsal depth. Hind femur with complete rows of long strong anteroventral and posteroventral setae, all much longer than femoral depth. Hind tibia with 45 posterodorsals, 34 strong anterodorsals interspersed with 45 shorter setae, 45 anteroventrals, and 45 fine posteroventrals; 1 dorsal and 1 short anterodorsal preapical seta; ventral tip slightly produced, with 1 anteroventral and 1 posteroventral apical seta (
<figureCitation id="136C08C0FFF4C877FBE9BD54FB17F8F3" box="[1139,1222,1807,1833]" captionStart="FIGURES 1216" captionStartId="11.[151,264,1886,1910]" captionTargetBox="[206,1381,286,1845]" captionTargetId="figure-57@11.[189,1398,273,1863]" captionTargetPageId="11" captionText="FIGURES 1216. Drymeia fratercula sp.nov. 12. ♂ head, lateral view. 13. ♂ hind tibia and tarsomere 1, posterior view. 14. ♀ fore tarsomeres, anterior view. 15. ♂ sternite 5, dorsal view. 16. ♀ tergite 8. Scale-line 0.25 mm (15), 0.5 mm (16). (Georgia, Ka- zbegi, Mt Koltesh, meadow/heath, 2300 m, 8.vii.1983 (12, 13) and 30.vi.1983 (14, 16); Mt Koltesh, heath, 2500 m, 30.vi.1983 (15).)" figureDoi="http://doi.org/10.5281/zenodo.6538213" httpUri="https://zenodo.org/record/6538213/files/figure.png" pageId="10" pageNumber="365">Fig. 13</figureCitation>
). Hind tarsomere 1 with a short ventral seta at base.
</paragraph>
<paragraph id="8BE81445FFF4C877FF5DBD0CFC01F863" blockId="10.[151,1437,799,2013]" pageId="10" pageNumber="365">
<emphasis id="B923C857FFF4C877FF5DBD0CFED0F8AA" box="[199,257,1879,1904]" italics="true" pageId="10" pageNumber="365">Wing</emphasis>
. Brown, especially at base (
<figureCitation id="136C08C0FFF4C877FDA3BD0CFD5DF8AB" box="[569,652,1879,1905]" captionStart="FIGURES 1011" captionStartId="9.[151,262,1964,1988]" captionTargetBox="[479,1110,237,1933]" captionTargetId="figure-23@9.[477,1110,237,1940]" captionTargetPageId="9" captionText="FIGURES 1011. Drymeia fratercula sp. nov., ♂ habitus. 10. Lateral view. 11. Dorsal view. (holotype; Georgia, Kazbegi, Mt Koltesh, meadow/heath, 2300 m, 8.vii.1983.)" figureDoi="http://doi.org/10.5281/zenodo.6538211" httpUri="https://zenodo.org/record/6538211/files/figure.png" pageId="10" pageNumber="365">Fig. 10</figureCitation>
). Basicosta and tegula black. Costa with weak spinules, without costal spine. Cross-vein r-m distinctly beyond the point where subcosta enters costa. Cross-vein dm-cu oblique, almost straight. Calypters creamy, margins deeper yellow. Knob of haltere black.
</paragraph>
<paragraph id="8BE81445FFF4C876FF5DBD98FBDEFF0F" blockId="10.[151,1437,799,2013]" lastBlockId="11.[151,1436,150,249]" lastPageId="11" lastPageNumber="366" pageId="10" pageNumber="365">
<emphasis id="B923C857FFF4C877FF5DBD98FEE0F807" box="[199,305,1987,2013]" italics="true" pageId="10" pageNumber="365">Abdomen</emphasis>
. Ground-colour black. When viewed from above, appearing matt with grey dust at sides of syntergite 1+2 and forming broad antero-lateral patches on tergites 35. Setae strong and erect laterally and medially on all tergites, except medially on syntergite 1+2. Sternite 1 bare. Sternite 5 (
<figureCitation id="136C08C0FFF5C876FC35BAE0FBD3FF0F" box="[943,1026,186,213]" captionStart="FIGURES 1216" captionStartId="11.[151,264,1886,1910]" captionTargetBox="[206,1381,286,1845]" captionTargetId="figure-57@11.[189,1398,273,1863]" captionTargetPageId="11" captionText="FIGURES 1216. Drymeia fratercula sp.nov. 12. ♂ head, lateral view. 13. ♂ hind tibia and tarsomere 1, posterior view. 14. ♀ fore tarsomeres, anterior view. 15. ♂ sternite 5, dorsal view. 16. ♀ tergite 8. Scale-line 0.25 mm (15), 0.5 mm (16). (Georgia, Ka- zbegi, Mt Koltesh, meadow/heath, 2300 m, 8.vii.1983 (12, 13) and 30.vi.1983 (14, 16); Mt Koltesh, heath, 2500 m, 30.vi.1983 (15).)" figureDoi="http://doi.org/10.5281/zenodo.6538213" httpUri="https://zenodo.org/record/6538213/files/figure.png" pageId="11" pageNumber="366">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="8BE81445FFF5C876FF5DBA84FC79FF23" blockId="11.[151,1436,150,249]" box="[199,936,223,249]" pageId="11" pageNumber="366">
<emphasis id="B923C857FFF5C876FF5DBA84FE90FF23" box="[199,321,223,249]" italics="true" pageId="11" pageNumber="366">Terminalia</emphasis>
(
<figureCitation id="136C08C0FFF5C876FECABA84FE03FF23" box="[336,466,223,249]" captionStart="FIGURES 1718" captionStartId="12.[151,264,1909,1933]" captionTargetBox="[220,1383,196,1871]" captionTargetId="figure-38@12.[204,1383,181,1884]" captionTargetPageId="12" captionText="FIGURES 1718. Drymeia fratercula sp.nov., ♂ terminalia. 17. Cercal plate, dorsal view. 18. Hypopygium, lateral view. Scale lines 0.25 mm. (Georgia, Kazbegi, Mt Koltesh, heath, 2500 m, 30.vi.1983.)" figureDoi="http://doi.org/10.5281/zenodo.6538215" httpUri="https://zenodo.org/record/6538215/files/figure.png" pageId="11" pageNumber="366">Figs 1718</figureCitation>
). Syntergite 6+7+8 with long dense setae.
</paragraph>
<caption id="DF2844CDFFF5C876FF0DBD05FF00F839" ID-DOI="http://doi.org/10.5281/zenodo.6538213" ID-Zenodo-Dep="6538213" httpUri="https://zenodo.org/record/6538213/files/figure.png" pageId="11" pageNumber="366" startId="11.[151,264,1886,1910]" targetBox="[206,1381,286,1845]" targetPageId="11">
<paragraph id="8BE81445FFF5C876FF0DBD05FF00F839" blockId="11.[151,1436,1886,2019]" pageId="11" pageNumber="366">
<emphasis id="B923C857FFF5C876FF0DBD05FE85F8AD" bold="true" box="[151,340,1886,1911]" pageId="11" pageNumber="366">FIGURES 1216.</emphasis>
<taxonomicName id="4C576FC6FFF5C876FEC0BD04FDCCF8AD" authority="Pont, 2022" authorityName="Pont" authorityYear="2022" box="[346,541,1887,1911]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="366" phylum="Arthropoda" rank="species" species="fratercula" status="sp. nov.">
<emphasis id="B923C857FFF5C876FEC0BD04FDCCF8AD" box="[346,541,1887,1911]" italics="true" pageId="11" pageNumber="366">Drymeia fratercula</emphasis>
</taxonomicName>
<emphasis id="B923C857FFF5C876FDBEBD04FD45F8AD" bold="true" box="[548,660,1887,1911]" pageId="11" pageNumber="366">
<taxonomicNameLabel id="A210752CFFF5C876FDBEBD04FDBEF8AD" box="[548,623,1887,1911]" pageId="11" pageNumber="366" rank="species">sp.nov.</taxonomicNameLabel>
12.
</emphasis>
♂ head, lateral view.
<emphasis id="B923C857FFF5C876FCE8BD04FC43F8AD" bold="true" box="[882,914,1887,1911]" pageId="11" pageNumber="366">13.</emphasis>
♂ hind tibia and tarsomere 1, posterior view.
<emphasis id="B923C857FFF5C876FAF9BD04FA52F8AD" bold="true" box="[1379,1411,1887,1911]" pageId="11" pageNumber="366">14.</emphasis>
♀ fore tarsomeres, anterior view.
<emphasis id="B923C857FFF5C876FE54BDD8FE3CF841" bold="true" box="[462,493,1923,1947]" pageId="11" pageNumber="366">15.</emphasis>
♂ sternite 5, dorsal view.
<emphasis id="B923C857FFF5C876FD6BBDD8FCC0F841" bold="true" box="[753,785,1923,1947]" pageId="11" pageNumber="366">16.</emphasis>
♀ tergite 8. Scale-line 0.25 mm (15), 0.5 mm (16). (Georgia, Kazbegi, Mt Koltesh, meadow/heath, 2300 m, 8.vii.1983 (12, 13) and 30.vi.1983 (14, 16); Mt Koltesh, heath, 2500 m, 30.vi.1983 (15).)
</paragraph>
</caption>
<caption id="DF2844CDFFF2C871FF0DBD2EFC42F86B" ID-DOI="http://doi.org/10.5281/zenodo.6538215" ID-Zenodo-Dep="6538215" httpUri="https://zenodo.org/record/6538215/files/figure.png" pageId="12" pageNumber="367" startId="12.[151,264,1909,1933]" targetBox="[220,1383,196,1871]" targetPageId="12">
<paragraph id="8BE81445FFF2C871FF0DBD2EFC42F86B" blockId="12.[151,1436,1909,1969]" pageId="12" pageNumber="367">
<emphasis id="B923C857FFF2C871FF0DBD2EFE82F857" bold="true" box="[151,339,1909,1933]" pageId="12" pageNumber="367">FIGURES 1718.</emphasis>
<taxonomicName id="4C576FC6FFF2C871FEC3BD2DFDCAF857" authority="Pont, 2022" authorityName="Pont" authorityYear="2022" box="[345,539,1909,1933]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="367" phylum="Arthropoda" rank="species" species="fratercula" status="sp. nov.">
<emphasis id="B923C857FFF2C871FEC3BD2DFDCAF857" box="[345,539,1909,1933]" italics="true" pageId="12" pageNumber="367">Drymeia fratercula</emphasis>
</taxonomicName>
<emphasis id="B923C857FFF2C871FDBBBD2EFDBDF857" bold="true" box="[545,620,1909,1933]" pageId="12" pageNumber="367">
<taxonomicNameLabel id="A210752CFFF2C871FDBBBD2EFDBDF857" box="[545,620,1909,1933]" pageId="12" pageNumber="367" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
, ♂ terminalia.
<emphasis id="B923C857FFF2C871FC9EBD2EFCF2F857" bold="true" box="[772,803,1909,1933]" pageId="12" pageNumber="367">17.</emphasis>
Cercal plate, dorsal view.
<emphasis id="B923C857FFF2C871FBB7BD2EFB9CF857" bold="true" box="[1069,1101,1909,1933]" pageId="12" pageNumber="367">18.</emphasis>
Hypopygium, lateral view. Scale lines 0.25 mm. (Georgia, Kazbegi, Mt Koltesh, heath, 2500 m, 30.vi.1983.)
</paragraph>
</caption>
<paragraph id="8BE81445FFF3C870FF5DBACCFC7DFF6B" blockId="13.[151,1437,150,897]" box="[199,940,150,177]" pageId="13" pageNumber="368">
<emphasis id="B923C857FFF3C870FF5DBACCFEB8FF6A" box="[199,361,151,176]" italics="true" pageId="13" pageNumber="368">Measurements</emphasis>
. Length of body, 6.0 mm. Length of wing,
<quantity id="4CAFB9A0FFF3C870FCD4BACDFC76FF6B" box="[846,935,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="13" pageNumber="368" unit="mm" value="4.5">4.5 mm</quantity>
.
</paragraph>
<paragraph id="8BE81445FFF3C870FF5DBAE0FD8EFF0F" blockId="13.[151,1437,150,897]" box="[199,607,187,213]" pageId="13" pageNumber="368">
<specimenCount id="9D51DFCCFFF3C870FF5DBAE0FF0CFF0F" box="[199,221,187,213]" pageId="13" pageNumber="368" type="female"></specimenCount>
. Differs from the male as follows:
</paragraph>
<paragraph id="8BE81445FFF3C870FF5DBA84FD61FE53" blockId="13.[151,1437,150,897]" pageId="13" pageNumber="368">
<emphasis id="B923C857FFF3C870FF5DBA84FED5FF23" box="[199,260,223,249]" italics="true" pageId="13" pageNumber="368">Head</emphasis>
. Dichoptic. Frons at middle 0.4 of head-width at this point, and here each fronto-orbital plate 0.3 of frontal vitta. Frontal triangle inconspicuous, not reaching the crossed setae on frontal vitta. 67 pairs of inclinate frontal setae, including several interstitials; 3 pairs of orbital setae, the upper two pairs reclinate and exclinate, the lower pair proclinate and exclinate. Frontal vitta with a pair of strong crossed setae. Fronto-orbital plate bare apart from the setae. Occipital setae shorter and less dense.
</paragraph>
<paragraph id="8BE81445FFF3C870FF5DBBC8FAB1FE0B" blockId="13.[151,1437,150,897]" pageId="13" pageNumber="368">
<emphasis id="B923C857FFF3C870FF5DBBC8FF2DFE76" box="[199,252,403,428]" italics="true" pageId="13" pageNumber="368">Legs</emphasis>
. Fore tarsomeres 14 (
<figureCitation id="136C08C0FFF3C870FD9DBBC8FD8AFE77" box="[519,603,403,429]" captionStart="FIGURES 1216" captionStartId="11.[151,264,1886,1910]" captionTargetBox="[206,1381,286,1845]" captionTargetId="figure-57@11.[189,1398,273,1863]" captionTargetPageId="11" captionText="FIGURES 1216. Drymeia fratercula sp.nov. 12. ♂ head, lateral view. 13. ♂ hind tibia and tarsomere 1, posterior view. 14. ♀ fore tarsomeres, anterior view. 15. ♂ sternite 5, dorsal view. 16. ♀ tergite 8. Scale-line 0.25 mm (15), 0.5 mm (16). (Georgia, Ka- zbegi, Mt Koltesh, meadow/heath, 2300 m, 8.vii.1983 (12, 13) and 30.vi.1983 (14, 16); Mt Koltesh, heath, 2500 m, 30.vi.1983 (15).)" figureDoi="http://doi.org/10.5281/zenodo.6538213" httpUri="https://zenodo.org/record/6538213/files/figure.png" pageId="13" pageNumber="368">Fig. 14</figureCitation>
) with dense fine erect hairs on ventral surfaces, mostly as long as tarsal depth. Mid femur straight. Hind tibia with 45 anterodorsals and 45 anteroventrals; without posteroventrals.
</paragraph>
<paragraph id="8BE81445FFF3C870FF5DBB80FD16FDC3" blockId="13.[151,1437,150,897]" pageId="13" pageNumber="368">
<emphasis id="B923C857FFF3C870FF5DBB80FED0FE2E" box="[199,257,475,500]" italics="true" pageId="13" pageNumber="368">Wing</emphasis>
. Brownish, especially at base. Costa with strong spinules from humeral cross-vein almost to tip of vein R1. Calypters yellow, their margins bright yellow.
</paragraph>
<paragraph id="8BE81445FFF3C870FF5DB878FCC3FDBB" blockId="13.[151,1437,150,897]" pageId="13" pageNumber="368">
<emphasis id="B923C857FFF3C870FF5DB878FEE0FDE7" box="[199,305,547,573]" italics="true" pageId="13" pageNumber="368">Abdomen</emphasis>
. Grey dusting more extensive, tergites 35 each with a poorly defined black median vitta, very broad on tergite 3, to narrow on tergite 5. Without strong setae.
</paragraph>
<paragraph id="8BE81445FFF3C870FF5DB830FDC0FD17" blockId="13.[151,1437,150,897]" pageId="13" pageNumber="368">
<emphasis id="B923C857FFF3C870FF5DB830FEEEFD5F" box="[199,319,619,645]" italics="true" pageId="13" pageNumber="368">Ovipositor</emphasis>
. Long and slender, 1.7 times as long as length of abdomen. As in
<taxonomicName id="4C576FC6FFF3C870FBACB830FB6BFD5F" baseAuthorityName="Huckett" baseAuthorityYear="1965" box="[1078,1210,619,645]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="368" phylum="Arthropoda" rank="species" species="setibasis">
<emphasis id="B923C857FFF3C870FBACB830FB6BFD5F" box="[1078,1210,619,645]" italics="true" pageId="13" pageNumber="368">D. setibasis</emphasis>
</taxonomicName>
except that the two elongated rods of tergite 8 are fused into a single plate (
<figureCitation id="136C08C0FFF3C870FC9FB8D4FC89FD70" box="[773,856,655,682]" captionStart="FIGURES 1216" captionStartId="11.[151,264,1886,1910]" captionTargetBox="[206,1381,286,1845]" captionTargetId="figure-57@11.[189,1398,273,1863]" captionTargetPageId="11" captionText="FIGURES 1216. Drymeia fratercula sp.nov. 12. ♂ head, lateral view. 13. ♂ hind tibia and tarsomere 1, posterior view. 14. ♀ fore tarsomeres, anterior view. 15. ♂ sternite 5, dorsal view. 16. ♀ tergite 8. Scale-line 0.25 mm (15), 0.5 mm (16). (Georgia, Ka- zbegi, Mt Koltesh, meadow/heath, 2300 m, 8.vii.1983 (12, 13) and 30.vi.1983 (14, 16); Mt Koltesh, heath, 2500 m, 30.vi.1983 (15).)" figureDoi="http://doi.org/10.5281/zenodo.6538213" httpUri="https://zenodo.org/record/6538213/files/figure.png" pageId="13" pageNumber="368">Fig. 16</figureCitation>
), and the two lines of setulae at the tip of the tergite are placed on a single weak plate.
</paragraph>
<paragraph id="8BE81445FFF3C870FF5DB88CFC7DFD2B" blockId="13.[151,1437,150,897]" box="[199,940,726,753]" pageId="13" pageNumber="368">
<emphasis id="B923C857FFF3C870FF5DB88CFEB8FD2A" box="[199,361,727,752]" italics="true" pageId="13" pageNumber="368">Measurements</emphasis>
. Length of body, 7.0 mm. Length of wing,
<quantity id="4CAFB9A0FFF3C870FCD4B88DFC76FD2B" box="[846,935,726,753]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="13" pageNumber="368" unit="mm" value="5.5">5.5 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C34D47CEFFF3C873FF5DB8A0FC26FE77" lastPageId="14" lastPageNumber="369" pageId="13" pageNumber="368" type="discussion">
<paragraph id="8BE81445FFF3C870FF5DB8A0FA5BFC5B" blockId="13.[151,1437,150,897]" pageId="13" pageNumber="368">
<emphasis id="B923C857FFF3C870FF5DB8A0FEEAFCCF" bold="true" box="[199,315,763,789]" pageId="13" pageNumber="368">Remarks.</emphasis>
In
<bibRefCitation id="EFC669B4FFF3C870FEF8B8A0FDA0FCCF" author="Sorokina, V. S. &amp; Pont, A. C." box="[354,625,762,789]" pageId="13" pageNumber="368" pagination="151 - 206" refId="ref14383" refString="Sorokina, V. S. &amp; Pont, A. C. (2015) A review of the genus Drymeia Meigen, 1826 (Diptera: Muscidae) in Russia. Zootaxa, 4000 (2), 151 - 206. https: // doi. org / 10.11646 / zootaxa. 4000.2.1" type="journal article" year="2015">Sorokina &amp; Pont (2015)</bibRefCitation>
,
<taxonomicName id="4C576FC6FFF3C870FDE4B8A0FCDFFCCF" authorityName="Pont" authorityYear="2022" box="[638,782,763,789]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="368" phylum="Arthropoda" rank="species" species="fratercula">
<emphasis id="B923C857FFF3C870FDE4B8A0FCDFFCCF" box="[638,782,763,789]" italics="true" pageId="13" pageNumber="368">D. fratercula</emphasis>
</taxonomicName>
keys out to
<taxonomicName id="4C576FC6FFF3C870FC02B8A0FBC8FCCF" authorityName="" baseAuthorityName="Rondani" baseAuthorityYear="1866" box="[920,1049,763,789]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="368" phylum="Arthropoda" rank="species" species="glacialis">
<emphasis id="B923C857FFF3C870FC02B8A0FBC8FCCF" box="[920,1049,763,789]" italics="true" pageId="13" pageNumber="368">D. glacialis</emphasis>
</taxonomicName>
(as
<taxonomicName id="4C576FC6FFF3C870FBD0B8A0FAFCFCCC" authority="Rondani" authorityName="Rondani" authorityYear="1871" box="[1098,1325,763,790]" class="Insecta" family="Muscidae" genus="Pogonomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="368" phylum="Arthropoda" rank="species" species="alpicola">
<emphasis id="B923C857FFF3C870FBD0B8A0FB17FCCF" box="[1098,1222,763,789]" italics="true" pageId="13" pageNumber="368">D. alpicola</emphasis>
Rondani
</taxonomicName>
) from the European Alps from which it differs as stated in the diagnosis. The two species are extremely similar and are evidently sister-species. In the key by
<bibRefCitation id="EFC669B4FFF3C870FE51B918FD95FC87" author="Fan, Z." box="[459,580,835,861]" pageId="13" pageNumber="368" refId="ref13369" refString="Fan, Z. (Ed.) (2008) Fauna Sinica. Insecta. Vol. 49. Diptera Muscidae (I). Science Press, Beijing, xvi + 1186 pp." type="book" year="2008">Fan (2008)</bibRefCitation>
it runs to couplet 29 but differs from the species therein by the armature of the
<specimenCount id="9D51DFCCFFF3C870FF0DB93CFF7DFC5B" box="[151,172,871,897]" pageId="13" pageNumber="368" type="male"></specimenCount>
mid femur. In the key by
<bibRefCitation id="EFC669B4FFF3C870FE53B93CFD52FC5B" author="Xue, W. - Q. &amp; Pont, A. C. &amp; Wang, M. - F." box="[457,643,871,897]" pageId="13" pageNumber="368" pagination="493 - 503" refId="ref14638" refString="Xue, W. - Q., Pont, A. C. &amp; Wang, M. - F. (2008) A study of the genus Drymeia Meigen (Diptera: Muscidae) from the Tibetan Plateau and surrounding regions of P. R. China, with descriptions of three new species. Proceedings of the entomological Society of Washington, 110, 493 - 503. https: // doi. org / 10.4289 / 07 - 040.1" type="journal article" year="2008">Xue et al. (2008)</bibRefCitation>
it runs to couplet 39 and to
<taxonomicName id="4C576FC6FFF3C870FC2CB93CFB41FC5B" authority="Fan" authorityName="Fan" authorityYear="1993" box="[950,1168,871,897]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="368" phylum="Arthropoda" rank="species" species="melargentea">
<emphasis id="B923C857FFF3C870FC2CB93CFBB0FC5B" box="[950,1121,871,897]" italics="true" pageId="13" pageNumber="368">D. melargentea</emphasis>
Fan
</taxonomicName>
from
<collectingCountry id="F34054D5FFF3C870FB4EB93CFAC7FC5B" box="[1236,1302,871,897]" name="China" pageId="13" pageNumber="368">China</collectingCountry>
(
<collectingRegion id="4993DAA7FFF3C870FABEB93CFAAEFC5B" box="[1316,1407,871,897]" country="China" name="Yunnan" pageId="13" pageNumber="368">Yunnan</collectingRegion>
).
</paragraph>
<paragraph id="8BE81445FFF3C870FF0DB988FBB7FC37" blockId="13.[151,1126,979,1005]" box="[151,1126,979,1005]" pageId="13" pageNumber="368">
<taxonomicName id="4C576FC6FFF3C870FF0DB988FEA8FC37" authorityName="Pont" authorityYear="2022" box="[151,377,979,1005]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="368" phylum="Arthropoda" rank="species" species="fratercula">
<emphasis id="B923C857FFF3C870FF0DB988FEA8FC37" box="[151,377,979,1005]" italics="true" pageId="13" pageNumber="368">Drymeia fratercula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C576FC6FFF3C870FE28B988FDEAFC37" authorityName="" baseAuthorityName="Rondani" baseAuthorityYear="1866" box="[434,571,979,1005]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="368" phylum="Arthropoda" rank="species" species="glacialis">
<emphasis id="B923C857FFF3C870FE28B988FDEAFC37" box="[434,571,979,1005]" italics="true" pageId="13" pageNumber="368">D. glacialis</emphasis>
</taxonomicName>
can be separated by the following key couplet:
</paragraph>
<paragraph id="8BE81445FFF3C870FF0DBE43FA4DFB5C" blockId="13.[151,1436,1048,1158]" pageId="13" pageNumber="368">
- Larger and more robust species, length up to
<quantity id="4CAFB9A0FFF3C870FD14BE43FD16FBF7" box="[654,711,1048,1069]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="368" unit="mm" value="8.0">8 mm</quantity>
, with strong setae. Fore tibia with the apical posteroventral seta long, almost as long as fore tarsomere 1.
<specimenCount id="9D51DFCCFFF3C870FE70BE6EFE2EFB90" box="[490,511,1077,1098]" pageId="13" pageNumber="368" type="male"></specimenCount>
: fore tarsomere 1 with a distinct posteroventral apical thorn; frons at narrowest point usually equal in width to diameter of anterior ocellus; hind femur with no or few posteroventral setae in apical half.
<specimenCount id="9D51DFCCFFF3C870FB39BE08FB67FBB2" box="[1187,1206,1107,1128]" pageId="13" pageNumber="368" type="female"></specimenCount>
: the mat of ventral hairs on fore tarsomeres 24 much longer than the depth of the tarsomeres.............................
<taxonomicName id="4C576FC6FFF3C870FB53BE2BFA4DFB5C" authority="(Rondani)" authorityName="" baseAuthorityName="Rondani" baseAuthorityYear="1866" box="[1225,1436,1136,1158]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="368" phylum="Arthropoda" rank="species" species="glacialis">
<emphasis id="B923C857FFF3C870FB53BE2BFAE6FB5F" box="[1225,1335,1136,1157]" italics="true" pageId="13" pageNumber="368">D. glacialis</emphasis>
(Rondani)
</taxonomicName>
</paragraph>
<paragraph id="8BE81445FFF3C870FF0DBEF0FEFFFAEF" blockId="13.[151,1437,1194,1333]" pageId="13" pageNumber="368">
- Smaller and less robust species, length up to
<quantity id="4CAFB9A0FFF3C870FD12BEF0FD10FB1A" box="[648,705,1195,1216]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="13" pageNumber="368" unit="mm" value="7.0">7 mm</quantity>
, with weaker setae. Fore tibia with the apical posteroventral seta a little over half as long as length of fore tarsomere 1.
<specimenCount id="9D51DFCCFFF3C870FDE5BE93FD45FB07" box="[639,660,1224,1245]" pageId="13" pageNumber="368" type="male"></specimenCount>
: fore tarsomere 1 with the posteroventral apical thorn very inconspicuous; frons at narrowest point separated by a distance equal to 23 times diameter of anterior ocellus; hind femur with a complete row of posteroventral setae.
<specimenCount id="9D51DFCCFFF3C870FE37BF58FE11FAC2" box="[429,448,1283,1304]" pageId="13" pageNumber="368" type="female"></specimenCount>
: the mat of ventral hairs on fore tarsomeres 24 not as long as the depth of the tarsomeres
<taxonomicName id="4C576FC6FFF3C870FABABF58FA4DFACD" authority="Pont, 2022" authorityName="Pont" authorityYear="2022" box="[1312,1436,1282,1304]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="368" phylum="Arthropoda" rank="species" species="fratercula" status="sp. nov.">
<emphasis id="B923C857FFF3C870FABABF58FA4DFACD" box="[1312,1436,1282,1304]" italics="true" pageId="13" pageNumber="368">D. fratercula</emphasis>
</taxonomicName>
<emphasis id="B923C857FFF3C870FF78BF7BFEFFFAEF" bold="true" box="[226,302,1312,1333]" pageId="13" pageNumber="368">
<taxonomicNameLabel id="A210752CFFF3C870FF78BF7BFEFFFAEF" box="[226,302,1312,1333]" pageId="13" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BE81445FFF3C870FF5DBF06FCA4F94D" blockId="13.[151,1437,1373,2012]" pageId="13" pageNumber="368">
Some of these differential characters are slightly variable, but these two allopatric taxa appear to be species in the process of divergence.
<taxonomicName id="4C576FC6FFF3C870FE5FBFD9FD99FA41" authorityName="" baseAuthorityName="Rondani" baseAuthorityYear="1866" box="[453,584,1409,1435]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="368" phylum="Arthropoda" rank="species" species="glacialis">
<emphasis id="B923C857FFF3C870FE5FBFD9FD99FA41" box="[453,584,1409,1435]" italics="true" pageId="13" pageNumber="368">D. glacialis</emphasis>
</taxonomicName>
is found in the Western Palaearctic only in the European Alps, where it occupies the highest alpine biotopes, meadows and alpine heaths from
<quantity id="4CAFB9A0FFF3C870FC3FBFFEFB8CFA1A" box="[933,1117,1445,1472]" metricMagnitude="3" metricUnit="m" metricValue="1.975" metricValueMax="2.5" metricValueMin="1.45" pageId="13" pageNumber="368" unit="m" value="1975.0" valueMax="2500.0" valueMin="1450.0">1450 to 2500 m</quantity>
.
<taxonomicName id="4C576FC6FFF3C870FBF3BFFDFB2DFA65" authorityName="Pont" authorityYear="2022" box="[1129,1276,1445,1471]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="368" phylum="Arthropoda" rank="species" species="fratercula">
<emphasis id="B923C857FFF3C870FBF3BFFDFB2DFA65" box="[1129,1276,1445,1471]" italics="true" pageId="13" pageNumber="368">D. fratercula</emphasis>
</taxonomicName>
is endemic to the Caucasus Mountains and is found only in the zone of alpine heaths from
<quantity id="4CAFB9A0FFF3C870FC6DBF92FB78FA3E" box="[1015,1193,1481,1508]" metricMagnitude="3" metricUnit="m" metricValue="2.86" metricValueMax="3.42" metricValueMin="2.3" pageId="13" pageNumber="368" unit="m" value="2860.0" valueMax="3420.0" valueMin="2300.0">2300 to 3420 m</quantity>
asl. The Alps and the Caucasus are two segments of the late Coenozoic trans-Eurasian orogenic belt (
<bibRefCitation id="EFC669B4FFF3C870FBBDBFB6FADCF9D2" author="Sharkov, E. &amp; Lebedev, V. &amp; Chugaev, A. &amp; Zabarinskaya, L. &amp; Rodnikov, N. &amp; Sergeeva, N. &amp; Safonova, I." box="[1063,1293,1517,1544]" pageId="13" pageNumber="368" pagination="513 - 522" refId="ref14244" refString="Sharkov, E., Lebedev, V., Chugaev, A., Zabarinskaya, L., Rodnikov, N., Sergeeva, N. &amp; Safonova, I. (2015) The Caucasian- Arabian segment of the Alpine-Himalayan collisional belt: geology, volcanism and neotectonics. Geoscience Frontiers, 6, 513 - 522. https: // doi. org / 10.1016 / j. gsf. 2014.07.001" type="journal article" year="2015">Sharkov et al. 2015</bibRefCitation>
). During the Quaternary glaciations the snow-line in the Caucasus was up to
<quantity id="4CAFB9A0FFF3C870FCF7BC4AFC12F9F6" box="[877,963,1553,1580]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="368" unit="m" value="1300.0">1300 m</quantity>
lower than it is at present (
<bibRefCitation id="EFC669B4FFF3C870FB61BC4AFA49F9F1" author="Tielidze, L. G." box="[1275,1432,1553,1579]" pageId="13" pageNumber="368" pagination="517 - 532" refId="ref14595" refString="Tielidze, L. G. (2017) Late Pleistocene and Holocene glacier extent in the Georgian Caucasus. Open Journal of Geology, 7, 517 - 532. https: // doi. org / 10.4236 / ojg. 2017.74036" type="journal article" year="2017">Tielidze 2017</bibRefCitation>
,
<bibRefCitation id="EFC669B4FFF3C870FF0DBC6DFE8CF995" author="Revaz, K. &amp; Koba, K. &amp; Kukuri, T. &amp; Giorgio, C." box="[151,349,1589,1616]" pageId="13" pageNumber="368" pagination="56 - 64" refId="ref13977" refString="Revaz, K., Koba, K., Kukuri, T. &amp; Giorgio, C. (2018) Ancient glaciation of the Caucasus. Open Journal of Geology, 8, 56 - 64. https: // doi. org / 10.4236 / ojg. 2018.81004" type="journal article" year="2018">Revaz et al. 2018</bibRefCitation>
) and the foothills were in the tundra zone. During subsequent warming, the retreating tundra led to the faunistic isolation of these two ranges and the tundra fauna eventually occupied the highest zones where individual taxa have evidently begun to separate into sister-species.
</paragraph>
<paragraph id="8BE81445FFF3C870FF5DBCFAFE2CF86D" blockId="13.[151,1437,1373,2012]" pageId="13" pageNumber="368">
<emphasis id="B923C857FFF3C870FF5DBCFAFD75F961" bold="true" box="[199,676,1697,1724]" pageId="13" pageNumber="368">
Note on
<taxonomicName id="4C576FC6FFF3C870FEB7BCFAFD4EF961" authority="Rondani, 1866" authorityName="Rondani" authorityYear="1866" box="[301,671,1697,1724]" class="Magnoliopsida" family="Asteraceae" genus="Aspilia" kingdom="Plantae" order="Asterales" pageId="13" pageNumber="368" phylum="Tracheophyta" rank="species" species="glacialis">
<emphasis id="B923C857FFF3C870FEB7BCFAFE36F966" bold="true" box="[301,487,1697,1724]" italics="true" pageId="13" pageNumber="368">Aspilia glacialis</emphasis>
<bibRefCitation id="EFC669B4FFF3C870FE75BCFAFD4EF961" author="Rondani, C." box="[495,671,1697,1723]" pageId="13" pageNumber="368" pagination="68 - 217" refId="ref14028" refString="Rondani, C. (1866) Anthomyinae Italicae collectae distinctae et in ordinem dispositae [a prof. Camillo Rondani]. Atti della Societa italiana di Scienze naturali, 9, 68 - 217." type="journal article" year="1866">Rondani, 1866</bibRefCitation>
</taxonomicName>
.
</emphasis>
This species was described by Rondani from a single female from the Insubrian Alps of northern
<collectingCountry id="F34054D5FFF3C870FE68BC9EFDF4F905" box="[498,549,1733,1759]" name="Italy" pageId="13" pageNumber="368">Italy</collectingCountry>
(“unicum exemplar collectionis meae ... in Alpibus Insubriae captum et olim a Clar. De Cristofari Mediolanensi missum”). The name was tentatively synonymised with
<taxonomicName id="4C576FC6FFF3C870FB17BCB1FEC9F8FD" authority="(Rondani, 1870)" baseAuthorityName="Rondani" baseAuthorityYear="1870" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="368" phylum="Arthropoda" rank="species" species="alpicola">
<emphasis id="B923C857FFF3C870FB17BCB1FA82F8D9" box="[1165,1363,1769,1795]" italics="true" pageId="13" pageNumber="368">Drymeia alpicola</emphasis>
(Rondani, 1870)
</taxonomicName>
(as
<taxonomicName id="4C576FC6FFF3C870FED2BD55FE07F8FD" authorityName="Rondani" authorityYear="1871" box="[328,470,1806,1831]" class="Insecta" family="Muscidae" genus="Pogonomyia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="368" phylum="Arthropoda" rank="genus">
<emphasis id="B923C857FFF3C870FED2BD55FE07F8FD" box="[328,470,1806,1831]" italics="true" pageId="13" pageNumber="368">Pogonomyia</emphasis>
</taxonomicName>
) by Stein (in
<bibRefCitation id="EFC669B4FFF3C870FDE9BD56FC51F8F2" author="Becker, T. &amp; Bezzi, M. &amp; Kertesz, K. &amp; Stein, P." box="[627,896,1805,1832]" pageId="13" pageNumber="368" pagination="1 - 828" refId="ref13242" refString="Becker, T., Bezzi, M., Kertesz, K. &amp; Stein, P. (Eds.) (1907) Cyclorrapha Aschiza. Cyclorrapha Schizophora: Schizometopa. Katalog der palaarktischen Dipteren, Budapest, 3, 1 - 828." type="journal article" year="1907">
Becker
<emphasis id="B923C857FFF3C870FD50BD55FCD2F8FD" box="[714,771,1805,1831]" italics="true" pageId="13" pageNumber="368">et al.</emphasis>
1907: 631
</bibRefCitation>
). In the course of his revision of the Palaearctic
<taxonomicName id="4C576FC6FFF3C870FF0DBD6AFE69F896" authority=", Hennig (1961)" authorityName="Hennig" authorityYear="1961" box="[151,440,1841,1868]" class="Insecta" family="Muscidae" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="368" phylum="Arthropoda" rank="family">
Muscidae,
<bibRefCitation id="EFC669B4FFF3C870FE89BD6AFE69F896" author="Hennig, W." box="[275,440,1841,1868]" pageId="13" pageNumber="368" pagination="225 - 229" refId="ref13439" refString="Hennig, W. (1961) Bericht uber die Untersuchung der Typen einiger der von Rondani beschriebenen Arten aus der Familie Muscidae (Diptera). Beitrage zur Entomologie, 11, 225 - 229." type="journal article" year="1961">Hennig (1961)</bibRefCitation>
</taxonomicName>
studied Rondanis collection but did not find the
<typeStatus id="54ECAAE7FFF3C870FC76BD6AFB83F891" box="[1004,1106,1841,1867]" pageId="13" pageNumber="368" type="holotype">holotype</typeStatus>
. Later (
<bibRefCitation id="EFC669B4FFF3C870FB32BD6AFA84F896" author="Hennig, W." box="[1192,1365,1841,1868]" pageId="13" pageNumber="368" pagination="673 - 720" refId="ref13523" refString="Hennig, W. (1962 b) Muscidae. [Part, Lieferung 227.] In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region. 63 b. Schweizerbart, Stuttgart, pp. 673 - 720." type="book chapter" year="1962">Hennig, 1962b</bibRefCitation>
), after correspondence with several Italian Museums, he reported that the
<typeStatus id="54ECAAE7FFF3C870FC18BD0EFC35F8B5" box="[898,996,1877,1903]" pageId="13" pageNumber="368" type="holotype">holotype</typeStatus>
must be presumed lost and
<taxonomicName id="4C576FC6FFF3C870FA80BD0DFA4DF8B5" authorityName="Rondani" authorityYear="1866" box="[1306,1436,1877,1903]" class="Magnoliopsida" family="Asteraceae" genus="Aspilia" kingdom="Plantae" order="Asterales" pageId="13" pageNumber="368" phylum="Tracheophyta" rank="species" species="glacialis">
<emphasis id="B923C857FFF3C870FA80BD0DFA4DF8B5" box="[1306,1436,1877,1903]" italics="true" pageId="13" pageNumber="368">D. glacialis</emphasis>
</taxonomicName>
remained as an “unrecognised species” although the possibility of a synonymy with
<taxonomicName id="4C576FC6FFF3C870FBA4BD21FB6AF849" baseAuthorityName="Rondani" baseAuthorityYear="1870" box="[1086,1211,1913,1939]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="368" phylum="Arthropoda" rank="species" species="alpicola">
<emphasis id="B923C857FFF3C870FBA4BD21FB6AF849" box="[1086,1211,1913,1939]" italics="true" pageId="13" pageNumber="368">D. alpicola</emphasis>
</taxonomicName>
was left open (
<bibRefCitation id="EFC669B4FFF3C870FAF9BD22FE95F86D" author="Hennig, W." pageId="13" pageNumber="368" pagination="673 - 720" refId="ref13523" refString="Hennig, W. (1962 b) Muscidae. [Part, Lieferung 227.] In: Lindner, E. (Ed.), Die Fliegen der palaearktischen Region. 63 b. Schweizerbart, Stuttgart, pp. 673 - 720." type="book chapter" year="1962">Hennig 1962b: 677</bibRefCitation>
;
<bibRefCitation id="EFC669B4FFF3C870FECABDC5FE20F86D" author="Pont, A. C." box="[336,497,1949,1976]" pageId="13" pageNumber="368" refId="ref13885" refString="Pont, A. C. (1986) Family Muscidae. In: Soos, A. &amp; Papp, L. (Eds.), Catalogue of Palaearctic Diptera. Vol. 11. Scathophagidae - Hypodermatidae. Akademiai Kiado, Budapest, pp. 57 - 215." type="book" year="1986">Pont 1986: 73</bibRefCitation>
).
</paragraph>
<paragraph id="8BE81445FFF3C873FF5DBD9AFC26FE77" blockId="13.[151,1437,1373,2012]" lastBlockId="14.[151,1437,151,430]" lastPageId="14" lastPageNumber="369" pageId="13" pageNumber="368">
The
<typeStatus id="54ECAAE7FFF3C870FF66BD9AFE8DF801" box="[252,348,1985,2011]" pageId="13" pageNumber="368" type="holotype">holotype</typeStatus>
has recently been located in MZUF (
<bibRefCitation id="EFC669B4FFF3C870FC93BD9AFBAAF801" author="Sforzi, A. &amp; Sommaggio, D." box="[777,1147,1985,2011]" pageId="13" pageNumber="368" pagination="1 - 438" refId="ref14199" refString="Sforzi, A. &amp; Sommaggio, D. (2021) Catalog of the Diptera types described by Camillo Rondani. Zootaxa, 4989 (1), 1 - 438. https: // doi. org / 10.11646 / zootaxa. 4989.1.1" type="journal article" year="2021">Sforzi &amp; Sommaggio 2021: 140</bibRefCitation>
). It is a female, with two labels: (1) A white oval label, with “1162” printed in red ink; (2) A red label with the printed information “Museu “La Specola” / coll. Rondani /
<typeStatus id="54ECAAE7FFF0C873FE70BAE0FD44FF0F" box="[490,661,187,213]" pageId="14" pageNumber="369" type="holotype">HOLOTYPUS</typeStatus>
”. It is in fair condition (
<figureCitation id="136C08C0FFF0C873FC03BAE0FC3DFF0F" box="[921,1004,187,213]" captionStart="FIGURE 19" captionStartId="14.[151,250,1741,1765]" captionTargetBox="[151,1435,453,1717]" captionTargetId="figure-190@14.[151,1435,453,1717]" captionTargetPageId="14" captionText="FIGURE 19. Drymeia glacialis (Rondani), ♀ holotype, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6538217" httpUri="https://zenodo.org/record/6538217/files/figure.png" pageId="14" pageNumber="369">Fig. 19</figureCitation>
): slightly mouldy and dusty; both postpedicels and right hind leg missing; ptilinum partly extruded; right wing with part of costa missing; beetle damage to left side of syntergite 1+2; many setae on scutum rubbed off. Nevertheless, it can be recognised as
<taxonomicName id="4C576FC6FFF0C873FB40BB58FE9CFE9B" authority="(Rondani, 1871)" baseAuthorityName="Rondani" baseAuthorityYear="1871" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="369" phylum="Arthropoda" rank="species" species="alpicola">
<emphasis id="B923C857FFF0C873FB40BB58FA4DFEC7" box="[1242,1436,259,285]" italics="true" pageId="14" pageNumber="369">Drymeia alpicola</emphasis>
(
<bibRefCitation id="EFC669B4FFF0C873FF3ABB7CFE94FE9B" author="Rondani, C." box="[160,325,295,321]" pageId="14" pageNumber="369" pagination="317 - 338" refId="ref14065" refString="Rondani, C. (1871) Diptera Italica non vel minus cognita descripta aut annotata. Fasc. IV. Addenda Anthomyinis Prodr. Vol. VI. Bullettino della Societa entomologica italiana, 2, 317 - 338. [1870]" type="journal article" year="1871">Rondani, 1871</bibRefCitation>
)
</taxonomicName>
on account of the very enlarged parafacial, elongate proboscis, presence of a small snout between the antennal bases, and dense fine ventral setulae on fore tarsomeres 14, and, being an earlier name, must replace
<taxonomicName id="4C576FC6FFF0C873FF0DBB34FEC2FE53" baseAuthorityName="Rondani" baseAuthorityYear="1871" box="[151,275,367,393]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="369" phylum="Arthropoda" rank="species" species="alpicola">
<emphasis id="B923C857FFF0C873FF0DBB34FEC2FE53" box="[151,275,367,393]" italics="true" pageId="14" pageNumber="369">D. alpicola</emphasis>
</taxonomicName>
. This information was passed to Drs Jade Savage (Sherbrooke,
<collectingCountry id="F34054D5FFF0C873FC4BBB34FBF4FE50" box="[977,1061,367,394]" name="Canada" pageId="14" pageNumber="369">Canada</collectingCountry>
) and Vera Sorokina (
<collectingRegion id="4993DAA7FFF0C873FA97BB34FA49FE53" box="[1293,1432,367,393]" country="Russia" name="Novosibirsk" pageId="14" pageNumber="369">Novosibirsk</collectingRegion>
,
<collectingCountry id="F34054D5FFF0C873FF0DBBC8FF33FE74" box="[151,226,403,430]" name="Russia" pageId="14" pageNumber="369">Russia</collectingCountry>
) who formally published the synonymy (
<bibRefCitation id="EFC669B4FFF0C873FD3DBBC8FC3BFE77" author="Savage, J. &amp; Sorokina, V. S." box="[679,1002,402,429]" pageId="14" pageNumber="369" pagination="31 - 89" refId="ref14109" refString="Savage, J. &amp; Sorokina, V. S. (2021) Review of the North American fauna of Drymeia Meigen (Diptera, Muscidae) and evaluation of DNA barcodes for species-level identification in the genus. ZooKeys, 1024, 31 - 89. https: // doi. org / 10.3897 / zookeys. 1024.60393" type="journal article" year="2021">Savage &amp; Sorokina 2021: 51</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>