147 lines
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147 lines
23 KiB
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<document ID-DOI="10.7717/peerj.4417" ID-GBIF-Dataset="58039a1f-b9d0-4c93-b627-410c6decc440" ID-PMC="PMC5822849" ID-PubMed="29479504" ID-Zenodo-Dep="3097200" checkinTime="1519205475471" checkinUser="plazi" docAuthor="Jonathan P. Tennant, Alfio Alessandro Chiarenza & Matthew Baron" docDate="2018" docId="03FE9007FFB38307FE21FA1A7FB4D690" docLanguage="en" docName="peerj-4417.pdf" docOrigin="PeerJ 4417" docStyle="DocumentStyle{}" docTitle="Ornithischia" docType="treatment" docVersion="10" lastPageNumber="14" masterDocId="FFC7E87FFFBA830AFFD3FFB17D18D113" masterDocTitle="How has our knowledge of dinosaur diversity through geologic time changed through research history?" masterLastPageNumber="42" masterPageNumber="1" pageNumber="10" updateTime="1668138782670" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>How has our knowledge of dinosaur diversity through geologic time changed through research history?</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Jonathan P. Tennant</mods:namePart>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Alfio Alessandro Chiarenza</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Matthew Baron</mods:namePart>
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<mods:title>PeerJ</mods:title>
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<mods:date>2018</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2018-02-19</mods:number>
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<mods:detail type="volume">
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<mods:number>4417</mods:number>
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<mods:identifier type="DOI">10.7717/peerj.4417</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5614763" ID-GBIF-Taxon="141118103" ID-Zenodo-Dep="5614763" LSID="urn:lsid:plazi:treatment:03FE9007FFB38307FE21FA1A7FB4D690" httpUri="http://treatment.plazi.org/id/03FE9007FFB38307FE21FA1A7FB4D690" lastPageId="13" lastPageNumber="14" pageId="9" pageNumber="10">
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<subSubSection box="[498,700,1451,1479]" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph blockId="9.[498,1542,1406,1920]" box="[498,700,1451,1479]" pageId="9" pageNumber="10">
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<heading bold="true" box="[498,700,1451,1479]" fontSize="11" level="4" pageId="9" pageNumber="10" reason="6">
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<taxonomicName box="[498,700,1451,1479]" class="Reptilia" kingdom="Animalia" order="Ornithischia" pageId="9" pageNumber="10" phylum="Chordata" rank="order">
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<emphasis bold="true" box="[498,700,1451,1479]" italics="true" pageId="9" pageNumber="10">Ornithischians</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="13" lastPageNumber="14" pageId="9" pageNumber="10" type="discussion">
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<paragraph blockId="9.[498,1542,1406,1920]" pageId="9" pageNumber="10">
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Raw ‘global’ ornithischian diversity (
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<figureCitation box="[936,1021,1495,1522]" captionStart="Figure 5" captionStartId="11.[116,180,1302,1324]" captionTargetBox="[91,1471,235,1273]" captionTargetId="figure@11.[90,1472,234,1274]" captionTargetPageId="11" captionText="Figure 5 RaW Ornithischian diVersity at (A) glObal and (B–F) regiOnal leVels (EurOpe, Africa, Asia, NOrth America, and SOuth America, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-5" httpUri="https://zenodo.org/record/3097224/files/figure.png" pageId="9" pageNumber="10">Fig. 5A</figureCitation>
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) is constant and stable throughout publication history. The apparent magnitude of longer-term trends is obscured by the relative over-sampling of the Campanian and Maastrichtian, which are almost an order of magnitude higher than any other Jurassic or Cretaceous stage interval. Indeed, the Campanian shows no sign of slowing down in increasing diversity, and is the highest and most rapidly increasing of any time interval. In spite of this, the overall trends in raw diversity remain, with steadily increasing Middle to Late Jurassic diversity, a small earliest Cretaceous decline followed by a ‘middle’ Cretaceous peak in the Aptian, a shallow decline into the early Late Cretaceous, and an increase in the Campanian.
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</paragraph>
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<paragraph blockId="9.[498,1542,1406,1920]" lastBlockId="10.[498,1541,1636,1902]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
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Raw diversity in Europe shows increasing diversity across the J/K transition before an earliest Cretaceous decline (Valanginian–Hauterivian), constant ‘middle’ Cretaceous diversity, and an increase from the Campanian to Maastrichtian (
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<figureCitation box="[1278,1361,1636,1663]" captionStart="Figure 5" captionStartId="11.[116,180,1302,1324]" captionTargetBox="[91,1471,235,1273]" captionTargetId="figure@11.[90,1472,234,1274]" captionTargetPageId="11" captionText="Figure 5 RaW Ornithischian diVersity at (A) glObal and (B–F) regiOnal leVels (EurOpe, Africa, Asia, NOrth America, and SOuth America, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-5" httpUri="https://zenodo.org/record/3097224/files/figure.png" pageId="10" pageNumber="11">Fig. 5B</figureCitation>
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). Raw African ornithischian diversity is too inconsistent to analyse any changes through geological time or publication time (
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<figureCitation box="[811,897,1715,1742]" captionStart="Figure 5" captionStartId="11.[116,180,1302,1324]" captionTargetBox="[91,1471,235,1273]" captionTargetId="figure@11.[90,1472,234,1274]" captionTargetPageId="11" captionText="Figure 5 RaW Ornithischian diVersity at (A) glObal and (B–F) regiOnal leVels (EurOpe, Africa, Asia, NOrth America, and SOuth America, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-5" httpUri="https://zenodo.org/record/3097224/files/figure.png" pageId="10" pageNumber="11">Fig. 5C</figureCitation>
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). Raw Asian diversity is fairly constant through the Cretaceous, until an apparent major Campanian peak and Maastrichtian decline (
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<figureCitation box="[509,597,1795,1822]" captionStart="Figure 5" captionStartId="11.[116,180,1302,1324]" captionTargetBox="[91,1471,235,1273]" captionTargetId="figure@11.[90,1472,234,1274]" captionTargetPageId="11" captionText="Figure 5 RaW Ornithischian diVersity at (A) glObal and (B–F) regiOnal leVels (EurOpe, Africa, Asia, NOrth America, and SOuth America, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-5" httpUri="https://zenodo.org/record/3097224/files/figure.png" pageId="10" pageNumber="11">Fig. 5D</figureCitation>
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). In North America, empirical diversity is flat and low throughout the Late Jurassic and most of the Cretaceous (
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<figureCitation box="[919,997,1835,1862]" captionStart="Figure 5" captionStartId="11.[116,180,1302,1324]" captionTargetBox="[91,1471,235,1273]" captionTargetId="figure@11.[90,1472,234,1274]" captionTargetPageId="11" captionText="Figure 5 RaW Ornithischian diVersity at (A) glObal and (B–F) regiOnal leVels (EurOpe, Africa, Asia, NOrth America, and SOuth America, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-5" httpUri="https://zenodo.org/record/3097224/files/figure.png" pageId="10" pageNumber="11">Fig. 5E</figureCitation>
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). There is a Campanian peak, and order of magnitude higher than any prior interval, which is rapidly increasing through publication time.
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</paragraph>
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<caption httpUri="https://zenodo.org/record/3097220/files/figure.png" pageId="10" pageNumber="11" startId="10.[524,588,1354,1376]" targetBox="[500,1536,236,1323]" targetPageId="10">
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<paragraph blockId="10.[524,1516,1354,1580]" pageId="10" pageNumber="11">
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<emphasis bold="true" box="[524,1396,1354,1376]" pageId="10" pageNumber="11">Figure 4 Total dinosaur ‘global’ diversity patterns for (A) raw and (B) subsampled data.</emphasis>
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The vertical red lines represent major interval boundaries. Time stage abbreviations (in chronological order). N, Norian; R, Rhaetian, He, Hettangian; S, Sinemurian; P, Pliensbachian; T, Toarcian; A, Aalenian; Bj, Bajocian; B, Bathonian; C, Callovian; O, Oxfordian; K, Kimmeridgian; Ti, Tithonian; Be, Berriasian; V, Valanginian; Ha, Hauterivian; Ba, Barremian; Ap, Aptian; Al, Albian; Ce, Cenomanian; Tu, Turonian; Co, Coniacian; Sa, Santonian; Cam, Campanian; M, Maastrichtian. Vertical dashed red lines indicate boundaries between different periods (Triassic/Jurassic, Jurassic/Cretaceous, and Cretaceous/Paleogene). Full-size DOI: 10.7717/peerj.4417/fig-4
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</paragraph>
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</caption>
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<caption httpUri="https://zenodo.org/record/3097224/files/figure.png" pageId="11" pageNumber="12" startId="11.[116,180,1302,1324]" targetBox="[91,1471,235,1273]" targetPageId="11">
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<paragraph blockId="11.[116,1515,1302,1353]" pageId="11" pageNumber="12">
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<emphasis bold="true" pageId="11" pageNumber="12">Figure 5 Raw ornithischian diversity at (A) global and (B–F) regional levels (Europe, Africa, Asia, North America, and South America, respectively) based on our published knowledge in 1991 and 2015.</emphasis>
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Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-5
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</paragraph>
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</caption>
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<paragraph blockId="11.[498,1542,1414,1919]" lastBlockId="12.[498,1542,1416,1921]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
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Diversity decreases from this into the Maastrichtian, in which diversity has remained relatively stable through publication time. Sampling in South America is also relatively poor, with apparent diversity remaining low and flat where a signal is obtained (
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<figureCitation box="[1444,1523,1494,1521]" captionStart="Figure 5" captionStartId="11.[116,180,1302,1324]" captionTargetBox="[91,1471,235,1273]" captionTargetId="figure@11.[90,1472,234,1274]" captionTargetPageId="11" captionText="Figure 5 RaW Ornithischian diVersity at (A) glObal and (B–F) regiOnal leVels (EurOpe, Africa, Asia, NOrth America, and SOuth America, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-5" httpUri="https://zenodo.org/record/3097224/files/figure.png" pageId="11" pageNumber="12">Fig. 5F</figureCitation>
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). Subsampled ‘global’ ornithischian diversity shows a distinctly different pattern from the raw curve, both in terms of overall trends, and in terms of the magnitude of the effect of publication history (
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<figureCitation box="[776,861,1613,1640]" captionStart="Figure6" captionStartId="12.[116,180,1302,1324]" captionTargetBox="[91,1502,235,1273]" captionTargetId="figure@12.[90,1503,234,1274]" captionTargetPageId="12" captionText="Figure6 SubsampledOrnithischiandiVersityat(A)glObal and(B–F)regiOnalleVels (EurOpe,Africa, Asia,NOrth America,and SOuthAmerica, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-6" httpUri="https://zenodo.org/record/3097226/files/figure.png" pageId="11" pageNumber="12">Fig. 6A</figureCitation>
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). The Jurassic is generally too poorly sampled to reveal a constant signal, but there is evidence of a decline through the J/K transition, which remains constant through publication time. This is followed by a middle-Cretaceous increase, in which ornithischian diversity is at its second highest level throughout their history. The magnitude of this Albian radiation has rapidly increased over publication time, the result being that originally what appeared to be increasing subsampled diversity over the Early/Late Cretaceous transition now shows a major decline from the Albian to Coniacian. Santonian subsampled diversity remains unknown, but when we see a signal emerge in the Campanian, diversity is higher than the Albian, reaching its highest level before declining by more than half into the Maastrichtian. This overall structure, besides the Albian, remains consistent throughout publication time with no major perturbations to the apparent ‘global’ curve.
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</paragraph>
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<caption httpUri="https://zenodo.org/record/3097226/files/figure.png" pageId="12" pageNumber="13" startId="12.[116,180,1302,1324]" targetBox="[91,1502,235,1273]" targetPageId="12">
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<paragraph blockId="12.[116,1515,1302,1353]" pageId="12" pageNumber="13">
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<emphasis bold="true" pageId="12" pageNumber="13">Figure 6 Subsampled ornithischian diversity at (A) global and (B–F) regional levels (Europe, Africa, Asia, North America, and South America, respectively) based on our published knowledge in 1991 and 2015.</emphasis>
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Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-6
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</paragraph>
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</caption>
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<paragraph blockId="12.[498,1542,1416,1921]" pageId="12" pageNumber="13">
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Subsampled European diversity reveals increasing diversity across the Tithonian/ Berriasian transition, followed by overall gradually decreasing diversity throughout the remainder of the Early Cretaceous (
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<figureCitation box="[915,996,1655,1682]" captionStart="Figure6" captionStartId="12.[116,180,1302,1324]" captionTargetBox="[91,1502,235,1273]" captionTargetId="figure@12.[90,1503,234,1274]" captionTargetPageId="12" captionText="Figure6 SubsampledOrnithischiandiVersityat(A)glObal and(B–F)regiOnalleVels (EurOpe,Africa, Asia,NOrth America,and SOuthAmerica, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-6" httpUri="https://zenodo.org/record/3097226/files/figure.png" pageId="12" pageNumber="13">Fig. 6B</figureCitation>
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). In Africa, the signal is too poor to reveal anything besides a Kimmeridgian/Tithonian subsampled diversity drop (
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<figureCitation box="[1344,1428,1695,1722]" captionStart="Figure6" captionStartId="12.[116,180,1302,1324]" captionTargetBox="[91,1502,235,1273]" captionTargetId="figure@12.[90,1503,234,1274]" captionTargetPageId="12" captionText="Figure6 SubsampledOrnithischiandiVersityat(A)glObal and(B–F)regiOnalleVels (EurOpe,Africa, Asia,NOrth America,and SOuthAmerica, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-6" httpUri="https://zenodo.org/record/3097226/files/figure.png" pageId="12" pageNumber="13">Fig. 6C</figureCitation>
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), and in Asia, there is evidence of a decline in subsampled diversity across the Albian/Cenomanian transition (
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<figureCitation box="[630,715,1774,1801]" captionStart="Figure6" captionStartId="12.[116,180,1302,1324]" captionTargetBox="[91,1502,235,1273]" captionTargetId="figure@12.[90,1503,234,1274]" captionTargetPageId="12" captionText="Figure6 SubsampledOrnithischiandiVersityat(A)glObal and(B–F)regiOnalleVels (EurOpe,Africa, Asia,NOrth America,and SOuthAmerica, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-6" httpUri="https://zenodo.org/record/3097226/files/figure.png" pageId="12" pageNumber="13">Fig. 6D</figureCitation>
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). In North America, subsampled diversity reveals a decline across the Early–Late Cretaceous transition, and a major decline from the Campanian to Maastrichtian, a pattern that remains stable through publication history (
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<figureCitation box="[1338,1417,1854,1881]" captionStart="Figure6" captionStartId="12.[116,180,1302,1324]" captionTargetBox="[91,1502,235,1273]" captionTargetId="figure@12.[90,1503,234,1274]" captionTargetPageId="12" captionText="Figure6 SubsampledOrnithischiandiVersityat(A)glObal and(B–F)regiOnalleVels (EurOpe,Africa, Asia,NOrth America,and SOuthAmerica, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-6" httpUri="https://zenodo.org/record/3097226/files/figure.png" pageId="12" pageNumber="13">Fig. 6E</figureCitation>
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). In South America, the subsampled signal is too poor to comment on ornithischian diversity (
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<figureCitation box="[1447,1524,1894,1921]" captionStart="Figure6" captionStartId="12.[116,180,1302,1324]" captionTargetBox="[91,1502,235,1273]" captionTargetId="figure@12.[90,1503,234,1274]" captionTargetPageId="12" captionText="Figure6 SubsampledOrnithischiandiVersityat(A)glObal and(B–F)regiOnalleVels (EurOpe,Africa, Asia,NOrth America,and SOuthAmerica, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-6" httpUri="https://zenodo.org/record/3097226/files/figure.png" pageId="12" pageNumber="13">Fig. 6F</figureCitation>
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).
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</paragraph>
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<caption httpUri="https://zenodo.org/record/3097230/files/figure.png" pageId="13" pageNumber="14" startId="13.[116,180,1302,1324]" targetBox="[91,1487,235,1272]" targetPageId="13">
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<paragraph blockId="13.[116,1515,1302,1383]" pageId="13" pageNumber="14">
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<caption httpUri="https://zenodo.org/record/3097238/files/figure.png" pageId="13" pageNumber="14" startId="13.[116,180,1302,1324]" targetBox="[91,1487,235,1272]" targetPageId="13">
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<emphasis bold="true" pageId="13" pageNumber="14">
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Figure 7 Good’s
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<emphasis bold="true" box="[290,303,1302,1324]" italics="true" pageId="13" pageNumber="14">u</emphasis>
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estimates for ornithischians at (A) global and (B–F) regional levels (Europe, Africa, Asia, North America, and South America, respectively) based on our published knowledge in 1991 and 2015.
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</emphasis>
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Abbreviations as Fig. 4.
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</caption>
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Full-size DOI: 10.7717/peerj.4417/fig-7
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</paragraph>
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</caption>
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<paragraph blockId="13.[498,1541,1457,1923]" pageId="13" pageNumber="14">
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If we look at how coverage has changed through publication history (based on Good’s
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<emphasis box="[589,604,1498,1523]" italics="true" pageId="13" pageNumber="14">u</emphasis>
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), we should expect that subsampled diversity patterns are reflective of this pattern. At a global level, coverage in the Cretaceous is much better than the Jurassic (
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<figureCitation box="[509,593,1577,1604]" captionStart="Figure 7" captionStartId="13.[116,180,1302,1324]" captionTargetBox="[91,1487,235,1272]" captionTargetId="figure@13.[90,1488,234,1274]" captionTargetPageId="13" captionText="Figure 7 GOOd’s u estimates fOr Ornithischians at (A) glObal and (B–F) regiOnal leVels (EurOpe, Africa, Asia, NOrth America, and SOuth America, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-7" httpUri="https://zenodo.org/record/3097238/files/figure.png" pageId="13" pageNumber="14">Fig. 7A</figureCitation>
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). Much of this, however, is based on patchy regional records. In Europe, we find that coverage increases across the J/K interval (
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<figureCitation box="[1057,1141,1617,1644]" captionStart="Figure 7" captionStartId="13.[116,180,1302,1324]" captionTargetBox="[91,1487,235,1272]" captionTargetId="figure@13.[90,1488,234,1274]" captionTargetPageId="13" captionText="Figure 7 GOOd’s u estimates fOr Ornithischians at (A) glObal and (B–F) regiOnal leVels (EurOpe, Africa, Asia, NOrth America, and SOuth America, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-7" httpUri="https://zenodo.org/record/3097238/files/figure.png" pageId="13" pageNumber="14">Fig. 7B</figureCitation>
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), and is the only place where a consistently reliable record here can be obtained. In Africa, coverage is generally poor, besides in the latest Jurassic (
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<figureCitation box="[846,932,1696,1723]" captionStart="Figure 7" captionStartId="13.[116,180,1302,1324]" captionTargetBox="[91,1487,235,1272]" captionTargetId="figure@13.[90,1488,234,1274]" captionTargetPageId="13" captionText="Figure 7 GOOd’s u estimates fOr Ornithischians at (A) glObal and (B–F) regiOnal leVels (EurOpe, Africa, Asia, NOrth America, and SOuth America, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-7" httpUri="https://zenodo.org/record/3097238/files/figure.png" pageId="13" pageNumber="14">Fig. 7C</figureCitation>
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). In Asia, coverage is poor up until the late Early Cretaceous (
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<figureCitation box="[647,734,1736,1763]" captionStart="Figure 7" captionStartId="13.[116,180,1302,1324]" captionTargetBox="[91,1487,235,1272]" captionTargetId="figure@13.[90,1488,234,1274]" captionTargetPageId="13" captionText="Figure 7 GOOd’s u estimates fOr Ornithischians at (A) glObal and (B–F) regiOnal leVels (EurOpe, Africa, Asia, NOrth America, and SOuth America, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-7" httpUri="https://zenodo.org/record/3097238/files/figure.png" pageId="13" pageNumber="14">Fig. 7D</figureCitation>
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). In North America, coverage is good in the latest Jurassic and ‘middle’ to Late Cretaceous, but non-existent in Early to Middle Jurassic and earliest Cretaceous (
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<figureCitation box="[647,729,1816,1843]" captionStart="Figure 7" captionStartId="13.[116,180,1302,1324]" captionTargetBox="[91,1487,235,1272]" captionTargetId="figure@13.[90,1488,234,1274]" captionTargetPageId="13" captionText="Figure 7 GOOd’s u estimates fOr Ornithischians at (A) glObal and (B–F) regiOnal leVels (EurOpe, Africa, Asia, NOrth America, and SOuth America, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-7" httpUri="https://zenodo.org/record/3097238/files/figure.png" pageId="13" pageNumber="14">Fig. 7E</figureCitation>
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). Coverage is generally poor for the entire South American ornithischian record (
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<figureCitation box="[759,840,1856,1883]" captionStart="Figure 7" captionStartId="13.[116,180,1302,1324]" captionTargetBox="[91,1487,235,1272]" captionTargetId="figure@13.[90,1488,234,1274]" captionTargetPageId="13" captionText="Figure 7 GOOd’s u estimates fOr Ornithischians at (A) glObal and (B–F) regiOnal leVels (EurOpe, Africa, Asia, NOrth America, and SOuth America, respectiVely) based On Our published knOWledge in 1991 and 2015. Abbreviations as Fig. 4. Full-size DOI: 10.7717/peerj.4417/fig-7" httpUri="https://zenodo.org/record/3097238/files/figure.png" pageId="13" pageNumber="14">Fig. 7F</figureCitation>
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), explaining why obtaining a subsampled diversity signal here is difficult.
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||
</paragraph>
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||
</subSubSection>
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||
</treatment>
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</document> |