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<document id="7735D95376811CF1ABD0CA189CD98F8D" ID-ISSN="1175-5326" ID-ZooBank="7F7A7C35-7E0F-4264-A053-BF06095F3F08" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629810134680" checkinUser="felipe" docAuthor="Wells, A. &amp; Johanson, K. A." docDate="2012" docId="03FD87B2FFEFFFF2FCFA8D75FB78FCD3" docLanguage="en" docName="zt03478p344.pdf" docOrigin="Zootaxa 3478" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Paroxyethira asymmetrica Wells &amp; Johanson 2012, new species" docType="treatment" docVersion="3" lastPageNumber="335" masterDocId="FFC4FFCAFFEBFFF7FC6D8A25FF90FFA9" masterDocTitle="Review of the New Caledonian species of Paroxyethira Mosely, 1924 (Trichoptera: Hydroptilidae)" masterLastPageNumber="344" masterPageNumber="330" pageNumber="334" updateTime="1699115451624" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="1E7807D41477678B218907D42A247BA2">Review of the New Caledonian species of Paroxyethira Mosely, 1924 (Trichoptera: Hydroptilidae)</mods:title>
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<mods:name id="4407A4DCC1AB38F837FD3C47B207A509" type="personal">
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<mods:namePart id="E26B9626EB6F0A94049E1100937B24FD">Wells, A.</mods:namePart>
<mods:affiliation id="FA2B3F0726DEC7FE60B33C9663B9CFB3">Australian Biological Resources Study, PO Box 787, Canberra, ACT 2601 Australia. E-mail: alice. wells @ environment. gov. au Entomology Department, Swedish Museum of Natural History, Box 50007, SE- 104 05 Stockholm, Sweden.</mods:affiliation>
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<mods:namePart id="EE0E4F3A13078F72136BF66552BAE40A">Johanson, K. A.</mods:namePart>
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<mods:typeOfResource id="FA003E8E7B38B43274BAD5C5F11A83CE">text</mods:typeOfResource>
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<mods:part id="6826B7B64CE5C5CB86FE37AA18363BB9">
<mods:date id="6F42E7021E54C5AFB18A1332B6BBA6DB">2012</mods:date>
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<treatment id="03FD87B2FFEFFFF2FCFA8D75FB78FCD3" ID-DOI="http://doi.org/10.5281/zenodo.5278895" ID-Zenodo-Dep="5278895" LSID="urn:lsid:plazi:treatment:03FD87B2FFEFFFF2FCFA8D75FB78FCD3" httpUri="http://treatment.plazi.org/id/03FD87B2FFEFFFF2FCFA8D75FB78FCD3" lastPageId="5" lastPageNumber="335" pageId="4" pageNumber="334">
<subSubSection id="C34E652FFFEFFFF3FCFA8D75FDE3F8C3" box="[151,627,1872,1898]" pageId="4" pageNumber="334" type="nomenclature">
<paragraph id="8BEB36A4FFEFFFF3FCFA8D75FDE3F8C3" blockId="4.[151,627,1872,1931]" box="[151,627,1872,1898]" pageId="4" pageNumber="334">
<heading id="D0A381C8FFEFFFF3FCFA8D75FDE3F8C3" bold="true" box="[151,627,1872,1898]" fontSize="11" level="1" pageId="4" pageNumber="334" reason="1">
<emphasis id="B920EAB6FFEFFFF3FCFA8D75FDE3F8C3" bold="true" box="[151,627,1872,1898]" pageId="4" pageNumber="334">
<taxonomicName id="4C544D27FFEFFFF3FCFA8D75FE45F8C3" authority="Wells &amp; Johanson, 2012" authorityName="Wells &amp; Johanson" authorityYear="2012" box="[151,469,1872,1898]" class="Insecta" family="Hydroptilidae" genus="Paroxyethira" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="asymmetrica" status="sp. nov.">
<emphasis id="B920EAB6FFEFFFF3FCFA8D75FE45F8C3" bold="true" box="[151,469,1872,1898]" italics="true" pageId="4" pageNumber="334">Paroxyethira asymmetrica</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21357CDFFEFFFF3FD8E8D75FDE3F8C3" box="[483,627,1872,1898]" pageId="4" pageNumber="334" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E652FFFEFFFF2FCFA8D56FC2AFD43" lastPageId="5" lastPageNumber="335" pageId="4" pageNumber="334" type="description">
<paragraph id="8BEB36A4FFEFFFF3FCFA8D56FEF7F822" blockId="4.[151,627,1872,1931]" box="[151,359,1907,1931]" pageId="4" pageNumber="334">
<figureCitation id="136F2A21FFEFFFF3FCFA8D56FE86F822" box="[151,278,1907,1931]" captionStart="FIGURES 612" captionStartId="4.[151,264,1705,1728]" captionTargetBox="[173,1436,185,1700]" captionTargetId="figure-105@4.[151,1436,185,1700]" captionTargetPageId="4" captionText="FIGURES 612. 69: Paroxyethira koegi, new species, holotype, male. 6—genitalia, ventral; 7—genitalia, left lateral, ventral; 8—phallic sheath; 9—phallic apparatus, dorsal view. 1012: Paroxyethira asymmetrica, new species, holotype, male. 10 —genitalia, ventral; 11—genitalia, dorsal (left gonopod in part only); 12—genitalia, right lateral." pageId="4" pageNumber="334">Figs 1012</figureCitation>
,
<figureCitation id="136F2A21FFEFFFF3FD4F8D56FEF7F822" box="[290,359,1907,1931]" captionStart="FIGURES 3342" captionStartId="12.[151,264,1885,1908]" captionTargetBox="[151,1436,185,1864]" captionTargetId="figure-119@12.[151,1436,185,1864]" captionTargetPageId="12" captionText="FIGURES 3342. Paroxyethira species, female terminalia. 33—P. dumagnes Kelley, ventral; 34—P. asymmetrica, new species, ventral; 35—P. asymmetrica, new species, dorsal; 36—P. asymmetrica, new species, left lateral; 37—P. atypica, new species, ventral; 38—P.atypica, new species, dorsal view; 39—P. dzumac, new species, ventral; 40—P. serrata, new species, ventral; 41—P. serrata, new species, right lateral; 42—P. anomala, new species, ventral. In Figs 37-39 arrow indicating position and shape of process on tergite VIII; in Figs 40, 41 indicating lateral extension of tergum VIII." pageId="4" pageNumber="334">3436</figureCitation>
</paragraph>
<paragraph id="8BEB36A4FFEFFFF2FCFA8D9CFCE9FEBE" blockId="4.[151,1436,1976,2035]" lastBlockId="5.[151,1436,151,891]" lastPageId="5" lastPageNumber="335" pageId="4" pageNumber="334">
Most closely similar to
<taxonomicName id="4C544D27FFEFFFF3FDDC8D9CFDA1F879" authority="Wells &amp; Johanson, 2012" authorityName="Wells &amp; Johanson" authorityYear="2012" box="[433,561,1977,2000]" class="Insecta" family="Hydroptilidae" genus="Paroxyethira" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="opposita" status="sp. nov.">
<emphasis id="B920EAB6FFEFFFF3FDDC8D9CFDA1F879" box="[433,561,1977,2000]" italics="true" pageId="4" pageNumber="334">P. opposita</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A21357CDFFEFFFF3FE288D9CFD42F879" box="[581,722,1976,2000]" pageId="4" pageNumber="334" rank="species">new species</taxonomicNameLabel>
, with which it shares the feature of male gonopods strongly asymmetric; however, with the right gonopod shorter than the left, in contrast to
<taxonomicName id="4C544D27FFEFFFF3F8228DF9FB5EF85A" authorityName="Wells &amp; Johanson" authorityYear="2012" box="[1103,1230,2012,2035]" class="Insecta" family="Hydroptilidae" genus="Paroxyethira" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="334" phylum="Arthropoda" rank="species" species="opposita">
<emphasis id="B920EAB6FFEFFFF3F8228DF9FB5EF85A" box="[1103,1230,2012,2035]" italics="true" pageId="4" pageNumber="334">P. opposita</emphasis>
</taxonomicName>
in which the left gonopod is shorter than the right.
<taxonomicName id="4C544D27FFEEFFF2FE498ABDFCC0FF06" authorityName="Wells &amp; Johanson" authorityYear="2012" box="[548,848,152,176]" class="Insecta" family="Hydroptilidae" genus="Paroxyethira" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="asymmetrica">
<emphasis id="B920EAB6FFEEFFF2FE498ABDFCC0FF06" box="[548,848,152,176]" italics="true" pageId="5" pageNumber="335">Paroxyethira asymmetrica</emphasis>
</taxonomicName>
is distinguished from other species with markedly unequal gonopods by the very short phallic sheath. The female of
<taxonomicName id="4C544D27FFEEFFF2FF118A9EFBB7FF7B" authorityName="Wells &amp; Johanson" authorityYear="2012" box="[892,1063,187,210]" class="Insecta" family="Hydroptilidae" genus="Paroxyethira" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="asymmetrica">
<emphasis id="B920EAB6FFEEFFF2FF118A9EFC1AFF7B" box="[892,906,187,210]" italics="true" pageId="5" pageNumber="335">P</emphasis>
.
<emphasis id="B920EAB6FFEEFFF2FFF58A9EFBB7FF7B" box="[920,1063,187,210]" italics="true" pageId="5" pageNumber="335">asymmetrica</emphasis>
</taxonomicName>
has terminalia more symmetrical than other congeners, regularly conical in ventral view; the surface is noticeably pocked by setal bases; and in dorsal view there is a prominent pocket subapically on the right.
</paragraph>
<paragraph id="8BEB36A4FFEEFFF2FCFA8B07FB0DFE93" blockId="5.[151,1436,151,891]" box="[151,1181,290,314]" pageId="5" pageNumber="335">Male antennae each with 2122 flagellomeres; female antennae each with 19 flagellomeres.</paragraph>
<paragraph id="8BEB36A4FFEEFFF2FCAA8B63FC4CFEF7" blockId="5.[151,1436,151,891]" box="[199,988,326,350]" pageId="5" pageNumber="335">
Forewing length. Male
<quantity id="4CAC9B41FFEEFFF2FDA38B63FDC3FEF7" box="[462,595,326,350]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.4" metricValueMin="1.7" pageId="5" pageNumber="335" unit="mm" value="2.05" valueMax="2.4" valueMin="1.7">1.72.4 mm</quantity>
(n=5); female 1.62.0 mm (n=10).
</paragraph>
<paragraph id="8BEB36A4FFEEFFF2FCAA8B4FFCD4FDF3" blockId="5.[151,1436,151,891]" pageId="5" pageNumber="335">
Male genitalia (
<figureCitation id="136F2A21FFEEFFF2FD108B4FFD94FE2B" box="[381,516,362,386]" captionStart="FIGURES 612" captionStartId="4.[151,264,1705,1728]" captionTargetBox="[173,1436,185,1700]" captionTargetId="figure-105@4.[151,1436,185,1700]" captionTargetPageId="4" captionText="FIGURES 612. 69: Paroxyethira koegi, new species, holotype, male. 6—genitalia, ventral; 7—genitalia, left lateral, ventral; 8—phallic sheath; 9—phallic apparatus, dorsal view. 1012: Paroxyethira asymmetrica, new species, holotype, male. 10 —genitalia, ventral; 11—genitalia, dorsal (left gonopod in part only); 12—genitalia, right lateral." pageId="5" pageNumber="335">Figs 1012</figureCitation>
). Sternite VII bearing short sharp spur medially. Median process on abdominal sternite VIII stoutly spatulate; length about 3x width at middle. Abdominal segment IX with broadly rounded posterolateral lobes; dorsally with distal margin deeply concave. Gonopods unequal, left elongate, slender, ~2.5x length of right; both bearing mesal brushes of stout black setae. Phallic apparatus as for
<taxonomicName id="4C544D27FFEEFFF2F8078BF2FB66FE47" authorityName="Kelley. One" authorityYear="1989" box="[1130,1270,470,494]" class="Insecta" family="Hydroptilidae" genus="Paroxyethira" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="335" phylum="Arthropoda" rank="species" species="dumagnes">
<emphasis id="B920EAB6FFEEFFF2F8078BF2FB66FE47" box="[1130,1270,470,494]" italics="true" pageId="5" pageNumber="335">P. dumagnes</emphasis>
</taxonomicName>
(
<figureCitation id="136F2A21FFEEFFF2F9688BF3FAD8FE47" box="[1285,1352,470,494]" captionStart="FIGURES 15" captionStartId="2.[151,263,1950,1973]" captionTargetBox="[166,1436,800,1935]" captionTargetId="figure-965@2.[151,1436,788,1945]" captionTargetPageId="2" captionText="FIGURES 15. Paroxyethira dumagnes Kelley, male specimen from Ouenghi River, Boulouparis. 1—right wings, dorsal; 2—genitalia, ventral; 3—genitalia, dorsal (gonopods in part only); 4—genitalia, left lateral; 5—phallic apparatus, dorsal view. Numbers in parentheses in Fig. 1 refer to fork numbers." pageId="5" pageNumber="335">Fig. 5</figureCitation>
), being slender and straight but with distal portion proportionally shorter, constricted at 1/4 length, at which constriction short, threadlike titillator arising. Phallic sheath short, broad; with seta at each apico-lateral angle; on left 2 setaebearing papillae and proximally anteriorly directed apodeme.
</paragraph>
<paragraph id="8BEB36A4FFEEFFF2FCAA8843FC2AFD43" blockId="5.[151,1436,151,891]" pageId="5" pageNumber="335">
Female (
<figureCitation id="136F2A21FFEEFFF2FD408843FE26FDD7" box="[301,438,614,638]" captionStart="FIGURES 3342" captionStartId="12.[151,264,1885,1908]" captionTargetBox="[151,1436,185,1864]" captionTargetId="figure-119@12.[151,1436,185,1864]" captionTargetPageId="12" captionText="FIGURES 3342. Paroxyethira species, female terminalia. 33—P. dumagnes Kelley, ventral; 34—P. asymmetrica, new species, ventral; 35—P. asymmetrica, new species, dorsal; 36—P. asymmetrica, new species, left lateral; 37—P. atypica, new species, ventral; 38—P.atypica, new species, dorsal view; 39—P. dzumac, new species, ventral; 40—P. serrata, new species, ventral; 41—P. serrata, new species, right lateral; 42—P. anomala, new species, ventral. In Figs 37-39 arrow indicating position and shape of process on tergite VIII; in Figs 40, 41 indicating lateral extension of tergum VIII." pageId="5" pageNumber="335">Figs 3436</figureCitation>
). Abdominal sternites VIIVIII fused, broad-based, constricted laterally at about 1/3 length, conical in distal 2/3, almost symmetrical. Tergites VIIVIII dorsally discrete; VIII with prominent pocket subapically on right; surface of VIIVIII disrupted by pronounced sockets of scattered fine setae; apex rounded ventrally, concave dorsally. Segment X divided distally to form 2 lobes.
</paragraph>
</subSubSection>
<subSubSection id="C34E652FFFEEFFF2FCAA88D2FC70FCFF" pageId="5" pageNumber="335" type="materials_examined">
<paragraph id="8BEB36A4FFEEFFF2FCAA88D2FB6CFC9B" blockId="5.[151,1436,151,891]" pageId="5" pageNumber="335">
<materialsCitation id="3B3C3CF9FFEEFFF2FCAA88D2FB6CFC9B" collectingDate="2003-11-29" collectionCode="MNHP" collectorName="over Dathio River &amp; K. A. Johanson" county="Xwe Pemou Stream" elevation="300" latitude="-21.58835" location="Ate" longLatPrecision="1" longitude="166.15117" municipality="Dathio River" pageId="5" pageNumber="335" specimenCount="1" stateProvince="Province Sud" typeStatus="holotype">
<emphasis id="B920EAB6FFEEFFF2FCAA88D2FEEEFCA6" bold="true" box="[199,382,759,783]" pageId="5" pageNumber="335">
<typeStatus id="54EF8806FFEEFFF2FCAA88D2FEA4FCA6" box="[199,308,759,783]" pageId="5" pageNumber="335" type="holotype">Holotype</typeStatus>
male:
</emphasis>
<collectingRegion id="4990F846FFEEFFF2FDEB88D3FD8FFCA7" box="[390,543,758,782]" country="New Caledonia" name="Province Sud" pageId="5" pageNumber="335">Province Sud</collectingRegion>
,
<collectingCounty id="628A4E28FFEEFFF2FE4688D3FC80FCA7" box="[555,784,758,782]" pageId="5" pageNumber="335">Xwé Pemöu Stream</collectingCounty>
,
<quantity id="4CAC9B41FFEEFFF2FF7188D3FCF3FCA7" box="[796,867,758,782]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="335" unit="m" value="300.0">
<elevation id="0079D197FFEEFFF2FF7188D3FCF3FCA7" box="[796,867,758,782]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="335" unit="m" value="300.0">300 m</elevation>
</quantity>
N bridge
<collectorName id="26A15372FFEEFFF2FFBB88D2FB31FCA7" box="[982,1185,758,782]" pageId="5" pageNumber="335">
over
<collectingMunicipality id="6B8FACDEFFEEFFF2F86288D3FB31FCA7" box="[1039,1185,758,782]" pageId="5" pageNumber="335">Dathio River</collectingMunicipality>
</collectorName>
at
<location id="8E8B607FFFEEFFF2F8A888D3FB60FCA7" LSID="urn:lsid:plazi:treatment:03FD87B2FFEFFFF2FCFA8D75FB78FCD3:8E8B607FFFEEFFF2F8A888D3FB60FCA7" box="[1221,1264,758,782]" county="Xwe Pemou Stream" latitude="-21.58835" longLatPrecision="1" longitude="166.15117" municipality="Dathio River" name="Ate" pageId="5" pageNumber="335" stateProvince="Province Sud">Atè</location>
,
<locationDeviation id="9A592F86FFEEFFF2F89088D3FF5EFC9B" location="Thio" pageId="5" pageNumber="335">
<quantity id="4CAC9B41FFEEFFF2F89088D3FADBFCA7" box="[1277,1355,758,782]" metricMagnitude="3" metricUnit="m" metricValue="6.2" pageId="5" pageNumber="335" unit="km" value="6.2">6.2 km</quantity>
WNW Thio
</locationDeviation>
,
<geoCoordinate id="EE605063FFEEFFF2FCB4893FFEF0FC9B" box="[217,352,794,818]" degrees="21.58835" direction="south" orientation="latitude" pageId="5" pageNumber="335" precision="1" value="-21.58835">21.58835°S</geoCoordinate>
,
<geoCoordinate id="EE605063FFEEFFF2FD06893FFD91FC9B" box="[363,513,794,818]" degrees="166.15117" direction="east" orientation="longitude" pageId="5" pageNumber="335" precision="1" value="166.15117">166.15117°E</geoCoordinate>
,
<quantity id="4CAC9B41FFEEFFF2FE66893FFDD6FC9B" box="[523,582,794,818]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="335" unit="m" value="13.0">
<elevation id="0079D197FFEEFFF2FE66893FFDD6FC9B" box="[523,582,794,818]" metricMagnitude="1" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="335" unit="m" value="13.0">13 m</elevation>
</quantity>
,
<date id="FFEA1064FFEEFFF2FE3F893FFD5CFC9B" box="[594,716,794,818]" pageId="5" pageNumber="335" value="2003-11-29">
<collectingDate id="EFAEE98CFFEEFFF2FE3F893FFD5CFC9B" box="[594,716,794,818]" pageId="5" pageNumber="335" value="2003-11-29">29.xi.2003</collectingDate>
</date>
,
<collectingMethod id="52154EB3FFEEFFF2FEB5893FFCD0FC9B" box="[728,832,794,818]" pageId="5" pageNumber="335">light trap</collectingMethod>
, loc#056, leg.
<collectorName id="26A15372FFEEFFF2FF89893FFB1AFC9B" box="[996,1162,794,818]" pageId="5" pageNumber="335">K.A. Johanson</collectorName>
(
<collectionCode id="ED45AE61FFEEFFF2F8F4893FFB60FC9B" box="[1177,1264,794,818]" country="USA" httpUri="http://grbio.org/cool/3kvn-7rck" name="Princeton University" pageId="5" pageNumber="335">MNHP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BEB36A4FFEEFFF2FCAA891AFC70FCFF" blockId="5.[151,1436,151,891]" box="[199,992,830,855]" pageId="5" pageNumber="335">
<materialsCitation id="3B3C3CF9FFEEFFF2FCAA891AFC4CFCFF" box="[199,988,830,855]" collectingDate="2003-11-29" collectionCode="SMNH" collectorName="over Dathio River &amp; K. A. Johanson" county="Xwe Pemou Stream" elevation="300" latitude="-21.58835" location="Ate" longLatPrecision="1" longitude="166.15117" municipality="Dathio River" pageId="5" pageNumber="335" specimenCount="27" specimenCount-female="22" specimenCount-male="5" stateProvince="Province Sud" typeStatus="paratype">
<emphasis id="B920EAB6FFEEFFF2FCAA891AFED8FCFE" bold="true" box="[199,328,831,855]" pageId="5" pageNumber="335">
<typeStatus id="54EF8806FFEEFFF2FCAA891AFED3FCFE" box="[199,323,831,855]" pageId="5" pageNumber="335" type="paratype">Paratypes</typeStatus>
:
</emphasis>
Same data as for holotype
<specimenCount id="9D52FD2DFFEEFFF2FEE3891BFD77FCFF" box="[654,743,830,854]" pageId="5" pageNumber="335" type="male">5 males</specimenCount>
,
<specimenCount id="9D52FD2DFFEEFFF2FE9F891BFCFCFCFF" box="[754,876,830,854]" pageId="5" pageNumber="335" type="female">22 females</specimenCount>
(
<collectionCode id="ED45AE61FFEEFFF2FF16891BFC44FCFF" box="[891,980,830,854]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="5" pageNumber="335">SMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E652FFFEEFFF2FCAA8946FB78FCD3" box="[199,1256,866,891]" pageId="5" pageNumber="335" type="etymology">
<paragraph id="8BEB36A4FFEEFFF2FCAA8946FB78FCD3" blockId="5.[151,1436,151,891]" box="[199,1256,866,891]" pageId="5" pageNumber="335">
<emphasis id="B920EAB6FFEEFFF2FCAA8946FEC1FCD2" bold="true" box="[199,337,867,891]" pageId="5" pageNumber="335">Etymology:</emphasis>
<emphasis id="B920EAB6FFEEFFF2FD358946FE7AFCD2" box="[344,490,867,891]" italics="true" pageId="5" pageNumber="335">Asymmetrica</emphasis>
, named for the strongly asymmetric gonopods of the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>