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<document id="654A6FC9BC093639ECA7C696C1E4ACF5" ID-DOI="10.11646/zootaxa.5447.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="11150249" ID-ZooBank="367106D8-6248-49DA-8ABE-3125FC46BE08" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1715181318044" checkinUser="plazi" docAuthor="Chen, Jia-Heng, Yin, Wen-Qi &amp; Shi, Hong-Liang" docDate="2024" docId="03FD87AC6549966CFF1FFC632CCF539E" docLanguage="en" docName="zootaxa.5447.4.1.pdf" docOrigin="Zootaxa 5447 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5360.3.8" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Wraseiellus) idiopterus Chen &amp; Yin &amp; Shi 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="463" masterDocId="FFC4FFD465439660FF88FFCD2D25501F" masterDocTitle="On the Pterostichus subgenus Wraseiellus from China: descriptions of five new species and supplementary notes on taxonomy (Coleoptera: Carabidae)" masterLastPageNumber="472" masterPageNumber="451" pageNumber="461" updateTime="1715181786403" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="234024F54DEA1453A634B6FB75CEF6C3" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8E02B6010E009C4419B5C1911EE045A3">
<mods:title id="8468DFE59656ED0EB0FDA9569F9691CE">On the Pterostichus subgenus Wraseiellus from China: descriptions of five new species and supplementary notes on taxonomy (Coleoptera: Carabidae)</mods:title>
</mods:titleInfo>
<mods:name id="9CFEA4290555103F723580F67079E2AF" type="personal">
<mods:role id="D77403CC79FA05ECE2237F3F9925A378">
<mods:roleTerm id="EBD8ED42632B3D7A0FE3B9F83522743B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="443E1731D16B4C26C65447D43C633482">Chen, Jia-Heng</mods:namePart>
<mods:affiliation id="E6A961B9F53C2C586CA1EE6EB661E2A1">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
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<mods:namePart id="23ECEEDD4D52E7934745EAEE5F0E8A6A">Yin, Wen-Qi</mods:namePart>
<mods:affiliation id="CC7E722ECD95BE0197804BA467FD95B9">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="49B10D8A55A8D83C993E5EEEF16B6648">Shi, Hong-Liang</mods:namePart>
<mods:affiliation id="4C4DD65FC94961BDEDA23F4EF46FF034">College of Forestry, Beijing Forestry University, Beijing, 100083, P. R. China</mods:affiliation>
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<mods:title id="AA1227E69E342B9A66C4D5DF01E8F51D">Zootaxa</mods:title>
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<mods:date id="FDFEBFF05398B163360E7F946EB51AE9">2024</mods:date>
<mods:detail id="E22671A04830EF251034783DC11FBEFB" type="pubDate">
<mods:number id="9CB42B4D344019C166B9F4DCE04C79CE">2024-05-08</mods:number>
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<mods:number id="72D630CDCEA833AD2B8E760B4DAAE8A3">5447</mods:number>
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<mods:number id="0E0CDFBD789BA05BE180AA4190ADF86C">4</mods:number>
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<mods:url id="D073E242DECEC18053F0819BBF129A9F">http://dx.doi.org/10.11646/zootaxa.5360.3.8</mods:url>
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<mods:classification id="F79995C79BF2C5EF4BD4BBA051CF9824">journal article</mods:classification>
<mods:identifier id="82239F97FACEE4F0E84433453A7F4CC6" type="DOI">10.11646/zootaxa.5447.4.1</mods:identifier>
<mods:identifier id="F712BB6DAAC4B5BAE6CE2C3D6D73AB3C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F72294EEAAEC60DA3B120DBA2DE7AE33" type="Zenodo-Dep">11150249</mods:identifier>
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</mods:mods>
<treatment id="03FD87AC6549966CFF1FFC632CCF539E" LSID="urn:lsid:plazi:treatment:03FD87AC6549966CFF1FFC632CCF539E" httpUri="http://treatment.plazi.org/id/03FD87AC6549966CFF1FFC632CCF539E" lastPageId="12" lastPageNumber="463" pageId="10" pageNumber="461">
<subSubSection id="C34E65316549966AFF1FFC632F4C53F5" pageId="10" pageNumber="461" type="nomenclature">
<paragraph id="8BEB36BA6549966AFF1FFC632F9D53D6" blockId="10.[151,696,942,1041]" box="[151,696,942,969]" pageId="10" pageNumber="461">
<heading id="D0A381D66549966AFF1FFC632F9D53D6" bold="true" box="[151,696,942,969]" fontSize="11" level="1" pageId="10" pageNumber="461" reason="1">
<emphasis id="B920EAA86549966AFF1FFC632F9D53D6" bold="true" box="[151,696,942,969]" pageId="10" pageNumber="461">
<taxonomicName id="4C544D396549966AFF1FFC632F7153D6" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[151,596,942,969]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="idiopterus" status="sp. nov." subGenus="Wraseiellus">
<emphasis id="B920EAA86549966AFF1FFC632C0853D6" bold="true" box="[151,301,942,969]" italics="true" pageId="10" pageNumber="461">Pterostichus</emphasis>
(
<emphasis id="B920EAA86549966AFEB6FC632CEF53D6" bold="true" box="[318,458,942,969]" italics="true" pageId="10" pageNumber="461">Wraseiellus</emphasis>
)
<emphasis id="B920EAA86549966AFE53FC632F7153D6" bold="true" box="[475,596,942,969]" italics="true" pageId="10" pageNumber="461">idiopterus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357D36549966AFDD3FC622F9D53D6" box="[603,696,943,969]" pageId="10" pageNumber="461" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEB36BA6549966AFF1FFC1E2F4C53F5" blockId="10.[151,696,942,1041]" box="[151,617,979,1005]" pageId="10" pageNumber="461">
Chinese vernacular name:
<emphasis id="B920EAA86549966AFE49FC1E2F4C53F5" bold="true" box="[449,617,979,1002]" pageId="10" pageNumber="461">Ƃaeaeữfflm</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C34E65316549966AFF1FFC3A2CEC540E" box="[151,457,1014,1041]" pageId="10" pageNumber="461" type="description">
<paragraph id="8BEB36BA6549966AFF1FFC3A2CEC540E" blockId="10.[151,696,942,1041]" box="[151,457,1014,1041]" pageId="10" pageNumber="461">
<figureCitation id="136F2A3F6549966AFF1FFC3A2C3B540E" box="[151,286,1014,1041]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="11.[151,250,1418,1442]" captionStartId-1="12.[151,250,1854,1878]" captionTargetBox-0="[159,1427,189,1386]" captionTargetBox-1="[160,1427,930,1822]" captionTargetId-0="figure-17@11.[151,1436,181,1394]" captionTargetId-1="figure-380@12.[151,1436,921,1831]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURE 5. Habitus of Pterostichus idiopterus sp. nov. (A) holotype; (B) Paratype, female. Scale bar = 5.0 mm." captionText-1="FIGURE 6. Male genitalia of Pterostichus idiopterus sp. nov., holotype. (A) median lobe of aedeagus, left lateral view; (B) median lobe of aedeagus, dorsal view; (C) right paramere; (D) endophallus, right lateral view; (E) endophallus, left lateral view. Scale bars = 1.0 mm. Abbreviations: pa pre-apical lobe; gp gonopore." figureDoi-0="http://doi.org/10.5281/zenodo.11150273" figureDoi-1="http://doi.org/10.5281/zenodo.11150275" httpUri-0="https://zenodo.org/record/11150273/files/figure.png" httpUri-1="https://zenodo.org/record/11150275/files/figure.png" pageId="10" pageNumber="461">Figures 5–6</figureCitation>
,
<figureCitation id="136F2A3F6549966AFEA2FC3A2C6B540E" box="[298,334,1015,1041]" captionStart="FIGURE 9" captionStartId="15.[151,250,1411,1435]" captionTargetBox="[151,1436,201,1388]" captionTargetId="figure-17@15.[151,1436,201,1388]" captionTargetPageId="15" captionText="FIGURE 9. Pronotum posterior angle and basal area of Pterostichus (Wraseiellus) spp. (A) P. pseudandrewesi sp. nov.; (B) P. leishanensis sp. nov.; (C) P. strumatus sp. nov.; (D) P. idiopterus sp. nov.; (E) P. hemichlorus sp. nov.." figureDoi="http://doi.org/10.5281/zenodo.11150285" httpUri="https://zenodo.org/record/11150285/files/figure.png" pageId="10" pageNumber="461">9D</figureCitation>
,
<figureCitation id="136F2A3F6549966AFED2FC3A2CEC540E" box="[346,457,1015,1041]" captionStart="FIGURE 13" captionStartId="19.[151,250,1591,1615]" captionTargetBox="[176,1419,205,1547]" captionTargetId="figure-17@19.[151,1436,181,1567]" captionTargetPageId="19" captionText="FIGURE 13. Female genitalia of Pterostichus (Wraseiellus) spp. (A–B) female internal reproductive tracts; (C–E) gonocoxites of ovipositor, ventral view (left) and dorsal view (right, gonocoxite 2 only). (A, D) P. idiopterus sp. nov.; (B, E.) P. hemichlorus sp. nov.; (C) P. leishanensis sp. nov. Scale bars = 1.0 mm for A–B; = 0.5 mm for C–E. Abbreviations: bc bursa copulatrix; co common oviduct; sc seminal canal; rc receptaculum; g1 gonocoxite 1; g2 gonocoxite 2; es ensiform setae; ns nematiform setae." figureDoi="http://doi.org/10.5281/zenodo.11150301" httpUri="https://zenodo.org/record/11150301/files/figure.png" pageId="10" pageNumber="461">13A, 13D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34E65316549966AFF1FFBF22F0C552E" pageId="10" pageNumber="461" type="materials_examined">
<paragraph id="8BEB36BA6549966AFF1FFBF22F0C552E" blockId="10.[151,1437,1086,1905]" pageId="10" pageNumber="461">
<emphasis id="B920EAA86549966AFF1FFBF22C6B5447" bold="true" box="[151,334,1086,1113]" pageId="10" pageNumber="461">Type materials</emphasis>
.
<materialsCitation id="3B3C3CE76549966AFEE8FBF22FD754BE" collectingDate="2022-05-14" collectionCode="IZAS" country="China" county="Weixi County" determinerName="Chen" elevation="2771" latitude="27.240423" location="Zhonglu Township" longLatPrecision="1" longitude="99.019905" municipality="Diqing" pageId="10" pageNumber="461" specimenCount="2" stateProvince="Yunnan" typeStatus="holotype">
<typeStatus id="54EF88186549966AFEE8FBF22CE85446" box="[352,461,1087,1113]" pageId="10" pageNumber="461" type="holotype">Holotype</typeStatus>
: male, body length = 13.0 mm, (
<collectionCode id="ED45AE7F6549966AFCD3FBF22EBB5446" box="[859,926,1087,1113]" country="0" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="10" pageNumber="461">IZAS</collectionCode>
), “
<collectingCountry id="F343762A6549966AFC48FBF2293A5446" box="[960,1055,1087,1113]" name="China" pageId="10" pageNumber="461">CHINA</collectingCountry>
:
<collectingRegion id="4990F8586549966AFBA6FBF229A25446" box="[1070,1159,1087,1113]" country="China" name="Yunnan" pageId="10" pageNumber="461">Yunnan</collectingRegion>
,
<collectingMunicipality id="6B8FACC06549966AFB1EFBF229C25446" box="[1174,1255,1087,1113]" pageId="10" pageNumber="461">Diqing</collectingMunicipality>
,
<collectingCounty id="628A4E366549966AFB7EFBF228BC5446" box="[1270,1433,1087,1113]" pageId="10" pageNumber="461">Weixi County</collectingCounty>
,
<location id="8E8B60616549966AFF1FFBAE2C545462" LSID="urn:lsid:plazi:treatment:03FD87AC6549966CFF1FFC632CCF539E:8E8B60616549966AFF1FFBAE2C545462" box="[151,369,1123,1149]" country="China" county="Weixi County" latitude="27.240423" longLatPrecision="1" longitude="99.019905" municipality="Diqing" name="Zhonglu Township" pageId="10" pageNumber="461" stateProvince="Yunnan">Zhonglu Township</location>
,
<quantity id="4CAC9B5F6549966AFEF7FBAE2CEA5462" box="[383,463,1123,1149]" metricMagnitude="3" metricUnit="m" metricValue="2.771" pageId="10" pageNumber="461" unit="m" value="2771.0">
<elevation id="0079D1896549966AFEF7FBAE2CEA5462" box="[383,463,1123,1149]" metricMagnitude="3" metricUnit="m" metricValue="2.771" pageId="10" pageNumber="461" unit="m" value="2771.0">2771m</elevation>
</quantity>
,
<geoCoordinate id="EE60507D6549966AFE56FBAE2F525462" box="[478,631,1123,1149]" degrees="27.2404236" direction="north" orientation="latitude" pageId="10" pageNumber="461" precision="1" value="27.240423">N27.2404236</geoCoordinate>
<geoCoordinate id="EE60507D6549966AFD09FBAF2E325462" box="[641,791,1122,1149]" degrees="99.0199035" direction="east" orientation="longitude" pageId="10" pageNumber="461" precision="1" value="99.019905">E99.0199035</geoCoordinate>
<date id="FFEA107A6549966AFCA9FBAE2EBF5462" box="[801,922,1123,1149]" pageId="10" pageNumber="461" value="2022-05-14">
<collectingDate id="EFAEE9926549966AFCA9FBAE2EBF5462" box="[801,922,1123,1149]" pageId="10" pageNumber="461" value="2022-05-14">2022.V-14</collectingDate>
</date>
” // “
<typeStatus id="54EF88186549966AFC59FBAE294D5462" box="[977,1128,1123,1149]" pageId="10" pageNumber="461" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C544D396549966AFBFAFBAE2C2C54BE" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="idiopterus" status="sp. nov." subGenus="Wraseiellus">
<emphasis id="B920EAA86549966AFBFAFBAE29D85462" box="[1138,1277,1123,1149]" italics="true" pageId="10" pageNumber="461">Pterostichus</emphasis>
(
<emphasis id="B920EAA86549966AFA98FBAE28B65462" box="[1296,1427,1123,1149]" italics="true" pageId="10" pageNumber="461">Wraseiellus</emphasis>
)
<emphasis id="B920EAA86549966AFF1FFB4A2C2C54BE" box="[151,265,1159,1185]" italics="true" pageId="10" pageNumber="461">idiopterus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357D36549966AFE87FB4A2C4254BE" box="[271,359,1159,1185]" pageId="10" pageNumber="461" rank="species">sp. nov.</taxonomicNameLabel>
, det.
<determinerName id="7E37DA7E6549966AFE29FB4A2CF954BE" box="[417,476,1159,1185]" pageId="10" pageNumber="461">Chen</determinerName>
&amp; Shi. 2024” [red label]
</materialsCitation>
;
<typeStatus id="54EF88186549966AFD74FB4A2E4C54BE" box="[764,873,1159,1185]" pageId="10" pageNumber="461" type="paratype">Paratypes</typeStatus>
(a total of
<specimenCount id="9D52FD336549966AFC69FB4A296154BE" box="[993,1092,1159,1185]" count="87" pageId="10" pageNumber="461" type="male">87 males</specimenCount>
and
<specimenCount id="9D52FD336549966AFBF1FB4B29D054BE" box="[1145,1269,1158,1185]" count="57" pageId="10" pageNumber="461" type="female">57 females</specimenCount>
):
<materialsCitation id="3B3C3CE76549966AFA82FB4A2F8F54DA" collectingDate="2022-05-14" collectionCode="CCJH" country="China" county="Weixi County" determinerName="Chen" elevation="2771" latitude="27.240423" location="Zhonglu Township" longLatPrecision="1" longitude="99.019905" municipality="Diqing" pageId="10" pageNumber="461" specimenCount="22" specimenCount-female="10" specimenCount-male="12" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D52FD336549966AFA82FB4A284854BE" box="[1290,1389,1159,1185]" count="12" pageId="10" pageNumber="461" type="male">12 males</specimenCount>
and
<specimenCount id="9D52FD336549966AFF1FFB662C3754DA" box="[151,274,1195,1221]" count="10" pageId="10" pageNumber="461" type="female">10 females</specimenCount>
(
<collectionCode id="ED45AE7F6549966AFEABFB662C4854DA" box="[291,365,1195,1221]" pageId="10" pageNumber="461">CCJH</collectionCode>
), the same data as holotype
</materialsCitation>
;
<materialsCitation id="3B3C3CE76549966AFD3DFB662FF854F6" collectingDate="2022-06-08" collectionCode="IZAS, CBFU" country="China" county="Weixi county" location="Zanli village" municipality="Zhonglu" pageId="10" pageNumber="461" specimenCount="117" specimenCount-female="45" specimenCount-male="72" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D52FD336549966AFD3DFB662E3E54DA" box="[693,795,1195,1221]" count="72" pageId="10" pageNumber="461" type="male">72 males</specimenCount>
and
<specimenCount id="9D52FD336549966AFCDCFB672EEA54DA" box="[852,975,1194,1221]" count="45" pageId="10" pageNumber="461" type="female">45 females</specimenCount>
(
<collectionCode id="ED45AE7F6549966AFC57FB66290454DA" box="[991,1057,1195,1221]" country="0" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="10" pageNumber="461">IZAS</collectionCode>
,
<collectionCode id="ED45AE7F6549966AFBA6FB66295E54DA" box="[1070,1147,1195,1221]" pageId="10" pageNumber="461">CBFU</collectionCode>
), “
<collectingRegion id="4990F8586549966AFB12FB6629D054DA" box="[1178,1269,1195,1221]" country="China" name="Yunnan" pageId="10" pageNumber="461">Yunnan</collectingRegion>
,
<collectingCounty id="628A4E366549966AFA88FB6628BD54DA" box="[1280,1432,1195,1221]" pageId="10" pageNumber="461">Weixi county</collectingCounty>
,
<collectingMunicipality id="6B8FACC06549966AFF1FFB022DDC54F6" box="[151,249,1231,1257]" pageId="10" pageNumber="461">Zhonglu</collectingMunicipality>
,
<location id="8E8B60616549966AFE8DFB022CB254F6" LSID="urn:lsid:plazi:treatment:03FD87AC6549966CFF1FFC632CCF539E:8E8B60616549966AFE8DFB022CB254F6" box="[261,407,1231,1257]" country="China" county="Weixi county" municipality="Zhonglu" name="Zanli village" pageId="10" pageNumber="461" stateProvince="Yunnan">Zanli village</location>
,
<date id="FFEA107A6549966AFE2BFB022F3D54F6" box="[419,536,1231,1257]" pageId="10" pageNumber="461" value="2022-06-08">
<collectingDate id="EFAEE9926549966AFE2BFB022F3D54F6" box="[419,536,1231,1257]" pageId="10" pageNumber="461" value="2022-06-08">2022.VI.8</collectingDate>
</date>
, Yu Zhibiao lgt.”
</materialsCitation>
;
<materialsCitation id="3B3C3CE76549966AFD62FB022FCC5512" collectionCode="CLHY" country="China" county="Diqing" elevation="3000" location="Zhonglu" municipality="Weixi" pageId="10" pageNumber="461" specimenCount="1" specimenCount-male="1" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D52FD336549966AFD62FB022E1354F6" box="[746,822,1231,1257]" count="1" pageId="10" pageNumber="461" type="male">1 male</specimenCount>
(
<collectionCode id="ED45AE7F6549966AFCCEFB022EBD54F6" box="[838,920,1231,1257]" pageId="10" pageNumber="461">CLHY</collectionCode>
), “
<collectingRegion id="4990F8586549966AFC3EFB02295954F6" box="[950,1148,1231,1257]" country="China" name="Yunnan" pageId="10" pageNumber="461">Yunnan Province</collectingRegion>
,
<collectingCounty id="628A4E366549966AFB01FB0229FF54F6" box="[1161,1242,1231,1257]" pageId="10" pageNumber="461">Diqing</collectingCounty>
,
<collectingMunicipality id="6B8FACC06549966AFB6DFB02280F54F6" box="[1253,1322,1231,1257]" pageId="10" pageNumber="461">Weixi</collectingMunicipality>
,
<location id="8E8B60616549966AFABEFB0228BD54F6" LSID="urn:lsid:plazi:treatment:03FD87AC6549966CFF1FFC632CCF539E:8E8B60616549966AFABEFB0228BD54F6" box="[1334,1432,1231,1257]" country="China" county="Diqing" municipality="Weixi" name="Zhonglu" pageId="10" pageNumber="461" stateProvince="Yunnan">Zhonglu</location>
,
<location id="8E8B60616549966AFF1FFB3E2C305512" LSID="urn:lsid:plazi:treatment:03FD87AC6549966CFF1FFC632CCF539E:8E8B60616549966AFF1FFB3E2C305512" box="[151,277,1267,1293]" country="China" county="Diqing" municipality="Weixi" name="Dayingpan" pageId="10" pageNumber="461" stateProvince="Yunnan">Dayingpan</location>
,
<quantity id="4CAC9B5F6549966AFEABFB3E2C515512" box="[291,372,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="461" unit="m" value="3000.0">
<elevation id="0079D1896549966AFEABFB3E2C515512" box="[291,372,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="461" unit="m" value="3000.0">3000m</elevation>
</quantity>
, 2022.V, Feng Xiangzhong lgt.”
</materialsCitation>
;
<materialsCitation id="3B3C3CE76549966AFD70FB3E2F00552E" collectingDate="2022-06-20" collectionCode="CLYH" country="China" location="Zhonglu" municipality="Weixi" pageId="10" pageNumber="461" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Yunnan" typeStatus="paratype">
<specimenCount id="9D52FD336549966AFD70FB3E2E755512" box="[760,848,1267,1293]" count="2" pageId="10" pageNumber="461" type="male">2 males</specimenCount>
and
<specimenCount id="9D52FD336549966AFC04FB3E2EDE5512" box="[908,1019,1267,1293]" count="2" pageId="10" pageNumber="461" type="female">2 females</specimenCount>
(
<collectionCode id="ED45AE7F6549966AFB84FB3E29795512" box="[1036,1116,1267,1293]" pageId="10" pageNumber="461">CLYH</collectionCode>
), “
<collectingRegion id="4990F8586549966AFBF3FB3E29F35512" box="[1147,1238,1267,1293]" country="China" name="Yunnan" pageId="10" pageNumber="461">Yunnan</collectingRegion>
,
<collectingMunicipality id="6B8FACC06549966AFB6CFB3E280C5512" box="[1252,1321,1267,1293]" pageId="10" pageNumber="461">Weixi</collectingMunicipality>
,
<location id="8E8B60616549966AFABEFB3E28BD5512" LSID="urn:lsid:plazi:treatment:03FD87AC6549966CFF1FFC632CCF539E:8E8B60616549966AFABEFB3E28BD5512" box="[1334,1432,1267,1293]" country="China" municipality="Weixi" name="Zhonglu" pageId="10" pageNumber="461" stateProvince="Yunnan">Zhonglu</location>
, Zanli,
<date id="FFEA107A6549966AFF68FADA2C46552E" box="[224,355,1303,1329]" pageId="10" pageNumber="461" value="2022-06-20">
<collectingDate id="EFAEE9926549966AFF68FADA2C46552E" box="[224,355,1303,1329]" pageId="10" pageNumber="461" value="2022-06-20">2022.VI.20</collectingDate>
</date>
, Yu Zhibiao lgt.”
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E65316549966AFF4FFAF72F71576E" pageId="10" pageNumber="461" type="diagnosis">
<paragraph id="8BEB36BA6549966AFF4FFAF72FC15616" blockId="10.[151,1437,1086,1905]" pageId="10" pageNumber="461">
<emphasis id="B920EAA86549966AFF4FFAF72C1F554B" bold="true" box="[199,314,1338,1364]" pageId="10" pageNumber="461">Diagnosis</emphasis>
. Frontal furrows shallow and short, exceeding level of anterior margin of eyes; vertex nearly impunctate; temporae short (TL/EYL 0.31–0.33). Maximum width of pronotum near middle; lateral margins nearly straight before posterior angles; posterior angles obtuse, without denticle; basal foveal area sparsely punctate, outer groove short but well-defined. Elytral intervals heterogeneous, on elytra median region, odd intervals about twice width as even ones. Apical lamella of aedeagus laminar, apex not thickened or turned dorsally (
<figureCitation id="136F2A3F6549966AFB7AFA06286A55FA" box="[1266,1359,1483,1509]" captionStart="FIGURE 6" captionStartId="12.[151,250,1854,1878]" captionTargetBox="[160,1427,930,1822]" captionTargetId="figure-380@12.[151,1436,921,1831]" captionTargetPageId="12" captionText="FIGURE 6. Male genitalia of Pterostichus idiopterus sp. nov., holotype. (A) median lobe of aedeagus, left lateral view; (B) median lobe of aedeagus, dorsal view; (C) right paramere; (D) endophallus, right lateral view; (E) endophallus, left lateral view. Scale bars = 1.0 mm. Abbreviations: pa pre-apical lobe; gp gonopore." figureDoi="http://doi.org/10.5281/zenodo.11150275" httpUri="https://zenodo.org/record/11150275/files/figure.png" pageId="10" pageNumber="461">Fig. 6A</figureCitation>
); right paramere short and laminar, apex rounded (
<figureCitation id="136F2A3F6549966AFDF6FA222FF25616" box="[638,727,1519,1545]" captionStart="FIGURE 6" captionStartId="12.[151,250,1854,1878]" captionTargetBox="[160,1427,930,1822]" captionTargetId="figure-380@12.[151,1436,921,1831]" captionTargetPageId="12" captionText="FIGURE 6. Male genitalia of Pterostichus idiopterus sp. nov., holotype. (A) median lobe of aedeagus, left lateral view; (B) median lobe of aedeagus, dorsal view; (C) right paramere; (D) endophallus, right lateral view; (E) endophallus, left lateral view. Scale bars = 1.0 mm. Abbreviations: pa pre-apical lobe; gp gonopore." figureDoi="http://doi.org/10.5281/zenodo.11150275" httpUri="https://zenodo.org/record/11150275/files/figure.png" pageId="10" pageNumber="461">Fig. 6C</figureCitation>
).
</paragraph>
<paragraph id="8BEB36BA6549966AFF4FF9DF2CF556FE" blockId="10.[151,1437,1086,1905]" pageId="10" pageNumber="461">
<emphasis id="B920EAA86549966AFF4FF9DF2C435633" bold="true" box="[199,358,1554,1580]" pageId="10" pageNumber="461">Comparisons</emphasis>
. This new species can be easily distinguished from all other species of the subgenus by its heterogeneous elytral intervals. Besides the elytral characters,
<taxonomicName id="4C544D396549966AFCC2F9FA2EF3564E" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[842,982,1591,1617]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="idiopterus" status="sp. n.">
<emphasis id="B920EAA86549966AFCC2F9FA2EF3564E" box="[842,982,1591,1617]" italics="true" pageId="10" pageNumber="461">P. idiopterus</emphasis>
</taxonomicName>
<emphasis id="B920EAA86549966AFC53F9FA293C564E" bold="true" box="[987,1049,1591,1617]" pageId="10" pageNumber="461">
<taxonomicNameLabel id="A21357D36549966AFC53F9FA293C564E" box="[987,1049,1591,1617]" pageId="10" pageNumber="461" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is also different from other species of the
<taxonomicName id="4C544D396549966AFF6BF9962C65566A" baseAuthorityName="Fairmaire" baseAuthorityYear="1886" box="[227,320,1627,1653]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B920EAA86549966AFF6BF9962C65566A" box="[227,320,1627,1653]" italics="true" pageId="10" pageNumber="461">diversus</emphasis>
</taxonomicName>
species-group by the combination of: (1) pronotum with posterior margin narrower than anterior margin; (2) pronotal posterior angles obtuse; (3) females with slightly stronger isodiametric elytral microsculpture than males, but not granular; (4) in dorsal view, apical lamella of aedeagus indistinctly bent to left; (5) right paramere short, apex widely rounded.
</paragraph>
<paragraph id="8BEB36BA6549966AFF4FF9262F71576E" blockId="10.[151,1437,1086,1905]" pageId="10" pageNumber="461">
Despite their conspicuous differences on elytra, the new species is inferred to be closest to
<taxonomicName id="4C544D396549966AFB27F9262853571A" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[1199,1398,1771,1797]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="pseudodiversus">
<emphasis id="B920EAA86549966AFB27F9262853571A" box="[1199,1398,1771,1797]" italics="true" pageId="10" pageNumber="461">P. pseudodiversus</emphasis>
</taxonomicName>
for their similarities on male genitalia: apical lamella short and slightly twisted, apex not turned dorsally. Comparing to the latter species, the male genitalia of
<taxonomicName id="4C544D396549966AFDC2F8FE2FFD5752" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[586,728,1843,1869]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="461" phylum="Arthropoda" rank="species" species="idiopterus" status="sp. n.">
<emphasis id="B920EAA86549966AFDC2F8FE2FFD5752" box="[586,728,1843,1869]" italics="true" pageId="10" pageNumber="461">P. idiopterus</emphasis>
</taxonomicName>
<emphasis id="B920EAA86549966AFD56F8FE2E385752" bold="true" box="[734,797,1843,1869]" pageId="10" pageNumber="461">
<taxonomicNameLabel id="A21357D36549966AFD56F8FE2E385752" box="[734,797,1843,1869]" pageId="10" pageNumber="461" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is different in the apical lamella only very weakly turned to left and right paramere much shorter.
</paragraph>
</subSubSection>
<caption id="DF2B66326548966BFF1FFA47282B55BD" ID-DOI="http://doi.org/10.5281/zenodo.11150273" ID-Zenodo-Dep="11150273" box="[151,1294,1418,1443]" httpUri="https://zenodo.org/record/11150273/files/figure.png" pageId="11" pageNumber="462" startId="11.[151,250,1418,1442]" targetBox="[159,1427,189,1386]" targetPageId="11" targetType="figure">
<paragraph id="8BEB36BA6548966BFF1FFA47282B55BD" blockId="11.[151,1294,1418,1443]" box="[151,1294,1418,1443]" pageId="11" pageNumber="462">
<emphasis id="B920EAA86548966BFF1FFA472C3655BD" bold="true" box="[151,275,1418,1442]" pageId="11" pageNumber="462">FIGURE 5.</emphasis>
Habitus of
<taxonomicName id="4C544D396548966BFE02FA472F5055BD" authority="Chen &amp; Yin &amp; Shi, 2024" authorityName="Chen &amp; Yin &amp; Shi" authorityYear="2024" box="[394,629,1418,1442]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="462" phylum="Arthropoda" rank="species" species="idiopterus" status="sp. nov.">
<emphasis id="B920EAA86548966BFE02FA472F5055BD" box="[394,629,1418,1442]" italics="true" pageId="11" pageNumber="462">Pterostichus idiopterus</emphasis>
</taxonomicName>
<emphasis id="B920EAA86548966BFDF3FA472FE955BD" bold="true" box="[635,716,1418,1442]" pageId="11" pageNumber="462">
<taxonomicNameLabel id="A21357D36548966BFDF3FA472FE955BD" box="[635,716,1418,1442]" pageId="11" pageNumber="462" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A) holotype; (B) Paratype, female. Scale bar = 5.0 mm.
</paragraph>
</caption>
<subSubSection id="C34E65316548966CFF4FFA1D285852B6" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462" type="description">
<paragraph id="8BEB36BA6548966CFF4FFA1D28B85102" blockId="11.[151,1437,1488,2019]" lastBlockId="12.[151,1437,151,897]" lastPageId="12" lastPageNumber="463" pageId="11" pageNumber="462">
<emphasis id="B920EAA86548966BFF4FFA1D2C7455F5" bold="true" box="[199,337,1488,1514]" pageId="11" pageNumber="462">Description</emphasis>
. BL
<quantity id="4CAC9B5F6548966BFE19FA1C2F1F55F4" box="[401,570,1489,1515]" metricMagnitude="-2" metricUnit="m" metricValue="1.345" metricValueMax="1.42" metricValueMin="1.27" pageId="11" pageNumber="462" unit="mm" value="13.45" valueMax="14.2" valueMin="12.7">12.7–14.2 mm</quantity>
; dorsal surface dark brown to black, without metallic luster, elytral lateral margin and epipleura reddish brown (
<figureCitation id="136F2A3F6548966BFDCAFA392FAD5610" box="[578,648,1524,1551]" captionStart="FIGURE 5" captionStartId="11.[151,250,1418,1442]" captionTargetBox="[159,1427,189,1386]" captionTargetId="figure-17@11.[151,1436,181,1394]" captionTargetPageId="11" captionText="FIGURE 5. Habitus of Pterostichus idiopterus sp. nov. (A) holotype; (B) Paratype, female. Scale bar = 5.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11150273" httpUri="https://zenodo.org/record/11150273/files/figure.png" pageId="11" pageNumber="462">Fig. 5</figureCitation>
).
<emphasis id="B920EAA86548966BFD16FA382FFB5610" bold="true" box="[670,734,1525,1551]" pageId="11" pageNumber="462">Head</emphasis>
. Frons finely and sparsely punctate, frontal furrows short and shallow, exceeding level of eyes’ anterior margin; vertex impunctate or with very few fine punctures; temporae short (TL/EYL 0.31–0.33), not swollen behind eyes; paraocular grooves deep, impunctate and nearly straight, reaching posterior margin of eyes; antennomere 3 without accessory setae.
<emphasis id="B920EAA86548966BFC0EF9AC2EDA5664" bold="true" box="[902,1023,1633,1659]" pageId="11" pageNumber="462">Pronotum</emphasis>
sub-cordiform (PW/ PL 1.31–1.35); anterior and posterior margins almost straight, lateral margins evenly curved from anterior angles to posterior third, nearly straight or weakly sinuate before posterior angles; anterior margin slightly wider than posterior margin; anterior angles obtusely rounded, slightly projecting forwardly; posterior angles obtuse, apex a little distinct, not forming prominent denticle; discal area glabrous with faint transverse wrinkles; basal foveae relatively short and wide, deeply depressed and areal punctate, outer groove short but well-defined, area between inner and outer basal foveal grooves rugose and depressed, mid-basal area between two basal foveae sparsely and finely punctate, ridge between outer groove and lateral margin distinct; one mid-lateral seta and one basal-lateral seta present on each side.
<emphasis id="B920EAA86548966BFF1FF84C2DC65784" bold="true" box="[151,227,1921,1947]" pageId="11" pageNumber="462">Elytra</emphasis>
oblong (EL/EW 1.48–1.52); humeral teeth small; intervals heterogeneous and slightly convex, odd intervals (1, 3, 5 and 7) obviously wider than even ones, about twice width as even ones on elytra middle, slightly convex similarly to even ones; striae relatively shallow, without punctures; parascutellar striae short, apex joined to stria 1, stria 1 usually entire at base; umbilicate series on interval 9 composed of approximately 13–16 pores, sparse in middle and dense in anterior and posterior areas; elytral microsculpture shallowly isodiametric in males, stronger in females but not granular as other species of the subgenus.
<emphasis id="B920EAA8654F966CFC9EFF122E8150E7" bold="true" box="[790,932,222,249]" pageId="12" pageNumber="463">Ventral side</emphasis>
. Proepisterna rugose, usually with coarse and dense punctures on anterior half; metaepisterna coarsely punctate; sternite VII of males without sexual structure.
</paragraph>
<paragraph id="8BEB36BA654F966CFF1FFEEA285852B6" blockId="12.[151,1437,151,897]" pageId="12" pageNumber="463">
<emphasis id="B920EAA8654F966CFF1FFEEA2DEA515E" bold="true" box="[151,207,295,321]" pageId="12" pageNumber="463">Legs</emphasis>
. Tarsomere 5 without setae beneath.
<emphasis id="B920EAA8654F966CFDFFFEEA2E07515F" bold="true" box="[631,802,294,321]" pageId="12" pageNumber="463">Male genitalia</emphasis>
(
<figureCitation id="136F2A3F654F966CFCBBFEEA2E5D515E" box="[819,888,295,321]" captionStart="FIGURE 6" captionStartId="12.[151,250,1854,1878]" captionTargetBox="[160,1427,930,1822]" captionTargetId="figure-380@12.[151,1436,921,1831]" captionTargetPageId="12" captionText="FIGURE 6. Male genitalia of Pterostichus idiopterus sp. nov., holotype. (A) median lobe of aedeagus, left lateral view; (B) median lobe of aedeagus, dorsal view; (C) right paramere; (D) endophallus, right lateral view; (E) endophallus, left lateral view. Scale bars = 1.0 mm. Abbreviations: pa pre-apical lobe; gp gonopore." figureDoi="http://doi.org/10.5281/zenodo.11150275" httpUri="https://zenodo.org/record/11150275/files/figure.png" pageId="12" pageNumber="463">Fig. 6</figureCitation>
). Median lobe of aedeagus relatively slender; in lateral view, ventral margin almost straight near middle, weakly curved to the ventral side at apical third, apical orifice large, slightly turned to left side, apical lamella evenly thin, not apically thickened or dorsally turned; in dorsal view, apical lamella short (AL/AW 1.09–1.19), gradually narrowed to apex, apex rounded-truncated, very weakly bent to left; ventral surface of apical lamella simple. Right paramere laminar, apical portion (area distal to the articulation) very short, apex rounded. Endophallus (
<figureCitation id="136F2A3F654F966CFC80FE162EA451EA" box="[776,897,475,501]" captionStart="FIGURE 6" captionStartId="12.[151,250,1854,1878]" captionTargetBox="[160,1427,930,1822]" captionTargetId="figure-380@12.[151,1436,921,1831]" captionTargetPageId="12" captionText="FIGURE 6. Male genitalia of Pterostichus idiopterus sp. nov., holotype. (A) median lobe of aedeagus, left lateral view; (B) median lobe of aedeagus, dorsal view; (C) right paramere; (D) endophallus, right lateral view; (E) endophallus, left lateral view. Scale bars = 1.0 mm. Abbreviations: pa pre-apical lobe; gp gonopore." figureDoi="http://doi.org/10.5281/zenodo.11150275" httpUri="https://zenodo.org/record/11150275/files/figure.png" pageId="12" pageNumber="463">Figs 6D, E</figureCitation>
) bent to ventral side across left side of aedeagus, major parts of endophallus located on ventral side of aedeagus (in lateral view); gonopore (gp) opened to ventral-basal direction of aedeagus; surface with fine scales, denser near gonopore, without coarse setae; pre-apical lobe (pa) rounded, visible in both lateral views.
<emphasis id="B920EAA8654F966CFD0CFD8A2E6E527F" bold="true" box="[644,843,582,609]" pageId="12" pageNumber="463">Female genitalia</emphasis>
(
<figureCitation id="136F2A3F654F966CFCD3FD8A2ECE527E" box="[859,1003,583,609]" captionStart="FIGURE 13" captionStartId="19.[151,250,1591,1615]" captionTargetBox="[176,1419,205,1547]" captionTargetId="figure-17@19.[151,1436,181,1567]" captionTargetPageId="19" captionText="FIGURE 13. Female genitalia of Pterostichus (Wraseiellus) spp. (A–B) female internal reproductive tracts; (C–E) gonocoxites of ovipositor, ventral view (left) and dorsal view (right, gonocoxite 2 only). (A, D) P. idiopterus sp. nov.; (B, E.) P. hemichlorus sp. nov.; (C) P. leishanensis sp. nov. Scale bars = 1.0 mm for A–B; = 0.5 mm for C–E. Abbreviations: bc bursa copulatrix; co common oviduct; sc seminal canal; rc receptaculum; g1 gonocoxite 1; g2 gonocoxite 2; es ensiform setae; ns nematiform setae." figureDoi="http://doi.org/10.5281/zenodo.11150301" httpUri="https://zenodo.org/record/11150301/files/figure.png" pageId="12" pageNumber="463">Figs 13A, D</figureCitation>
). Spermatheca with the seminal canal about five times long as receptaculum; seminal canal inserted at base of common oviduct; base of seminal canal sclerotized. Gonocoxite 2 with two or three ensiform setae at the outer margin and three or four at inner margin.
</paragraph>
</subSubSection>
<subSubSection id="C34E6531654F966CFF4FFD7E282B52EE" pageId="12" pageNumber="463" type="etymology">
<paragraph id="8BEB36BA654F966CFF4FFD7E282B52EE" blockId="12.[151,1437,151,897]" pageId="12" pageNumber="463">
<emphasis id="B920EAA8654F966CFF4FFD7E2C6D52D2" bold="true" box="[199,328,691,717]" pageId="12" pageNumber="463">Etymology</emphasis>
. The scientific name of the new species is the combination of the Greek root “
<emphasis id="B920EAA8654F966CFB32FD7E29C352D2" box="[1210,1254,691,717]" italics="true" pageId="12" pageNumber="463">idio</emphasis>
-”, which means pertaining to self, personal, and the Greek root “
<emphasis id="B920EAA8654F966CFD3CFD1A2FC852EF" box="[692,749,727,752]" italics="true" pageId="12" pageNumber="463">Pter-</emphasis>
”, referring to its heterogeneous elytral intervals.
</paragraph>
</subSubSection>
<subSubSection id="C34E6531654F966CFF4FFD372CCF539E" pageId="12" pageNumber="463" type="distribution">
<paragraph id="8BEB36BA654F966CFF4FFD372CCF539E" blockId="12.[151,1437,151,897]" pageId="12" pageNumber="463">
<emphasis id="B920EAA8654F966CFF4FFD372C7C530B" bold="true" box="[199,345,762,788]" pageId="12" pageNumber="463">Distribution</emphasis>
. Only known from the mountain area western to Zhonglu, Weixi Lisu Autonomous County in Northwest
<collectingRegion id="4990F858654F966CFE9AFCD22C4E5326" box="[274,363,799,825]" country="China" name="Yunnan" pageId="12" pageNumber="463">Yunnan</collectingRegion>
. This new species was collected at the Biluoxueshan mountain range, western to Lancang Jiang (= Mekong River); while the more widely distributed species
<taxonomicName id="4C544D39654F966CFCD8FC8E29895342" authority="Shi &amp; Sciaky" authorityName="Shi &amp; Sciaky" authorityYear="2013" box="[848,1196,835,861]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="463" phylum="Arthropoda" rank="species" species="pseudodiverus">
<emphasis id="B920EAA8654F966CFCD8FC8E292B5342" box="[848,1038,835,861]" italics="true" pageId="12" pageNumber="463">P. pseudodiverus</emphasis>
Shi &amp; Sciaky
</taxonomicName>
is only distributed in area eastern to Lancang Jiang.
</paragraph>
</subSubSection>
</treatment>
</document>
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