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<document id="251BCF320F953B0F8557F460B9CE0DE3" ID-CLB-Dataset="2470" ID-DOI="10.21248/contrib.entomol.67.2.213-246" ID-GBIF-Dataset="43c6cab4-bf7d-43c9-b9bf-b22c4ebf7924" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742339" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638267510724" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03FD87A1F11FFFEF761AFF2350DF7B79" docLanguage="en" docName="ContribEntomol.67.2.213-246.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) matangensis Cameron 1943" docType="treatment" docVersion="10" lastPageNumber="232" masterDocId="FFC4FFD9F10CFFFC7525FFD054747F38" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="246" masterPageNumber="213" pageNumber="232" updateTime="1699471357828" updateUser="ExternalLinkService">
<mods:mods id="0542E05E5ED308A491A0C99DA563B501" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="A1A207327EA5D75686CFB0DC59515C2B">On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
</mods:titleInfo>
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<mods:namePart id="8990519F540A33FB37FBD33146A11305">Assing, Volker</mods:namePart>
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<mods:title id="96902C9CDA79167A0F18FA3F778B64B6">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="56A092E4D00BF45DCF5471280E8041DC">
<mods:date id="7707ADCE41096321C45728E22A23A24B">2017</mods:date>
<mods:detail id="D2711E7B0982034840E13B52AA913B25" type="pubDate">
<mods:number id="A5749ACA5A446AE931EBC3DDF3CE6B95">2017-12-08</mods:number>
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<mods:number id="04879365E8B4E6E712F499594F4FC593">67</mods:number>
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<mods:classification id="3AE4D81CC970CAB11A73543F5E48CDAA">journal article</mods:classification>
<mods:identifier id="65BC742ACBB6E56B2CD74B2EB1BE4ACA" type="CLB-Dataset">2470</mods:identifier>
<mods:identifier id="4018850604A1EA9396BED1B3E7871D9D" type="DOI">10.21248/contrib.entomol.67.2.213-246</mods:identifier>
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</mods:mods>
<treatment id="03FD87A1F11FFFEF761AFF2350DF7B79" ID-DOI="http://doi.org/10.5281/zenodo.5742272" ID-GBIF-Taxon="192500446" ID-Zenodo-Dep="5742272" LSID="urn:lsid:plazi:treatment:03FD87A1F11FFFEF761AFF2350DF7B79" httpUri="http://treatment.plazi.org/id/03FD87A1F11FFFEF761AFF2350DF7B79" lastPageNumber="232" pageId="19" pageNumber="232">
<subSubSection id="C34E653CF11FFFEF761AFF2350827E36" box="[831,1270,243,270]" pageId="19" pageNumber="232" type="nomenclature">
<paragraph id="8BEB36B7F11FFFEF761AFF2350827E36" blockId="19.[831,1270,243,303]" box="[831,1270,243,270]" pageId="19" pageNumber="232">
<heading id="D0A381DBF11FFFEF761AFF2350827E36" box="[831,1270,243,270]" centered="true" fontSize="12" level="2" pageId="19" pageNumber="232" reason="2">
<taxonomicName id="4C544D34F11FFFEF761AFF2350827E36" ID-CoL="5DDDR" authority="CAMERON, 1943" authorityName="Cameron" authorityYear="1943" box="[831,1270,243,270]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="matangensis" subGenus="Zyras">
<emphasis id="B920EAA5F11FFFEF761AFF2357017E36" box="[831,885,243,270]" italics="true" pageId="19" pageNumber="232">Zyras</emphasis>
(
<emphasis id="B920EAA5F11FFFEF76A6FF2357CD7E36" box="[899,953,243,270]" italics="true" pageId="19" pageNumber="232">Zyras</emphasis>
)
<emphasis id="B920EAA5F11FFFEF76E2FF23503F7E36" box="[967,1099,243,270]" italics="true" pageId="19" pageNumber="232">matangensis</emphasis>
<bibRefCitation id="EFC54B46F11FFFEF7174FF2350827E36" author="CAMERON, M." box="[1105,1270,243,270]" pageId="19" pageNumber="232" pagination="139 - 143" refId="ref26208" refString="CAMERON, M. 1943: New species of Staphylinidae (Col.) from Borneo. - Entomologist's Monthly Magazine 79: 39 - 42, 139 - 143, 183." type="journal article" year="1943">CAMERON, 1943</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11FFFEF761AFECB57937E17" box="[831,999,283,303]" pageId="19" pageNumber="232" type="description">
<paragraph id="8BEB36B7F11FFFEF761AFECB57937E17" blockId="19.[831,1270,243,303]" box="[831,999,283,303]" pageId="19" pageNumber="232">
(
<figureCitation id="136F2A32F11FFFEF7666FECB57FB7E17" box="[835,911,283,303]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="19" pageNumber="232">Figs 5, 26</figureCitation>
,
<figureCitation id="136F2A32F11FFFEF76B3FECB57DF7E17" box="[918,939,283,303]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="19" pageNumber="232">41</figureCitation>
,
<figureCitation id="136F2A32F11FFFEF7697FECB57967E17" box="[946,994,283,303]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="19" pageNumber="232">63–65</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11FFFEF763FFE9C516F7E58" box="[794,1307,331,353]" pageId="19" pageNumber="232" type="reference_group">
<paragraph id="8BEB36B7F11FFFEF763FFE9C516F7E58" blockId="19.[794,1307,331,353]" box="[794,1307,331,353]" pageId="19" pageNumber="232">
<treatmentCitationGroup id="AB441199F11FFFEF763FFE9C516F7E58" box="[794,1307,331,353]" pageId="19" pageNumber="232">
<treatmentCitation id="0AF510A6F11FFFEF763FFE9C50DD7E59" author="PACE, R." box="[794,1193,331,353]" page="152" pageId="19" pageNumber="232" year="2008">
<taxonomicName id="4C544D34F11FFFEF763FFE9C50DD7E59" authority="PACE, 2008: 152" authorityName="PACE" authorityPageNumber="152" authorityYear="2008" box="[794,1193,331,353]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="daiaccorum" subGenus="Zyras">
<emphasis id="B920EAA5F11FFFEF763FFE9C573A7E59" box="[794,846,332,353]" italics="true" pageId="19" pageNumber="232">Zyras</emphasis>
(
<emphasis id="B920EAA5F11FFFEF7679FE9C57E47E59" box="[860,912,332,353]" italics="true" pageId="19" pageNumber="232">Zyras</emphasis>
)
<emphasis id="B920EAA5F11FFFEF76B8FE9B50787E58" box="[925,1036,331,352]" italics="true" pageId="19" pageNumber="232">daiaccorum</emphasis>
<bibRefCitation id="EFC54B46F11FFFEF7137FE9B50DD7E59" author="PACE, R." box="[1042,1193,331,353]" pageId="19" pageNumber="232" pagination="107 - 156" refId="ref27259" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">
PACE, 2008:
<specimenCount id="9D52FD3EF11FFFEF71A3FE9B50DD7E59" box="[1158,1193,331,353]" pageId="19" pageNumber="232" type="generic">152</specimenCount>
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
f.;
<emphasis id="B920EAA5F11FFFEF71E3FE9B516F7E58" bold="true" box="[1222,1307,331,352]" pageId="19" pageNumber="232">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11FFFEF763FFE5950877C39" pageId="19" pageNumber="232" type="materials_examined">
<paragraph id="8BEB36B7F11FFFEF763FFE5957AB7DB9" blockId="19.[794,1399,393,641]" pageId="19" pageNumber="232">
<emphasis id="B920EAA5F11FFFEF763FFE59504B7E99" bold="true" box="[794,1087,393,417]" pageId="19" pageNumber="232">Type material examined</emphasis>
:
<materialsCitation id="3B3C3CEAF11FFFEF717CFE5951007D79" ID-GBIF-Occurrence="3450187313" collectingDate="1988-08-10" collectionCode="MHNG" collectorName="A. Smetana" country="BORNEO" determinerName="V. Assing" elevation="1490" location="Liwago River" municipality="Mt. Kinabalu Nat. Pk." pageId="19" pageNumber="232" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<typeStatus id="54EF8815F11FFFEF717CFE5950C87E99" box="[1113,1212,393,417]" pageId="19" pageNumber="232" type="holotype">Holotype</typeStatus>
<specimenCount id="9D52FD3EF11FFFEF71F4FE5A50927E99" box="[1233,1254,394,417]" pageId="19" pageNumber="232">♀</specimenCount>
: “
<collectingCountry id="F3437627F11FFFEF702FFE5951027E99" box="[1290,1398,393,417]" pageId="19" pageNumber="232">BORNEO</collectingCountry>
<collectingRegion id="4990F855F11FFFEF763FFE79572E7EF9" box="[794,858,425,449]" country="Malaysia" name="Sabah" pageId="19" pageNumber="232">Sabah</collectingRegion>
<collectingMunicipality id="6B8FACCDF11FFFEF7647FE7A50257EF9" box="[866,1105,425,449]" pageId="19" pageNumber="232">Mt. Kinabalu Nat. Pk.</collectingMunicipality>
, HQ
<location id="8E8B606CF11FFFEF71A8FE7951687EF9" LSID="urn:lsid:plazi:treatment:03FD87A1F11FFFEF761AFF2350DF7B79:8E8B606CF11FFFEF71A8FE7951687EF9" box="[1165,1308,425,449]" country="BORNEO" municipality="Mt. Kinabalu Nat. Pk." name="Liwago River" pageId="19" pageNumber="232" stateProvince="Sabah">Liwago River</location>
,
<quantity id="4CAC9B52F11FFFEF7002FE7951037EF9" box="[1319,1399,425,449]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="19" pageNumber="232" unit="m" value="1490.0">
<elevation id="0079D184F11FFFEF7002FE7951037EF9" box="[1319,1399,425,449]" metricMagnitude="3" metricUnit="m" metricValue="1.49" pageId="19" pageNumber="232" unit="m" value="1490.0">1490 m</elevation>
</quantity>
<date id="FFEA1077F11FFFEF763FFE1957FD7ED9" box="[794,905,457,481]" pageId="19" pageNumber="232" value="1988-08-10">
<collectingDate id="EFAEE99FF11FFFEF763FFE1957FD7ED9" box="[794,905,457,481]" pageId="19" pageNumber="232" value="1988-08-10">10.VIII.88</collectingDate>
</date>
,
<collectorName id="26A15361F11FFFEF76BDFE1950667ED9" box="[920,1042,457,481]" pageId="19" pageNumber="232">A. Smetana</collectorName>
[B97] /
<originalDetermination id="A84A8E4BF11FFFEF7152FE1950287D39" pageId="19" pageNumber="232">
Holotypus
<taxonomicName id="4C544D34F11FFFEF71D4FE1A57297D39" authorityName="PACE" authorityYear="2008" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="daiaccorum">
<emphasis id="B920EAA5F11FFFEF71D4FE1A57297D39" italics="true" pageId="19" pageNumber="232">Zyras daiaccorum</emphasis>
</taxonomicName>
mihi, det. R. Pace 2000
</originalDetermination>
/
<originalDetermination id="A84A8E4BF11FFFEF7151FE3A57A87D19" pageId="19" pageNumber="232">
<taxonomicName id="4C544D34F11FFFEF7151FE3A51457D39" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[1140,1329,489,513]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="daiaccorum" status="sp. nov.">
<emphasis id="B920EAA5F11FFFEF7151FE3A51457D39" box="[1140,1329,489,513]" italics="true" pageId="19" pageNumber="232">Zyras daiaccorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357DEF11FFFEF701CFE3A51067D39" box="[1337,1394,490,513]" pageId="19" pageNumber="232" rank="species">n. sp.</taxonomicNameLabel>
, det. R. Pace 2000
</originalDetermination>
/ MHNG ENTO 00010004 /
<taxonomicName id="4C544D34F11FFFEF7018FDDA50797D79" authority="Cameron" authorityName="Cameron" authorityYear="1943" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="matangensis">
<emphasis id="B920EAA5F11FFFEF7018FDDA57EF7D79" italics="true" pageId="19" pageNumber="232">Zyras matangensis</emphasis>
Cameron
</taxonomicName>
, det.
<determinerName id="7E37DA73F11FFFEF7177FDFA50C07D79" box="[1106,1204,553,577]" pageId="19" pageNumber="232">
V.
<bibRefCitation id="EFC54B46F11FFFEF7148FDF950C07D79" author="ASSING, V." box="[1133,1204,553,577]" pageId="19" pageNumber="232" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing</bibRefCitation>
</determinerName>
2017” (
<collectionCode id="ED45AE72F11FFFEF7036FDF951187D79" box="[1299,1388,553,577]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="19" pageNumber="232" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B3C3CEAF11FFFEF763FFD9A57A87DB9" ID-GBIF-Occurrence="3450187327" collectingDate="1988-09-03" collectionCode="MHNG" collectorName="A. Smetana" determinerName="R. Pace" elevation="1490" municipality="Mt. Kinabalu Nat. Pk." pageId="19" pageNumber="232" specimenCount="1" stateProvince="Sabah" typeStatus="paratype">
<typeStatus id="54EF8815F11FFFEF763FFD9A57037D5A" box="[794,887,586,610]" pageId="19" pageNumber="232" type="paratype">Paratype</typeStatus>
<specimenCount id="9D52FD3EF11FFFEF76A6FD9A57EC7D59" box="[899,920,586,609]" pageId="19" pageNumber="232">♀</specimenCount>
: same data as holotype, but “
<date id="FFEA1077F11FFFEF71D4FD99512C7D59" box="[1265,1368,585,609]" pageId="19" pageNumber="232" value="1988-09-03">
<collectingDate id="EFAEE99FF11FFFEF71D4FD99512C7D59" box="[1265,1368,585,609]" pageId="19" pageNumber="232" value="1988-09-03">3.IX.1988</collectingDate>
</date>
... [B174]” (
<collectionCode id="ED45AE72F11FFFEF765EFDB957A07DB9" box="[891,980,617,641]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="19" pageNumber="232" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEB36B7F11FFFEF763FFD7950877C39" blockId="19.[794,1398,681,769]" pageId="19" pageNumber="232">
<emphasis id="B920EAA5F11FFFEF763FFD7950177DF9" bold="true" box="[794,1123,681,705]" pageId="19" pageNumber="232">Additional material examined</emphasis>
:
<materialsCitation id="3B3C3CEAF11FFFEF714AFD79509B7C39" ID-GBIF-Occurrence="3450187302" collectingDate="1987-05-18" collectionCode="MHNG" collectorName="Burckhardt &amp; Lobl" country="Malaysia" elevation="1600" location="Crocker Range" municipality="Crocker Range" pageId="19" pageNumber="232" specimenCount="1" stateProvince="Sabah">
<emphasis id="B920EAA5F11FFFEF714AFD7950A47DF9" bold="true" box="[1135,1232,681,705]" pageId="19" pageNumber="232">
<collectingCountry id="F3437627F11FFFEF714AFD7950A47DF9" box="[1135,1232,681,705]" name="Malaysia" pageId="19" pageNumber="232">Malaysia</collectingCountry>
</emphasis>
:
<specimenCount id="9D52FD3EF11FFFEF71F9FD7951757DF9" box="[1244,1281,681,705]" pageId="19" pageNumber="232">1 ♂</specimenCount>
[identified as
<taxonomicName id="4C544D34F11FFFEF7610FD1A57CB7DD9" baseAuthorityName="PACE" baseAuthorityYear="1987" box="[821,959,713,737]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="adulescens">
<emphasis id="B920EAA5F11FFFEF7610FD1A57CB7DD9" box="[821,959,713,737]" italics="true" pageId="19" pageNumber="232">Z. adulescens</emphasis>
</taxonomicName>
by Pace],
<collectingRegion id="4990F855F11FFFEF710EFD1950197DD9" box="[1067,1133,713,737]" country="Malaysia" name="Sabah" pageId="19" pageNumber="232">Sabah</collectingRegion>
,
<collectingMunicipality id="6B8FACCDF11FFFEF7153FD19516C7DDA" box="[1142,1304,713,738]" country="Malaysia" name="Crocker Range" pageId="19" pageNumber="232" stateProvince="Sabah">Crocker Range</collectingMunicipality>
,
<quantity id="4CAC9B52F11FFFEF7004FD1951077DD9" box="[1313,1395,713,737]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="19" pageNumber="232" unit="m" value="1600.0">
<elevation id="0079D184F11FFFEF7004FD1951077DD9" box="[1313,1395,713,737]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="19" pageNumber="232" unit="m" value="1600.0">1600 m</elevation>
</quantity>
,
<date id="FFEA1077F11FFFEF763FFD3957F77C39" box="[794,899,745,769]" pageId="19" pageNumber="232" value="1987-05-18">
<collectingDate id="EFAEE99FF11FFFEF763FFD3957F77C39" box="[794,899,745,769]" pageId="19" pageNumber="232" value="1987-05-18">18.V.1987</collectingDate>
</date>
, leg.
<collectorName id="26A15361F11FFFEF769DFD3950457C39" box="[952,1073,745,769]" pageId="19" pageNumber="232">Burckhardt</collectorName>
&amp;
<collectorName id="26A15361F11FFFEF7175FD3950F47C39" box="[1104,1152,745,769]" pageId="19" pageNumber="232">Löbl</collectorName>
(
<collectionCode id="ED45AE72F11FFFEF71ABFD3950937C39" box="[1166,1255,745,769]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="19" pageNumber="232" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11FFFEF763FFCF950DF7B79" pageId="19" pageNumber="232" type="discussion">
<paragraph id="8BEB36B7F11FFFEF763FFCF950DF7B79" blockId="19.[794,1399,809,1089]" pageId="19" pageNumber="232">
<emphasis id="B920EAA5F11FFFEF763FFCF957FD7C79" bold="true" box="[794,905,809,833]" pageId="19" pageNumber="232">Comment</emphasis>
: The original description of
<taxonomicName id="4C544D34F11FFFEF71FAFCFA51027C79" authorityName="Assing" authorityYear="2017" box="[1247,1398,809,833]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="daiaccorum">
<emphasis id="B920EAA5F11FFFEF71FAFCFA51027C79" box="[1247,1398,809,833]" italics="true" pageId="19" pageNumber="232">Z. daiaccorum</emphasis>
</taxonomicName>
is based on a female
<typeStatus id="54EF8815F11FFFEF713DFC9950027C59" box="[1048,1142,841,865]" pageId="19" pageNumber="232" type="holotype">holotype</typeStatus>
from “Borneo,
<collectingRegion id="4990F855F11FFFEF7014FC9951077C59" box="[1329,1395,841,865]" country="Malaysia" name="Sabah" pageId="19" pageNumber="232">Sabah</collectingRegion>
, Mt. Kanabalu Nat. Pd., HQ Liwagu River” and
<specimenCount id="9D52FD3EF11FFFEF7065FCB9571E7C99" pageId="19" pageNumber="232" type="female">three females</specimenCount>
from the same locality (
<bibRefCitation id="EFC54B46F11FFFEF715CFC5950847C99" author="PACE, R." box="[1145,1264,905,929]" pageId="19" pageNumber="232" pagination="107 - 156" refId="ref27259" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">PACE 2008</bibRefCitation>
). An examination of the
<typeStatus id="54EF8815F11FFFEF768CFC7950737CF9" box="[937,1031,937,961]" pageId="19" pageNumber="232" type="holotype">holotype</typeStatus>
and a
<typeStatus id="54EF8815F11FFFEF7168FC7A50DE7CFA" box="[1101,1194,938,962]" pageId="19" pageNumber="232" type="paratype">paratype</typeStatus>
revealed that these specimens are conspecific with the
<typeStatus id="54EF8815F11FFFEF71AAFC1950997CD9" box="[1167,1261,969,993]" pageId="19" pageNumber="232" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C544D34F11FFFEF7028FC1A57237B39" authorityName="Cameron" authorityYear="1943" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="matangensis">
<emphasis id="B920EAA5F11FFFEF7028FC1A57237B39" italics="true" pageId="19" pageNumber="232">Z. matangensis</emphasis>
</taxonomicName>
, which was revised by
<bibRefCitation id="EFC54B46F11FFFEF716FFC3950937B39" author="ASSING, V." box="[1098,1255,1001,1025]" pageId="19" pageNumber="232" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING (2017)</bibRefCitation>
. The external and sexual characters of the
<typeStatus id="54EF8815F11FFFEF7170FBD950C77B19" box="[1109,1203,1033,1057]" pageId="19" pageNumber="232" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C544D34F11FFFEF71FFFBDA51037B19" authorityName="Cameron" authorityYear="1943" box="[1242,1399,1034,1057]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="matangensis">
<emphasis id="B920EAA5F11FFFEF71FFFBDA51037B19" box="[1242,1399,1034,1057]" italics="true" pageId="19" pageNumber="232">Z. matangensis</emphasis>
</taxonomicName>
are illustrated in
<figureCitation id="136F2A32F11FFFEF76E8FBF950407B79" box="[973,1076,1065,1089]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="19" pageNumber="232">Figs 5, 26</figureCitation>
,
<figureCitation id="136F2A32F11FFFEF7118FBF9502C7B79" box="[1085,1112,1065,1089]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="19" pageNumber="232">41</figureCitation>
,
<figureCitation id="136F2A32F11FFFEF7146FBF950D37B79" box="[1123,1191,1065,1089]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="19" pageNumber="232">63–65</figureCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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