437 lines
58 KiB
XML
437 lines
58 KiB
XML
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<mods:title id="AA46CAB99754923D7347580A88F8EB1E">On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<treatment id="03FD87A1F11FFFE8761AFB4350BC7AB9" ID-DOI="http://doi.org/10.5281/zenodo.5888802" ID-GBIF-Taxon="192500447" ID-Zenodo-Dep="5888802" LSID="urn:lsid:plazi:treatment:03FD87A1F11FFFE8761AFB4350BC7AB9" httpUri="http://treatment.plazi.org/id/03FD87A1F11FFFE8761AFB4350BC7AB9" lastPageId="20" lastPageNumber="233" pageId="19" pageNumber="232">
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<subSubSection id="C34E653CF11FFFEF761AFB4350CD7B96" box="[831,1209,1171,1198]" pageId="19" pageNumber="232" type="nomenclature">
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<paragraph id="8BEB36B7F11FFFEF761AFB4350CD7B96" blockId="19.[831,1209,1171,1231]" box="[831,1209,1171,1198]" pageId="19" pageNumber="232">
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<heading id="D0A381DBF11FFFEF761AFB4350CD7B96" box="[831,1209,1171,1198]" centered="true" fontSize="12" level="2" pageId="19" pageNumber="232" reason="2">
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<taxonomicName id="4C544D34F11FFFEF761AFB4350CD7B96" ID-CoL="5DDBB" authority="CAMERON, 1943" authorityName="Cameron" authorityYear="1943" box="[831,1209,1171,1198]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="bryanti" subGenus="Zyras">
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<emphasis id="B920EAA5F11FFFEF761AFB4357017B96" box="[831,885,1171,1198]" italics="true" pageId="19" pageNumber="232">Zyras</emphasis>
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(
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<emphasis id="B920EAA5F11FFFEF76A6FB4357CD7B96" box="[899,953,1171,1198]" italics="true" pageId="19" pageNumber="232">Zyras</emphasis>
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)
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<emphasis id="B920EAA5F11FFFEF76E2FB43507A7B96" box="[967,1038,1171,1198]" italics="true" pageId="19" pageNumber="232">bryanti</emphasis>
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<bibRefCitation id="EFC54B46F11FFFEF7130FB4350CD7B96" author="CAMERON, M." box="[1045,1209,1171,1198]" pageId="19" pageNumber="232" pagination="139 - 143" refId="ref26208" refString="CAMERON, M. 1943: New species of Staphylinidae (Col.) from Borneo. - Entomologist's Monthly Magazine 79: 39 - 42, 139 - 143, 183." type="journal article" year="1943">CAMERON, 1943</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E653CF11FFFEF761AFB6B57937BF7" box="[831,999,1211,1231]" pageId="19" pageNumber="232" type="description">
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<paragraph id="8BEB36B7F11FFFEF761AFB6B57937BF7" blockId="19.[831,1209,1171,1231]" box="[831,999,1211,1231]" pageId="19" pageNumber="232">
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(
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<figureCitation id="136F2A32F11FFFEF7666FB6B57FB7BF7" box="[835,911,1211,1231]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="19" pageNumber="232">Figs 6, 23</figureCitation>
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,
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<figureCitation id="136F2A32F11FFFEF76B3FB6B57DF7BF7" box="[918,939,1211,1231]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="19" pageNumber="232">38</figureCitation>
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,
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<figureCitation id="136F2A32F11FFFEF7697FB6B57977BF7" box="[946,995,1211,1231]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="19" pageNumber="232">59–62</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E653CF11FFFEF763FFB3C51757A78" pageId="19" pageNumber="232" type="reference_group">
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<paragraph id="8BEB36B7F11FFFEF763FFB3C50B77A39" blockId="19.[794,1281,1259,1345]" box="[794,1219,1259,1281]" pageId="19" pageNumber="232">
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<treatmentCitationGroup id="AB441199F11FFFEF763FFB3C50B77A39" box="[794,1219,1259,1281]" pageId="19" pageNumber="232">
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<treatmentCitation id="0AF510A6F11FFFEF763FFB3C50C57A38" author="CAMERON, M." box="[794,1201,1259,1281]" page="141" pageId="19" pageNumber="232" year="1943">
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<taxonomicName id="4C544D34F11FFFEF763FFB3C50C57A38" ID-CoL="5DDBB" authority="CAMERON, 1943: 141" authorityName="Cameron" authorityPageNumber="141" authorityYear="1943" box="[794,1201,1259,1281]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="bryanti" subGenus="Zyras">
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<emphasis id="B920EAA5F11FFFEF763FFB3C573A7A39" box="[794,846,1260,1281]" italics="true" pageId="19" pageNumber="232">Zyras</emphasis>
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(
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||
<emphasis id="B920EAA5F11FFFEF7679FB3C57E47A39" box="[860,912,1260,1281]" italics="true" pageId="19" pageNumber="232">Zyras</emphasis>
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)
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<emphasis id="B920EAA5F11FFFEF76B8FB3B57967A38" box="[925,994,1259,1280]" italics="true" pageId="19" pageNumber="232">bryanti</emphasis>
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<bibRefCitation id="EFC54B46F11FFFEF76C2FB3B50C57A38" author="CAMERON, M." box="[999,1201,1259,1280]" pageId="19" pageNumber="232" pagination="139 - 143" refId="ref26208" refString="CAMERON, M. 1943: New species of Staphylinidae (Col.) from Borneo. - Entomologist's Monthly Magazine 79: 39 - 42, 139 - 143, 183." type="journal article" year="1943">
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||
CAMERON, 1943:
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<specimenCount id="9D52FD3EF11FFFEF71ABFB3B50C57A38" box="[1166,1201,1259,1280]" pageId="19" pageNumber="232" type="generic">141</specimenCount>
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</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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f.
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||
</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="8BEB36B7F11FFFEF763FFADC50897A18" blockId="19.[794,1281,1259,1345]" box="[794,1277,1291,1313]" pageId="19" pageNumber="232">
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<treatmentCitationGroup id="AB441199F11FFFEF763FFADC50897A18" box="[794,1277,1291,1313]" pageId="19" pageNumber="232">
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<treatmentCitation id="0AF510A6F11FFFEF763FFADC50EB7A18" author="PACE, R." box="[794,1183,1291,1313]" page="99" pageId="19" pageNumber="232" year="1990">
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<taxonomicName id="4C544D34F11FFFEF763FFADC50EB7A18" authority="PACE, 1990: 99" authorityName="PACE" authorityPageNumber="99" authorityYear="1990" box="[794,1183,1291,1313]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="mortuorum" subGenus="Zyras">
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||
<emphasis id="B920EAA5F11FFFEF763FFADC573A7A19" box="[794,846,1292,1313]" italics="true" pageId="19" pageNumber="232">Zyras</emphasis>
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||
(
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||
<emphasis id="B920EAA5F11FFFEF7679FADC57E47A19" box="[860,912,1292,1313]" italics="true" pageId="19" pageNumber="232">Zyras</emphasis>
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||
)
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||
<emphasis id="B920EAA5F11FFFEF76B8FADC507F7A19" box="[925,1035,1292,1313]" italics="true" pageId="19" pageNumber="232">mortuorum</emphasis>
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||
<bibRefCitation id="EFC54B46F11FFFEF7134FADB50EB7A18" author="PACE, R." box="[1041,1183,1291,1312]" pageId="19" pageNumber="232" pagination="57 - 113" refId="ref26872" refString="PACE, R. 1990: Aleocharinae delle Filippine. 82 ° contributo alla conoscenza delle Aleocharinae. - BERTI, N. (ed.): Miscellanees sur les Staphylins. - Memoires du Museum National d'Histoire Naturelle (A, Zoologie) 147: 57 - 113." type="journal article" year="1990">PACE, 1990: 99</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
;
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<emphasis id="B920EAA5F11FFFEF718DFADB50897A18" bold="true" box="[1192,1277,1291,1312]" pageId="19" pageNumber="232">syn. nov.</emphasis>
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</treatmentCitationGroup>
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</paragraph>
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||
<paragraph id="8BEB36B7F11FFFEF763FFAFC51757A78" blockId="19.[794,1281,1259,1345]" box="[794,1281,1323,1345]" pageId="19" pageNumber="232">
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<treatmentCitationGroup id="AB441199F11FFFEF763FFAFC51757A78" box="[794,1281,1323,1345]" pageId="19" pageNumber="232">
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<treatmentCitation id="0AF510A6F11FFFEF763FFAFC50D67A79" author="PACE, R." box="[794,1186,1323,1345]" page="153" pageId="19" pageNumber="232" year="2008">
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<taxonomicName id="4C544D34F11FFFEF763FFAFC50D67A79" authority="PACE, 2008: 153" authorityName="PACE" authorityPageNumber="153" authorityYear="2008" box="[794,1186,1323,1345]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="paederinus" subGenus="Zyras">
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||
<emphasis id="B920EAA5F11FFFEF763FFAFC573A7A79" box="[794,846,1324,1345]" italics="true" pageId="19" pageNumber="232">Zyras</emphasis>
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(
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||
<emphasis id="B920EAA5F11FFFEF7679FAFC57E47A79" box="[860,912,1324,1345]" italics="true" pageId="19" pageNumber="232">Zyras</emphasis>
|
||
)
|
||
<emphasis id="B920EAA5F11FFFEF76B8FAFB50707A78" box="[925,1028,1323,1344]" italics="true" pageId="19" pageNumber="232">paederinus</emphasis>
|
||
<bibRefCitation id="EFC54B46F11FFFEF712FFAFB50D67A79" author="PACE, R." box="[1034,1186,1323,1345]" pageId="19" pageNumber="232" pagination="107 - 156" refId="ref27259" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">PACE, 2008: 153</bibRefCitation>
|
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</taxonomicName>
|
||
</treatmentCitation>
|
||
;
|
||
<emphasis id="B920EAA5F11FFFEF7189FAFB51757A78" bold="true" box="[1196,1281,1323,1344]" pageId="19" pageNumber="232">syn. nov.</emphasis>
|
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</treatmentCitationGroup>
|
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</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E653CF11FFFEF763FFAB9504678B9" pageId="19" pageNumber="232" type="materials_examined">
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<paragraph id="8BEB36B7F11FFFEF763FFAB957F77939" blockId="19.[794,1399,1385,1921]" pageId="19" pageNumber="232">
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||
<emphasis id="B920EAA5F11FFFEF763FFAB950357AB9" bold="true" box="[794,1089,1385,1409]" pageId="19" pageNumber="232">Type material examined</emphasis>
|
||
:
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||
<taxonomicName id="4C544D34F11FFFEF717BFABA50B27AB9" authorityName="Cameron" authorityYear="1943" box="[1118,1222,1385,1409]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="bryanti">
|
||
<emphasis id="B920EAA5F11FFFEF717BFABA50B27AB9" box="[1118,1222,1385,1409]" italics="true" pageId="19" pageNumber="232">Z. bryanti</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<materialsCitation id="3B3C3CEAF11FFFEF71C7FAB957F77939" ID-GBIF-Occurrence="3450187338" collectingDate="2014-01-22" collectionCode="BMNH" collectorName="M. Cameron & Bequest. B. M." determinerName="V. Assing" elevation="610" location="Matang" municipality="Matang" pageId="19" pageNumber="232" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54EF8815F11FFFEF71C7FAB951317AB9" box="[1250,1349,1385,1409]" pageId="19" pageNumber="232" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D52FD3EF11FFFEF7079FABA51057AB9" box="[1372,1393,1386,1409]" pageId="19" pageNumber="232" type="generic">♂</specimenCount>
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: “
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<collectingMunicipality id="6B8FACCDF11FFFEF7606FA5957037A99" box="[803,887,1417,1441]" pageId="19" pageNumber="232">Matang</collectingMunicipality>
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,
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<date id="FFEA1077F11FFFEF76A1FA5957BA7A99" box="[900,974,1417,1441]" pageId="19" pageNumber="232" value="2014-01-22">
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||
<collectingDate id="EFAEE99FF11FFFEF76A1FA5957BA7A99" box="[900,974,1417,1441]" pageId="19" pageNumber="232" value="2014-01-22">22.I.14</collectingDate>
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||
</date>
|
||
,
|
||
<quantity id="4CAC9B52F11FFFEF76F9FA5950517A99" box="[988,1061,1417,1441]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="19" pageNumber="232" unit="ft" value="2000.0">
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<elevation id="0079D184F11FFFEF76F9FA5950517A99" box="[988,1061,1417,1441]" metricMagnitude="2" metricUnit="m" metricValue="6.096" pageId="19" pageNumber="232" unit="ft" value="2000.0">2000 ft</elevation>
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</quantity>
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/ G 28 /
|
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<taxonomicName id="4C544D34F11FFFEF71ABFA5A514E7A99" authority="Cam." authorityName="Cameron" authorityYear="1943" box="[1166,1338,1417,1441]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="bryanti">
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<emphasis id="B920EAA5F11FFFEF71ABFA5A50827A99" box="[1166,1270,1417,1441]" italics="true" pageId="19" pageNumber="232">Z. bryanti</emphasis>
|
||
Cam.
|
||
</taxonomicName>
|
||
Type / Holotype / M. Cameron. Bequest. B.M. 1955-147 / Holotypus
|
||
<taxonomicName id="4C544D34F11FFFEF76B7FA1A50E57AD9" authority="Cameron" authorityName="Cameron" authorityYear="1943" box="[914,1169,1481,1505]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="bryanti">
|
||
<emphasis id="B920EAA5F11FFFEF76B7FA1A50557AD9" box="[914,1057,1481,1505]" italics="true" pageId="19" pageNumber="232">Zyras bryanti</emphasis>
|
||
Cameron
|
||
</taxonomicName>
|
||
, rev.
|
||
<determinerName id="7E37DA73F11FFFEF71E8FA1A515B7AD9" box="[1229,1327,1481,1505]" pageId="19" pageNumber="232">
|
||
V.
|
||
<bibRefCitation id="EFC54B46F11FFFEF71CDFA19515B7AD9" author="ASSING, V." box="[1256,1327,1481,1505]" pageId="19" pageNumber="232" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing</bibRefCitation>
|
||
</determinerName>
|
||
2017” (
|
||
<collectionCode id="ED45AE72F11FFFEF7604FA3957037939" box="[801,887,1513,1537]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="19" pageNumber="232">BMNH</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F11FFFEF763FF9DA503679F9" blockId="19.[794,1399,1385,1921]" pageId="19" pageNumber="232">
|
||
<materialsCitation id="3B3C3CEAF11FFFEF763FF9DA504B79F9" ID-GBIF-Occurrence="3450194302" collectingDate="1979-12-19" collectorName="L. Deharveng" country="Philippines" location="SAGADA" pageId="19" pageNumber="232" specimenCount="1" typeStatus="holotype">
|
||
<taxonomicName id="4C544D34F11FFFEF763FF9DA57C57919" authorityName="PACE" authorityYear="1990" box="[794,945,1546,1569]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="mortuorum">
|
||
<emphasis id="B920EAA5F11FFFEF763FF9DA57C57919" box="[794,945,1546,1569]" italics="true" pageId="19" pageNumber="232">
|
||
<originalDetermination id="A84A8E4BF11FFFEF763FF9DA57C57919" box="[794,945,1546,1569]" pageId="19" pageNumber="232">Z. mortuorum</originalDetermination>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<typeStatus id="54EF8815F11FFFEF76E5F9D950577919" box="[960,1059,1545,1569]" pageId="19" pageNumber="232" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="9D52FD3EF11FFFEF710BF9DA50377919" box="[1070,1091,1546,1569]" pageId="19" pageNumber="232">♂</specimenCount>
|
||
: “
|
||
<collectingCountry id="F3437627F11FFFEF7179F9D9508E7919" box="[1116,1274,1545,1569]" name="Philippines" pageId="19" pageNumber="232">PHILIPPINES</collectingCountry>
|
||
,
|
||
<location id="8E8B606CF11FFFEF7022F9D951067919" LSID="urn:lsid:plazi:treatment:03FD87A1F11FFFE8761AFB4350BC7AB9:8E8B606CF11FFFEF7022F9D951067919" box="[1287,1394,1545,1569]" country="Philippines" name="SAGADA" pageId="19" pageNumber="232">SAGADA</location>
|
||
,
|
||
<date id="FFEA1077F11FFFEF763FF9F9570E7979" box="[794,890,1577,1601]" pageId="19" pageNumber="232" value="1979-12-19">
|
||
<collectingDate id="EFAEE99FF11FFFEF763FF9F9570E7979" box="[794,890,1577,1601]" pageId="19" pageNumber="232" value="1979-12-19">19/12/79</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26A15361F11FFFEF76A6F9FA50677979" box="[899,1043,1577,1601]" pageId="19" pageNumber="232">L. Deharveng</collectorName>
|
||
/ En Aval Village, lavage d’humus / Holotypus
|
||
<taxonomicName id="4C544D34F11FFFEF768CF99A501D7959" authorityName="PACE" authorityYear="1990" box="[937,1129,1610,1633]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="mortuorum">
|
||
<emphasis id="B920EAA5F11FFFEF768CF99A501D7959" box="[937,1129,1610,1633]" italics="true" pageId="19" pageNumber="232">Zyras mortuorum</emphasis>
|
||
</taxonomicName>
|
||
m., det. R. Pace 1985 /
|
||
<taxonomicName id="4C544D34F11FFFEF763FF9BA57A879B9" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[794,988,1642,1665]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="mortuorum" status="sp. nov.">
|
||
<emphasis id="B920EAA5F11FFFEF763FF9BA57A879B9" box="[794,988,1642,1665]" italics="true" pageId="19" pageNumber="232">Zyras mortuorum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21357DEF11FFFEF76CDF9BA505479B9" box="[1000,1056,1642,1665]" pageId="19" pageNumber="232" rank="species">sp. n.</taxonomicNameLabel>
|
||
, det. R. Pace 1985 / Museum Paris 1998, Coll. J. Orousset /
|
||
<taxonomicName id="4C544D34F11FFFEF7175F95A51327999" authority="Cameron" authorityName="Cameron" authorityYear="1943" box="[1104,1350,1673,1697]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="bryanti">
|
||
<emphasis id="B920EAA5F11FFFEF7175F95A50AF7999" box="[1104,1243,1673,1697]" italics="true" pageId="19" pageNumber="232">Zyras bryanti</emphasis>
|
||
Cameron
|
||
</taxonomicName>
|
||
, det. V.
|
||
<bibRefCitation id="EFC54B46F11FFFEF7610F97957C379F9" author="ASSING, V." box="[821,951,1705,1729]" pageId="19" pageNumber="232" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing 2017</bibRefCitation>
|
||
” (MNHNP)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F11FFFEF763FF91A504678B9" blockId="19.[794,1399,1385,1921]" pageId="19" pageNumber="232">
|
||
<materialsCitation id="3B3C3CEAF11FFFEF763FF91A504678B9" ID-GBIF-Occurrence="3450194301" collectingDate="1987-05-09" collectionCode="MHNG" collectorName="A. Smetana /" country="BORNEO" determinerName="V. Assing" elevation="520" location="Kinabalu N. P." pageId="19" pageNumber="232" specimenCount="1" typeStatus="holotype">
|
||
<taxonomicName id="4C544D34F11FFFEF763FF91A57DC79D9" authorityName="PACE" authorityYear="2008" box="[794,936,1737,1761]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="paederinus">
|
||
<emphasis id="B920EAA5F11FFFEF763FF91A57DC79D9" box="[794,936,1737,1761]" italics="true" pageId="19" pageNumber="232">
|
||
<originalDetermination id="A84A8E4BF11FFFEF763FF91A57DC79D9" box="[794,936,1737,1761]" pageId="19" pageNumber="232">Z. paederinus</originalDetermination>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<typeStatus id="54EF8815F11FFFEF76E5F919505779D9" box="[960,1059,1737,1761]" pageId="19" pageNumber="232" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="9D52FD3EF11FFFEF7113F91A503F79D9" box="[1078,1099,1738,1761]" pageId="19" pageNumber="232">♂</specimenCount>
|
||
: “
|
||
<collectingCountry id="F3437627F11FFFEF7149F91950AC79D9" box="[1132,1240,1737,1761]" pageId="19" pageNumber="232">BORNEO</collectingCountry>
|
||
SABAH Mt.
|
||
<location id="8E8B606CF11FFFEF763FF93957C27839" LSID="urn:lsid:plazi:treatment:03FD87A1F11FFFE8761AFB4350BC7AB9:8E8B606CF11FFFEF763FF93957C27839" box="[794,950,1769,1793]" country="BORNEO" name="Kinabalu N. P." pageId="19" pageNumber="232">Kinabalu N. P.</location>
|
||
,
|
||
<location id="8E8B606CF11FFFEF76E5F93950A17839" LSID="urn:lsid:plazi:treatment:03FD87A1F11FFFE8761AFB4350BC7AB9:8E8B606CF11FFFEF76E5F93950A17839" box="[960,1237,1769,1793]" country="BORNEO" name="above Poring Hot Springs" pageId="19" pageNumber="232">above Poring Hot Springs</location>
|
||
,
|
||
<quantity id="4CAC9B52F11FFFEF71C5F93951507839" box="[1248,1316,1769,1793]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="19" pageNumber="232" unit="m" value="520.0">
|
||
<elevation id="0079D184F11FFFEF71C5F93951507839" box="[1248,1316,1769,1793]" metricMagnitude="2" metricUnit="m" metricValue="5.2" pageId="19" pageNumber="232" unit="m" value="520.0">520 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFEA1077F11FFFEF700AF93951067839" box="[1327,1394,1769,1793]" pageId="19" pageNumber="232" value="1987-05-09">
|
||
<collectingDate id="EFAEE99FF11FFFEF700AF93951067839" box="[1327,1394,1769,1793]" pageId="19" pageNumber="232" value="1987-05-09">9.V.87</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26A15361F11FFFEF763FF8D957D87819" box="[794,940,1801,1825]" pageId="19" pageNumber="232">A. Smetana /</collectorName>
|
||
Holotypus
|
||
<taxonomicName id="4C544D34F11FFFEF711EF8DA50837819" authorityName="PACE" authorityYear="2008" box="[1083,1271,1801,1825]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="paederinus">
|
||
<emphasis id="B920EAA5F11FFFEF711EF8DA50837819" box="[1083,1271,1801,1825]" italics="true" pageId="19" pageNumber="232">Zyras paederinus</emphasis>
|
||
</taxonomicName>
|
||
mihi, det. R. Pace 2000 /
|
||
<taxonomicName id="4C544D34F11FFFEF7699F8FA50047879" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[956,1136,1833,1857]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="paederinus" status="sp. nov.">
|
||
<emphasis id="B920EAA5F11FFFEF7699F8FA50047879" box="[956,1136,1833,1857]" italics="true" pageId="19" pageNumber="232">Zyras paederinus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21357DEF11FFFEF715DF8FA50C77879" box="[1144,1203,1834,1857]" pageId="19" pageNumber="232" rank="species">n. sp.</taxonomicNameLabel>
|
||
, det. R. Pace 2000 / MHNG ENTO 00010006 /
|
||
<taxonomicName id="4C544D34F11FFFEF7169F89A51307859" authority="Cameron" authorityName="Cameron" authorityYear="1943" box="[1100,1348,1865,1889]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="bryanti">
|
||
<emphasis id="B920EAA5F11FFFEF7169F89A50AC7859" box="[1100,1240,1865,1889]" italics="true" pageId="19" pageNumber="232">Zyras bryanti</emphasis>
|
||
Cameron
|
||
</taxonomicName>
|
||
, det.
|
||
<determinerName id="7E37DA73F11FFFEF763FF8BA570878B9" box="[794,892,1897,1921]" pageId="19" pageNumber="232">
|
||
V.
|
||
<bibRefCitation id="EFC54B46F11FFFEF7610F8B9570878B9" author="ASSING, V." box="[821,892,1897,1921]" pageId="19" pageNumber="232" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing</bibRefCitation>
|
||
</determinerName>
|
||
2017” (
|
||
<collectionCode id="ED45AE72F11FFFEF76EBF8B9505378B9" box="[974,1063,1897,1921]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="19" pageNumber="232" type="Museum">MHNG</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E653CF11FFFE8763FF87955A17C19" lastPageId="20" lastPageNumber="233" pageId="19" pageNumber="232" type="discussion">
|
||
<paragraph id="8BEB36B7F11FFFE8763FF87955947D59" blockId="19.[794,1399,1961,2082]" lastBlockId="20.[189,794,233,609]" lastPageId="20" lastPageNumber="233" pageId="19" pageNumber="232">
|
||
<emphasis id="B920EAA5F11FFFEF763FF87957FD78F9" bold="true" box="[794,905,1961,1985]" pageId="19" pageNumber="232">Comment</emphasis>
|
||
: The original description of
|
||
<taxonomicName id="4C544D34F11FFFEF7190F87A516978F9" authorityName="Cameron" authorityYear="1943" box="[1205,1309,1961,1985]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="bryanti">
|
||
<emphasis id="B920EAA5F11FFFEF7190F87A516978F9" box="[1205,1309,1961,1985]" italics="true" pageId="19" pageNumber="232">Z. bryanti</emphasis>
|
||
</taxonomicName>
|
||
is based on a unique
|
||
<typeStatus id="54EF8815F11FFFEF768EF819507D78D9" box="[939,1033,1993,2017]" pageId="19" pageNumber="232" type="holotype">holotype</typeStatus>
|
||
from “W.
|
||
<collectingCountry id="F3437627F11FFFEF71A6F81950AB78D9" box="[1155,1247,1993,2017]" name="Malaysia" pageId="19" pageNumber="232">Sarawak</collectingCountry>
|
||
: Mt. Matang” (
|
||
<bibRefCitation id="EFC54B46F11FFFEF7606F83957BA7739" author="CAMERON, M." box="[803,974,2025,2049]" pageId="19" pageNumber="232" pagination="139 - 143" refId="ref26208" refString="CAMERON, M. 1943: New species of Staphylinidae (Col.) from Borneo. - Entomologist's Monthly Magazine 79: 39 - 42, 139 - 143, 183." type="journal article" year="1943">CAMERON 1943</bibRefCitation>
|
||
), that of
|
||
<taxonomicName id="4C544D34F11FFFEF710BF83A50B27739" authorityName="Assing" authorityYear="2017" box="[1070,1222,2026,2049]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="232" phylum="Arthropoda" rank="species" species="mortuorum">
|
||
<emphasis id="B920EAA5F11FFFEF710BF83A50B27739" box="[1070,1222,2026,2049]" italics="true" pageId="19" pageNumber="232">Z. mortuorum</emphasis>
|
||
</taxonomicName>
|
||
on a male
|
||
<typeStatus id="54EF8815F11FFFEF7018F839573C771A" pageId="19" pageNumber="232" type="holotype">holotype</typeStatus>
|
||
and a male
|
||
<typeStatus id="54EF8815F11FFFEF76C7F7DA504B771A" box="[994,1087,2058,2082]" pageId="19" pageNumber="232" type="paratype">paratype</typeStatus>
|
||
from “environs de Sagada” (
|
||
<bibRefCitation id="EFC54B46F118FFE875E3FF39554D7E39" author="PACE, R." box="[198,313,233,257]" pageId="20" pageNumber="233" pagination="57 - 113" refId="ref26872" refString="PACE, R. 1990: Aleocharinae delle Filippine. 82 ° contributo alla conoscenza delle Aleocharinae. - BERTI, N. (ed.): Miscellanees sur les Staphylins. - Memoires du Museum National d'Histoire Naturelle (A, Zoologie) 147: 57 - 113." type="journal article" year="1990">PACE 1990</bibRefCitation>
|
||
), and that of
|
||
<taxonomicName id="4C544D34F118FFE874ECFF3A56237E39" authorityName="Assing" authorityYear="2017" box="[457,599,233,257]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="paederinus">
|
||
<emphasis id="B920EAA5F118FFE874ECFF3A56237E39" box="[457,599,233,257]" italics="true" pageId="20" pageNumber="233">Z. paederinus</emphasis>
|
||
</taxonomicName>
|
||
on a unique male from “
|
||
<collectingRegion id="4990F855F118FFE87420FED955337E19" box="[261,327,265,289]" country="Malaysia" name="Sabah" pageId="20" pageNumber="233">Sabah</collectingRegion>
|
||
, Mt. Kinabalu, above Poring Hot Springs” (
|
||
<bibRefCitation id="EFC54B46F118FFE875E3FEF9554E7E79" author="PACE, R." box="[198,314,297,321]" pageId="20" pageNumber="233" pagination="107 - 156" refId="ref27259" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">PACE 2008</bibRefCitation>
|
||
). An examination of the
|
||
<typeStatus id="54EF8815F118FFE87747FEF956BD7E79" box="[610,713,297,321]" pageId="20" pageNumber="233" type="holotype">holotypes</typeStatus>
|
||
of the three names revealed that they are conspecific. Hence the synonymies proposed above. The
|
||
<typeStatus id="54EF8815F118FFE87702FEB956F17EB9" box="[551,645,361,385]" pageId="20" pageNumber="233" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C544D34F118FFE87782FEBA549E7E99" authorityName="Assing" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="mortuorum">
|
||
<emphasis id="B920EAA5F118FFE87782FEBA549E7E99" italics="true" pageId="20" pageNumber="233">Z. mortuorum</emphasis>
|
||
</taxonomicName>
|
||
has the antennae somewhat darker (antennomeres III–X dark-brown, distinctly contrasting with the reddish-yellow antennomere XI), more extensively infuscate tergites VI and VII, and the ventral process of the aedeagus slightly more acute in ventral view, but otherwise no evidence was found suggesting that it should represent a distinct species.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F118FFE87598FD5955A17C19" blockId="20.[189,794,649,805]" pageId="20" pageNumber="233">
|
||
<emphasis id="B920EAA5F118FFE87598FD5956637D99" bold="true" box="[189,535,649,673]" pageId="20" pageNumber="233">Additional material examined</emphasis>
|
||
:
|
||
<materialsCitation id="3B3C3CEAF118FFE8770EFD59576E7DD9" ID-GBIF-Occurrence="3450187333" collectingDate="1996-01-01" collectingDateMax="1996-01-04" collectingDateMin="1996-01-01" collectorName="Schulz & Vock" country="Thailand" elevation="300" location="Thale Ban National Park" municipality="Thale Ban National Park" pageId="20" pageNumber="233" specimenCount="1" stateProvince="Satun Province">
|
||
<emphasis id="B920EAA5F118FFE8770EFD5956FB7D99" bold="true" box="[555,655,649,673]" pageId="20" pageNumber="233">
|
||
<collectingCountry id="F3437627F118FFE8770EFD5956FB7D99" box="[555,655,649,673]" name="Thailand" pageId="20" pageNumber="233">Thailand</collectingCountry>
|
||
</emphasis>
|
||
: 1 ♀,
|
||
<collectingRegion id="4990F855F118FFE877F8FD5955697DF9" country="Thailand" name="Satun" pageId="20" pageNumber="233">Satun Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B8FACCDF118FFE87417FD79562E7DF9" box="[306,602,681,705]" country="Thailand" name="Thale Ban National Park" pageId="20" pageNumber="233" stateProvince="Satun">Thale Ban National Park</collectingMunicipality>
|
||
,
|
||
<locationDeviation id="9A592F95F118FFE8774BFD7956A57DF9" bearing="90.0" box="[622,721,681,705]" distance="20000.0" pageId="20" pageNumber="233">
|
||
<quantity id="4CAC9B52F118FFE8774BFD7956C57DF9" box="[622,689,681,705]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="20" pageNumber="233" unit="km" value="20.0">20 km</quantity>
|
||
E
|
||
</locationDeviation>
|
||
<collectingRegion id="4990F855F118FFE877F2FD7957627DF9" box="[727,790,681,705]" country="Thailand" name="Satun" pageId="20" pageNumber="233">Satun</collectingRegion>
|
||
,
|
||
<quantity id="4CAC9B52F118FFE87598FD1955427DD9" box="[189,310,713,737]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="20" pageNumber="233" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">
|
||
<elevation id="0079D184F118FFE87598FD1955427DD9" box="[189,310,713,737]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="20" pageNumber="233" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">200–400 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFEA1077F118FFE87469FD1955C97DD9" box="[332,445,713,737]" pageId="20" pageNumber="233" value="1996-01-01" valueMax="1996-01-04" valueMin="1996-01-01">
|
||
<collectingDate id="EFAEE99FF118FFE87469FD1955C97DD9" box="[332,445,713,737]" pageId="20" pageNumber="233" value="1996-01-01" valueMax="1996-01-04" valueMin="1996-01-01">1–4.I.1996</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26A15361F118FFE8772EFD1956257DD9" box="[523,593,713,737]" pageId="20" pageNumber="233">Schulz</collectorName>
|
||
&
|
||
<collectorName id="26A15361F118FFE877ADFD1956CA7DD9" box="[648,702,713,737]" pageId="20" pageNumber="233">Vock</collectorName>
|
||
(cAss).
|
||
</materialsCitation>
|
||
<materialsCitation id="3B3C3CEAF118FFE87598FD3955A57C19" ID-GBIF-Occurrence="3450187318" collectingDate="2006-06-03" collectingDateMax="1996-01-04" collectingDateMin="1996-01-01" collectorName="Ciampor" country="Malaysia" elevation="300" location="Gua Musang" municipality="Gua Musang" pageId="20" pageNumber="233" specimenCount="1" stateProvince="Kelantan">
|
||
<emphasis id="B920EAA5F118FFE87598FD39556A7C39" bold="true" box="[189,286,745,769]" pageId="20" pageNumber="233">
|
||
<collectingCountry id="F3437627F118FFE87598FD39556A7C39" box="[189,286,745,769]" name="Malaysia" pageId="20" pageNumber="233">Malaysia</collectingCountry>
|
||
</emphasis>
|
||
: 1 ♂, 1 ♀,
|
||
<collectingRegion id="4990F855F118FFE874B9FD3955897C39" box="[412,509,745,769]" country="Malaysia" name="Kelantan" pageId="20" pageNumber="233">Kelantan</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6B8FACCDF118FFE87728FD3956E87C3A" box="[525,668,745,770]" country="Malaysia" name="Gua Musang" pageId="20" pageNumber="233" stateProvince="Kelantan">Gua Musang</collectingMunicipality>
|
||
,
|
||
<date id="FFEA1077F118FFE87789FD3957627C39" box="[684,790,745,769]" pageId="20" pageNumber="233" value="2006-06-03">
|
||
<collectingDate id="EFAEE99FF118FFE87789FD3957627C39" box="[684,790,745,769]" pageId="20" pageNumber="233" value="2006-06-03">3.VI.2006</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="26A15361F118FFE875CDFCDA55337C1D" box="[232,327,778,805]" pageId="20" pageNumber="233">Čiampor</collectorName>
|
||
(cHla, cAss)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E653CF118FFE87598FC99514A7A39" pageId="20" pageNumber="233" type="description">
|
||
<paragraph id="8BEB36B7F118FFE87598FC9955137A39" blockId="20.[189,794,841,2081]" pageId="20" pageNumber="233">
|
||
<emphasis id="B920EAA5F118FFE87598FC99552E7C59" bold="true" box="[189,346,841,865]" pageId="20" pageNumber="233">Redescription</emphasis>
|
||
: Small species; body length
|
||
<quantity id="4CAC9B52F118FFE877B7FC9957617C59" box="[658,789,841,865]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" metricValueMax="4.8" metricValueMin="3.8" pageId="20" pageNumber="233" unit="mm" value="4.3" valueMax="4.8" valueMin="3.8">3.8–4.8 mm</quantity>
|
||
; length of forebody
|
||
<quantity id="4CAC9B52F118FFE8748EFCB9565B7CB9" box="[427,559,873,897]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="20" pageNumber="233" unit="mm" value="1.9" valueMax="2.1" valueMin="1.7">1.7–2.1 mm</quantity>
|
||
. Coloration (
|
||
<figureCitation id="136F2A32F118FFE877F4FCB954AD7C99" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="20" pageNumber="233">Figs 6, 23</figureCitation>
|
||
,
|
||
<figureCitation id="136F2A32F118FFE875C1FC5955767C99" box="[228,258,905,929]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="20" pageNumber="233">38</figureCitation>
|
||
): head brown to blackish-brown; pronotum pale-reddish; elytra yellowish red with the posterolateral portions more or less extensively and more or less distinctly, diffusely infuscate (sometimes leaving only the anterior margin and the scutellar portion paler); abdomen yellowish-red with most to nearly all of tergite VI (except for the anterior and antero-lateral portions) and the posterior two-fifths to four-fifths of tergite VII infuscate; antennae yellowish-red to darkbrown with the basal antennomeres dark-yellowish to reddish and antennomere XI reddish-yellow; maxillary palpi yellowish.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F118FFE87598FAD955367979" blockId="20.[189,794,841,2081]" pageId="20" pageNumber="233">
|
||
Head (
|
||
<figureCitation id="136F2A32F118FFE87428FAD9552E7A19" box="[269,346,1289,1313]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="20" pageNumber="233">Fig. 23</figureCitation>
|
||
) distinctly transverse, median portion extensively impunctate; punctures in lateral portions rather dense and moderately coarse. Eyes large, approximately twice as long as postocular region in dorsal view. Antenna (
|
||
<figureCitation id="136F2A32F118FFE87414FA5955047A99" box="[305,368,1417,1441]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="20" pageNumber="233">Fig. 6</figureCitation>
|
||
) long and very slender,
|
||
<quantity id="4CAC9B52F118FFE877BDFA59576E7A99" box="[664,794,1417,1441]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" pageId="20" pageNumber="233" unit="mm" value="1.9" valueMax="2.1" valueMin="1.7">1.7–2.1 mm</quantity>
|
||
long; antennomeres IV–VI approximately twice as long as broad (or nearly so), VII–X distinctly oblong, of gradually decreasing length, and decreasingly oblong, and XI approximately as long as the combined length of IX and X.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F118FFE87598F99A56A579F9" blockId="20.[189,794,841,2081]" pageId="20" pageNumber="233">
|
||
Pronotum (
|
||
<figureCitation id="136F2A32F118FFE8741DF99955F77959" box="[312,387,1609,1633]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="20" pageNumber="233">Fig. 23</figureCitation>
|
||
) 1.08–1.14 times as broad as long and 1.08–1.17 times as broad as head, broadest near anterior angles, weakly convex in cross-section; punctation very dense and fine; midline without impunctate band.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F118FFE87598F91956F67879" blockId="20.[189,794,841,2081]" pageId="20" pageNumber="233">
|
||
Elytra (
|
||
<figureCitation id="136F2A32F118FFE87429F919552379D9" box="[268,343,1737,1761]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="20" pageNumber="233">Fig. 23</figureCitation>
|
||
) nearly 0.83–0.89 times as long as pronotum; punctation very dense, slightly coarser than that of pronotum. Hind wings present. Metatarsomere I slender, shorter than the combined length of II–IV.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F118FFE87598F89957D47ED9" blockId="20.[189,794,841,2081]" lastBlockId="20.[831,1437,233,737]" pageId="20" pageNumber="233">
|
||
Abdomen (
|
||
<figureCitation id="136F2A32F118FFE87410F89955F47859" box="[309,384,1865,1889]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="20" pageNumber="233">Fig. 38</figureCitation>
|
||
) narrower than elytra, with deep anterior impressions on tergites III–V; anterior impressions of tergites III–V each with a transverse row of few very large, but weakly defined puncture-like impressions, with a lateral puncture on either side, and with four (tergites III and IV) or six (tergite V) fine setiferous punctures at posterior margin; tergite VI without non-setiferous punctures anteriorly, with a lateral setiferous puncture on either side, and with six setiferous punctures at posterior margin; tergite VII without non-setiferous punctures anteriorly, with two transverse rows each composed of four setiferous punctures posteriorly, posterior margin with palisade fringe; tergite VIII with setiferous punctation only in posterior fourth, posterior margin convex (
|
||
<figureCitation id="136F2A32F118FFE87662FE1957E07ED9" box="[839,916,457,481]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="20" pageNumber="233">Fig. 61</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F118FFE8761AFE3A57AD7D99" blockId="20.[831,1437,233,737]" pageId="20" pageNumber="233">
|
||
♂: posterior margin of sternite VIII convex (
|
||
<figureCitation id="136F2A32F118FFE87061FE3951E47D39" box="[1348,1424,489,513]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="20" pageNumber="233">Fig. 62</figureCitation>
|
||
); median lobe of aedeagus (
|
||
<figureCitation id="136F2A32F118FFE87155FDD950937D19" box="[1136,1255,521,545]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="20" pageNumber="233">Figs 59–60</figureCitation>
|
||
) nearly
|
||
<quantity id="4CAC9B52F118FFE8706EFDD951E97D19" box="[1355,1437,521,545]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="20" pageNumber="233" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long; ventral process slender, ventrally with or without a median carina; paramere slightly longer than median lobe, apical lobe very long and with conspicuously long subapical seta.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F118FFE8761AFD7A57CD7DD9" blockId="20.[831,1437,233,737]" pageId="20" pageNumber="233">♀: posterior margin of sternite VIII weakly concave in the middle.</paragraph>
|
||
<paragraph id="8BEB36B7F118FFE8761AFCD9514A7A39" blockId="20.[831,1436,777,1281]" pageId="20" pageNumber="233">
|
||
<emphasis id="B920EAA5F118FFE8761AFCD9507A7C19" bold="true" box="[831,1038,777,801]" pageId="20" pageNumber="233">Comparative notes</emphasis>
|
||
: Males of
|
||
<taxonomicName id="4C544D34F118FFE87153FCDA50AA7C19" authorityName="Cameron" authorityYear="1943" box="[1142,1246,777,801]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="bryanti">
|
||
<emphasis id="B920EAA5F118FFE87153FCDA50AA7C19" box="[1142,1246,777,801]" italics="true" pageId="20" pageNumber="233">Z. bryanti</emphasis>
|
||
</taxonomicName>
|
||
are readily distinguished from all other
|
||
<taxonomicName id="4C544D34F118FFE8711BFCFA500C7C79" authorityName="STEPHENS" authorityYear="1833" box="[1086,1144,810,833]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B920EAA5F118FFE8711BFCFA500C7C79" box="[1086,1144,810,833]" italics="true" pageId="20" pageNumber="233">Zyras</emphasis>
|
||
</taxonomicName>
|
||
sensu strictu species from the Palaearctic and Oriental regions by the morphology of the aedeagus. In addition, this species is characterized by the coloration, small body size, dense punctation of the forebody, the punctation pattern of the abdomen, and the long seta on the apical lobe of the paramere. As can be inferred from the similarly derived punctation of the forebody and the abdomen, as well as from the similarly slender antennae and habitus,
|
||
<taxonomicName id="4C544D34F118FFE871AFFBFA50867B79" authorityName="Cameron" authorityYear="1943" box="[1162,1266,1065,1089]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="bryanti">
|
||
<emphasis id="B920EAA5F118FFE871AFFBFA50867B79" box="[1162,1266,1065,1089]" italics="true" pageId="20" pageNumber="233">Z. bryanti</emphasis>
|
||
</taxonomicName>
|
||
is closely allied to
|
||
<taxonomicName id="4C544D34F118FFE87641FB9A50EB7B59" authority="ASSING, 2017" authorityName="ASSING" authorityYear="2017" box="[868,1183,1097,1121]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="densissimus">
|
||
<emphasis id="B920EAA5F118FFE87641FB9A578F7B59" box="[868,1019,1097,1121]" italics="true" pageId="20" pageNumber="233">Z. densissimus</emphasis>
|
||
<bibRefCitation id="EFC54B46F118FFE8712CFB9950EB7B59" author="ASSING, V." box="[1033,1183,1097,1121]" pageId="20" pageNumber="233" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">ASSING, 2017</bibRefCitation>
|
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</taxonomicName>
|
||
from
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<collectingRegion id="4990F855F118FFE871D4FB9951EC7B59" box="[1265,1432,1097,1121]" country="Indonesia" name="Sulawesi Utara" pageId="20" pageNumber="233">Sulawesi Utara</collectingRegion>
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, from which it differs by different coloration (
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<taxonomicName id="4C544D34F118FFE8700DFBBA57187B99" authorityName="ASSING" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="densissimus">
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<emphasis id="B920EAA5F118FFE8700DFBBA57187B99" italics="true" pageId="20" pageNumber="233">Z. densissimus</emphasis>
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</taxonomicName>
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||
: whole body of much darker coloration; pronotum blackish), less dense and less fine punctation of the forebody, and a less convex (cross-section) and posteriorly less strongly tapering pronotum (cross-section).
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E653CF118FFE87665FAF950BC7AB9" pageId="20" pageNumber="233" type="distribution">
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||
<paragraph id="8BEB36B7F118FFE87665FAF950BC7AB9" blockId="20.[832,1436,1321,1409]" pageId="20" pageNumber="233">
|
||
<emphasis id="B920EAA5F118FFE87665FAF957BD7A79" bold="true" box="[832,969,1321,1345]" pageId="20" pageNumber="233">Distribution</emphasis>
|
||
:
|
||
<taxonomicName id="4C544D34F118FFE876F0FAFA50167A79" authorityName="Cameron" authorityYear="1943" box="[981,1122,1321,1345]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="bryanti">
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<emphasis id="B920EAA5F118FFE876F0FAFA50167A79" box="[981,1122,1321,1345]" italics="true" pageId="20" pageNumber="233">Zyras bryanti</emphasis>
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</taxonomicName>
|
||
is currently known from one locality in Borneo, one in
|
||
<collectingCountry id="F3437627F118FFE8714DFA9950BE7A59" box="[1128,1226,1353,1377]" name="Thailand" pageId="20" pageNumber="233">Thailand</collectingCountry>
|
||
, one in Peninsular
|
||
<collectingCountry id="F3437627F118FFE87665FAB957D47AB9" box="[832,928,1385,1409]" name="Malaysia" pageId="20" pageNumber="233">Malaysia</collectingCountry>
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||
, and one in the
|
||
<collectingCountry id="F3437627F118FFE8716CFAB950B07AB9" box="[1097,1220,1385,1409]" name="Philippines" pageId="20" pageNumber="233">Philippines</collectingCountry>
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.
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</paragraph>
|
||
</subSubSection>
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||
</treatment>
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||
</document> |