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<document id="FFED8722A20A0A34AC9E7D667EF4D976" ID-CLB-Dataset="2470" ID-DOI="10.21248/contrib.entomol.67.2.213-246" ID-GBIF-Dataset="43c6cab4-bf7d-43c9-b9bf-b22c4ebf7924" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742339" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638267510724" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03FD87A1F11CFFED75C6FC4356D87E59" docLanguage="en" docName="ContribEntomol.67.2.213-246.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) latibasalis Assing 2017, spec. nov." docType="treatment" docUuid="35276B1A-6492-4F55-8836-91058719625C" docUuidSource="ZooBank" docVersion="12" lastPageNumber="230" masterDocId="FFC4FFD9F10CFFFC7525FFD054747F38" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="246" masterPageNumber="213" pageNumber="229" updateTime="1699471357828" updateUser="ExternalLinkService">
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<mods:title id="1515D15B40A80620FED6E6C4D1366A35">On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="C339AA10EBF1AD4539A9EF32A98E9E32">Assing, Volker</mods:namePart>
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<mods:title id="92C184B2089B0084BA1396F98ED593E0">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<mods:part id="CF6528626E2A62AB13D6EFCC67805A24">
<mods:date id="2C4A053191D63700D401FC24902B59F6">2017</mods:date>
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<mods:number id="94F3D4EA7E3C2DDD114C2B057B319F6E">2017-12-08</mods:number>
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<treatment id="03FD87A1F11CFFED75C6FC4356D87E59" ID-DOI="http://doi.org/10.5281/zenodo.5888796" ID-GBIF-Taxon="192500454" ID-Zenodo-Dep="5888796" LSID="urn:lsid:zoobank.org:act:35276B1A-6492-4F55-8836-91058719625C" httpUri="http://treatment.plazi.org/id/03FD87A1F11CFFED75C6FC4356D87E59" lastPageId="17" lastPageNumber="230" pageId="16" pageNumber="229">
<subSubSection id="C34E653CF11CFFEC75C6FC4356967CF7" pageId="16" pageNumber="229" type="nomenclature">
<paragraph id="8BEB36B7F11CFFEC75C6FC4356357C96" blockId="16.[227,738,915,1007]" box="[227,577,915,942]" pageId="16" pageNumber="229">
<heading id="D0A381DBF11CFFEC75C6FC4356357C96" box="[227,577,915,942]" fontSize="12" level="2" pageId="16" pageNumber="229" reason="2">
<taxonomicName id="4C544D34F11CFFEC75C6FC4355A57C96" ID-CoL="5DDDD" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[227,465,915,942]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="229" phylum="Arthropoda" rank="species" species="latibasalis" status="sp. nov." subGenus="Zyras">
<emphasis id="B920EAA5F11CFFEC75C6FC43556D7C96" box="[227,281,915,942]" italics="true" pageId="16" pageNumber="229">Zyras</emphasis>
(
<emphasis id="B920EAA5F11CFFEC7402FC4355297C96" box="[295,349,915,942]" italics="true" pageId="16" pageNumber="229">Zyras</emphasis>
)
<emphasis id="B920EAA5F11CFFEC744FFC4355A57C96" box="[362,465,915,942]" italics="true" pageId="16" pageNumber="229">latibasalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357DEF11CFFEC74FDFC4356357C96" box="[472,577,915,942]" pageId="16" pageNumber="229" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
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<subSubSection id="C34E653CF11CFFEC75C6FC0B55DC7CD7" box="[227,424,987,1007]" pageId="16" pageNumber="229" type="description">
<paragraph id="8BEB36B7F11CFFEC75C6FC0B55DC7CD7" blockId="16.[227,738,915,1007]" box="[227,424,987,1007]" pageId="16" pageNumber="229">
(
<figureCitation id="136F2A32F11CFFEC75C2FC0B55467CD7" box="[231,306,987,1007]" captionStart="Figs 126" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 126: Antenna (118) and forebody (1926) of Zyras spp. (117, 1926) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (1112; 11: holotype; 12 female paratype); hlavaci (1314; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="16" pageNumber="229">Figs 3, 25</figureCitation>
,
<figureCitation id="136F2A32F11CFFEC741CFC0B55187CD7" box="[313,364,987,1007]" captionStart-0="Figs 2739" captionStart-1="Figs 4052" captionStartId-0="9.[151,191,1970,1992]" captionStartId-1="10.[190,231,1972,1994]" captionTargetBox-0="[154,1396,232,1963]" captionTargetBox-1="[192,1434,232,1963]" captionTargetId-0="figure-145@9.[154,1396,232,1963]" captionTargetId-1="figure-129@10.[192,1434,232,1963]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." captionText-1="Figs 4052: Abdomen (4049, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (4050) and Myrmedonota modiglianii (5152): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (4950). Scale bars:1.0 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5742345" figureDoi-1="http://doi.org/10.5281/zenodo.5742347" httpUri-0="https://zenodo.org/record/5742345/files/figure.png" httpUri-1="https://zenodo.org/record/5742347/files/figure.png" pageId="16" pageNumber="229">3940</figureCitation>
,
<figureCitation id="136F2A32F11CFFEC7456FC0B55D77CD7" box="[371,419,987,1007]" captionStart="Figs 5371" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 5371: Zyras elegantulus, lectotype (5355), Z. facundus, lectotype (5658), Z. bryanti (5962), Z. matangensis, holotype (6365), Z. granulipennis, holotype (6669), and Z. latibasalis (7071): median lobe of aedeagus in lateral and in ventral view (5354, 5657, 5960, 6364, 6667, 7071); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="16" pageNumber="229">7071</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11CFFEC7598FBD956167A19" pageId="16" pageNumber="229" type="materials_examined">
<paragraph id="8BEB36B7F11CFFEC7598FBD956B77B99" blockId="16.[189,795,1033,1186]" pageId="16" pageNumber="229">
<emphasis id="B920EAA5F11CFFEC7598FBD955A67B19" bold="true" box="[189,466,1033,1057]" pageId="16" pageNumber="229">Type material examined</emphasis>
:
<materialsCitation id="3B3C3CEAF11CFFEC74C1FBD956AC7BB9" ID-GBIF-Occurrence="3450187345" collectingDate="2004-01-29" collectingDateMax="2004-02-04" collectingDateMin="2004-01-29" collectionCode="FIT" collectorName="Maruyama M. et al." country="Indonesia" county="Cianjur" determinerName="V. Assing" elevation="1400" location="Cibodas" municipality="Kebun Raya" pageId="16" pageNumber="229" specimenCount="1" stateProvince="Jawa Barat" typeStatus="holotype">
<typeStatus id="54EF8815F11CFFEC74C1FBD956337B19" box="[484,583,1033,1057]" pageId="16" pageNumber="229" type="holotype">Holotype</typeStatus>
<specimenCount id="9D52FD3EF11CFFEC7771FBDA561A7B19" box="[596,622,1034,1057]" pageId="16" pageNumber="229" type="generic">♂:</specimenCount>
<collectingCountry id="F3437627F11CFFEC77A1FBD957637B19" box="[644,791,1033,1057]" name="Indonesia" pageId="16" pageNumber="229">INDONESIA</collectingCountry>
:
<collectingRegion id="4990F855F11CFFEC7598FBFA55457B79" box="[189,305,1066,1089]" country="Indonesia" name="Jawa Barat" pageId="16" pageNumber="229">Jawa Barat</collectingRegion>
,
<collectingCounty id="628A4E3BF11CFFEC7418FBF955FB7B79" box="[317,399,1065,1089]" pageId="16" pageNumber="229">Cianjur</collectingCounty>
,
<collectingMunicipality id="6B8FACCDF11CFFEC74BEFBF9566B7B7A" box="[411,543,1065,1090]" pageId="16" pageNumber="229">Kebun Raya</collectingMunicipality>
,
<location id="8E8B606CF11CFFEC7709FBF956F07B79" LSID="urn:lsid:plazi:treatment:03FD87A1F11CFFED75C6FC4356D87E59:8E8B606CF11CFFEC7709FBF956F07B79" box="[556,644,1065,1089]" country="Indonesia" county="Cianjur" municipality="Kebun Raya" name="Cibodas" pageId="16" pageNumber="229" stateProvince="Jawa Barat">Cibodas</location>
(
<quantity id="4CAC9B52F11CFFEC77B6FBF956907B79" box="[659,740,1065,1089]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="16" pageNumber="229" unit="m" value="1400.0">
<elevation id="0079D184F11CFFEC77B6FBF956907B79" box="[659,740,1065,1089]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="16" pageNumber="229" unit="m" value="1400.0">1400 m</elevation>
</quantity>
alt.),
<date id="FFEA1077F11CFFEC7598FB99552C7B59" box="[189,344,1097,1121]" pageId="16" pageNumber="229" value="2004-01-29" valueMax="2004-02-04" valueMin="2004-01-29">
<collectingDate id="EFAEE99FF11CFFEC7598FB99552C7B59" box="[189,344,1097,1121]" pageId="16" pageNumber="229" value="2004-01-29" valueMax="2004-02-04" valueMin="2004-01-29">29.I.4.II.2004</collectingDate>
</date>
(FIT),
<collectorName id="26A15361F11CFFEC7483FB9A560D7B59" box="[422,633,1097,1122]" pageId="16" pageNumber="229">Maruyama M. et al.</collectorName>
/ Holotypus ♂
<taxonomicName id="4C544D34F11CFFEC7598FBBA55117BB9" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[189,357,1129,1153]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="229" phylum="Arthropoda" rank="species" species="latibasalis" status="sp. nov.">
<emphasis id="B920EAA5F11CFFEC7598FBBA55117BB9" box="[189,357,1129,1153]" italics="true" pageId="16" pageNumber="229">Zyras latibasalis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357DEF11CFFEC7449FBBA55D07BB9" box="[364,420,1130,1153]" pageId="16" pageNumber="229" rank="species">sp. n.</taxonomicNameLabel>
, det.
<determinerName id="7E37DA73F11CFFEC74F8FBBA564B7BB9" box="[477,575,1129,1153]" pageId="16" pageNumber="229">
V.
<bibRefCitation id="EFC54B46F11CFFEC74DDFBB9564B7BB9" author="ASSING, V." box="[504,575,1129,1153]" pageId="16" pageNumber="229" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing</bibRefCitation>
</determinerName>
2017” (cMar)
</materialsCitation>
.
<materialsCitation id="3B3C3CEAF11CFFEC77C4FBBA56CB7B99" ID-GBIF-Occurrence="3450187339" collectingDate="2004-01-29" collectingDateMax="2004-02-04" collectingDateMin="2004-01-29" collectionCode="FIT" country="Indonesia" county="Cianjur" elevation="1400" municipality="Kebun Raya" pageId="16" pageNumber="229" specimenCount="9" stateProvince="Jawa Barat" typeStatus="paratype">
<typeStatus id="54EF8815F11CFFEC77C4FBBA548C7B9A" pageId="16" pageNumber="229" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D52FD3EF11CFFEC7424FB59554B7B99" box="[257,319,1161,1185]" pageId="16" pageNumber="229" type="generic">9 exs.</specimenCount>
: same data as holotype (cMar, cAss)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEB36B7F11CFFEC7598FB1956167A19" blockId="16.[189,794,1225,1313]" pageId="16" pageNumber="229">
<emphasis id="B920EAA5F11CFFEC7598FB1955E77BD9" bold="true" box="[189,403,1225,1249]" pageId="16" pageNumber="229">Additional material</emphasis>
:
<materialsCitation id="3B3C3CEAF11CFFEC74BBFB19562A7A19" ID-GBIF-Occurrence="3450187306" collectingDate="2001-04-02" collectingDateMax="2001-05-11" collectingDateMin="2001-04-02" collectionCode="NMP" collectorName="D. Hauck" county="Pu Phan mts." elevation="1750" latitude="20.25" location="Hu Phan prov." longLatPrecision="1268" longitude="104.03333" municipality="Ban Saluei" pageId="16" pageNumber="229" specimenCount="1">
1 ♀: “NE-LAOS:
<location id="8E8B606CF11CFFEC776AFB1A569C7BD9" LSID="urn:lsid:plazi:treatment:03FD87A1F11CFFED75C6FC4356D87E59:8E8B606CF11CFFEC776AFB1A569C7BD9" box="[591,744,1225,1249]" county="Pu Phan mts." latitude="20.25" longLatPrecision="1268" longitude="104.03333" municipality="Ban Saluei" name="Hu Phan prov." pageId="16" pageNumber="229">Hu Phan prov.</location>
,
<collectingMunicipality id="6B8FACCDF11CFFEC77D5FB1A548B7A39" pageId="16" pageNumber="229">Ban Saluei</collectingMunicipality>
,
<collectingCounty id="628A4E3BF11CFFEC7429FB3A55D07A39" box="[268,420,1257,1281]" pageId="16" pageNumber="229">Pu Phan mts.</collectingCounty>
,
<geoCoordinate id="EE605070F11CFFEC7497FB39567C7A39" box="[434,520,1257,1281]" degrees="20" direction="north" minutes="15" orientation="latitude" pageId="16" pageNumber="229" precision="925" value="20.25">20°15'N</geoCoordinate>
,
<geoCoordinate id="EE605070F11CFFEC7732FB3956017A39" box="[535,629,1257,1281]" degrees="104" direction="east" minutes="02" orientation="longitude" pageId="16" pageNumber="229" precision="925" value="104.03333">104°02'E</geoCoordinate>
,
<quantity id="4CAC9B52F11CFFEC77A6FB3957627A39" box="[643,790,1257,1281]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="16" pageNumber="229" unit="m" value="1750.0" valueMax="2000.0" valueMin="1500.0">
<elevation id="0079D184F11CFFEC77A6FB3957627A39" box="[643,790,1257,1281]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="16" pageNumber="229" unit="m" value="1750.0" valueMax="2000.0" valueMin="1500.0">15002000 m</elevation>
</quantity>
,
<date id="FFEA1077F11CFFEC7598FAD955127A19" box="[189,358,1289,1313]" pageId="16" pageNumber="229" value="2001-04-02" valueMax="2001-05-11" valueMin="2001-04-02">
<collectingDate id="EFAEE99FF11CFFEC7598FAD955127A19" box="[189,358,1289,1313]" pageId="16" pageNumber="229" value="2001-04-02" valueMax="2001-05-11" valueMin="2001-04-02">2.IV.11.V.2001</collectingDate>
</date>
, leg.
<collectorName id="26A15361F11CFFEC74BEFADA558B7A19" box="[411,511,1289,1313]" pageId="16" pageNumber="229">D. Hauck</collectorName>
” (
<collectionCode id="ED45AE72F11CFFEC7733FAD956227A19" box="[534,598,1289,1313]" collectionName="Czech Republic, Prague, National Museum (Natural History)" country="Czech Republic" httpUri="http://grbio.org/cool/fx6k-enrh" name="National Museum (Prague)" pageId="16" pageNumber="229">NMP</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11CFFEC7598FA9955777AD9" pageId="16" pageNumber="229" type="etymology">
<paragraph id="8BEB36B7F11CFFEC7598FA9955777AD9" blockId="16.[189,794,1353,1505]" pageId="16" pageNumber="229">
<emphasis id="B920EAA5F11CFFEC7598FA9955407A59" bold="true" box="[189,308,1353,1377]" pageId="16" pageNumber="229">Etymology</emphasis>
: The specific epithet (with broad base) is an adjective alluding to the basally broad ventral process of the aedeagus in ventral view, the most prominent character distinguishing this species from the similar
<taxonomicName id="4C544D34F11CFFEC7598FA1A54897AD9" authorityName="PACE" authorityYear="1993" box="[189,253,1482,1505]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="229" phylum="Arthropoda" rank="species" species="wei">
<emphasis id="B920EAA5F11CFFEC7598FA1A54897AD9" box="[189,253,1482,1505]" italics="true" pageId="16" pageNumber="229">Z. wei</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11CFFEC7598F9D950207719" pageId="16" pageNumber="229" type="description">
<paragraph id="8BEB36B7F11CFFEC7598F9D955977879" blockId="16.[189,794,1545,2081]" pageId="16" pageNumber="229">
<emphasis id="B920EAA5F11CFFEC7598F9D9554B7919" bold="true" box="[189,319,1545,1569]" pageId="16" pageNumber="229">Description</emphasis>
: Body length 5.57.0 mm; length of forebody 2.53.0 mm. Coloration (
<figureCitation id="136F2A32F11CFFEC7731F9F956087979" box="[532,636,1577,1601]" captionStart="Figs 126" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 126: Antenna (118) and forebody (1926) of Zyras spp. (117, 1926) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (1112; 11: holotype; 12 female paratype); hlavaci (1314; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="16" pageNumber="229">Figs 3, 25</figureCitation>
,
<figureCitation id="136F2A32F11CFFEC77A3F9F956D77979" box="[646,675,1577,1601]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="16" pageNumber="229">39</figureCitation>
): forebody blackish with the humeral and anterior portions of the elytra sometimes narrowly and diffusely paler; abdomen black with the posterior margins of the tergites more or less distinctly paler reddish; legs pale-yellowish; antennae blackish with antennomeres IXX often paler brown (X more so than IX) and antennomere XI dark-yellowish to pale-reddish; maxillary palpi pale-brown with the apical palpomere yellowish.
</paragraph>
<paragraph id="8BEB36B7F11CFFEC7598F89957DD7E19" blockId="16.[189,794,1545,2081]" lastBlockId="16.[831,1437,233,1537]" pageId="16" pageNumber="229">
Head (
<figureCitation id="136F2A32F11CFFEC7423F89955277859" box="[262,339,1865,1889]" captionStart="Figs 126" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 126: Antenna (118) and forebody (1926) of Zyras spp. (117, 1926) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (1112; 11: holotype; 12 female paratype); hlavaci (1314; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="16" pageNumber="229">Fig. 25</figureCitation>
) distinctly transverse; punctation moderately coarse and moderately dense; median dorsal portion extensively impunctate. Eyes large, much longer than postocular region in dorsal view. Antenna (
<figureCitation id="136F2A32F11CFFEC77F7F879576478F9" box="[722,784,1961,1985]" captionStart="Figs 126" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 126: Antenna (118) and forebody (1926) of Zyras spp. (117, 1926) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (1112; 11: holotype; 12 female paratype); hlavaci (1314; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="16" pageNumber="229">Fig. 3</figureCitation>
)
<quantity id="4CAC9B52F11CFFEC7598F819554A78D9" box="[189,318,1993,2017]" metricMagnitude="-3" metricUnit="m" metricValue="2.05" metricValueMax="2.2" metricValueMin="1.9" pageId="16" pageNumber="229" unit="mm" value="2.05" valueMax="2.2" valueMin="1.9">1.92.2 mm</quantity>
long and moderately massive; antennomeres IVVI approximately as long as broad, VIIX weakly transverse, X less than 1.5 times as broad as long, and XI approximately as long as the combined length of IX and X.
</paragraph>
<paragraph id="8BEB36B7F11CFFEC761AFEFA50187D39" blockId="16.[831,1437,233,1537]" pageId="16" pageNumber="229">
Pronotum (
<figureCitation id="136F2A32F11CFFEC769BFEF9507C7E79" box="[958,1032,297,321]" captionStart="Figs 126" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 126: Antenna (118) and forebody (1926) of Zyras spp. (117, 1926) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (1112; 11: holotype; 12 female paratype); hlavaci (1314; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="16" pageNumber="229">Fig. 25</figureCitation>
) strongly transverse, 1.221.24 times as broad as long and 1.291.36 times as broad as head, broadest near anterior angles; posterior angles weakly, obtusely marked; lateral margins not sinuate in posterior half in dorsal view; punctation moderately coarse and moderately dense, nearly regularly distributed; midline rather narrowly impunctate.
</paragraph>
<paragraph id="8BEB36B7F11CFFEC7665FDD9510B7D99" blockId="16.[831,1437,233,1537]" pageId="16" pageNumber="229">
Elytra (
<figureCitation id="136F2A32F11CFFEC76AEFDD957A17D19" box="[907,981,521,545]" captionStart="Figs 126" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 126: Antenna (118) and forebody (1926) of Zyras spp. (117, 1926) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (1112; 11: holotype; 12 female paratype); hlavaci (1314; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="16" pageNumber="229">Fig. 25</figureCitation>
) approximately 0.85 times as long as pronotum; punctation rather dense and coarse, slightly denser near scutellum and near suture than in posterior and lateral portions. Hind wings present. Metatarsomere I approximately as long as the combined length of IIIV.
</paragraph>
<paragraph id="8BEB36B7F11CFFEC7665FD7951597A19" blockId="16.[831,1437,233,1537]" pageId="16" pageNumber="229">
Abdomen (
<figureCitation id="136F2A32F11CFFEC7698FD79507D7DF9" box="[957,1033,681,705]" captionStart="Figs 4052" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 4052: Abdomen (4049, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (4050) and Myrmedonota modiglianii (5152): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (4950). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="16" pageNumber="229">Fig. 40</figureCitation>
) slightly narrower than elytra, with moderately deep anterior impressions on tergites IIIV; tergites IIIV each with a transverse row or band of rather fine and often sparse non-setiferous punctures in anterior impressions, with a lateral setiferous puncture on either side and with four (tergite III) or 68 (tergites IVV) setiferous punctures at posterior margins; tergite VI with a narrow transverse band or a transverse row of rather fine non-setiferous punctures anteriorly, with a lateral setiferous puncture on either side, and with usually eight setiferous punctures at posterior margin (individual punctures may be missing, or additional punctures may be present); tergite VII with a transverse band of fine non-setiferous punctures anteriorly and with two transverse rows of setiferous punctures posteriorly, posterior margin with palisade fringe; tergite VIII with setiferous punctures bearing long dark setae only in posterior fourth, anteriorly with a transverse band of oblong microsculpture (microstriae), posterior margin with shallow concavity in the middle (
<figureCitation id="136F2A32F11CFFEC71F1FAD951557A19" box="[1236,1313,1289,1313]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="16" pageNumber="229">Fig. 39</figureCitation>
).
</paragraph>
<paragraph id="8BEB36B7F11CFFEC7665FAFA501A7AF9" blockId="16.[831,1437,233,1537]" pageId="16" pageNumber="229">
♂: posterior margin of sternite VIII broadly truncate in the middle; median lobe of aedeagus (
<figureCitation id="136F2A32F11CFFEC703BFA9951E17A59" box="[1310,1429,1353,1377]" captionStart="Figs 5371" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 5371: Zyras elegantulus, lectotype (5355), Z. facundus, lectotype (5658), Z. bryanti (5962), Z. matangensis, holotype (6365), Z. granulipennis, holotype (6669), and Z. latibasalis (7071): median lobe of aedeagus in lateral and in ventral view (5354, 5657, 5960, 6364, 6667, 7071); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="16" pageNumber="229">Figs 7071</figureCitation>
)
<quantity id="4CAC9B52F11CFFEC7665FAB957AC7AB9" box="[832,984,1385,1409]" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="7.5" metricValueMin="7.0" pageId="16" pageNumber="229" unit="mm" value="0.725" valueMax="0.75" valueMin="0.7">0.700.75 mm</quantity>
long; ventral process broadly triangular in ventral view; parameres approximately 1.0 mm long, with long and slender apical lobe.
</paragraph>
<paragraph id="8BEB36B7F11CFFEC7665FA1A57CD7939" blockId="16.[831,1437,233,1537]" pageId="16" pageNumber="229">♀: posterior margin of sternite VIII shallowly concave in the middle.</paragraph>
<paragraph id="8BEB36B7F11CFFEC7665F9F9513379B9" blockId="16.[832,1437,1577,1665]" pageId="16" pageNumber="229">
<emphasis id="B920EAA5F11CFFEC7665F9F950457979" bold="true" box="[832,1073,1577,1601]" pageId="16" pageNumber="229">Intraspecific variation</emphasis>
:
<location id="8E8B606CF11CFFEC711EF9F950117979" LSID="urn:lsid:plazi:treatment:03FD87A1F11CFFED75C6FC4356D87E59:8E8B606CF11CFFEC711EF9F950117979" box="[1083,1125,1577,1601]" country="Laos" name="The" pageId="16" pageNumber="229">The</location>
female from
<collectingCountry id="F3437627F11CFFEC71C8F9FA516B7979" box="[1261,1311,1578,1601]" name="Laos" pageId="16" pageNumber="229">Laos</collectingCountry>
is similar to the
<typeStatus id="54EF8815F11CFFEC7643F99A57E0795A" box="[870,916,1610,1634]" pageId="16" pageNumber="229">type</typeStatus>
series in size, habitus, coloration, and punctation, except that the apical antennomeres are blackish.
</paragraph>
<paragraph id="8BEB36B7F11CFFEC7665F97950207719" blockId="16.[832,1436,1705,2081]" pageId="16" pageNumber="229">
<emphasis id="B920EAA5F11CFFEC7665F979506479F9" bold="true" box="[832,1040,1705,1729]" pageId="16" pageNumber="229">Comparative notes</emphasis>
: In external characters (size, habitus, punctation, coloration), in the lateral aspect of the ventral process of the aedeagus, and regarding the long apical lobe of the paramere,
<taxonomicName id="4C544D34F11CFFEC7100F8DA50DD7819" authorityName="Assing" authorityYear="2017" box="[1061,1193,1801,1825]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="229" phylum="Arthropoda" rank="species" species="latibasalis">
<emphasis id="B920EAA5F11CFFEC7100F8DA50DD7819" box="[1061,1193,1801,1825]" italics="true" pageId="16" pageNumber="229">Z. latibasalis</emphasis>
</taxonomicName>
strongly resembles the widespread
<taxonomicName id="4C544D34F11CFFEC76E6F8FA50777879" authorityName="PACE" authorityYear="1993" box="[963,1027,1834,1857]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="229" phylum="Arthropoda" rank="species" species="wei">
<emphasis id="B920EAA5F11CFFEC76E6F8FA50777879" box="[963,1027,1834,1857]" italics="true" pageId="16" pageNumber="229">Z. wei</emphasis>
</taxonomicName>
. It is distinguished from this species (and from the similar
<taxonomicName id="4C544D34F11CFFEC7175F89A509B7859" authorityName="Assing" authorityYear="2017" box="[1104,1263,1865,1889]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="229" phylum="Arthropoda" rank="species" species="parvilobatus">
<emphasis id="B920EAA5F11CFFEC7175F89A509B7859" box="[1104,1263,1865,1889]" italics="true" pageId="16" pageNumber="229">Z. parvilobatus</emphasis>
</taxonomicName>
) by the darkyellowish antennomere XI, by the presence of an anterior transverse band of microstriae on tergite VIII, and by the morphology of the aedeagus (
<taxonomicName id="4C544D34F11CFFEC7188F87A509978F9" authorityName="PACE" authorityYear="1993" box="[1197,1261,1962,1985]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="229" phylum="Arthropoda" rank="species" species="wei">
<emphasis id="B920EAA5F11CFFEC7188F87A509978F9" box="[1197,1261,1962,1985]" italics="true" pageId="16" pageNumber="229">Z. wei</emphasis>
</taxonomicName>
: ventral process much more slender and basally somewhat constricted in ventral view, and smoothly curved in lateral view; crista apicalis more prominent).
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF11DFFED75B2FF3956D87E59" pageId="17" pageNumber="230" type="distribution">
<paragraph id="8BEB36B7F11DFFED75B2FF3956D87E59" blockId="17.[151,756,233,353]" pageId="17" pageNumber="230">
<emphasis id="B920EAA5F11DFFED75B2FF3955547E39" bold="true" box="[151,288,233,257]" pageId="17" pageNumber="230">Distribution</emphasis>
: The
<typeStatus id="54EF8815F11DFFED7444FF3A55FB7E3A" box="[353,399,234,258]" pageId="17" pageNumber="230">type</typeStatus>
locality is situated in Jawa Barat at an altitude of
<quantity id="4CAC9B52F11DFFED746DFED955EE7E19" box="[328,410,265,289]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="17" pageNumber="230" unit="m" value="1400.0">1400 m</quantity>
. The tentative record from
<collectingCountry id="F3437627F11DFFED77E7FEDA56807E19" box="[706,756,266,289]" name="Laos" pageId="17" pageNumber="230">Laos</collectingCountry>
requires confirmation based on males. The
<typeStatus id="54EF8815F11DFFED7758FEFA56DF7E7A" box="[637,683,298,322]" pageId="17" pageNumber="230">type</typeStatus>
specimens were collected with flight interception traps.
</paragraph>
</subSubSection>
</treatment>
</document>