377 lines
53 KiB
XML
377 lines
53 KiB
XML
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<mods:title id="463B8AAA60D7D8064CB549B7A84F563C">On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:title id="67BE0A4124846937D103DEF7CF9948CF">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
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<treatment id="03FD87A1F118FFE97640FA03516F7A99" ID-DOI="http://doi.org/10.5281/zenodo.5888804" ID-GBIF-Taxon="192500461" ID-Zenodo-Dep="5888804" LSID="urn:lsid:plazi:treatment:03FD87A1F118FFE97640FA03516F7A99" httpUri="http://treatment.plazi.org/id/03FD87A1F118FFE97640FA03516F7A99" lastPageId="21" lastPageNumber="234" pageId="20" pageNumber="233">
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<subSubSection id="C34E653CF118FFE87640FA0351507AD6" box="[869,1316,1491,1518]" pageId="20" pageNumber="233" type="nomenclature">
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<paragraph id="8BEB36B7F118FFE87640FA0351507AD6" blockId="20.[869,1316,1491,1551]" box="[869,1316,1491,1518]" pageId="20" pageNumber="233">
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<heading id="D0A381DBF118FFE87640FA0351507AD6" box="[869,1316,1491,1518]" centered="true" fontSize="12" level="2" pageId="20" pageNumber="233" reason="2">
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<taxonomicName id="4C544D34F118FFE87640FA0351507AD6" ID-CoL="5DDCQ" authority="CAMERON, 1930" authorityName="Cameron" authorityYear="1930" box="[869,1316,1491,1518]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="granulipennis" subGenus="Zyras">
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<emphasis id="B920EAA5F118FFE87640FA0357EF7AD6" box="[869,923,1491,1518]" italics="true" pageId="20" pageNumber="233">Zyras</emphasis>
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(
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<emphasis id="B920EAA5F118FFE8768CFA0357AB7AD6" box="[937,991,1491,1518]" italics="true" pageId="20" pageNumber="233">Zyras</emphasis>
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)
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<emphasis id="B920EAA5F118FFE876C8FA03500D7AD6" box="[1005,1145,1491,1518]" italics="true" pageId="20" pageNumber="233">granulipennis</emphasis>
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<bibRefCitation id="EFC54B46F118FFE8715AFA0351507AD6" author="CAMERON, M." box="[1151,1316,1491,1518]" pageId="20" pageNumber="233" pagination="160 - 168" refId="ref26094" refString="CAMERON, M. 1930: Staphylinidae from British North Borneo, with descriptions of new species. - Journal of the Federated Malay States Museums 16: 160 - 168." type="journal article" year="1930">CAMERON, 1930</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E653CF118FFE87640FA2B50787937" box="[869,1036,1531,1551]" pageId="20" pageNumber="233" type="description">
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<paragraph id="8BEB36B7F118FFE87640FA2B50787937" blockId="20.[869,1316,1491,1551]" box="[869,1036,1531,1551]" pageId="20" pageNumber="233">
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(
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<figureCitation id="136F2A32F118FFE8764CFA2B57EC7937" box="[873,920,1531,1551]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="20" pageNumber="233">Figs 7</figureCitation>
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,
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<figureCitation id="136F2A32F118FFE87685FA2B57C17937" box="[928,949,1531,1551]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="20" pageNumber="233">30</figureCitation>
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,
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<figureCitation id="136F2A32F118FFE87699FA2B57A57937" box="[956,977,1531,1551]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="20" pageNumber="233">46</figureCitation>
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,
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<figureCitation id="136F2A32F118FFE876FDFA2B507C7937" box="[984,1032,1531,1551]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="20" pageNumber="233">66–69</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C34E653CF118FFE8761AF9FC50877958" pageId="20" pageNumber="233" type="reference_group">
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<paragraph id="8BEB36B7F118FFE8761AF9FC50AE7979" blockId="20.[831,1267,1579,1633]" box="[831,1242,1579,1601]" pageId="20" pageNumber="233">
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<treatmentCitationGroup id="AB441199F118FFE8761AF9FC50AE7979" box="[831,1242,1579,1601]" pageId="20" pageNumber="233">
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<treatmentCitation id="0AF510A6F118FFE8761AF9FC50B37979" author="CAMERON, M." box="[831,1223,1579,1601]" page="167" pageId="20" pageNumber="233" year="1930">
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<taxonomicName id="4C544D34F118FFE8761AF9FC50B37979" ID-CoL="5DDCQ" authority="CAMERON, 1930: 167" authorityName="Cameron" authorityPageNumber="167" authorityYear="1930" box="[831,1223,1579,1601]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="granulipennis">
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<emphasis id="B920EAA5F118FFE8761AF9FC578D7978" box="[831,1017,1579,1601]" italics="true" pageId="20" pageNumber="233">Zyras granulipennis</emphasis>
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<bibRefCitation id="EFC54B46F118FFE876DBF9FB50B37979" author="CAMERON, M." box="[1022,1223,1579,1601]" pageId="20" pageNumber="233" pagination="160 - 168" refId="ref26094" refString="CAMERON, M. 1930: Staphylinidae from British North Borneo, with descriptions of new species. - Journal of the Federated Malay States Museums 16: 160 - 168." type="journal article" year="1930">
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||
CAMERON, 1930:
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<specimenCount id="9D52FD3EF118FFE87181F9FB50B37979" box="[1188,1223,1579,1601]" pageId="20" pageNumber="233" type="generic">167</specimenCount>
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</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
f.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="8BEB36B7F118FFE8761AF99C50877958" blockId="20.[831,1267,1579,1633]" box="[831,1267,1611,1633]" pageId="20" pageNumber="233">
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||
<treatmentCitationGroup id="AB441199F118FFE8761AF99C50877958" box="[831,1267,1611,1633]" pageId="20" pageNumber="233">
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||
<treatmentCitation id="0AF510A6F118FFE8761AF99C50E07959" author="PACE, R." box="[831,1172,1611,1633]" page="150" pageId="20" pageNumber="233" year="2008">
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||
<taxonomicName id="4C544D34F118FFE8761AF99C50E07959" authority="PACE, 2008: 150" authorityName="PACE" authorityPageNumber="150" authorityYear="2008" box="[831,1172,1611,1633]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="pervariolosus">
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<emphasis id="B920EAA5F118FFE8761AF99C57827958" box="[831,1014,1611,1633]" italics="true" pageId="20" pageNumber="233">Zyras pervariolosus</emphasis>
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<bibRefCitation id="EFC54B46F118FFE876DEF99B50E07959" author="PACE, R." box="[1019,1172,1611,1633]" pageId="20" pageNumber="233" pagination="107 - 156" refId="ref27259" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">PACE, 2008: 150</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
;
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<emphasis id="B920EAA5F118FFE871B8F99B50877958" bold="true" box="[1181,1267,1611,1632]" pageId="20" pageNumber="233">syn. nov.</emphasis>
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C34E653CF118FFE9761AF959553E7E59" lastPageId="21" lastPageNumber="234" pageId="20" pageNumber="233" type="materials_examined">
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<paragraph id="8BEB36B7F118FFE8761AF95950857819" blockId="20.[831,1437,1673,2082]" pageId="20" pageNumber="233">
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<emphasis id="B920EAA5F118FFE8761AF95950397999" bold="true" box="[831,1101,1673,1697]" pageId="20" pageNumber="233">Type material examined</emphasis>
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:
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<taxonomicName id="4C544D34F118FFE87179F95A51727999" authorityName="Cameron" authorityYear="1930" box="[1116,1286,1673,1697]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="granulipennis">
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<emphasis id="B920EAA5F118FFE87179F95A51727999" box="[1116,1286,1673,1697]" italics="true" pageId="20" pageNumber="233">Z. granulipennis</emphasis>
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</taxonomicName>
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||
:
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<materialsCitation id="3B3C3CEAF118FFE87030F95950857819" ID-GBIF-Occurrence="3450187309" collectingDate="1927-07-24" collectionCode="BMNH" collectorName="M. Cameron & Bequest. B. M." determinerName="V. Assing" location="Bettotan" municipality="N. Borneo" pageId="20" pageNumber="233" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54EF8815F118FFE87030F959510C7999" box="[1301,1400,1673,1697]" pageId="20" pageNumber="233" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D52FD3EF118FFE870A4F95A51E27999" box="[1409,1430,1674,1697]" pageId="20" pageNumber="233" type="generic">♂</specimenCount>
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: “
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<collectingMunicipality id="6B8FACCDF118FFE8766FF97957CA79F9" box="[842,958,1705,1729]" name="N. Borneo" pageId="20" pageNumber="233">N. Borneo</collectingMunicipality>
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.
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||
<location id="8E8B606CF118FFE876E9F97A505D79F9" LSID="urn:lsid:plazi:treatment:03FD87A1F118FFE97640FA03516F7A99:8E8B606CF118FFE876E9F97A505D79F9" box="[972,1065,1706,1729]" municipality="N. Borneo" name="Bettotan" pageId="20" pageNumber="233">Bettotan</location>
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||
, /
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||
<location id="8E8B606CF118FFE8716FF97A509479F9" LSID="urn:lsid:plazi:treatment:03FD87A1F118FFE97640FA03516F7A99:8E8B606CF118FFE8716FF97A509479F9" box="[1098,1248,1705,1729]" municipality="N. Borneo" name="Nr. Sandakan" pageId="20" pageNumber="233">Nr. Sandakan</location>
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||
.
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||
<date id="FFEA1077F118FFE871C8F97951F279F9" box="[1261,1414,1703,1729]" pageId="20" pageNumber="233" value="1927-07-24">
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||
<collectingDate id="EFAEE99FF118FFE871C8F97951F279F9" box="[1261,1414,1703,1729]" pageId="20" pageNumber="233" value="1927-07-24">
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||
July 24
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||
<superScript id="7C219BFFF118FFE8701CF9775132798D" attach="left" box="[1337,1350,1703,1717]" fontSize="6" pageId="20" pageNumber="233">th</superScript>
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||
1927
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||
</collectingDate>
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||
</date>
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||
. /
|
||
<taxonomicName id="4C544D34F118FFE8761AF91A503879D9" authority="Cam." authorityName="Cameron" authorityYear="1930" box="[831,1100,1737,1761]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="granulipennis">
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||
<emphasis id="B920EAA5F118FFE8761AF91A507979D9" box="[831,1037,1737,1761]" italics="true" pageId="20" pageNumber="233">Zyras granulipennis</emphasis>
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||
Cam.
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||
</taxonomicName>
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||
Type /
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||
<typeStatus id="54EF8815F118FFE871BDF919508F79D9" box="[1176,1275,1737,1761]" pageId="20" pageNumber="233" type="holotype">Holotype</typeStatus>
|
||
/
|
||
<collectorName id="26A15361F118FFE8702BF91A51ED79D9" box="[1294,1433,1737,1761]" pageId="20" pageNumber="233">M. Cameron</collectorName>
|
||
.
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||
<collectorName id="26A15361F118FFE8761AF93A57A17839" box="[831,981,1770,1793]" pageId="20" pageNumber="233">Bequest. B.M.</collectorName>
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||
1955-147 /
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||
<typeStatus id="54EF8815F118FFE87172F93950B27839" box="[1111,1222,1769,1793]" pageId="20" pageNumber="233" type="holotype">Holotypus</typeStatus>
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||
<taxonomicName id="4C544D34F118FFE871E8F93A57D17819" authority="Cameron" authorityName="Cameron" authorityYear="1930" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="granulipennis">
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||
<emphasis id="B920EAA5F118FFE871E8F93A51E87839" box="[1229,1436,1769,1793]" italics="true" pageId="20" pageNumber="233">Zyras granulipennis</emphasis>
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||
Cameron
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||
</taxonomicName>
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||
, rev.
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||
<determinerName id="7E37DA73F118FFE876FEF8DA50497819" box="[987,1085,1801,1825]" pageId="20" pageNumber="233">
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V.
|
||
<bibRefCitation id="EFC54B46F118FFE876D3F8D950497819" author="ASSING, V." box="[1014,1085,1801,1825]" pageId="20" pageNumber="233" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing</bibRefCitation>
|
||
</determinerName>
|
||
2017” (
|
||
<collectionCode id="ED45AE72F118FFE871AAF8D950917819" box="[1167,1253,1801,1825]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="20" pageNumber="233">BMNH</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F118FFE9761AF8FA553E7E59" blockId="20.[831,1437,1673,2082]" lastBlockId="21.[151,757,233,353]" lastPageId="21" lastPageNumber="234" pageId="20" pageNumber="233">
|
||
<materialsCitation id="3B3C3CEAF118FFE8761AF8FA514578D9" ID-GBIF-Occurrence="3450198301" collectingDate="1987-04-30" collectionCode="MHNG, ENTO" collectorName="A. Smetana" country="BORNEO" determinerName="V. Assing" elevation="1500" location="HQ at Liwagu Rv." municipality="Mt. Kinabalu Nat. Pk." pageId="20" pageNumber="233" specimenCount="1" stateProvince="SABAH" typeStatus="holotype">
|
||
<taxonomicName id="4C544D34F118FFE8761AF8FA579E7879" authorityName="PACE" authorityYear="2008" box="[831,1002,1833,1857]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="pervariolosus">
|
||
<emphasis id="B920EAA5F118FFE8761AF8FA579E7879" box="[831,1002,1833,1857]" italics="true" pageId="20" pageNumber="233">
|
||
<originalDetermination id="A84A8E4BF118FFE8761AF8FA579E7879" box="[831,1002,1833,1857]" pageId="20" pageNumber="233">Z. pervariolosus</originalDetermination>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<typeStatus id="54EF8815F118FFE876D9F8F950157879" box="[1020,1121,1833,1857]" pageId="20" pageNumber="233" type="holotype">Holotype</typeStatus>
|
||
<specimenCount id="9D52FD3EF118FFE87149F8FA50F37879" box="[1132,1159,1834,1857]" pageId="20" pageNumber="233" type="generic">♂:</specimenCount>
|
||
“
|
||
<collectingCountry id="F3437627F118FFE871B9F8F9517E7879" box="[1180,1290,1833,1857]" pageId="20" pageNumber="233">BORNEO</collectingCountry>
|
||
<collectingRegion id="4990F855F118FFE87030F8F9511F7879" box="[1301,1387,1833,1857]" country="Malaysia" name="Sabah" pageId="20" pageNumber="233">SABAH</collectingRegion>
|
||
<collectingMunicipality id="6B8FACCDF118FFE87052F8FA578B7859" name="Mt. Kinabalu Nat. Pk." pageId="20" pageNumber="233" stateProvince="Sabah">Mt. Kinabalu Nat. Pk.</collectingMunicipality>
|
||
,
|
||
<location id="8E8B606CF118FFE87122F89950B17859" LSID="urn:lsid:plazi:treatment:03FD87A1F118FFE97640FA03516F7A99:8E8B606CF118FFE87122F89950B17859" box="[1031,1221,1865,1889]" country="BORNEO" municipality="Mt. Kinabalu Nat. Pk." name="HQ at Liwagu Rv." pageId="20" pageNumber="233" stateProvince="SABAH">HQ at Liwagu Rv.</location>
|
||
,
|
||
<quantity id="4CAC9B52F118FFE871E8F899516A7859" box="[1229,1310,1865,1889]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="233" unit="m" value="1500.0">
|
||
<elevation id="0079D184F118FFE871E8F899516A7859" box="[1229,1310,1865,1889]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="20" pageNumber="233" unit="m" value="1500.0">1500 m</elevation>
|
||
</quantity>
|
||
<date id="FFEA1077F118FFE87006F89951EC7859" box="[1315,1432,1865,1889]" pageId="20" pageNumber="233" value="1987-04-30">
|
||
<collectingDate id="EFAEE99FF118FFE87006F89951EC7859" box="[1315,1432,1865,1889]" pageId="20" pageNumber="233" value="1987-04-30">30.IV.1987</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26A15361F118FFE8761AF8B957C878B9" box="[831,956,1897,1921]" pageId="20" pageNumber="233">A. Smetana</collectorName>
|
||
/ Holotypus
|
||
<taxonomicName id="4C544D34F118FFE87171F8BA515378B9" authorityName="PACE" authorityYear="2008" box="[1108,1319,1897,1921]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="pervariolosus">
|
||
<emphasis id="B920EAA5F118FFE87171F8BA515378B9" box="[1108,1319,1897,1921]" italics="true" pageId="20" pageNumber="233">Zyras pervariolosus</emphasis>
|
||
</taxonomicName>
|
||
mihi, det. R. Pace 2000 /
|
||
<taxonomicName id="4C544D34F118FFE876CBF85A50B77899" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[1006,1219,1929,1953]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="pervariolosus" status="sp. nov.">
|
||
<emphasis id="B920EAA5F118FFE876CBF85A50B77899" box="[1006,1219,1929,1953]" italics="true" pageId="20" pageNumber="233">Zyras pervariolosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21357DEF118FFE871EBF85A517D7899" box="[1230,1289,1930,1953]" pageId="20" pageNumber="233" rank="species">n. sp.</taxonomicNameLabel>
|
||
, det. R. Pace 2000 /
|
||
<collectionCode id="ED45AE72F118FFE876B2F879579F78F9" box="[919,1003,1961,1985]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" pageId="20" pageNumber="233">MHNG</collectionCode>
|
||
<collectionCode id="ED45AE72F118FFE876DDF879503478F9" box="[1016,1088,1961,1985]" collectionName="ENTO" pageId="20" pageNumber="233">ENTO</collectionCode>
|
||
00010008 /
|
||
<taxonomicName id="4C544D34F118FFE871FDF87A57A478D9" authority="Cameron" authorityName="Cameron" authorityYear="1930" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="233" phylum="Arthropoda" rank="species" species="granulipennis">
|
||
<emphasis id="B920EAA5F118FFE871FDF87A572A78D9" italics="true" pageId="20" pageNumber="233">Zyras granulipennis</emphasis>
|
||
Cameron
|
||
</taxonomicName>
|
||
, det.
|
||
<determinerName id="7E37DA73F118FFE87135F81A500078D9" box="[1040,1140,1993,2017]" pageId="20" pageNumber="233">
|
||
V.
|
||
<bibRefCitation id="EFC54B46F118FFE8710EF819500078D9" author="ASSING, V." box="[1067,1140,1993,2017]" pageId="20" pageNumber="233" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing</bibRefCitation>
|
||
</determinerName>
|
||
2017” (
|
||
<collectionCode id="ED45AE72F118FFE871EBF819515C78D9" box="[1230,1320,1993,2017]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="20" pageNumber="233" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B3C3CEAF118FFE97018F81A55337E59" ID-GBIF-Occurrence="3450187320" collectingDate="1987-05-24" collectionCode="MHNG" collectorName="Burckhard & Lobl" determinerName="V. Assing" elevation="1150" lastPageId="21" lastPageNumber="234" location="E Mt. Kinabalu" pageId="20" pageNumber="233" specimenCount="1" stateProvince="SABAH" typeStatus="paratype">
|
||
<typeStatus id="54EF8815F118FFE87018F81A51E878DA" box="[1341,1436,1994,2018]" pageId="20" pageNumber="233" type="paratype">Paratype</typeStatus>
|
||
♀: “
|
||
<collectingRegion id="4990F855F118FFE87648F83957B37739" box="[877,967,2025,2049]" country="Malaysia" name="Sabah" pageId="20" pageNumber="233">SABAH</collectingRegion>
|
||
:
|
||
<location id="8E8B606CF118FFE876F1F83A50F47739" LSID="urn:lsid:plazi:treatment:03FD87A1F118FFE97640FA03516F7A99:8E8B606CF118FFE876F1F83A50F47739" box="[980,1152,2025,2049]" name="E Mt. Kinabalu" pageId="20" pageNumber="233" stateProvince="SABAH">E Mt. Kinabalu</location>
|
||
,
|
||
<quantity id="4CAC9B52F118FFE871ABF83950947739" box="[1166,1248,2025,2049]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="20" pageNumber="233" unit="m" value="1150.0">
|
||
<elevation id="0079D184F118FFE871ABF83950947739" box="[1166,1248,2025,2049]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="20" pageNumber="233" unit="m" value="1150.0">1150 m</elevation>
|
||
</quantity>
|
||
,
|
||
<location id="8E8B606CF118FFE871CBF83A57D17719" LSID="urn:lsid:plazi:treatment:03FD87A1F118FFE97640FA03516F7A99:8E8B606CF118FFE871CBF83A57D17719" name="rte Ranau - Kota Kinabalu" pageId="20" pageNumber="233" stateProvince="SABAH">rte Ranau–Kota Kinabalu</location>
|
||
,
|
||
<date id="FFEA1077F118FFE8769CF7D950517719" box="[953,1061,2057,2081]" pageId="20" pageNumber="233" value="1987-05-24">
|
||
<collectingDate id="EFAEE99FF118FFE8769CF7D950517719" box="[953,1061,2057,2081]" pageId="20" pageNumber="233" value="1987-05-24">24.V.1987</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26A15361F118FFE8711FF7D950D97719" box="[1082,1197,2057,2081]" pageId="20" pageNumber="233">Burckhard</collectorName>
|
||
-
|
||
<collectorName id="26A15361F118FFE871F2F7D9517C7719" box="[1239,1288,2057,2081]" pageId="20" pageNumber="233">Löbl</collectorName>
|
||
/ Paratypus
|
||
<taxonomicName id="4C544D34F119FFE975B2FF3A551B7E39" authorityName="Assing" authorityYear="2017" box="[151,367,233,257]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="234" phylum="Arthropoda" rank="species" species="pervariolosus">
|
||
<emphasis id="B920EAA5F119FFE975B2FF3A551B7E39" box="[151,367,233,257]" italics="true" pageId="21" pageNumber="234">Zyras pervariolosus</emphasis>
|
||
</taxonomicName>
|
||
mihi, det. R. Pace 2000 /
|
||
<taxonomicName id="4C544D34F119FFE9779CFF3A55517E19" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="234" phylum="Arthropoda" rank="species" species="pervariolosus" status="sp. nov.">
|
||
<emphasis id="B920EAA5F119FFE9779CFF3A55517E19" italics="true" pageId="21" pageNumber="234">Zyras pervariolosus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21357DEF119FFE97409FEDA55127E19" box="[300,358,266,289]" pageId="21" pageNumber="234" rank="species">n. sp.</taxonomicNameLabel>
|
||
, det. R. Pace 2000 / MHNG ENTO 00010009 /
|
||
<taxonomicName id="4C544D34F119FFE97432FEFA56207E79" authority="(Bernhauer)" baseAuthorityName="Bernhauer" baseAuthorityYear="1915" box="[279,596,297,321]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="234" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="B920EAA5F119FFE97432FEFA55B07E79" box="[279,452,298,321]" italics="true" pageId="21" pageNumber="234">Zyras montanus</emphasis>
|
||
(Bernhauer)
|
||
</taxonomicName>
|
||
, det.
|
||
<determinerName id="7E37DA73F119FFE977AAFEFA56807E79" box="[655,756,297,321]" pageId="21" pageNumber="234">
|
||
V.
|
||
<bibRefCitation id="EFC54B46F119FFE9778EFEF956807E79" author="ASSING, V." box="[683,756,297,321]" pageId="21" pageNumber="234" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing</bibRefCitation>
|
||
</determinerName>
|
||
2017” (
|
||
<collectionCode id="ED45AE72F119FFE975C1FE99554A7E59" box="[228,318,329,353]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="21" pageNumber="234" type="Museum">MHNG</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E653CF119FFE975B2FE5955887DF9" pageId="21" pageNumber="234" type="discussion">
|
||
<paragraph id="8BEB36B7F119FFE975B2FE5955887DF9" blockId="21.[151,757,393,705]" pageId="21" pageNumber="234">
|
||
<emphasis id="B920EAA5F119FFE975B2FE5955727E99" bold="true" box="[151,262,393,417]" pageId="21" pageNumber="234">Comment</emphasis>
|
||
: The original description of
|
||
<taxonomicName id="4C544D34F119FFE97711FE5A56AA7E99" authorityName="Cameron" authorityYear="1930" box="[564,734,393,417]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="234" phylum="Arthropoda" rank="species" species="granulipennis">
|
||
<emphasis id="B920EAA5F119FFE97711FE5A56AA7E99" box="[564,734,393,417]" italics="true" pageId="21" pageNumber="234">Z. granulipennis</emphasis>
|
||
</taxonomicName>
|
||
is based on a unique
|
||
<typeStatus id="54EF8815F119FFE97447FE7955B47EF9" box="[354,448,425,449]" pageId="21" pageNumber="234" type="holotype">holotype</typeStatus>
|
||
from “
|
||
<collectingCountry id="F3437627F119FFE97729FE7956D27EF9" box="[524,678,425,449]" name="Malaysia" pageId="21" pageNumber="234">North Borneo</collectingCountry>
|
||
; Bettotan” (
|
||
<bibRefCitation id="EFC54B46F119FFE975F7FE19550A7ED9" author="CAMERON, M." box="[210,382,457,481]" pageId="21" pageNumber="234" pagination="160 - 168" refId="ref26094" refString="CAMERON, M. 1930: Staphylinidae from British North Borneo, with descriptions of new species. - Journal of the Federated Malay States Museums 16: 160 - 168." type="journal article" year="1930">CAMERON 1930</bibRefCitation>
|
||
), that of
|
||
<taxonomicName id="4C544D34F119FFE974F8FE1A56F07ED9" authorityName="Assing" authorityYear="2017" box="[477,644,457,481]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="234" phylum="Arthropoda" rank="species" species="pervariolosus">
|
||
<emphasis id="B920EAA5F119FFE974F8FE1A56F07ED9" box="[477,644,457,481]" italics="true" pageId="21" pageNumber="234">Z. pervariolosus</emphasis>
|
||
</taxonomicName>
|
||
on a male
|
||
<typeStatus id="54EF8815F119FFE975B2FE3954817D39" box="[151,245,489,513]" pageId="21" pageNumber="234" type="holotype">holotype</typeStatus>
|
||
from “
|
||
<collectingRegion id="4990F855F119FFE97469FE3955FA7D39" box="[332,398,489,513]" country="Malaysia" name="Sabah" pageId="21" pageNumber="234">Sabah</collectingRegion>
|
||
, Mt. Kinabalu Nat. Pk., HQ at Liwagu Rv.” and
|
||
<specimenCount id="9D52FD3EF119FFE97460FDDA55CD7D19" box="[325,441,521,545]" pageId="21" pageNumber="234" type="female">two female</specimenCount>
|
||
<typeStatus id="54EF8815F119FFE9749BFDDA56507D1A" box="[446,548,522,546]" pageId="21" pageNumber="234" type="paratype">paratypes</typeStatus>
|
||
from “
|
||
<collectingRegion id="4990F855F119FFE9774BFDD956C47D19" box="[622,688,521,545]" country="Malaysia" name="Sabah" pageId="21" pageNumber="234">Sabah</collectingRegion>
|
||
, E Mt. Kinabalu, ..., rte. Ranau–Kota Kinabalu” (
|
||
<bibRefCitation id="EFC54B46F119FFE9776AFDF956B77D79" author="PACE, R." box="[591,707,553,577]" pageId="21" pageNumber="234" pagination="107 - 156" refId="ref27259" refString="PACE, R. 2008: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">PACE 2008</bibRefCitation>
|
||
). An examination of the
|
||
<specimenCount id="9D52FD3EF119FFE9744EFD9A56767D59" box="[363,514,585,609]" pageId="21" pageNumber="234" type="generic" typeStatus="holotypes">two holotypes</specimenCount>
|
||
revealed that they are conspecific; hence the synonymy proposed above. The examined
|
||
<typeStatus id="54EF8815F119FFE97423FD5A55177D9A" box="[262,355,650,674]" pageId="21" pageNumber="234" type="paratype">paratype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C544D34F119FFE974A6FD5A565E7D99" authorityName="Assing" authorityYear="2017" box="[387,554,649,673]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="234" phylum="Arthropoda" rank="species" species="pervariolosus">
|
||
<emphasis id="B920EAA5F119FFE974A6FD5A565E7D99" box="[387,554,649,673]" italics="true" pageId="21" pageNumber="234">Z. pervariolosus</emphasis>
|
||
</taxonomicName>
|
||
is conspecific with
|
||
<taxonomicName id="4C544D34F119FFE975B2FD7A558C7DF9" authority="(BERNHAUER, 1915)" baseAuthorityName="Bernhauer" baseAuthorityYear="1915" box="[151,504,681,705]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="234" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="B920EAA5F119FFE975B2FD7A556F7DF9" box="[151,283,682,705]" italics="true" pageId="21" pageNumber="234">Z. montanus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFC54B46F119FFE9740EFD7955847DF9" author="BERNHAUER, M." box="[299,496,681,705]" pageId="21" pageNumber="234" pagination="134 - 158" refId="ref25952" refString="BERNHAUER, M. 1915: Neue Staphyliniden der indomalaiischen Fauna, insbesondere der Sunda-Insel Borneo (9. Beitrag). - Verhandlungen der Kaiserlich- Koniglichen Zoologisch-Botanischen Gesellschaft in Wien 65: 134 - 158." type="journal article" year="1915">BERNHAUER, 1915</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E653CF119FFE975B2FD39513D7A79" pageId="21" pageNumber="234" type="description">
|
||
<paragraph id="8BEB36B7F119FFE975B2FD3955E87BB9" blockId="21.[151,756,745,2081]" pageId="21" pageNumber="234">
|
||
<emphasis id="B920EAA5F119FFE975B2FD3955447C39" bold="true" box="[151,304,745,769]" pageId="21" pageNumber="234">Redescription</emphasis>
|
||
: Rather small species; body length
|
||
<quantity id="4CAC9B52F119FFE975B2FCD9556C7C19" box="[151,280,777,801]" metricMagnitude="-3" metricUnit="m" metricValue="6.3" metricValueMax="6.4" metricValueMin="6.2" pageId="21" pageNumber="234" unit="mm" value="6.3" valueMax="6.4" valueMin="6.2">6.2–6.4 mm</quantity>
|
||
; length of forebody
|
||
<quantity id="4CAC9B52F119FFE974D6FCD956007C19" box="[499,628,777,801]" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="2.9" metricValueMin="2.8" pageId="21" pageNumber="234" unit="mm" value="2.8499999999999996" valueMax="2.9" valueMin="2.8">2.8–2.9 mm</quantity>
|
||
. Coloration (
|
||
<figureCitation id="136F2A32F119FFE975BBFCF954947C79" box="[158,224,809,833]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="21" pageNumber="234">Figs 7</figureCitation>
|
||
,
|
||
<figureCitation id="136F2A32F119FFE975D4FCF955797C79" box="[241,269,809,833]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="21" pageNumber="234">30</figureCitation>
|
||
,
|
||
<figureCitation id="136F2A32F119FFE97438FCF9554F7C79" box="[285,315,809,833]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="21" pageNumber="234">46</figureCitation>
|
||
): forebody dark-brown to black; abdomen: tergites III–VI dark-brown with the anterior and antero-lateral portions paler, tergites VII–VIII reddishbrown; legs with the profemora, the protibiae, and the mesofemora brown, metafemora distinctly bicoloured with the basal two-fifths pale yellowish and the apical three-fifths dark-brown to blackish, meso- and metatibiae pale yellowish with the base narrowly blackish and the apex slightly darker, and the tarsi yellowish; antennae pale-reddish; maxillary palpi dark reddish with the apical palpomere pale-reddish.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F119FFE975B2FB5956137A99" blockId="21.[151,756,745,2081]" pageId="21" pageNumber="234">
|
||
Head (
|
||
<figureCitation id="136F2A32F119FFE975C6FB59555A7B99" box="[227,302,1161,1185]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="21" pageNumber="234">Fig. 30</figureCitation>
|
||
) relatively weakly transverse; punctation conspicuously coarsely granulose, along middle of dorsal surface with narrow and irregular glossy elevation. Eyes moderately large, barely longer than postocular region in dorsal view. Antenna (
|
||
<figureCitation id="136F2A32F119FFE974A4FAD955CB7A19" box="[385,447,1289,1313]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="21" pageNumber="234">Fig. 7</figureCitation>
|
||
) approximately 2.0 mm long; antennomeres IV–V oblong, VI approximately as long as broad, VII–X increasingly transverse, X approximately 1.5 times as broad as long, and XI of conical shape, barely as long as the combined length of IX and X.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F119FFE975B2FA7A554B79D9" blockId="21.[151,756,745,2081]" pageId="21" pageNumber="234">
|
||
Pronotum (
|
||
<figureCitation id="136F2A32F119FFE9743DFA7955117AF9" box="[280,357,1449,1473]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="21" pageNumber="234">Fig. 30</figureCitation>
|
||
) very weakly transverse, 1.03 times as broad as long and 1.18–1.23 times as broad as head, broadest near anterior angles, posteriorly distinctly tapering, strongly convex in cross-section; posterior angles marked; lateral margins sinuate in posterior two-thirds in dorsal view; punctation conspicuously coarsely granulose, in the middle and in lateral portions with irregularly elevated glossy elevations; at lateral and anterior margins with very long and rather stout dark and erect setae.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F119FFE975B2F939548778B9" blockId="21.[151,756,745,2081]" pageId="21" pageNumber="234">
|
||
Elytra (
|
||
<figureCitation id="136F2A32F119FFE975D4F939554A7839" box="[241,318,1769,1793]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="21" pageNumber="234">Fig. 30</figureCitation>
|
||
) approximately 0.8 times as long as pronotum; punctation extremely coarsely granulose and very dense. Hind wings present. Metatarsomere I slender, approximately as long as the combined length of II–IV.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F119FFE975B2F85951777D79" blockId="21.[151,756,745,2081]" lastBlockId="21.[794,1399,233,737]" pageId="21" pageNumber="234">
|
||
Abdomen (
|
||
<figureCitation id="136F2A32F119FFE97437F85955297899" box="[274,349,1929,1953]" captionStart="Figs 40–52" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 40–52: Abdomen (40–49, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (40–50) and Myrmedonota modiglianii (51–52): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (49–50). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="21" pageNumber="234">Fig. 46</figureCitation>
|
||
) nearly as broad as elytra, with deep anterior impressions on tergites III–V; anterior impressions of tergites III–V each with a transverse row of very fine and weakly defined puncture-like impressions, with a lateral puncture on either side, and with a transverse series of eight setiferous punctures near posterior margin; tergite VI with a transverse row of few fine non-setiferous punctures anteriorly, with a lateral setiferous puncture on either side, and with eight setiferous punctures near posterior margin; tergite VII with sparse non-setiferous punctures in antero-median portion and with two transverse rows each composed of few setiferous punctures posteriorly, posterior margin with palisade fringe; tergite VIII with setiferous punctation bearing long and rather stout black setae in posterior third, posterior margin weakly convex in the middle (
|
||
<figureCitation id="136F2A32F119FFE9718FFDF950827D79" box="[1194,1270,553,577]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="21" pageNumber="234">Fig. 68</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F119FFE9763FFD9A570A7DF9" blockId="21.[794,1399,233,737]" pageId="21" pageNumber="234">
|
||
♂: sternite VIII with strongly convex posterior margin (
|
||
<figureCitation id="136F2A32F119FFE97604FDB9571A7DB9" box="[801,878,617,641]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="21" pageNumber="234">Fig. 69</figureCitation>
|
||
); median lobe of aedeagus
|
||
<quantity id="4CAC9B52F119FFE9718CFDB9517C7DB9" box="[1193,1288,617,641]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" pageId="21" pageNumber="234" unit="mm" value="0.72">0.72 mm</quantity>
|
||
long and shaped as in
|
||
<figureCitation id="136F2A32F119FFE97697FD5950537D99" box="[946,1063,649,673]" captionStart="Figs 53–71" captionStartId="11.[151,192,1971,1993]" captionTargetBox="[155,1387,245,1958]" captionTargetId="figure-129@11.[154,1396,232,1963]" captionText="Figs 53–71: Zyras elegantulus, lectotype (53–55), Z. facundus, lectotype (56–58), Z. bryanti (59–62), Z. matangensis, holotype (63–65), Z. granulipennis, holotype (66–69), and Z. latibasalis (70–71): median lobe of aedeagus in lateral and in ventral view (53–54, 56–57, 59–60, 63–64, 66–67, 70–71); paramere (55, 58); male tergite VIII (61, 65, 68); male sternite VIII (62, 69). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="21" pageNumber="234">Figs 66–67</figureCitation>
|
||
; parameres malformed in the
|
||
<typeStatus id="54EF8815F119FFE9763FFD79570E7DF9" box="[794,890,681,705]" pageId="21" pageNumber="234" type="holotype">holotype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB36B7F119FFE9763FFD1A57DC7DD9" blockId="21.[794,1399,233,737]" box="[794,936,713,737]" pageId="21" pageNumber="234">♀: unknown.</paragraph>
|
||
<paragraph id="8BEB36B7F119FFE9763FFCD9513D7A79" blockId="21.[794,1399,777,1345]" pageId="21" pageNumber="234">
|
||
<emphasis id="B920EAA5F119FFE9763FFCD957857C19" bold="true" box="[794,1009,777,801]" pageId="21" pageNumber="234">Comparative notes</emphasis>
|
||
: This highly distinctive species is easily distinguished from all its consubgeners based on the conspicuous punctation of the forebody alone, particularly the strongly granulose punctation of the elytra. It is additionally characterized by the shape of the pronotum, the coloration of the legs, the punctation pattern of the abdomen, and by the shape of the median lobe of the aedeagus. Among the described species recorded from the Oriental region,
|
||
<taxonomicName id="4C544D34F119FFE9710EFBDA50A17B19" authorityName="Cameron" authorityYear="1930" box="[1067,1237,1033,1057]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="234" phylum="Arthropoda" rank="species" species="granulipennis">
|
||
<emphasis id="B920EAA5F119FFE9710EFBDA50A17B19" box="[1067,1237,1033,1057]" italics="true" pageId="21" pageNumber="234">Z. granulipennis</emphasis>
|
||
</taxonomicName>
|
||
is most similar to the sympatric
|
||
<taxonomicName id="4C544D34F119FFE976FFFBFA502A7B79" baseAuthorityName="Bernhauer" baseAuthorityYear="1915" box="[986,1118,1066,1089]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="234" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="B920EAA5F119FFE976FFFBFA502A7B79" box="[986,1118,1066,1089]" italics="true" pageId="21" pageNumber="234">Z. montanus</emphasis>
|
||
</taxonomicName>
|
||
, from which it additionally differs by smaller body size, more slender head and pronotum, a distinctly more convex and less transverse pronotum with coarser punctation and more pronounced glossy elevations, by shorter antennae with shorter and more transverse antennomeres IV–X and darker apical antennomeres (
|
||
<taxonomicName id="4C544D34F119FFE976E0FB3A503D7A39" baseAuthorityName="Bernhauer" baseAuthorityYear="1915" box="[965,1097,1258,1281]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="234" phylum="Arthropoda" rank="species" species="montanus">
|
||
<emphasis id="B920EAA5F119FFE976E0FB3A503D7A39" box="[965,1097,1258,1281]" italics="true" pageId="21" pageNumber="234">Z. montanus</emphasis>
|
||
</taxonomicName>
|
||
: antennomeres X–XI paleyellowish), fewer setiferous punctures on the abdomen, and by the shape of the median lobe of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E653CF119FFE9763FFAB9516F7A99" pageId="21" pageNumber="234" type="distribution">
|
||
<paragraph id="8BEB36B7F119FFE9763FFAB9516F7A99" blockId="21.[794,1399,1385,1441]" pageId="21" pageNumber="234">
|
||
<emphasis id="B920EAA5F119FFE9763FFAB957D77AB9" bold="true" box="[794,931,1385,1409]" pageId="21" pageNumber="234">Distribution</emphasis>
|
||
:
|
||
<taxonomicName id="4C544D34F119FFE9768BFABA50087AB9" authorityName="Cameron" authorityYear="1930" box="[942,1148,1385,1409]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="234" phylum="Arthropoda" rank="species" species="granulipennis">
|
||
<emphasis id="B920EAA5F119FFE9768BFABA50087AB9" box="[942,1148,1385,1409]" italics="true" pageId="21" pageNumber="234">Zyras granulipennis</emphasis>
|
||
</taxonomicName>
|
||
has been recorded only from two localities in
|
||
<collectingRegion id="4990F855F119FFE97127FA5950367A99" box="[1026,1090,1417,1441]" country="Malaysia" name="Sabah" pageId="21" pageNumber="234">Sabah</collectingRegion>
|
||
(Borneo),
|
||
<collectingCountry id="F3437627F119FFE97193FA5951637A99" box="[1206,1303,1417,1441]" name="Malaysia" pageId="21" pageNumber="234">Malaysia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |