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<document id="CA79935EA900A0C0339923598D925FCF" ID-CLB-Dataset="2470" ID-DOI="10.21248/contrib.entomol.67.2.213-246" ID-GBIF-Dataset="43c6cab4-bf7d-43c9-b9bf-b22c4ebf7924" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742339" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638267510724" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03FD87A1F115FFE6761AFB4351E97B19" docLanguage="en" docName="ContribEntomol.67.2.213-246.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) hlavaci Assing 2017, spec. nov." docType="treatment" docUuid="F5734436-CE49-4064-8A52-BC7DF9A0EE67" docUuidSource="ZooBank" docVersion="13" lastPageNumber="239" masterDocId="FFC4FFD9F10CFFFC7525FFD054747F38" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="246" masterPageNumber="213" pageNumber="238" updateTime="1699471357828" updateUser="ExternalLinkService">
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<mods:title id="FBDD69368F2A0C7FB842E88C6426F644">On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:date id="DC9F98559B9CED8B1F468647FB526F5C">2017</mods:date>
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<taxonomicName id="4C544D34F115FFE5761AFB43507B7B96" ID-CoL="5DDD2" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[831,1039,1171,1198]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="species" species="hlavaci" status="sp. nov." subGenus="Zyras">
<emphasis id="B920EAA5F115FFE5761AFB4357017B96" box="[831,885,1171,1198]" italics="true" pageId="25" pageNumber="238">Zyras</emphasis>
(
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)
<emphasis id="B920EAA5F115FFE576E2FB43507B7B96" box="[967,1039,1171,1198]" italics="true" pageId="25" pageNumber="238">hlavaci</emphasis>
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(
<figureCitation id="136F2A32F115FFE57666FB0B57EE7BD7" box="[835,922,1243,1263]" captionStart="Figs 126" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 126: Antenna (118) and forebody (1926) of Zyras spp. (117, 1926) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (1112; 11: holotype; 12 female paratype); hlavaci (1314; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="25" pageNumber="238">Figs 1314</figureCitation>
,
<figureCitation id="136F2A32F115FFE57684FB0B57C27BD7" box="[929,950,1243,1263]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="25" pageNumber="238">33</figureCitation>
,
<figureCitation id="136F2A32F115FFE57698FB0B57A67BD7" box="[957,978,1243,1263]" captionStart="Figs 4052" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 4052: Abdomen (4049, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (4050) and Myrmedonota modiglianii (5152): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (4950). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="25" pageNumber="238">48</figureCitation>
,
<figureCitation id="136F2A32F115FFE576FCFB0B507D7BD7" box="[985,1033,1243,1263]" captionStart="Figs 87102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87102: Zyras parvilobatus (8789), Z. hlavaci (9095; 9092: holotype; 9395: paratype from Selangor), Z. hirtiventris, holotype (9698), Z. lunatus (99101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (8788, 9091, 9394, 9697, 99100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="25" pageNumber="238">9095</figureCitation>
)
</paragraph>
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<emphasis id="B920EAA5F115FFE5763FFAD950407A19" bold="true" box="[794,1076,1289,1313]" pageId="25" pageNumber="238">Type material examined</emphasis>
:
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<typeStatus id="54EF8815F115FFE5716CFAD950D87A19" box="[1097,1196,1289,1313]" pageId="25" pageNumber="238" type="holotype">Holotype</typeStatus>
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: “
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,
<collectingRegion id="4990F855F115FFE5763FFAF957D27A79" box="[794,934,1321,1345]" country="Malaysia" name="Pahang" pageId="25" pageNumber="238">Pahang state</collectingRegion>
,
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,
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,
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4.
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,
<collectorName id="26A15361F115FFE5712EFA9950107A59" box="[1035,1124,1353,1377]" pageId="25" pageNumber="238">Martinů</collectorName>
leg. /
<typeStatus id="54EF8815F115FFE5718CFA99516C7A59" box="[1193,1304,1353,1377]" pageId="25" pageNumber="238" type="holotype">Holotypus</typeStatus>
<taxonomicName id="4C544D34F115FFE57018FA9A571C7AB9" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="species" species="lunatus" status="sp. nov.">
<emphasis id="B920EAA5F115FFE57018FA9A571C7AB9" italics="true" pageId="25" pageNumber="238">Zyras lunatus</emphasis>
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, det.
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V.
<bibRefCitation id="EFC54B46F115FFE576DCFAB950347AB9" author="ASSING, V." box="[1017,1088,1385,1409]" pageId="25" pageNumber="238" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing</bibRefCitation>
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2017” (cAss)
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.
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<typeStatus id="54EF8815F115FFE571FCFABA51377ABA" box="[1241,1347,1386,1410]" pageId="25" pageNumber="238" type="paratype">Paratypes</typeStatus>
: 1 ♀: “
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,
<collectingMunicipality id="6B8FACCDF115FFE57169FA59515D7A99" box="[1100,1321,1417,1441]" pageId="25" pageNumber="238">Cameron Highlands</collectingMunicipality>
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,
<collectorName id="26A15361F115FFE571BBFA7951307AF9" box="[1182,1348,1449,1473]" pageId="25" pageNumber="238">Cechovsky Petr</collectorName>
leg.” (cHla);
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2 ♂♂: “
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:
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(
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<collectionCode id="ED45AE72F115FFE571E7FA39509A7939" box="[1218,1262,1513,1537]" pageId="25" pageNumber="238">FIT</collectionCode>
),
<location id="8E8B606CF115FFE57027FA3A574C7919" LSID="urn:lsid:plazi:treatment:03FD87A1F115FFE6761AFB4351E97B19:8E8B606CF115FFE57027FA3A574C7919" country="Malaysia" county="West" municipality="Cameron Highlands" name="Maruyama M." pageId="25" pageNumber="238" stateProvince="Pahang">Maruyama M.</location>
et al.” (cMar, cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF115FFE5763FF999509F7839" pageId="25" pageNumber="238" type="etymology">
<paragraph id="8BEB36B7F115FFE5763FF999509F7839" blockId="25.[794,1399,1609,1794]" pageId="25" pageNumber="238">
<emphasis id="B920EAA5F115FFE5763FF99957E57959" bold="true" box="[794,913,1609,1633]" pageId="25" pageNumber="238">Etymology</emphasis>
: This species is dedicated to my friend and colleague Peter Hlaváč (Prague), who provided, or arranged the loan of, substantial material not only for the present study, but also for previous revisions of Palaearctic and Oriental
<taxonomicName id="4C544D34F115FFE5768CF91A579779D9" authorityName="STEPHENS" authorityYear="1833" box="[937,995,1738,1761]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="238" phylum="Arthropoda" rank="genus">
<emphasis id="B920EAA5F115FFE5768CF91A579779D9" box="[937,995,1738,1761]" italics="true" pageId="25" pageNumber="238">Zyras</emphasis>
</taxonomicName>
sensu strictu, and to whom I owe the generous gift of the
<typeStatus id="54EF8815F115FFE576C8F939503F7839" box="[1005,1099,1769,1793]" pageId="25" pageNumber="238" type="holotype">holotype</typeStatus>
of this species.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF115FFE6763FF8F9508D7C99" lastPageId="26" lastPageNumber="239" pageId="25" pageNumber="238" type="description">
<paragraph id="8BEB36B7F115FFE5763FF8F951277719" blockId="25.[794,1399,1833,2081]" pageId="25" pageNumber="238">
<emphasis id="B920EAA5F115FFE5763FF8F957EF7879" bold="true" box="[794,923,1833,1857]" pageId="25" pageNumber="238">Description</emphasis>
: Body length
<quantity id="4CAC9B52F115FFE57109F8F950DF7879" box="[1068,1195,1833,1857]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" metricValueMax="8.5" metricValueMin="6.9" pageId="25" pageNumber="238" unit="mm" value="7.7" valueMax="8.5" valueMin="6.9">6.98.5 mm</quantity>
; length of forebody 3.0
<quantity id="4CAC9B52F115FFE57662F89957EE7859" box="[839,922,1865,1889]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="25" pageNumber="238" unit="mm" value="3.5">3.5 mm</quantity>
. Coloration (
<figureCitation id="136F2A32F115FFE57114F89950D27859" box="[1073,1190,1865,1889]" captionStart="Figs 126" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 126: Antenna (118) and forebody (1926) of Zyras spp. (117, 1926) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (1112; 11: holotype; 12 female paratype); hlavaci (1314; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="25" pageNumber="238">Figs 1314</figureCitation>
,
<figureCitation id="136F2A32F115FFE57192F89950A67859" box="[1207,1234,1865,1889]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="25" pageNumber="238">33</figureCitation>
,
<figureCitation id="136F2A32F115FFE571C6F89951747859" box="[1251,1280,1865,1889]" captionStart="Figs 4052" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 4052: Abdomen (4049, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (4050) and Myrmedonota modiglianii (5152): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (4950). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="25" pageNumber="238">48</figureCitation>
): forebody dark-brown to blackish; abdomen dark-reddish, with or without the antero-median portion of tergite VI diffusely infuscate and the antero-median portion of tergite VII indistinctly darker; legs yellowish to pale yellowish-brown; antennae brown to dark-brown; maxillary palpi reddishyellow to reddish with the apical palpomere yellowish.
</paragraph>
<paragraph id="8BEB36B7F116FFE67598FF39558D7D79" blockId="26.[189,794,233,1985]" pageId="26" pageNumber="239">
Head (
<figureCitation id="136F2A32F116FFE67422FF3955267E39" box="[263,338,233,257]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="26" pageNumber="239">Fig. 33</figureCitation>
) distinctly transverse; punctation moderately coarse to coarse and rather dense, median dorsal portion extensively impunctate; pubescence long, suberect to erect, and pale to brown. Eyes moderately large, as long as postocular region in dorsal view, or slightly longer. Antenna (
<figureCitation id="136F2A32F116FFE674A5FE5955827E99" box="[384,502,393,417]" captionStart="Figs 126" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 126: Antenna (118) and forebody (1926) of Zyras spp. (117, 1926) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (1112; 11: holotype; 12 female paratype); hlavaci (1314; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="26" pageNumber="239">Figs 1314</figureCitation>
) 2.0
<quantity id="4CAC9B52F116FFE6771DFE5956FD7E99" box="[568,649,393,417]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="26" pageNumber="239" unit="mm" value="2.3">2.3 mm</quantity>
long; antennomeres IV approximately as long as broad or weakly oblong, VX of gradually increasing width and increasingly transverse, X approximately twice as broad as long, and XI of conical shape and short, shorter than the combined length of IX and X.
</paragraph>
<paragraph id="8BEB36B7F116FFE67598FD9A56217CB9" blockId="26.[189,794,233,1985]" pageId="26" pageNumber="239">
Pronotum (
<figureCitation id="136F2A32F116FFE6741DFD9955F77D59" box="[312,387,585,609]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="26" pageNumber="239">Fig. 33</figureCitation>
) 1.141.15 times as broad as long and 1.211.25 times as broad as head, broadest in anterior half, moderately convex in cross-section; posterior angles obtusely marked; lateral margins weakly sinuate in posterior half in dorsal view; punctation coarse, dense, and somewhat irregularly distributed, laterally and near posterior margin with impunctate patches; midline narrowly to broadly impunctate; pubescence long, pale, and suberect; lateral and anterior margins with numerous longer, stouter, and distinctly erect dark setae.
</paragraph>
<paragraph id="8BEB36B7F116FFE67598FC5956447B79" blockId="26.[189,794,233,1985]" pageId="26" pageNumber="239">
Elytra (
<figureCitation id="136F2A32F116FFE67437FC5955287C99" box="[274,348,905,929]" captionStart="Figs 2739" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 2739: Forebody (2733), abdomen (3438), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 2738: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="26" pageNumber="239">Fig. 33</figureCitation>
) 0.850.89 times as long as pronotum; punctation moderately dense and moderately fine to moderately coarse, nearly regularly distributed, and defined; pubescence long, pale, and sub-erect. Hind wings fully developed. Metatarsomere I slightly shorter than the combined length of IIIV.
</paragraph>
<paragraph id="8BEB36B7F116FFE67598FB99563879D9" blockId="26.[189,794,233,1985]" pageId="26" pageNumber="239">
Abdomen (
<figureCitation id="136F2A32F116FFE67464FB9955FF7B59" box="[321,395,1097,1121]" captionStart="Figs 4052" captionStartId="10.[190,231,1972,1994]" captionTargetBox="[192,1434,232,1963]" captionTargetId="figure-129@10.[192,1434,232,1963]" captionTargetPageId="10" captionText="Figs 4052: Abdomen (4049, 52), abdominal tergite VI (50), and forebody (51) of Zyras spp. (4050) and Myrmedonota modiglianii (5152): latibasalis (40); matangensis, holotype (41); flavorufus, lectotype (42); parvilobatus (43); kinabaluensis, holotype (44); rutrilobatus (45); granulipennis, holotype (46); lunatus (47); hlavaci, holotype (48); hirtiventris, holotype (4950). Scale bars:1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="26" pageNumber="239">Fig. 48</figureCitation>
) nearly as broad as elytra, with moderately deep anterior impressions on tergites IIIV; tergites IIIV with non-setiferous punctation only in lateral portions of anterior impressions, with a transverse row of 48 setiferous punctures in posterior portions, with usually eight setiferous punctures at posterior margins, and with or without additional non-setiferous punctures on discs; tergite VI with dense and coarse non-setiferous punctation in anterior half, sparser nonsetiferous punctation in posterior half, with some lateral setiferous punctures, and with some setiferous punctures at posterior margin; tergite VII with dense non-setiferous punctation anteriorly, with sparser non-setiferous punctation on remainder of disc, and with two indistinct transverse rows of setiferous punctures posteriorly, posterior margin with palisade fringe; tergite VIII with setiferous punctures bearing a mix of yellowish and long black setae in posterior third and non-setiferous punctures in median third, posterior margin truncate or very weakly concave in the middle; all sternites with long and dense pubescence in posterior halves.
</paragraph>
<paragraph id="8BEB36B7F116FFE67598F93A561778B9" blockId="26.[189,794,233,1985]" pageId="26" pageNumber="239">
♂: sternite VIII with convex posterior margin; median lobe of aedeagus
<quantity id="4CAC9B52F116FFE674A5F8D9566D7819" box="[384,537,1801,1825]" metricMagnitude="-4" metricUnit="m" metricValue="8.65" metricValueMax="9.0" metricValueMin="8.299999999999999" pageId="26" pageNumber="239" unit="mm" value="0.865" valueMax="0.9" valueMin="0.83">0.830.90 mm</quantity>
long and shaped as in
<figureCitation id="136F2A32F116FFE67598F8F955097879" box="[189,381,1833,1857]" captionStart="Figs 87102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87102: Zyras parvilobatus (8789), Z. hlavaci (9095; 9092: holotype; 9395: paratype from Selangor), Z. hirtiventris, holotype (9698), Z. lunatus (99101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (8788, 9091, 9394, 9697, 99100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="26" pageNumber="239">Figs 9091, 9394</figureCitation>
; ventral process slender and subapically distinctly angled; paramere (
<figureCitation id="136F2A32F116FFE674D2F89956047859" box="[503,624,1865,1889]" captionStart="Figs 87102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87102: Zyras parvilobatus (8789), Z. hlavaci (9095; 9092: holotype; 9395: paratype from Selangor), Z. hirtiventris, holotype (9698), Z. lunatus (99101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (8788, 9091, 9394, 9697, 99100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="26" pageNumber="239">Figs 92, 95</figureCitation>
) approximately
<quantity id="4CAC9B52F116FFE67598F8B9557B78B9" box="[189,271,1897,1921]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="26" pageNumber="239" unit="mm" value="0.9">0.9 mm</quantity>
long and with short apical lobe.
</paragraph>
<paragraph id="8BEB36B7F116FFE67598F85A554378F9" blockId="26.[189,794,233,1985]" pageId="26" pageNumber="239">♀: posterior margin of sternite VIII weakly concave in the middle.</paragraph>
<paragraph id="8BEB36B7F116FFE67598F83951557E99" blockId="26.[189,794,2025,2081]" lastBlockId="26.[831,1437,233,417]" pageId="26" pageNumber="239">
<emphasis id="B920EAA5F116FFE67598F83955C57739" bold="true" box="[189,433,2025,2049]" pageId="26" pageNumber="239">Intraspecific variation</emphasis>
: The
<typeStatus id="54EF8815F116FFE674CBF83A5620773A" box="[494,596,2026,2050]" pageId="26" pageNumber="239" type="paratype">paratypes</typeStatus>
from
<collectingRegion id="4990F855F116FFE677B3F83956867739" box="[662,754,2025,2049]" country="Malaysia" name="Selangor" pageId="26" pageNumber="239">Selangor</collectingRegion>
are distinguished from the
<typeStatus id="54EF8815F116FFE6749EF7D9566D7719" box="[443,537,2057,2081]" pageId="26" pageNumber="239" type="holotype">holotype</typeStatus>
by slightly smaller size, and by a slightly smaller aedeagus with an apically more acute ventral process (both in lateral and in ventral view) (
<figureCitation id="136F2A32F116FFE67662FEF950667E79" box="[839,1042,297,321]" captionStart="Figs 87102" captionStartId="13.[151,192,1969,1991]" captionTargetBox="[162,1396,242,1953]" captionTargetId="figure-120@13.[154,1396,234,1965]" captionTargetPageId="13" captionText="Figs 87102: Zyras parvilobatus (8789), Z. hlavaci (9095; 9092: holotype; 9395: paratype from Selangor), Z. hirtiventris, holotype (9698), Z. lunatus (99101), and Myrmedonota modiglanii (102): median lobe of aedeagus in lateral and in ventral view (8788, 9091, 9394, 9697, 99100); paramere (89, 92, 95, 98, 101); spermatheca (102). Scale bars: 87101: 0.5 mm; 102: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="26" pageNumber="239">Figs 9091, 9394</figureCitation>
). Since no additional evidence was found suggesting that the material from
<collectingRegion id="4990F855F116FFE671C3FE9951417E59" box="[1254,1333,329,353]" country="Malaysia" name="Pahang" pageId="26" pageNumber="239">Pahang</collectingRegion>
and from
<collectingRegion id="4990F855F116FFE6761AFEB957EF7EB9" box="[831,923,361,385]" country="Malaysia" name="Selangor" pageId="26" pageNumber="239">Selangor</collectingRegion>
should represent distinct species, these differences are attributed to intraspecific variation.
</paragraph>
<paragraph id="8BEB36B7F116FFE6761AFE19508D7C99" blockId="26.[831,1437,457,929]" pageId="26" pageNumber="239">
<emphasis id="B920EAA5F116FFE6761AFE1950627ED9" bold="true" box="[831,1046,457,481]" pageId="26" pageNumber="239">Comparative notes</emphasis>
: This species, too, belongs to the
<taxonomicName id="4C544D34F116FFE6761AFE3A57EE7D39" baseAuthorityName="KRAATZ" baseAuthorityYear="1859" box="[831,922,489,513]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="239" phylum="Arthropoda" rank="species" species="hirtus">
<emphasis id="B920EAA5F116FFE6761AFE3A57EE7D39" box="[831,922,489,513]" italics="true" pageId="26" pageNumber="239">Z. hirtus</emphasis>
</taxonomicName>
group. It is distinguished from the similar
<taxonomicName id="4C544D34F116FFE6761AFDDA57DE7D19" authorityName="Assing" authorityYear="2017" box="[831,938,521,545]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="239" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B920EAA5F116FFE6761AFDDA57DE7D19" box="[831,938,521,545]" italics="true" pageId="26" pageNumber="239">Z. lunatus</emphasis>
</taxonomicName>
particularly by larger body size, darker coloration, by the punctation pattern of the abdomen (especially more extensive and denser non-setiferous punctation on the posterior tergites, also on tergite VIII), and by the more robust median lobe of the aedeagus with a more pronounced apex (ventral view). It differs from
<taxonomicName id="4C544D34F116FFE67066FD7A57A47DD9" authority="ASSING, 2016" authorityName="ASSING" authorityYear="2016" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="239" phylum="Arthropoda" rank="species" species="flexus">
<emphasis id="B920EAA5F116FFE67066FD7A51E87DF9" box="[1347,1436,681,705]" italics="true" pageId="26" pageNumber="239">Z. flexus</emphasis>
ASSING, 2016
</taxonomicName>
(
<collectingCountry id="F3437627F116FFE676C4FD1950537DD9" box="[993,1063,713,737]" name="China" pageId="26" pageNumber="239">China</collectingCountry>
:
<collectingRegion id="4990F855F116FFE67116FD19500D7DD9" box="[1075,1145,713,737]" country="China" name="Fujian" pageId="26" pageNumber="239">Fujian</collectingRegion>
) by somewhat larger body size, uniformly dark antennae (
<taxonomicName id="4C544D34F116FFE671AFFD3A50977C39" authorityName="ASSING" authorityYear="2016" box="[1162,1251,745,769]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="239" phylum="Arthropoda" rank="species" species="flexus">
<emphasis id="B920EAA5F116FFE671AFFD3A50977C39" box="[1162,1251,745,769]" italics="true" pageId="26" pageNumber="239">Z. flexus</emphasis>
</taxonomicName>
: antennomere XI yellowish), the presence of an impunctate median band on the less convex pronotum, less dense punctation of the elytra, denser and more extensive non-setiferous punctation on the abdomen, and by a distinctly larger median lobe of the aedeagus (
<taxonomicName id="4C544D34F116FFE67103FC5A500B7C99" authorityName="ASSING" authorityYear="2016" box="[1062,1151,905,929]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="239" phylum="Arthropoda" rank="species" species="flexus">
<emphasis id="B920EAA5F116FFE67103FC5A500B7C99" box="[1062,1151,905,929]" italics="true" pageId="26" pageNumber="239">Z. flexus</emphasis>
</taxonomicName>
:
<quantity id="4CAC9B52F116FFE671AEFC59509A7C99" box="[1163,1262,905,929]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="26" pageNumber="239" unit="mm" value="0.75">0.75 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF116FFE6761AFC1951E97B19" pageId="26" pageNumber="239" type="distribution">
<paragraph id="8BEB36B7F116FFE6761AFC1951E97B19" blockId="26.[831,1437,969,1057]" pageId="26" pageNumber="239">
<emphasis id="B920EAA5F116FFE6761AFC1950DA7CD9" bold="true" box="[831,1198,969,993]" pageId="26" pageNumber="239">Distribution and natural history</emphasis>
: The
<typeStatus id="54EF8815F116FFE671D0FC1A51577CDA" box="[1269,1315,970,994]" pageId="26" pageNumber="239">type</typeStatus>
specimens were collected in three localities in
<collectingRegion id="4990F855F116FFE6719DFC3951727B39" box="[1208,1286,1001,1025]" country="Malaysia" name="Pahang" pageId="26" pageNumber="239">Pahang</collectingRegion>
and
<collectingRegion id="4990F855F116FFE67018FC3951ED7B39" box="[1341,1433,1001,1025]" country="Malaysia" name="Selangor" pageId="26" pageNumber="239">Selangor</collectingRegion>
, Peninsular
<collectingCountry id="F3437627F116FFE67690FBD950607B19" box="[949,1044,1033,1057]" name="Malaysia" pageId="26" pageNumber="239">Malaysia</collectingCountry>
, at altitudes between 250 and
<quantity id="4CAC9B52F116FFE6706DFBD951EC7B19" box="[1352,1432,1033,1057]" metricMagnitude="3" metricUnit="m" metricValue="1.65" pageId="26" pageNumber="239" unit="m" value="1650.0">1650 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>