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<document id="04983ED74F9EA3DFF189309C6E09A867" ID-CLB-Dataset="2470" ID-DOI="10.21248/contrib.entomol.67.2.213-246" ID-GBIF-Dataset="43c6cab4-bf7d-43c9-b9bf-b22c4ebf7924" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742339" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638267510724" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="03FD87A1F10BFFF2761AFA0350067EF9" docLanguage="en" docName="ContribEntomol.67.2.213-246.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (2)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Zyras (Zyras) punctipennis Cameron 1939" docType="treatment" docVersion="11" lastPageNumber="227" masterDocId="FFC4FFD9F10CFFFC7525FFD054747F38" masterDocTitle="On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)" masterLastPageNumber="246" masterPageNumber="213" pageNumber="220" updateTime="1699471357828" updateUser="ExternalLinkService">
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<mods:title id="BB1FB7599C57CB6EFD29900CAD9C5022">On Zyras sensu strictu in the East Palaearctic and Oriental regions III, with a focus on the Southeast of Continental Asia and the Sunda Islands (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini)</mods:title>
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<mods:namePart id="86DF006F0821153B0413D7CA659AC42E">Assing, Volker</mods:namePart>
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<mods:typeOfResource id="974F17B3626F2C0A7CD6EEB320C44258">text</mods:typeOfResource>
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<mods:title id="AF06E1538B235EEA37945C67247E38A7">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="5ED922B4AD0E28339F00995818B1C44C">
<mods:date id="5BA3E0A26E15020C140460F2897A3D62">2017</mods:date>
<mods:detail id="14F4C3E2A572EB4C138F865D98C47439" type="pubDate">
<mods:number id="57F0EFAAC1308CF29DF8ED25BB12DB25">2017-12-08</mods:number>
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<mods:detail id="00F18B8D060FBACA36C9CAF58BBB7ADD" type="volume">
<mods:number id="07D7B28B25F07D6A6A1282DA04A8DF9A">67</mods:number>
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<mods:classification id="F7ABA9FBC41D15720989B772B356AF2A">journal article</mods:classification>
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<mods:identifier id="82159A7A3760C82C17AA11F56FAAF3DE" type="DOI">10.21248/contrib.entomol.67.2.213-246</mods:identifier>
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<treatment id="03FD87A1F10BFFF2761AFA0350067EF9" ID-DOI="http://doi.org/10.5281/zenodo.5888790" ID-GBIF-Taxon="192500466" ID-Zenodo-Dep="5888790" LSID="urn:lsid:plazi:treatment:03FD87A1F10BFFF2761AFA0350067EF9" httpUri="http://treatment.plazi.org/id/03FD87A1F10BFFF2761AFA0350067EF9" lastPageId="14" lastPageNumber="227" pageId="7" pageNumber="220">
<subSubSection id="C34E653CF10BFFFB761AFA0350837AD6" box="[831,1271,1491,1518]" pageId="7" pageNumber="220" type="nomenclature">
<paragraph id="8BEB36B7F10BFFFB761AFA0350837AD6" blockId="7.[831,1271,1491,1551]" box="[831,1271,1491,1518]" pageId="7" pageNumber="220">
<heading id="D0A381DBF10BFFFB761AFA0350837AD6" box="[831,1271,1491,1518]" centered="true" fontSize="12" level="2" pageId="7" pageNumber="220" reason="2">
<taxonomicName id="4C544D34F10BFFFB761AFA0350837AD6" ID-CoL="5DDF7" authority="CAMERON, 1939" authorityName="Cameron" authorityYear="1939" box="[831,1271,1491,1518]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="220" phylum="Arthropoda" rank="species" species="punctipennis" subGenus="Zyras">
<emphasis id="B920EAA5F10BFFFB761AFA0357017AD6" box="[831,885,1491,1518]" italics="true" pageId="7" pageNumber="220">Zyras</emphasis>
(
<emphasis id="B920EAA5F10BFFFB76A6FA0357CD7AD6" box="[899,953,1491,1518]" italics="true" pageId="7" pageNumber="220">Zyras</emphasis>
)
<emphasis id="B920EAA5F10BFFFB76E2FA0350387AD6" box="[967,1100,1491,1518]" italics="true" pageId="7" pageNumber="220">punctipennis</emphasis>
CAMERON, 1939
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF10BFFFB761AFA2B57DA7937" box="[831,942,1531,1551]" pageId="7" pageNumber="220" type="description">
<paragraph id="8BEB36B7F10BFFFB761AFA2B57DA7937" blockId="7.[831,1271,1491,1551]" box="[831,942,1531,1551]" pageId="7" pageNumber="220">
(
<figureCitation id="136F2A32F10BFFFB7666FA2B57FB7937" box="[835,911,1531,1551]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="7" pageNumber="220">Figs 1, 19</figureCitation>
,
<figureCitation id="136F2A32F10BFFFB76B3FA2B57DE7937" box="[918,938,1531,1551]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="7" pageNumber="220">34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF10BFFFB763FF9FC50837979" box="[794,1271,1579,1601]" pageId="7" pageNumber="220" type="reference_group">
<paragraph id="8BEB36B7F10BFFFB763FF9FC50837979" blockId="7.[794,1271,1579,1601]" box="[794,1271,1579,1601]" pageId="7" pageNumber="220">
<treatmentCitationGroup id="AB441199F10BFFFB763FF9FC50837979" box="[794,1271,1579,1601]" pageId="7" pageNumber="220">
<treatmentCitation id="0AF510A6F10BFFFB763FF9FC50907978" author="CAMERON, M." box="[794,1252,1579,1601]" page="18" pageId="7" pageNumber="220" year="1939">
<taxonomicName id="4C544D34F10BFFFB763FF9FC50907978" ID-CoL="5DDF7" authority="CAMERON, 1939 b: 18" authorityName="Cameron" authorityPageNumber="18" authorityYear="1939" box="[794,1252,1579,1601]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="220" phylum="Arthropoda" rank="species" species="punctipennis" subGenus="Zyras">
<emphasis id="B920EAA5F10BFFFB763FF9FC573A7979" box="[794,846,1580,1601]" italics="true" pageId="7" pageNumber="220">Zyras</emphasis>
(
<emphasis id="B920EAA5F10BFFFB7679F9FC57E47979" box="[860,912,1580,1601]" italics="true" pageId="7" pageNumber="220">Zyras</emphasis>
)
<emphasis id="B920EAA5F10BFFFB76B8F9FC50617979" box="[925,1045,1580,1601]" italics="true" pageId="7" pageNumber="220">punctipennis</emphasis>
<bibRefCitation id="EFC54B46F10BFFFB713EF9FB50907978" author="CAMERON, M." box="[1051,1252,1579,1601]" pageId="7" pageNumber="220" pagination="1 - 29" refId="ref26173" refString="CAMERON, M. 1939 b: Fauna Javanica. The Staphylinidae collected by Mr. F. C. Drescher. Part III. - Tijdschrift voor Entomologie 82: 1 - 29." type="journal article" year="1939">
CAMERON, 1939b:
<specimenCount id="9D52FD3EF10BFFFB71E8F9FB50907978" box="[1229,1252,1579,1600]" pageId="7" pageNumber="220" type="generic">18</specimenCount>
</bibRefCitation>
</taxonomicName>
</treatmentCitation>
f.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF10BFFFB763FF9B950447719" pageId="7" pageNumber="220" type="materials_examined">
<paragraph id="8BEB36B7F10BFFFB763FF9B951227839" blockId="7.[794,1399,1641,1793]" pageId="7" pageNumber="220">
<emphasis id="B920EAA5F10BFFFB763FF9B9505D79B9" bold="true" box="[794,1065,1641,1665]" pageId="7" pageNumber="220">Type material examined</emphasis>
:
<materialsCitation id="3B3C3CEAF10BFFFB711CF9B951227839" ID-GBIF-Occurrence="3450187322" collectingDate="1933-09" collectionCode="BMNH" collectorName="M. Cameron. &amp; Bequest." determinerName="V. Assing" elevation="0" location="Tangkoeban Prahoe" municipality="Voet" pageId="7" pageNumber="220" specimenCount="1" stateProvince="Jawa Timur" typeStatus="holotype">
<typeStatus id="54EF8815F10BFFFB711CF9B950E879B9" box="[1081,1180,1641,1665]" pageId="7" pageNumber="220" type="holotype">Holotype</typeStatus>
<specimenCount id="9D52FD3EF10BFFFB7183F9BA50CF79B9" box="[1190,1211,1642,1665]" pageId="7" pageNumber="220" type="generic">♀</specimenCount>
: “F. C. Drescher, G.
<location id="8E8B606CF10BFFFB761CF95950547999" LSID="urn:lsid:plazi:treatment:03FD87A1F10BFFF2761AFA0350067EF9:8E8B606CF10BFFFB761CF95950547999" box="[825,1056,1673,1697]" municipality="Voet" name="Tangkoeban Prahoe" pageId="7" pageNumber="220" stateProvince="Jawa Timur">Tangkoeban Prahoe</location>
,
<elevation id="0079D184F10BFFFB711FF95950DA7999" box="[1082,1198,1673,1697]" pageId="7" pageNumber="220">4000–5000</elevation>
Voet, Preanger,
<location id="8E8B606CF10BFFFB763FF97A573C79F9" LSID="urn:lsid:plazi:treatment:03FD87A1F10BFFF2761AFA0350067EF9:8E8B606CF10BFFFB763FF97A573C79F9" box="[794,840,1706,1729]" municipality="Voet" name="Java" pageId="7" pageNumber="220" stateProvince="Jawa Timur">Java</location>
,
<date id="FFEA1077F10BFFFB7671F97957DD79F9" box="[852,937,1705,1729]" pageId="7" pageNumber="220" value="1933-09">
<collectingDate id="EFAEE99FF10BFFFB7671F97957DD79F9" box="[852,937,1705,1729]" pageId="7" pageNumber="220" value="1933-09">IX.1933</collectingDate>
</date>
/
<taxonomicName id="4C544D34F10BFFFB76E1F97A50DD79F9" authority="Cam." authorityName="Cameron" authorityYear="1939" box="[964,1193,1705,1729]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="220" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B920EAA5F10BFFFB76E1F97A501179F9" box="[964,1125,1706,1729]" italics="true" pageId="7" pageNumber="220">Z. punctipennis</emphasis>
Cam.
</taxonomicName>
Type / Holotype /
<collectorName id="26A15361F10BFFFB763FF91A507E79D9" box="[794,1034,1737,1761]" pageId="7" pageNumber="220">M. Cameron. Bequest.</collectorName>
B.M. 1955-147 / Holotypus
<taxonomicName id="4C544D34F10BFFFB7018F91A507F7839" authority="Cameron" authorityName="Cameron" authorityYear="1939" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="220" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B920EAA5F10BFFFB7018F91A57EB7839" italics="true" pageId="7" pageNumber="220">Zyras punctipennis</emphasis>
Cameron
</taxonomicName>
, rev.
<determinerName id="7E37DA73F10BFFFB7164F93A50D77839" box="[1089,1187,1769,1793]" pageId="7" pageNumber="220">
V.
<bibRefCitation id="EFC54B46F10BFFFB7179F93950D77839" author="ASSING, V." box="[1116,1187,1769,1793]" pageId="7" pageNumber="220" pagination="117 - 192" refId="ref25856" refString="ASSING, V. 2017: On Zyras sensu strictu in the East Palaearctic and Oriental regions, with a focus on the faunas of the Himalaya, India, Sri Lanka, Thailand, and Sulawesi (Coleoptera: Staphylinidae: Aleocharinae: Lomechusini). - Contributions to Entomology 67 (1): 117 - 192 - DOI: 10.21248 / contrib. entomol. 67.1.117 - 192." type="journal article" year="2017">Assing</bibRefCitation>
</determinerName>
2017” (
<collectionCode id="ED45AE72F10BFFFB71D0F939513F7839" box="[1269,1355,1769,1793]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="7" pageNumber="220">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BEB36B7F10BFFFB763FF8F9571E78B9" blockId="7.[794,1399,1833,1921]" pageId="7" pageNumber="220">
<emphasis id="B920EAA5F10BFFFB763FF8F957FD7879" bold="true" box="[794,905,1833,1857]" pageId="7" pageNumber="220">Comment</emphasis>
: The original description is based on a unique holoype from “G. Tangkoeban Prahoe” (
<bibRefCitation id="EFC54B46F10BFFFB702CF899572B78B9" author="CAMERON, M." pageId="7" pageNumber="220" pagination="1 - 29" refId="ref26173" refString="CAMERON, M. 1939 b: Fauna Javanica. The Staphylinidae collected by Mr. F. C. Drescher. Part III. - Tijdschrift voor Entomologie 82: 1 - 29." type="journal article" year="1939">CAMERON 1939b</bibRefCitation>
).
</paragraph>
<paragraph id="8BEB36B7F10BFFFB763FF87950447719" blockId="7.[794,1399,1961,2081]" pageId="7" pageNumber="220">
<emphasis id="B920EAA5F10BFFFB763FF879501F78F9" bold="true" box="[794,1131,1961,1985]" pageId="7" pageNumber="220">Additional material examined</emphasis>
:
<materialsCitation id="3B3C3CEAF10BFFFB715EF879505F7719" ID-GBIF-Occurrence="3450187340" collectingDate="2001-08" collectingMethod="flight interception trap" collectionCode="BMNH" collectorName="Brendell &amp; Mendel" country="Indonesia" latitude="-0.05" location="Busang-Rekut" longLatPrecision="1308" longitude="113.98333" municipality="Busang-Rekut" pageId="7" pageNumber="220" specimenCount="1" stateProvince="Kalimantan Tengah">
<emphasis id="B920EAA5F10BFFFB715EF879509C78F9" bold="true" box="[1147,1256,1961,1985]" pageId="7" pageNumber="220">
<collectingCountry id="F3437627F10BFFFB715EF879509C78F9" box="[1147,1256,1961,1985]" name="Indonesia" pageId="7" pageNumber="220">Indonesia</collectingCountry>
</emphasis>
: 5 ♀♀,
<collectingRegion id="4990F855F10BFFFB7067F87957A778D9" country="Indonesia" name="Kalimantan Tengah" pageId="7" pageNumber="220">Kalimantan Tengah</collectingRegion>
,
<collectingMunicipality id="6B8FACCDF10BFFFB76CAF81950F178D9" box="[1007,1157,1993,2017]" country="Indonesia" latitude="-0.05" longLatPrecision="1308" longitude="113.98333" name="Busang-Rekut" pageId="7" pageNumber="220" stateProvince="Kalimantan Tengah">Busang-Rekut</collectingMunicipality>
confluence,
<geoCoordinate id="EE605070F10BFFFB7015F819510678D9" box="[1328,1394,1993,2017]" degrees="0" direction="south" minutes="03" orientation="latitude" pageId="7" pageNumber="220" precision="925" value="-0.05">0°03'S</geoCoordinate>
,
<geoCoordinate id="EE605070F10BFFFB763FF839570C7739" box="[794,888,2025,2049]" degrees="113" direction="east" minutes="59" orientation="longitude" pageId="7" pageNumber="220" precision="925" value="113.98333">113°59'E</geoCoordinate>
,
<collectingMethod id="52154EA0F10BFFFB76A7F83950017739" box="[898,1141,2025,2049]" pageId="7" pageNumber="220">flight interception trap</collectingMethod>
,
<date id="FFEA1077F10BFFFB715AF839509C7739" box="[1151,1256,2025,2049]" pageId="7" pageNumber="220" value="2001-08">
<collectingDate id="EFAEE99FF10BFFFB715AF839509C7739" box="[1151,1256,2025,2049]" pageId="7" pageNumber="220" value="2001-08">VIII.2001</collectingDate>
</date>
, leg.
<collectorName id="26A15361F10BFFFB7038F83951037739" box="[1309,1399,2025,2049]" pageId="7" pageNumber="220">Brendell</collectorName>
&amp;
<collectorName id="26A15361F10BFFFB7616F7D957F07719" box="[819,900,2057,2081]" pageId="7" pageNumber="220">Mendel</collectorName>
(
<collectionCode id="ED45AE72F10BFFFB76B4F7D957917719" box="[913,997,2057,2081]" pageId="7" pageNumber="220">BMNH</collectionCode>
, cAss)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF2B663FF104FFF47598F846574C7719" ID-DOI="http://doi.org/10.5281/zenodo.5742343" ID-Zenodo-Dep="5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="8" pageNumber="221" startId="8.[189,229,1942,1964]" targetBox="[189,1436,238,1930]" targetPageId="8">
<paragraph id="8BEB36B7F104FFF47598F846574C7719" blockId="8.[189,1436,1942,2081]" pageId="8" pageNumber="221">
<emphasis id="B920EAA5F104FFF47598F84655697893" bold="true" box="[189,285,1942,1964]" pageId="8" pageNumber="221">Figs 1–26</emphasis>
: Antenna (
<emphasis id="B920EAA5F104FFF474A8F84655C87893" bold="true" box="[397,444,1942,1963]" pageId="8" pageNumber="221">1–18</emphasis>
) and forebody (
<emphasis id="B920EAA5F104FFF4777BF84656ED7893" bold="true" box="[606,665,1942,1963]" pageId="8" pageNumber="221">19–26</emphasis>
) of
<taxonomicName id="4C544D34F104FFF477E1F847575C7893" box="[708,808,1942,1964]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B920EAA5F104FFF477E1F847568C7894" box="[708,760,1943,1964]" italics="true" pageId="8" pageNumber="221">Zyras</emphasis>
spp.
</taxonomicName>
(
<emphasis id="B920EAA5F104FFF47613F84657DA7893" bold="true" box="[822,942,1942,1963]" pageId="8" pageNumber="221">1–17, 19–26</emphasis>
) and
<taxonomicName id="4C544D34F104FFF476CFF84650967893" authorityName="Assing" authorityYear="2017" baseAuthorityName="Cameron" baseAuthorityYear="1925" box="[1002,1250,1942,1963]" class="Insecta" family="Staphylinidae" genus="Myrmedonota" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="modiglianii">
<emphasis id="B920EAA5F104FFF476CFF84650967893" box="[1002,1250,1942,1963]" italics="true" pageId="8" pageNumber="221">Myrmedonota modiglianii</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F104FFF471D7F846517E7893" bold="true" box="[1266,1290,1942,1963]" pageId="8" pageNumber="221">18</emphasis>
):
<taxonomicName id="4C544D34F104FFF4703AF84751E37894" authorityName="Cameron" authorityYear="1939" box="[1311,1431,1943,1964]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B920EAA5F104FFF4703AF84751E37894" box="[1311,1431,1943,1964]" italics="true" pageId="8" pageNumber="221">punctipennis</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F104FFF47406F863552178F0" bold="true" box="[291,341,1971,1992]" pageId="8" pageNumber="221">1, 19</emphasis>
);
<taxonomicName id="4C544D34F104FFF47449F86355B578F0" authorityName="Last" authorityYear="1969" box="[364,449,1971,1992]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="facundus">
<emphasis id="B920EAA5F104FFF47449F86355B578F0" box="[364,449,1971,1992]" italics="true" pageId="8" pageNumber="221">facundus</emphasis>
</taxonomicName>
, lectotype (
<emphasis id="B920EAA5F104FFF4771DF863561F78F0" bold="true" box="[568,619,1971,1992]" pageId="8" pageNumber="221">2, 24</emphasis>
);
<taxonomicName id="4C544D34F104FFF477A7F863569478F0" authorityName="Assing" authorityYear="2017" box="[642,736,1971,1992]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="latibasalis">
<emphasis id="B920EAA5F104FFF477A7F863569478F0" box="[642,736,1971,1992]" italics="true" pageId="8" pageNumber="221">latibasalis</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F104FFF477D4F863575078F0" bold="true" box="[753,804,1971,1992]" pageId="8" pageNumber="221">3, 25</emphasis>
);
<taxonomicName id="4C544D34F104FFF4761FF86357CD78F0" authorityName="Pace" authorityYear="2008" box="[826,953,1971,1992]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B920EAA5F104FFF4761FF86357CD78F0" box="[826,953,1971,1992]" italics="true" pageId="8" pageNumber="221">kinabaluensis</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F104FFF4710BF863501578F0" bold="true" box="[1070,1121,1971,1992]" pageId="8" pageNumber="221">4, 21</emphasis>
);
<taxonomicName id="4C544D34F104FFF47152F864509F78F1" authorityName="Cameron" authorityYear="1943" box="[1143,1259,1972,1993]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="matangensis">
<emphasis id="B920EAA5F104FFF47152F864509F78F1" box="[1143,1259,1972,1993]" italics="true" pageId="8" pageNumber="221">matangensis</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F104FFF47045F86351FB78F0" bold="true" box="[1376,1423,1971,1992]" pageId="8" pageNumber="221">5, 26</emphasis>
);
<taxonomicName id="4C544D34F104FFF47598F801557678DE" authorityName="Cameron" authorityYear="1943" box="[189,258,2001,2022]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="B920EAA5F104FFF47598F801557678DE" box="[189,258,2001,2022]" italics="true" pageId="8" pageNumber="221">bryanti</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F104FFF4745DF80055D878DD" bold="true" box="[376,428,2000,2021]" pageId="8" pageNumber="221">6, 23</emphasis>
);
<taxonomicName id="4C544D34F104FFF474E1F801563078DE" authorityName="Cameron" authorityYear="1930" box="[452,580,2001,2022]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="granulipennis">
<emphasis id="B920EAA5F104FFF474E1F801563078DE" box="[452,580,2001,2022]" italics="true" pageId="8" pageNumber="221">granulipennis</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F104FFF4779FF80156B278DE" bold="true" box="[698,710,2001,2022]" pageId="8" pageNumber="221">7</emphasis>
);
<taxonomicName id="4C544D34F104FFF477FBF801574978DE" authorityName="Cameron" authorityYear="1939" box="[734,829,2001,2022]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="flavorufus">
<emphasis id="B920EAA5F104FFF477FBF801574978DE" box="[734,829,2001,2022]" italics="true" pageId="8" pageNumber="221">flavorufus</emphasis>
</taxonomicName>
, lectotype (
<emphasis id="B920EAA5F104FFF47692F80057B778DD" bold="true" box="[951,963,2000,2021]" pageId="8" pageNumber="221">8</emphasis>
);
<taxonomicName id="4C544D34F104FFF476FEF801503F78DE" authorityName="Assing" authorityYear="2017" box="[987,1099,2001,2022]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="rutrilobatus">
<emphasis id="B920EAA5F104FFF476FEF801503F78DE" box="[987,1099,2001,2022]" italics="true" pageId="8" pageNumber="221">rutrilobatus</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F104FFF47178F800501D78DD" bold="true" box="[1117,1129,2000,2021]" pageId="8" pageNumber="221">9</emphasis>
);
<taxonomicName id="4C544D34F104FFF471A4F801508278DE" authorityName="Assing" authorityYear="2017" box="[1153,1270,2001,2022]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="parvilobatus">
<emphasis id="B920EAA5F104FFF471A4F801508278DE" box="[1153,1270,2001,2022]" italics="true" pageId="8" pageNumber="221">parvilobatus</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F104FFF47021F800516878DD" bold="true" box="[1284,1308,2000,2021]" pageId="8" pageNumber="221">10</emphasis>
);
<taxonomicName id="4C544D34F104FFF47011F80151E878DE" authorityName="Assing" authorityYear="2017" box="[1332,1436,2001,2022]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="hirtiventris">
<emphasis id="B920EAA5F104FFF47011F80151E878DE" box="[1332,1436,2001,2022]" italics="true" pageId="8" pageNumber="221">hirtiventris</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F104FFF475E0F83E5575773B" bold="true" box="[197,257,2030,2051]" pageId="8" pageNumber="221">11–12</emphasis>
;
<emphasis id="B920EAA5F104FFF47429F83E5550773B" bold="true" box="[268,292,2030,2051]" pageId="8" pageNumber="221">11</emphasis>
: holotype;
<emphasis id="B920EAA5F104FFF474B7F83E55DE773B" bold="true" box="[402,426,2030,2051]" pageId="8" pageNumber="221">12</emphasis>
female paratype);
<taxonomicName id="4C544D34F104FFF47743F83E56DD773B" authorityName="Assing" authorityYear="2017" box="[614,681,2030,2051]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="hlavaci">
<emphasis id="B920EAA5F104FFF47743F83E56DD773B" box="[614,681,2030,2051]" italics="true" pageId="8" pageNumber="221">hlavaci</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F104FFF4779EF83E5682773B" bold="true" box="[699,758,2030,2051]" pageId="8" pageNumber="221">13–14</emphasis>
;
<emphasis id="B920EAA5F104FFF47623F83E576A773B" bold="true" box="[774,798,2030,2051]" pageId="8" pageNumber="221">13</emphasis>
holotype;
<emphasis id="B920EAA5F104FFF476AFF83E57D6773B" bold="true" box="[906,930,2030,2051]" pageId="8" pageNumber="221">14</emphasis>
paratype from Selangor);
<taxonomicName id="4C544D34F104FFF4718EF83E5085773B" authorityName="Assing" authorityYear="2017" box="[1195,1265,2030,2051]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B920EAA5F104FFF4718EF83E5085773B" box="[1195,1265,2030,2051]" italics="true" pageId="8" pageNumber="221">lunatus</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F104FFF471DAF83E5163773B" bold="true" box="[1279,1303,2030,2051]" pageId="8" pageNumber="221">15</emphasis>
);
<taxonomicName id="4C544D34F104FFF4700BF83E51E3773B" authorityName="Cameron" authorityYear="1939" box="[1326,1431,2030,2051]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="B920EAA5F104FFF4700BF83E51E3773B" box="[1326,1431,2030,2051]" italics="true" pageId="8" pageNumber="221">elegantulus</emphasis>
</taxonomicName>
, lectotype (
<emphasis id="B920EAA5F104FFF47407F7DB55297718" bold="true" box="[290,349,2059,2080]" pageId="8" pageNumber="221">16, 20</emphasis>
);
<taxonomicName id="4C544D34F104FFF47455F7DB55B17718" authorityName="Pace" authorityYear="2008" box="[368,453,2059,2080]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="221" phylum="Arthropoda" rank="species" species="pallipyga">
<emphasis id="B920EAA5F104FFF47455F7DB55B17718" box="[368,453,2059,2080]" italics="true" pageId="8" pageNumber="221">pallipyga</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F104FFF47717F7DB56187718" bold="true" box="[562,620,2059,2080]" pageId="8" pageNumber="221">17, 22</emphasis>
). Scale bars: 1.0 mm.
</paragraph>
</caption>
<caption id="DF2B663FF105FFF575B2F86256787718" ID-DOI="http://doi.org/10.5281/zenodo.5742345" ID-Zenodo-Dep="5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="9" pageNumber="222" startId="9.[151,191,1970,1992]" targetBox="[154,1396,232,1963]" targetPageId="9">
<paragraph id="8BEB36B7F105FFF575B2F86256787718" blockId="9.[151,1399,1970,2081]" pageId="9" pageNumber="222">
<emphasis id="B920EAA5F105FFF575B2F862557578FF" bold="true" box="[151,257,1970,1992]" pageId="9" pageNumber="222">Figs 27–39</emphasis>
: Forebody (
<emphasis id="B920EAA5F105FFF57453F86255C678FF" bold="true" box="[374,434,1970,1991]" pageId="9" pageNumber="222">27–33</emphasis>
), abdomen (
<emphasis id="B920EAA5F105FFF5770BF862561E78FF" bold="true" box="[558,618,1970,1991]" pageId="9" pageNumber="222">34–38</emphasis>
), and median portion of tergite VIII (
<emphasis id="B920EAA5F105FFF576F2F862579B78FF" bold="true" box="[983,1007,1970,1991]" pageId="9" pageNumber="222">39</emphasis>
) of
<taxonomicName id="4C544D34F105FFF57133F863500D78F0" box="[1046,1145,1971,1992]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="222" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B920EAA5F105FFF57133F863503E78F0" box="[1046,1098,1971,1992]" italics="true" pageId="9" pageNumber="222">Zyras</emphasis>
spp.
</taxonomicName>
:
<taxonomicName id="4C544D34F105FFF571A4F863509478F0" authorityName="Cameron" authorityYear="1939" box="[1153,1248,1971,1992]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="222" phylum="Arthropoda" rank="species" species="flavorufus">
<emphasis id="B920EAA5F105FFF571A4F863509478F0" box="[1153,1248,1971,1992]" italics="true" pageId="9" pageNumber="222">flavorufus</emphasis>
</taxonomicName>
, lectotype (
<emphasis id="B920EAA5F105FFF57074F862511D78FF" bold="true" box="[1361,1385,1970,1991]" pageId="9" pageNumber="222">27</emphasis>
);
<taxonomicName id="4C544D34F105FFF575B2F800557878DD" authorityName="Assing" authorityYear="2017" box="[151,268,2000,2021]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="222" phylum="Arthropoda" rank="species" species="parvilobatus">
<emphasis id="B920EAA5F105FFF575B2F800557878DD" box="[151,268,2000,2021]" italics="true" pageId="9" pageNumber="222">parvilobatus</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F105FFF5743FF800554678DD" bold="true" box="[282,306,2000,2021]" pageId="9" pageNumber="222">28</emphasis>
);
<taxonomicName id="4C544D34F105FFF57460F80055FF78DD" authorityName="Assing" authorityYear="2017" box="[325,395,2000,2021]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="222" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B920EAA5F105FFF57460F80055FF78DD" box="[325,395,2000,2021]" italics="true" pageId="9" pageNumber="222">lunatus</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F105FFF574BDF80055C478DD" bold="true" box="[408,432,2000,2021]" pageId="9" pageNumber="222">29</emphasis>
);
<taxonomicName id="4C544D34F105FFF574E6F800563778DD" authorityName="Cameron" authorityYear="1930" box="[451,579,2000,2021]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="222" phylum="Arthropoda" rank="species" species="granulipennis">
<emphasis id="B920EAA5F105FFF574E6F800563778DD" box="[451,579,2000,2021]" italics="true" pageId="9" pageNumber="222">granulipennis</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F105FFF5778BF80056B278DD" bold="true" box="[686,710,2000,2021]" pageId="9" pageNumber="222">30</emphasis>
);
<taxonomicName id="4C544D34F105FFF577FDF800573C78DD" authorityName="Assing" authorityYear="2017" box="[728,840,2000,2021]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="222" phylum="Arthropoda" rank="species" species="rutrilobatus">
<emphasis id="B920EAA5F105FFF577FDF800573C78DD" box="[728,840,2000,2021]" italics="true" pageId="9" pageNumber="222">rutrilobatus</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F105FFF57673F800571A78DD" bold="true" box="[854,878,2000,2021]" pageId="9" pageNumber="222">31</emphasis>
);
<taxonomicName id="4C544D34F105FFF576A5F800579C78DD" authorityName="Assing" authorityYear="2017" box="[896,1000,2000,2021]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="222" phylum="Arthropoda" rank="species" species="hirtiventris">
<emphasis id="B920EAA5F105FFF576A5F800579C78DD" box="[896,1000,2000,2021]" italics="true" pageId="9" pageNumber="222">hirtiventris</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F105FFF576D0F800507978DD" bold="true" box="[1013,1037,2000,2021]" pageId="9" pageNumber="222">32</emphasis>
);
<taxonomicName id="4C544D34F105FFF57105F800501778DD" authorityName="Assing" authorityYear="2017" box="[1056,1123,2000,2021]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="222" phylum="Arthropoda" rank="species" species="hlavaci">
<emphasis id="B920EAA5F105FFF57105F800501778DD" box="[1056,1123,2000,2021]" italics="true" pageId="9" pageNumber="222">hlavaci</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F105FFF571EAF800509378DD" bold="true" box="[1231,1255,2000,2021]" pageId="9" pageNumber="222">33</emphasis>
);
<taxonomicName id="4C544D34F105FFF571DCF801510578DE" authorityName="Cameron" authorityYear="1939" box="[1273,1393,2001,2022]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="222" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B920EAA5F105FFF571DCF801510578DE" box="[1273,1393,2001,2022]" italics="true" pageId="9" pageNumber="222">punctipennis</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F105FFF575DCF83D5565773A" bold="true" box="[249,273,2029,2050]" pageId="9" pageNumber="222">34</emphasis>
);
<taxonomicName id="4C544D34F105FFF57401F83D55F9773A" authorityName="Cameron" authorityYear="1939" box="[292,397,2029,2050]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="222" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="B920EAA5F105FFF57401F83D55F9773A" box="[292,397,2029,2050]" italics="true" pageId="9" pageNumber="222">elegantulus</emphasis>
</taxonomicName>
, lectotype (
<emphasis id="B920EAA5F105FFF574DBF83D5662773A" bold="true" box="[510,534,2029,2050]" pageId="9" pageNumber="222">35</emphasis>
);
<taxonomicName id="4C544D34F105FFF5770CF83D560A773A" authorityName="Last" authorityYear="1969" box="[553,638,2029,2050]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="222" phylum="Arthropoda" rank="species" species="facundus">
<emphasis id="B920EAA5F105FFF5770CF83D560A773A" box="[553,638,2029,2050]" italics="true" pageId="9" pageNumber="222">facundus</emphasis>
</taxonomicName>
, lectotype (
<emphasis id="B920EAA5F105FFF577CAF83D5773773A" bold="true" box="[751,775,2029,2050]" pageId="9" pageNumber="222">36</emphasis>
);
<taxonomicName id="4C544D34F105FFF5763FF83D571B773A" authorityName="Pace" authorityYear="2008" box="[794,879,2029,2050]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="222" phylum="Arthropoda" rank="species" species="pallipyga">
<emphasis id="B920EAA5F105FFF5763FF83D571B773A" box="[794,879,2029,2050]" italics="true" pageId="9" pageNumber="222">pallipyga</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F105FFF576F8F83D5781773A" bold="true" box="[989,1013,2029,2050]" pageId="9" pageNumber="222">37</emphasis>
);
<taxonomicName id="4C544D34F105FFF5712DF83D5039773A" authorityName="Cameron" authorityYear="1943" box="[1032,1101,2029,2050]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="222" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="B920EAA5F105FFF5712DF83D5039773A" box="[1032,1101,2029,2050]" italics="true" pageId="9" pageNumber="222">bryanti</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F105FFF5719FF83D50A6773A" bold="true" box="[1210,1234,2029,2050]" pageId="9" pageNumber="222">38</emphasis>
);
<taxonomicName id="4C544D34F105FFF571C0F83D5137773A" authorityName="Assing" authorityYear="2017" box="[1253,1347,2029,2050]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="222" phylum="Arthropoda" rank="species" species="latibasalis">
<emphasis id="B920EAA5F105FFF571C0F83D5137773A" box="[1253,1347,2029,2050]" italics="true" pageId="9" pageNumber="222">latibasalis</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F105FFF57074F83D511D773A" bold="true" box="[1361,1385,2029,2050]" pageId="9" pageNumber="222">39</emphasis>
). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm.
</paragraph>
</caption>
<caption id="DF2B663FF106FFF6759BF86455077719" ID-DOI="http://doi.org/10.5281/zenodo.5742347" ID-Zenodo-Dep="5742347" httpUri="https://zenodo.org/record/5742347/files/figure.png" pageId="10" pageNumber="223" startId="10.[190,231,1972,1994]" targetBox="[192,1434,232,1963]" targetPageId="10">
<paragraph id="8BEB36B7F106FFF6759BF86455077719" blockId="10.[190,1437,1972,2082]" pageId="10" pageNumber="223">
<emphasis id="B920EAA5F106FFF6759BF864555978F1" bold="true" box="[190,301,1972,1994]" pageId="10" pageNumber="223">Figs 40–52</emphasis>
: Abdomen (
<emphasis id="B920EAA5F106FFF67495F864566178F1" bold="true" box="[432,533,1972,1993]" pageId="10" pageNumber="223">40–49, 52</emphasis>
), abdominal tergite VI (
<emphasis id="B920EAA5F106FFF6762AF864575378F1" bold="true" box="[783,807,1972,1993]" pageId="10" pageNumber="223">50</emphasis>
), and forebody (
<emphasis id="B920EAA5F106FFF676FDF864578478F1" bold="true" box="[984,1008,1972,1993]" pageId="10" pageNumber="223">51</emphasis>
) of
<taxonomicName id="4C544D34F106FFF67105F86550F378F1" box="[1056,1159,1972,1994]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="223" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B920EAA5F106FFF67105F865502178F2" box="[1056,1109,1973,1994]" italics="true" pageId="10" pageNumber="223">Zyras</emphasis>
spp.
</taxonomicName>
(
<emphasis id="B920EAA5F106FFF671BDF86450A178F1" bold="true" box="[1176,1237,1972,1993]" pageId="10" pageNumber="223">40–50</emphasis>
) and
<taxonomicName id="4C544D34F106FFF67033F864555E78DE" authorityName="Assing" authorityYear="2017" baseAuthorityName="Cameron" baseAuthorityYear="1925" class="Insecta" family="Staphylinidae" genus="Myrmedonota" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="223" phylum="Arthropoda" rank="species" species="modiglianii">
<emphasis id="B920EAA5F106FFF67033F864555E78DE" italics="true" pageId="10" pageNumber="223">Myrmedonota modiglianii</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F106FFF6741DF801550078DE" bold="true" box="[312,372,2001,2022]" pageId="10" pageNumber="223">51–52</emphasis>
):
<taxonomicName id="4C544D34F106FFF674A8F801559878DE" authorityName="Assing" authorityYear="2017" box="[397,492,2001,2022]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="223" phylum="Arthropoda" rank="species" species="latibasalis">
<emphasis id="B920EAA5F106FFF674A8F801559878DE" box="[397,492,2001,2022]" italics="true" pageId="10" pageNumber="223">latibasalis</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F106FFF67725F801566C78DE" bold="true" box="[512,536,2001,2022]" pageId="10" pageNumber="223">40</emphasis>
);
<taxonomicName id="4C544D34F106FFF67717F80256D378DF" authorityName="Cameron" authorityYear="1943" box="[562,679,2002,2023]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="223" phylum="Arthropoda" rank="species" species="matangensis">
<emphasis id="B920EAA5F106FFF67717F80256D378DF" box="[562,679,2002,2023]" italics="true" pageId="10" pageNumber="223">matangensis</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F106FFF67604F801574D78DE" bold="true" box="[801,825,2001,2022]" pageId="10" pageNumber="223">41</emphasis>
);
<taxonomicName id="4C544D34F106FFF67677F80157C678DE" authorityName="Cameron" authorityYear="1939" box="[850,946,2001,2022]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="223" phylum="Arthropoda" rank="species" species="flavorufus">
<emphasis id="B920EAA5F106FFF67677F80157C678DE" box="[850,946,2001,2022]" italics="true" pageId="10" pageNumber="223">flavorufus</emphasis>
</taxonomicName>
, lectotype (
<emphasis id="B920EAA5F106FFF6710AF801503378DE" bold="true" box="[1071,1095,2001,2022]" pageId="10" pageNumber="223">42</emphasis>
);
<taxonomicName id="4C544D34F106FFF67144F80150AC78DE" authorityName="Assing" authorityYear="2017" box="[1121,1240,2001,2022]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="223" phylum="Arthropoda" rank="species" species="parvilobatus">
<emphasis id="B920EAA5F106FFF67144F80150AC78DE" box="[1121,1240,2001,2022]" italics="true" pageId="10" pageNumber="223">parvilobatus</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F106FFF671C0F801508978DE" bold="true" box="[1253,1277,2001,2022]" pageId="10" pageNumber="223">43</emphasis>
);
<taxonomicName id="4C544D34F106FFF67032F80151EC78DE" authorityName="Pace" authorityYear="2008" box="[1303,1432,2001,2022]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="223" phylum="Arthropoda" rank="species" species="kinabaluensis">
<emphasis id="B920EAA5F106FFF67032F80151EC78DE" box="[1303,1432,2001,2022]" italics="true" pageId="10" pageNumber="223">kinabaluensis</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F106FFF67404F83F554D773C" bold="true" box="[289,313,2031,2052]" pageId="10" pageNumber="223">44</emphasis>
);
<taxonomicName id="4C544D34F106FFF67468F83F55CA773C" authorityName="Assing" authorityYear="2017" box="[333,446,2031,2052]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="223" phylum="Arthropoda" rank="species" species="rutrilobatus">
<emphasis id="B920EAA5F106FFF67468F83F55CA773C" box="[333,446,2031,2052]" italics="true" pageId="10" pageNumber="223">rutrilobatus</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F106FFF674E8F83F5591773C" bold="true" box="[461,485,2031,2052]" pageId="10" pageNumber="223">45</emphasis>
);
<taxonomicName id="4C544D34F106FFF674DFF83F560F773C" authorityName="Cameron" authorityYear="1930" box="[506,635,2031,2052]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="223" phylum="Arthropoda" rank="species" species="granulipennis">
<emphasis id="B920EAA5F106FFF674DFF83F560F773C" box="[506,635,2031,2052]" italics="true" pageId="10" pageNumber="223">granulipennis</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F106FFF677CEF83F5777773C" bold="true" box="[747,771,2031,2052]" pageId="10" pageNumber="223">46</emphasis>
);
<taxonomicName id="4C544D34F106FFF67632F83F572A773C" authorityName="Assing" authorityYear="2017" box="[791,862,2031,2052]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="223" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B920EAA5F106FFF67632F83F572A773C" box="[791,862,2031,2052]" italics="true" pageId="10" pageNumber="223">lunatus</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F106FFF67648F83F57F1773C" bold="true" box="[877,901,2031,2052]" pageId="10" pageNumber="223">47</emphasis>
);
<taxonomicName id="4C544D34F106FFF676BFF83F57AA773C" authorityName="Assing" authorityYear="2017" box="[922,990,2031,2052]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="223" phylum="Arthropoda" rank="species" species="hlavaci">
<emphasis id="B920EAA5F106FFF676BFF83F57AA773C" box="[922,990,2031,2052]" italics="true" pageId="10" pageNumber="223">hlavaci</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F106FFF6716BF83F5012773C" bold="true" box="[1102,1126,2031,2052]" pageId="10" pageNumber="223">48</emphasis>
);
<taxonomicName id="4C544D34F106FFF6715FF83F5097773C" authorityName="Assing" authorityYear="2017" box="[1146,1251,2031,2052]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="223" phylum="Arthropoda" rank="species" species="hirtiventris">
<emphasis id="B920EAA5F106FFF6715FF83F5097773C" box="[1146,1251,2031,2052]" italics="true" pageId="10" pageNumber="223">hirtiventris</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F106FFF67076F83F51FB773C" bold="true" box="[1363,1423,2031,2052]" pageId="10" pageNumber="223">49–50</emphasis>
). Scale bars:1.0 mm.
</paragraph>
</caption>
<caption id="DF2B663FF107FFF775B2F86355267719" ID-DOI="http://doi.org/10.5281/zenodo.5888738" ID-Zenodo-Dep="5888738" httpUri="https://zenodo.org/record/5888738/files/figure.png" pageId="11" pageNumber="224" startId="11.[151,192,1971,1993]" targetBox="[155,1387,245,1958]" targetPageId="11">
<paragraph id="8BEB36B7F107FFF775B2F86355267719" blockId="11.[151,1398,1971,2081]" pageId="11" pageNumber="224">
<emphasis id="B920EAA5F107FFF775B2F863557978F0" bold="true" box="[151,269,1971,1993]" pageId="11" pageNumber="224">Figs 53–71</emphasis>
:
<taxonomicName id="4C544D34F107FFF77401F86455A078F0" authorityName="Cameron" authorityYear="1939" box="[292,468,1971,1993]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="224" phylum="Arthropoda" rank="species" species="elegantulus">
<emphasis id="B920EAA5F107FFF77401F86455A078F0" box="[292,468,1971,1993]" italics="true" pageId="11" pageNumber="224">Zyras elegantulus</emphasis>
</taxonomicName>
,
<typeStatus id="54EF8815F107FFF774CEF863563778F1" box="[491,579,1971,1993]" pageId="11" pageNumber="224" type="lectotype">lectotype</typeStatus>
(
<emphasis id="B920EAA5F107FFF77778F86356EE78F0" bold="true" box="[605,666,1971,1992]" pageId="11" pageNumber="224">53–55</emphasis>
),
<taxonomicName id="4C544D34F107FFF7779CF864575D78F0" authorityName="Last" authorityYear="1969" box="[697,809,1971,1993]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="224" phylum="Arthropoda" rank="species" species="facundus">
<emphasis id="B920EAA5F107FFF7779CF864575D78F0" box="[697,809,1971,1993]" italics="true" pageId="11" pageNumber="224">Z. facundus</emphasis>
</taxonomicName>
,
<typeStatus id="54EF8815F107FFF77665F86357EC78F1" box="[832,920,1971,1993]" pageId="11" pageNumber="224" type="lectotype">lectotype</typeStatus>
(
<emphasis id="B920EAA5F107FFF77697F863579A78F0" bold="true" box="[946,1006,1971,1992]" pageId="11" pageNumber="224">56–58</emphasis>
),
<taxonomicName id="4C544D34F107FFF7712BF864501878F0" authorityName="Cameron" authorityYear="1943" box="[1038,1132,1971,1993]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="224" phylum="Arthropoda" rank="species" species="bryanti">
<emphasis id="B920EAA5F107FFF7712BF864501878F0" box="[1038,1132,1971,1993]" italics="true" pageId="11" pageNumber="224">Z. bryanti</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F107FFF771A3F86350B778F0" bold="true" box="[1158,1219,1971,1992]" pageId="11" pageNumber="224">59–62</emphasis>
),
<taxonomicName id="4C544D34F107FFF771C7F864510578F1" authorityName="Cameron" authorityYear="1943" box="[1250,1393,1972,1993]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="224" phylum="Arthropoda" rank="species" species="matangensis">
<emphasis id="B920EAA5F107FFF771C7F864510578F1" box="[1250,1393,1972,1993]" italics="true" pageId="11" pageNumber="224">Z. matangensis</emphasis>
</taxonomicName>
,
<typeStatus id="54EF8815F107FFF775B2F801549878DF" box="[151,236,2001,2023]" pageId="11" pageNumber="224" type="holotype">holotype</typeStatus>
(
<emphasis id="B920EAA5F107FFF775DFF800554378DD" bold="true" box="[250,311,2000,2021]" pageId="11" pageNumber="224">63–65</emphasis>
),
<taxonomicName id="4C544D34F107FFF7746FF801559178DE" authorityName="Cameron" authorityYear="1930" box="[330,485,2001,2022]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="224" phylum="Arthropoda" rank="species" species="granulipennis">
<emphasis id="B920EAA5F107FFF7746FF801559178DE" box="[330,485,2001,2022]" italics="true" pageId="11" pageNumber="224">Z. granulipennis</emphasis>
</taxonomicName>
,
<typeStatus id="54EF8815F107FFF774D5F801563178DF" box="[496,581,2001,2023]" pageId="11" pageNumber="224" type="holotype">holotype</typeStatus>
(
<emphasis id="B920EAA5F107FFF77776F80056FB78DD" bold="true" box="[595,655,2000,2021]" pageId="11" pageNumber="224">66–69</emphasis>
), and
<taxonomicName id="4C544D34F107FFF777E9F801573178DE" authorityName="Assing" authorityYear="2017" box="[716,837,2001,2022]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="224" phylum="Arthropoda" rank="species" species="latibasalis">
<emphasis id="B920EAA5F107FFF777E9F801573178DE" box="[716,837,2001,2022]" italics="true" pageId="11" pageNumber="224">Z. latibasalis</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F107FFF77676F80057FA78DD" bold="true" box="[851,910,2000,2021]" pageId="11" pageNumber="224">70–71</emphasis>
): median lobe of aedeagus in lateral and in ventral view (
<emphasis id="B920EAA5F107FFF775F7F83E560D773B" bold="true" box="[210,633,2030,2051]" pageId="11" pageNumber="224">53–54, 56–57, 59–60, 63–64, 66–67, 70–71</emphasis>
); paramere (
<emphasis id="B920EAA5F107FFF777DCF83E5741773B" bold="true" box="[761,821,2030,2051]" pageId="11" pageNumber="224">55, 58</emphasis>
); male tergite VIII (
<emphasis id="B920EAA5F107FFF776DEF83E502F773B" bold="true" box="[1019,1115,2030,2051]" pageId="11" pageNumber="224">61, 65, 68</emphasis>
); male sternite VIII (
<emphasis id="B920EAA5F107FFF77009F83E511D773B" bold="true" box="[1324,1385,2030,2051]" pageId="11" pageNumber="224">62, 69</emphasis>
). Scale bars:
<quantity id="4CAC9B52F107FFF77427F7DB553A7719" box="[258,334,2059,2081]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="224" unit="mm" value="0.2">0.2 mm</quantity>
.
</paragraph>
</caption>
<caption id="DF2B663FF100FFF07598F86355BC7719" ID-DOI="http://doi.org/10.5281/zenodo.5742353" ID-Zenodo-Dep="5742353" httpUri="https://zenodo.org/record/5742353/files/figure.png" pageId="12" pageNumber="225" startId="12.[189,229,1971,1993]" targetBox="[198,1434,242,1959]" targetPageId="12">
<paragraph id="8BEB36B7F100FFF07598F86355BC7719" blockId="12.[189,1437,1971,2081]" pageId="12" pageNumber="225">
<emphasis id="B920EAA5F100FFF07598F863555178F0" bold="true" box="[189,293,1971,1993]" pageId="12" pageNumber="225">Figs 72–86</emphasis>
:
<taxonomicName id="4C544D34F100FFF07415F86455B378F0" authorityName="Assing" authorityYear="2017" box="[304,455,1971,1993]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="225" phylum="Arthropoda" rank="species" species="latibasalis">
<emphasis id="B920EAA5F100FFF07415F86455B378F0" box="[304,455,1971,1993]" italics="true" pageId="12" pageNumber="225">Zyras latibasalis</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F100FFF074F1F863559878F0" bold="true" box="[468,492,1971,1992]" pageId="12" pageNumber="225">72</emphasis>
),
<taxonomicName id="4C544D34F100FFF074DBF864560278F0" authorityName="Cameron" authorityYear="1939" box="[510,630,1971,1993]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="225" phylum="Arthropoda" rank="species" species="flavorufus">
<emphasis id="B920EAA5F100FFF074DBF864560278F0" box="[510,630,1971,1993]" italics="true" pageId="12" pageNumber="225">Z. flavorufus</emphasis>
</taxonomicName>
, lectotype (
<emphasis id="B920EAA5F100FFF077C0F863575578F0" bold="true" box="[741,801,1971,1992]" pageId="12" pageNumber="225">73–75</emphasis>
),
<taxonomicName id="4C544D34F100FFF07616F864571878F1" authorityName="PACE" authorityYear="1993" box="[819,876,1972,1993]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="225" phylum="Arthropoda" rank="species" species="wei">
<emphasis id="B920EAA5F100FFF07616F864571878F1" box="[819,876,1972,1993]" italics="true" pageId="12" pageNumber="225">Z. wei</emphasis>
</taxonomicName>
from Laos (
<emphasis id="B920EAA5F100FFF076FAF863506F78F0" bold="true" box="[991,1051,1971,1992]" pageId="12" pageNumber="225">76–77</emphasis>
),
<taxonomicName id="4C544D34F100FFF07108F86450C378F1" authorityName="PACE" authorityYear="1998" box="[1069,1207,1972,1993]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="225" phylum="Arthropoda" rank="species" species="notaticornis">
<emphasis id="B920EAA5F100FFF07108F86450C378F1" box="[1069,1207,1972,1993]" italics="true" pageId="12" pageNumber="225">Z. notaticornis</emphasis>
</taxonomicName>
from Laos (
<emphasis id="B920EAA5F100FFF0700FF863511278F0" bold="true" box="[1322,1382,1971,1992]" pageId="12" pageNumber="225">78–80</emphasis>
), and
<taxonomicName id="4C544D34F100FFF07598F801553278DE" authorityName="Assing" authorityYear="2017" box="[189,326,2001,2022]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="225" phylum="Arthropoda" rank="species" species="rutrilobatus">
<emphasis id="B920EAA5F100FFF07598F801553278DE" box="[189,326,2001,2022]" italics="true" pageId="12" pageNumber="225">Z. rutrilobatus</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F100FFF07470F80055E578DD" bold="true" box="[341,401,2000,2021]" pageId="12" pageNumber="225">81–86</emphasis>
): paramere (
<emphasis id="B920EAA5F100FFF0772AF80056E778DD" bold="true" box="[527,659,2000,2022]" pageId="12" pageNumber="225">72, 75, 80, 83</emphasis>
); median lobe of aedeagus in lateral and in ventral view (
<emphasis id="B920EAA5F100FFF071E6F80051FA78DD" bold="true" box="[1219,1422,2000,2021]" pageId="12" pageNumber="225">73–74, 76–79, 81–82</emphasis>
); male tergite VIII (
<emphasis id="B920EAA5F100FFF0744BF83E55F2773B" bold="true" box="[366,390,2030,2051]" pageId="12" pageNumber="225">84</emphasis>
); male sternite VIII (
<emphasis id="B920EAA5F100FFF07744F83E560D773B" bold="true" box="[609,633,2030,2051]" pageId="12" pageNumber="225">85</emphasis>
); postero-median portion of male sternite VIII (
<emphasis id="B920EAA5F100FFF07141F83E5008773B" bold="true" box="[1124,1148,2030,2051]" pageId="12" pageNumber="225">86</emphasis>
). Scale bars: 84–85: 0.5 mm; 72–83: 0.2 mm; 86: 0.1 mm.
</paragraph>
</caption>
<caption id="DF2B663FF101FFF175B2F86155627727" ID-DOI="http://doi.org/10.5281/zenodo.5742357" ID-Zenodo-Dep="5742357" httpUri="https://zenodo.org/record/5742357/files/figure.png" pageId="13" pageNumber="226" startId="13.[151,192,1969,1991]" targetBox="[162,1396,242,1953]" targetPageId="13">
<paragraph id="8BEB36B7F101FFF175B2F86155627727" blockId="13.[151,1399,1969,2079]" pageId="13" pageNumber="226">
<emphasis id="B920EAA5F101FFF175B2F861556578FF" bold="true" box="[151,273,1969,1991]" pageId="13" pageNumber="226">Figs 87–102</emphasis>
:
<taxonomicName id="4C544D34F101FFF17405F86255A078FF" authorityName="Assing" authorityYear="2017" box="[288,468,1970,1991]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="226" phylum="Arthropoda" rank="species" species="parvilobatus">
<emphasis id="B920EAA5F101FFF17405F86255A078FF" box="[288,468,1970,1991]" italics="true" pageId="13" pageNumber="226">Zyras parvilobatus</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F101FFF174C3F862565678FF" bold="true" box="[486,546,1970,1991]" pageId="13" pageNumber="226">87–89</emphasis>
),
<taxonomicName id="4C544D34F101FFF1771CF86256E378FF" authorityName="Assing" authorityYear="2017" box="[569,663,1970,1991]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="226" phylum="Arthropoda" rank="species" species="hlavaci">
<emphasis id="B920EAA5F101FFF1771CF86256E378FF" box="[569,663,1970,1991]" italics="true" pageId="13" pageNumber="226">Z. hlavaci</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F101FFF1778DF862575B78FF" bold="true" box="[680,815,1970,1991]" pageId="13" pageNumber="226">90–95; 90–92</emphasis>
: holotype;
<emphasis id="B920EAA5F101FFF17687F86257AA78FF" bold="true" box="[930,990,1970,1991]" pageId="13" pageNumber="226">93–95</emphasis>
: paratype from Selangor),
<taxonomicName id="4C544D34F101FFF171CBF862510478FF" authorityName="Assing" authorityYear="2017" box="[1262,1392,1970,1991]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="226" phylum="Arthropoda" rank="species" species="hirtiventris">
<emphasis id="B920EAA5F101FFF171CBF862510478FF" box="[1262,1392,1970,1991]" italics="true" pageId="13" pageNumber="226">Z. hirtiventris</emphasis>
</taxonomicName>
, holotype (
<emphasis id="B920EAA5F101FFF175DFF81F554378DC" bold="true" box="[250,311,1999,2020]" pageId="13" pageNumber="226">96–98</emphasis>
),
<taxonomicName id="4C544D34F101FFF17468F80055DA78DC" authorityName="Assing" authorityYear="2017" box="[333,430,1999,2021]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="226" phylum="Arthropoda" rank="species" species="lunatus">
<emphasis id="B920EAA5F101FFF17468F80055DA78DC" box="[333,430,1999,2021]" italics="true" pageId="13" pageNumber="226">Z. lunatus</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F101FFF1749BF81F567278DC" bold="true" box="[446,518,1999,2020]" pageId="13" pageNumber="226">99–101</emphasis>
), and
<taxonomicName id="4C544D34F101FFF1776FF81F574B78DC" box="[586,831,1999,2020]" class="Insecta" family="Staphylinidae" genus="Myrmedonota" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="226" phylum="Arthropoda" rank="species" species="modiglanii">
<emphasis id="B920EAA5F101FFF1776FF81F574B78DC" box="[586,831,1999,2020]" italics="true" pageId="13" pageNumber="226">Myrmedonota modiglanii</emphasis>
</taxonomicName>
(
<emphasis id="B920EAA5F101FFF17675F81F570078DC" bold="true" box="[848,884,1999,2020]" pageId="13" pageNumber="226">102</emphasis>
): median lobe of aedeagus in lateral and in ventral view (
<emphasis id="B920EAA5F101FFF175F7F83C56317739" bold="true" box="[210,581,2028,2049]" pageId="13" pageNumber="226">87–88, 90–91, 93–94, 96–97, 99–100</emphasis>
); paramere (
<emphasis id="B920EAA5F101FFF177EDF83C57F17739" bold="true" box="[712,901,2028,2049]" pageId="13" pageNumber="226">89, 92, 95, 98, 101</emphasis>
); spermatheca (
<emphasis id="B920EAA5F101FFF17102F83C503F7739" bold="true" box="[1063,1099,2028,2049]" pageId="13" pageNumber="226">102</emphasis>
). Scale bars: 87–101: 0.5 mm; 102: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="C34E653CF102FFF27598FF3957D87E79" pageId="14" pageNumber="227" type="description">
<paragraph id="8BEB36B7F102FFF27598FF3955207D59" blockId="14.[189,794,233,1985]" pageId="14" pageNumber="227">
<emphasis id="B920EAA5F102FFF27598FF3955227E39" bold="true" box="[189,342,233,257]" pageId="14" pageNumber="227">Redescription</emphasis>
: Body length 6.5–8.0 mm; length of forebody 3.0–
<quantity id="4CAC9B52F102FFF27416FED955F37E19" box="[307,391,265,289]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="14" pageNumber="227" unit="mm" value="3.4">3.4 mm</quantity>
. Coloration: head and pronotum brown; elytra reddish with the postero-lateral portions somewhat infuscate; abdomen dark-brown anteriorly and blackish-brown to blackish posteriorly, with the anterior and posterior margins of the segments reddish; legs pale-yellow, sometimes with the apices of the meso-and metafemora narrowly infuscate; antennae with antennomeres I–II reddish, III–VII or III–VIII brown to blackish-brown, and IX–XI or VIII–XI yellowish; maxillary palpi reddish with the apical palpomere yellowish-red.
</paragraph>
<paragraph id="8BEB36B7F102FFF27598FDB955FF7CB9" blockId="14.[189,794,233,1985]" pageId="14" pageNumber="227">
Head (
<figureCitation id="136F2A32F102FFF2742BFDB9552D7DB9" box="[270,345,617,641]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="14" pageNumber="227">Fig. 19</figureCitation>
) distinctly transverse, median portion extensively impunctate; punctures in lateral portions sparse and rather coarse. Eyes longer than postocular region in dorsal view. Antenna (
<figureCitation id="136F2A32F102FFF2776FFD1956F27DD9" box="[586,646,713,737]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="14" pageNumber="227">Fig. 1</figureCitation>
)
<quantity id="4CAC9B52F102FFF277BFFD19576E7DD9" box="[666,794,713,737]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" metricValueMax="2.4" metricValueMin="2.2" pageId="14" pageNumber="227" unit="mm" value="2.3" valueMax="2.4" valueMin="2.2">2.2–2.4 mm</quantity>
long; antennomeres IV–VI approximately as long as broad, VII–X increasingly transverse and of gradually increasing width, X barely 1.5 times as broad as long, and XI rather short, slightly shorter than the combined length of IX and X.
</paragraph>
<paragraph id="8BEB36B7F102FFF27598FC5A55497B79" blockId="14.[189,794,233,1985]" pageId="14" pageNumber="227">
Pronotum (
<figureCitation id="136F2A32F102FFF2741EFC5955F17C99" box="[315,389,905,929]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="14" pageNumber="227">Fig. 19</figureCitation>
) approximately 1.1 times as broad as long and 1.3 times as broad as head, broadest in anterior half, strongly convex in cross-section; punctation rather coarse, sparse, and irregularly distributed, with impunctate areas on either side of midline; midline broadly impunctate.
</paragraph>
<paragraph id="8BEB36B7F102FFF27598FB9955157A39" blockId="14.[189,794,233,1985]" pageId="14" pageNumber="227">
Elytra (
<figureCitation id="136F2A32F102FFF27437FB9955287B59" box="[274,348,1097,1121]" captionStart="Figs 1–26" captionStartId="8.[189,229,1942,1964]" captionTargetBox="[189,1436,238,1930]" captionTargetId="figure-170@8.[189,1436,232,1933]" captionTargetPageId="8" captionText="Figs 1–26: Antenna (1–18) and forebody (19–26) of Zyras spp. (1–17, 19–26) and Myrmedonota modiglianii (18): punctipennis, holotype (1, 19); facundus, lectotype (2, 24); latibasalis (3, 25); kinabaluensis, holotype (4, 21); matangensis, holotype (5, 26); bryanti, holotype (6, 23); granulipennis, holotype (7); flavorufus, lectotype (8); rutrilobatus (9); parvilobatus (10); hirtiventris (11–12; 11: holotype; 12 female paratype); hlavaci (13–14; 13 holotype; 14 paratype from Selangor); lunatus (15); elegantulus, lectotype (16, 20); pallipyga, holotype (17, 22). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5742343" httpUri="https://zenodo.org/record/5742343/files/figure.png" pageId="14" pageNumber="227">Fig. 19</figureCitation>
) 0.75–0.80 times as long as pronotum; punctation very coarse, defined, very dense near scutellum, gradually becoming less dense towards posterior margin. Hind wings fully developed. Metatarsomere I very long and slender, slightly longer than the combined length of II–IV.
</paragraph>
<paragraph id="8BEB36B7F102FFF27598FAD956BC78B9" blockId="14.[189,794,233,1985]" pageId="14" pageNumber="227">
Abdomen (
<figureCitation id="136F2A32F102FFF27411FAD9550B7A19" box="[308,383,1289,1313]" captionStart="Figs 27–39" captionStartId="9.[151,191,1970,1992]" captionTargetBox="[154,1396,232,1963]" captionTargetId="figure-145@9.[154,1396,232,1963]" captionTargetPageId="9" captionText="Figs 27–39: Forebody (27–33), abdomen (34–38), and median portion of tergite VIII (39) of Zyras spp.: flavorufus, lectotype (27); parvilobatus (28); lunatus (29); granulipennis, holotype (30); rutrilobatus (31); hirtiventris (32); hlavaci, holotype (33); punctipennis, holotype (34); elegantulus, lectotype (35); facundus, lectotype (36); pallipyga, holotype (37); bryanti, holotype (38); latibasalis (39). Scale bars: 27–38: 1.0 mm; 39: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5742345" httpUri="https://zenodo.org/record/5742345/files/figure.png" pageId="14" pageNumber="227">Fig. 34</figureCitation>
) approximately as broad as elytra, with moderately deep anterior impressions on tergites III–V; anterior impressions of tergites III–V each with a transverse row of dense, coarse, and defined non-setiferous punctures; tergite III with one or two lateral punctures on either side and with twelve setiferous punctures at posterior margin; tergite IV–V with a median pair of punctures, with a short lateral transverse series of three setiferous punctures on either side, and with approximately 15 setiferous punctures at posterior margins; tergite VI with a narrow transverse band of coarse non-setiferous punctures anteriorly, with a median pair of punctures, with a lateral transverse series of three setiferous punctures on either side, and with approximately 15 setiferous punctures at posterior margin; tergite VII with a transverse band of non-setiferous punctures anteriorly and with two transverse series of setiferous punctures in posterior portion, posterior margin with palisade fringe; tergite VIII with setiferous punctures bearing long dark setae in posterior third, posterior margin convex.
</paragraph>
<paragraph id="8BEB36B7F102FFF27598F85A553F7899" blockId="14.[189,794,233,1985]" box="[189,331,1929,1953]" pageId="14" pageNumber="227">♂: unknown.</paragraph>
<paragraph id="8BEB36B7F102FFF27598F87A568578F9" blockId="14.[189,794,233,1985]" box="[189,753,1961,1985]" pageId="14" pageNumber="227">♀: posterior margin of sternite VIII broadly concave.</paragraph>
<paragraph id="8BEB36B7F102FFF27598F83957D87E79" blockId="14.[189,793,2025,2081]" lastBlockId="14.[831,1436,233,321]" pageId="14" pageNumber="227">
<emphasis id="B920EAA5F102FFF27598F83955E37739" bold="true" box="[189,407,2025,2049]" pageId="14" pageNumber="227">Comparative notes</emphasis>
: This species is characterized by relatively large size, its coloration (particularly of the antennae), conspicuously dense and coarse punctation of the elytra, and by the punctation pattern of the abdomen.
</paragraph>
</subSubSection>
<subSubSection id="C34E653CF102FFF2761AFEB950067EF9" pageId="14" pageNumber="227" type="distribution">
<paragraph id="8BEB36B7F102FFF2761AFEB950067EF9" blockId="14.[831,1437,361,449]" pageId="14" pageNumber="227">
<emphasis id="B920EAA5F102FFF2761AFEB957BC7EB9" bold="true" box="[831,968,361,385]" pageId="14" pageNumber="227">Distribution</emphasis>
:
<taxonomicName id="4C544D34F102FFF276F3FEBA50E87EB9" authorityName="Cameron" authorityYear="1939" box="[982,1180,362,385]" class="Insecta" family="Staphylinidae" genus="Zyras" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="227" phylum="Arthropoda" rank="species" species="punctipennis">
<emphasis id="B920EAA5F102FFF276F3FEBA50E87EB9" box="[982,1180,362,385]" italics="true" pageId="14" pageNumber="227">Zyras punctipennis</emphasis>
</taxonomicName>
has been recorded only from the
<typeStatus id="54EF8815F102FFF2768FFE5A57AC7E9A" box="[938,984,394,418]" pageId="14" pageNumber="227">type</typeStatus>
locality in
<collectingRegion id="4990F855F102FFF2717CFE5A50F07E99" box="[1113,1156,394,417]" country="Indonesia" name="Jawa Timur" pageId="14" pageNumber="227">Java</collectingRegion>
and from one locality in
<collectingRegion id="4990F855F102FFF2761AFE7950677EF9" box="[831,1043,425,449]" country="Indonesia" name="Kalimantan Tengah" pageId="14" pageNumber="227">Kalimantan Tengah</collectingRegion>
, Borneo.
</paragraph>
</subSubSection>
</treatment>
</document>
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