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<document id="7B20E838E2FBB0AE18505A7FDE69E9FA" ID-CLB-Dataset="106789" ID-DOI="10.11646/zootaxa.5169.1.1" ID-GBIF-Dataset="b76236d0-0336-4d17-804a-80f0344b097e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6911151" ID-ZooBank="3CB49021-90C5-46F0-AAAF-619EE8068F42" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658900973851" checkinUser="plazi" docAuthor="Grall, Elena &amp; Jäger, Peter" docDate="2022" docId="03FD87925834FFCEFF56FE7E84E34181" docLanguage="en" docName="zootaxa.5169.1.1.pdf" docOrigin="Zootaxa 5169 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Menarik kecil Grall &amp; Jäger 2022, spec. nov." docType="treatment" docVersion="6" lastPageNumber="12" masterDocId="FFC4FFEA5832FFC5FFC1FFA480644008" masterDocTitle="Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="7" updateTime="1699362128362" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="04EAACAB7C1270C153A6BD271EE218C9" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="64A1057DF8B3D6E9C52C60173DC979F1">
<mods:title id="524FDDBAC0CF3A25512818C5564BA5A5">Four new genera of Heteropodinae Thorell, 1873 from Malaysia, Brunei and Papua New Guinea (Araneae: Sparassidae)</mods:title>
</mods:titleInfo>
<mods:name id="B32E0EDDADE3825ABA28B41B545A95E2" type="personal">
<mods:role id="2E3DD635DB52CD4E0237800507CD581F">
<mods:roleTerm id="A52E6F15BE48075E0FA6BB1CC76BA9FC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6AAF03AB9B146E42BC8B1FCA04E533F3">Grall, Elena</mods:namePart>
<mods:nameIdentifier id="6D5DC79095F197A77A6035BE9EF1AC64" type="ORCID">0000-0002-8938-4105</mods:nameIdentifier>
<mods:affiliation id="74CC98738DA36069BD53F19472EC6C75">Arachnology, Senckenberg Research Institute, Mertonstrasse 17 - 21, 60325 Frankfurt. &amp; Institute of Ecology, Diversity and Evolution, Campus Riedberg, Goethe University, 60438 Frankfurt am Main, Germany; elena. grall @ senckenberg. de, https: // orcid. org / 0000 - 0002 - 8938 - 4105</mods:affiliation>
</mods:name>
<mods:name id="FC86F26FA6751CB4753C6FF7ABBEE7A7" type="personal">
<mods:role id="2EDC5EF7EC5F7C5DF45346B08FF10607">
<mods:roleTerm id="FFEFC65ABB76DA36C601FB631D2E9870">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6F5D3552ADF4182C2276ACEE938DFD41">Jäger, Peter</mods:namePart>
<mods:nameIdentifier id="855EB928A75BBD72DDF0E1A62F6A9368" type="ORCID">0000-0003-1563-0147</mods:nameIdentifier>
<mods:affiliation id="B647EE4634A1C711ECB4B9F0B4BFC8CB">Arachnology, Senckenberg Research Institute, Mertonstrasse 17 - 21, 60325 Frankfurt. &amp; peter. jaeger @ senckenberg. de, https: // orcid. org / 0000 - 0003 - 1563 - 0147</mods:affiliation>
</mods:name>
<mods:typeOfResource id="F5CCBE60E1587E90D7497BE81B1E684A">text</mods:typeOfResource>
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<mods:titleInfo id="57B3FE87EDB4F8DE10AA5C64AFBEB72E">
<mods:title id="28D94D2B9B01A759C5FFFFE7D72B87EA">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0A074442A95A5F11A755F707414BAD0C">
<mods:date id="0CC956F54018DAD7EF181130F254B5C2">2022</mods:date>
<mods:detail id="68CD3C57D1A530675160F628D29EDC67" type="pubDate">
<mods:number id="DD28E23BEAEE593C4CFFAE937DCA7B18">2022-07-27</mods:number>
</mods:detail>
<mods:detail id="0C5CE570617613D458C06851A531E739" type="volume">
<mods:number id="94A5BFA9524D7638048912C30AF57175">5169</mods:number>
</mods:detail>
<mods:detail id="943FCCC39C8FFE0DB83A341C9778AF0A" type="issue">
<mods:number id="BFC22227823085785462864BCF214061">1</mods:number>
</mods:detail>
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<mods:start id="8909F1140F680A87F2ACE4E9A86FE62C">1</mods:start>
<mods:end id="4DEC812E9FE497CFC1ACF519C936EDA3">25</mods:end>
</mods:extent>
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<mods:classification id="836B67DAAFFBE8276714717D8FD4036B">journal article</mods:classification>
<mods:identifier id="3BB38A9FFDD2C9BA88551BB86673510D" type="CLB-Dataset">106789</mods:identifier>
<mods:identifier id="03BEFFD4F14A541E4241C58FC1193E29" type="DOI">10.11646/zootaxa.5169.1.1</mods:identifier>
<mods:identifier id="7D07AE0D1CF73011979F211E512E265C" type="GBIF-Dataset">b76236d0-0336-4d17-804a-80f0344b097e</mods:identifier>
<mods:identifier id="1A61E18C04FF946F53A7E96421DA11C8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="361A3DC300EADD0278DAD0F47178B58A" type="Zenodo-Dep">6911151</mods:identifier>
<mods:identifier id="7E055A46DDB8CB163EC2E1ADB093A09E" type="ZooBank">3CB49021-90C5-46F0-AAAF-619EE8068F42</mods:identifier>
</mods:mods>
<treatment id="03FD87925834FFCEFF56FE7E84E34181" ID-DOI="http://doi.org/10.5281/zenodo.6911171" ID-GBIF-Taxon="197859758" ID-Zenodo-Dep="6911171" LSID="urn:lsid:plazi:treatment:03FD87925834FFCEFF56FE7E84E34181" httpUri="http://treatment.plazi.org/id/03FD87925834FFCEFF56FE7E84E34181" lastPageId="11" lastPageNumber="12" pageId="6" pageNumber="7">
<subSubSection id="C34E650F5834FFC3FF56FE7E81DB41FD" box="[151,447,474,501]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="8BEB36845834FFC3FF56FE7E81DB41FD" blockId="6.[151,447,474,537]" box="[151,447,474,501]" pageId="6" pageNumber="7">
<heading id="D0A381E85834FFC3FF56FE7E81DB41FD" bold="true" box="[151,447,474,501]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="B920EA965834FFC3FF56FE7E81DB41FD" bold="true" box="[151,447,474,501]" pageId="6" pageNumber="7">
<taxonomicName id="4C544D075834FFC3FF56FE7E812441FD" authority="Grall &amp; Jäger, 2022" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[151,320,474,501]" class="Arachnida" family="Sparassidae" genus="Menarik" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kecil" status="sp. nov.">
<emphasis id="B920EA965834FFC3FF56FE7E812441FD" bold="true" box="[151,320,474,501]" italics="true" pageId="6" pageNumber="7">Menarik kecil</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357ED5834FFC3FE86FE7F81DB41FD" box="[327,447,475,501]" pageId="6" pageNumber="7" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E650F5834FFC3FF56FE5B81F64211" box="[151,402,511,537]" pageId="6" pageNumber="7" type="description">
<paragraph id="8BEB36845834FFC3FF56FE5B81F64211" blockId="6.[151,447,474,537]" box="[151,402,511,537]" pageId="6" pageNumber="7">
<figureCitation id="136F2A015834FFC3FF56FE5B81734211" box="[151,279,511,537]" captionStart-0="FIGURES 17–22" captionStart-1="FIGURES 23–27" captionStart-2="FIGURES 28–39" captionStart-3="FIGURES 40–46" captionStartId-0="7.[151,264,1857,1881]" captionStartId-1="8.[151,264,1883,1907]" captionStartId-2="9.[151,264,1944,1968]" captionStartId-3="10.[151,264,1585,1609]" captionTargetBox-0="[151,1436,453,1834]" captionTargetBox-1="[222,1366,190,1859]" captionTargetBox-2="[152,1436,345,1921]" captionTargetBox-3="[151,1413,356,1562]" captionTargetId-0="figure-168@7.[151,1436,453,1834]" captionTargetId-1="figure-21@8.[222,1366,181,1859]" captionTargetId-2="figure-91@9.[151,1436,345,1921]" captionTargetId-3="figure-90@10.[151,1436,345,1562]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="FIGURES 17–22. Menarik kecil gen. &amp; spec. nov., males from Sarawak (17–20 holotype; 21–22 male, SD 1370). 17–19 Left palp (17 prolateral, 18 ventral, 19 retrolateral); 20–22 Palpal tibia (20, 22 dorsal, 21 retrolateral). aRTA—apical part of RTA, bRTA—basal part of RTA, dB—dorsal branch of RTA, ddA—disto-dorsal apophysis, E—embolus, S—spermophor, ST—subtegulum, TA—tegular apophysis." captionText-1="FIGURES 23–27. Menarik kecil gen. &amp; spec. nov. female paratypes (23–26) and male paratype (27) from Sarawak. 23–24 Epigyne, ventral; 25 Vulva, dorsal; 26 Schematic course of internal duct system, ventral; 27 Trilobate membrane, distal metatarsus of right leg II, dorsal.At—atrium, EF—epigynal field, FB—“fusion bubbles”, FD—fertilization duct, FW—first winding of internal duct system, GA—glandular appendages, MA—muscle attachment point, Sc—scar, SP—spermatheca." captionText-2="FIGURES 28–39. Menarik kecil gen. &amp; spec. nov. female paratypes (28–37) and egg-sac (38–39) from Sarawak. 28–36 Epigyne, ventral (28–35 not dissected, 36 dissected); 37 Vulva, dorsal; 38–39 two views of the same egg-sac. Arrows indicating scars at the anterior epigyne. Note the different shapes of the epigynal atrium." captionText-3="FIGURES 40–46. Menarik kecil gen. &amp; spec. nov. holotype male (40, 42, 45) and female paratype (41, 43–44, 46) from Sarawak. 40–41 Eyes, dorsal; 42–44 Claws (42, 44 right leg I, retrolateral claw, retrolateral; 43 left palp, prolateral); 45–46 Left chelicera, ventral." figureDoi-0="http://doi.org/10.5281/zenodo.6911167" figureDoi-1="http://doi.org/10.5281/zenodo.6911169" figureDoi-2="http://doi.org/10.5281/zenodo.6911173" figureDoi-3="http://doi.org/10.5281/zenodo.6911177" httpUri-0="https://zenodo.org/record/6911167/files/figure.png" httpUri-1="https://zenodo.org/record/6911169/files/figure.png" httpUri-2="https://zenodo.org/record/6911173/files/figure.png" httpUri-3="https://zenodo.org/record/6911177/files/figure.png" pageId="6" pageNumber="7">Figs 17–46</figureCitation>
,
<figureCitation id="136F2A015834FFC3FEE3FE5B810F4211" box="[290,363,511,537]" captionStart="FIGURES 82–97" captionStartId="19.[151,264,1899,1923]" captionTargetBox="[152,1435,182,1873]" captionTargetId="figure-19@19.[151,1436,181,1875]" captionTargetPageId="19" captionText="FIGURES 82–97. Heteropodinae gen. &amp; spp. nov., habitus (dorsal and ventral, respectively). 82–85 Borniella parva gen. &amp; spec. nov.; 86–89 Menarik kecil gen. &amp; spec. nov.; 90–93 Micropoda davisae gen. &amp; spec. nov.; 94–97 Tiomaniella ladam gen. &amp; spec. nov. (82–83, 86–87, 90–91, 94–95 holotype male, 84–85, 88–89, 92–93, 96–97 female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6911203" httpUri="https://zenodo.org/record/6911203/files/figure.png" pageId="6" pageNumber="7">86–89</figureCitation>
,
<figureCitation id="136F2A015834FFC3FEB7FE5B81F64211" box="[374,402,511,537]" captionStart="FIGURE 98" captionStartId="20.[151,250,1436,1460]" captionTargetBox="[151,1436,956,1413]" captionTargetId="figure-372@20.[151,1436,956,1413]" captionTargetPageId="20" captionText="FIGURE 98. Distribution of the new genera and species in Malaysia, Brunei and Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.6911205" httpUri="https://zenodo.org/record/6911205/files/figure.png" pageId="6" pageNumber="7">98</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34E650F5834FFC3FF56FDE3827D43E5" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="8BEB36845834FFC3FF56FDE382C8428D" blockId="6.[151,1437,582,2013]" pageId="6" pageNumber="7">
<emphasis id="B920EA965834FFC3FF56FDE381234268" bold="true" box="[151,327,582,609]" pageId="6" pageNumber="7">Type material.</emphasis>
<materialsCitation id="3B3C3CD95834FFC3FE90FDE282C8428D" ID-GBIF-Occurrence="3866517319" collectingDate="1984-01-04" collectingDateMax="1984-01-10" collectingDateMin="1984-01-04" collectionCode="RMNH" collectorName="C. L. &amp; P. R. Deeleman" country="Malaysia" location="Sarawak" pageId="6" pageNumber="7" specimenCode="SD 1366" specimenCount="1" stateProvince="Sarawak" typeStatus="holotype">
<emphasis id="B920EA965834FFC3FE90FDE282BA4269" bold="true" box="[337,734,582,609]" pageId="6" pageNumber="7">
<collectingCountry id="F34376145834FFC3FE90FDE281834268" box="[337,487,582,608]" name="Malaysia" pageId="6" pageNumber="7">MALAYSIA</collectingCountry>
:
<emphasis id="B920EA965834FFC3FE38FDE2820C4269" bold="true" box="[505,616,582,609]" italics="true" pageId="6" pageNumber="7">Sarawak:</emphasis>
<typeStatus id="54EF88265834FFC3FDB0FDE382BA4269" box="[625,734,583,609]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
</emphasis>
male (
<specimenCode id="DBF29EFF5834FFC3FCEEFDE383FF4269" box="[815,923,583,609]" collectionCode="SD" pageId="6" pageNumber="7">SD 1366</specimenCode>
): West
<collectingRegion id="4990F8665834FFC3FC31FDE384374269" box="[1008,1107,583,609]" country="Malaysia" name="Sarawak" pageId="6" pageNumber="7">Sarawak</collectingRegion>
, Semengoh arboretum,
<date id="FFEA10445834FFC3FAA5FDE38154428D" pageId="6" pageNumber="7" value="1984-01-04" valueMax="1984-01-10" valueMin="1984-01-04">
<collectingDate id="EFAEE9AC5834FFC3FAA5FDE38154428D" pageId="6" pageNumber="7" value="1984-01-04" valueMax="1984-01-10" valueMin="1984-01-04">4–10 January 1984</collectingDate>
</date>
,
<collectorName id="26A153525834FFC3FEFAFDCF8253428D" box="[315,567,619,645]" pageId="6" pageNumber="7">C.L. &amp; P.R. Deeleman</collectorName>
(
<collectionCode id="ED45AE415834FFC3FD84FDCF82C4428D" box="[581,672,619,645]" country="Netherlands" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="7" type="Museum">RMNH</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BEB36845834FFC3FF06FD2B840F42F9" blockId="6.[151,1437,582,2013]" pageId="6" pageNumber="7">
<emphasis id="B920EA965834FFC3FF06FD2B81B542A0" bold="true" box="[199,465,654,681]" pageId="6" pageNumber="7">
<typeStatus id="54EF88265834FFC3FF06FD2B815B42A1" box="[199,319,655,681]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D52FD0D5834FFC3FE8EFD2A81A042A0" box="[335,452,654,680]" pageId="6" pageNumber="7" type="female">9 females</specimenCount>
):
</emphasis>
<materialsCitation id="3B3C3CD95834FFC3FE19FD2B847042A1" ID-GBIF-Occurrence="3866517318" box="[472,1044,655,681]" collectingDate="1984-01-04" collectingDateMax="1984-01-10" collectingDateMin="1984-01-04" collectionCode="RMNH" collectorName="C. L. &amp; P. R. Deeleman" country="Malaysia" location="Sarawak" pageId="6" pageNumber="7" specimenCount="2" specimenCount-female="2" stateProvince="Sarawak" typeStatus="paratype">
<specimenCount id="9D52FD0D5834FFC3FE19FD2B822042A1" box="[472,580,655,681]" pageId="6" pageNumber="7" type="female">2 females</specimenCount>
with same data as for holotype (
<collectionCode id="ED45AE415834FFC3FC70FD2B846842A1" box="[945,1036,655,681]" country="Netherlands" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="7" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3C3CD95834FFC3FBDEFD2B824942C5" ID-GBIF-Occurrence="3866517330" collectingDate="1984-01-04" collectingDateMax="1984-01-10" collectingDateMin="1984-01-04" collectionCode="RMNH" collectorName="C. L. &amp; P. R. Deeleman" country="Malaysia" location="Sarawak" pageId="6" pageNumber="7" specimenCode="SD 1365" specimenCount="4" specimenCount-female="4" stateProvince="Sarawak" typeStatus="paratype">
<specimenCount id="9D52FD0D5834FFC3FBDEFD2B84EF42A1" box="[1055,1163,655,681]" pageId="6" pageNumber="7" type="female">4 females</specimenCount>
(
<specimenCode id="DBF29EFF5834FFC3FB5BFD2B849A42A1" box="[1178,1278,654,681]" collectionCode="SD" pageId="6" pageNumber="7">SD 1365</specimenCode>
&amp; 1561) with same data as for holotype (
<collectionCode id="ED45AE415834FFC3FE0BFD17824142C5" box="[458,549,691,717]" country="Netherlands" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="7" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3C3CD95834FFC3FDF8FD1785E642C5" ID-GBIF-Occurrence="3866517333" box="[569,1410,690,717]" collectingDate="1985-03-23" collectingDateMax="1984-01-10" collectingDateMin="1984-01-04" collectionCode="RMNH" collectorName="C. L. &amp; P. R. Deeleman" country="Malaysia" location="Sarawak" pageId="6" pageNumber="7" specimenCount="1" specimenCount-female="1" stateProvince="Sarawak" typeStatus="paratype">
<specimenCount id="9D52FD0D5834FFC3FDF8FD1782FF42C5" box="[569,667,691,717]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
with same data as for holotype except:
<date id="FFEA10445834FFC3FBA3FD17856B42C5" box="[1122,1295,690,717]" pageId="6" pageNumber="7" value="1985-03-23">
<collectingDate id="EFAEE9AC5834FFC3FBA3FD17856B42C5" box="[1122,1295,690,717]" pageId="6" pageNumber="7" value="1985-03-23">23 March 1985</collectingDate>
</date>
(
<collectionCode id="ED45AE415834FFC3FADEFD17851E42C5" box="[1311,1402,691,717]" country="Netherlands" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="7" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3C3CD95834FFC3FA4FFD17810F42F9" ID-GBIF-Occurrence="3866517321" collectionCode="SD" country="Malaysia" pageId="6" pageNumber="7" specimenCode="SD 1368" specimenCount="2" specimenCount-female="2" stateProvince="Sarawak" typeStatus="paratype">
<specimenCount id="9D52FD0D5834FFC3FA4FFD17808A42F9" pageId="6" pageNumber="7" type="female">2 females</specimenCount>
(
<specimenCode id="DBF29EFF5834FFC3FF3CFD73810642F9" box="[253,354,727,753]" collectionCode="SD" pageId="6" pageNumber="7">SD 1368</specimenCode>
)
</materialsCitation>
<materialsCitation id="3B3C3CD95834FFC3FEB3FD73840342F9" ID-GBIF-Occurrence="3866517341" box="[370,1127,726,753]" collectingDate="1985-03-27" collectingDateMax="1984-01-10" collectingDateMin="1984-01-04" collectionCode="SMF" collectorName="C. L. &amp; P. R. Deeleman" country="Malaysia" location="Sarawak" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Sarawak" typeStatus="paratype">
with same data as for holotype except: 24 &amp;
<date id="FFEA10445834FFC3FCA6FD73847642F9" box="[871,1042,726,753]" pageId="6" pageNumber="7" value="1985-03-27">
<collectingDate id="EFAEE9AC5834FFC3FCA6FD73847642F9" box="[871,1042,726,753]" pageId="6" pageNumber="7" value="1985-03-27">27 March 1985</collectingDate>
</date>
(
<collectionCode id="ED45AE415834FFC3FBE3FD73840442F9" box="[1058,1120,727,753]" country="Germany" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="7">SMF</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEB36845834FFC3FF06FD5E827D43E5" blockId="6.[151,1437,582,2013]" pageId="6" pageNumber="7">
<emphasis id="B920EA965834FFC3FF06FD5E81CA4331" bold="true" pageId="6" pageNumber="7">
<materialsCitation id="3B3C3CD95834FFC3FF06FD5E81C14330" ID-GBIF-Occurrence="3866517301" country="Malaysia" location="MALAYSIA" pageId="6" pageNumber="7" specimenCount="22" specimenCount-female="3" specimenCount-juvenile="17" specimenCount-male="1" specimenCount-subadult="1" stateProvince="Province">
Additional material examined (
<specimenCount id="9D52FD0D5834FFC3FD8DFD5E82C5431D" box="[588,673,762,789]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
,
<specimenCount id="9D52FD0D5834FFC3FD73FD5E834C431C" box="[690,808,762,788]" pageId="6" pageNumber="7" type="female">3 females</specimenCount>
,
<specimenCount id="9D52FD0D5834FFC3FCF8FD5E83DF431D" box="[825,955,762,789]" pageId="6" pageNumber="7" type="subadult">1 subadult</specimenCount>
male,
<specimenCount id="9D52FD0D5834FFC3FBD2FD5E84C3431C" box="[1043,1191,762,788]" pageId="6" pageNumber="7" type="juvenile">17 juveniles</specimenCount>
, 2 egg-sacs with 15 nymphs).
<collectingCountry id="F34376145834FFC3FECEFCBA81C14330" box="[271,421,798,824]" name="Malaysia" pageId="6" pageNumber="7">MALAYSIA</collectingCountry>
</materialsCitation>
:
</emphasis>
<materialsCitation id="3B3C3CD95834FFC3FE76FCBA84924331" ID-GBIF-Occurrence="3866517310" box="[439,1270,798,825]" collectingDate="1984-01-04" collectingDateMax="1984-01-10" collectingDateMin="1984-01-04" collectionCode="RMNH" collectorName="C. L. &amp; P. R. Deeleman" country="Malaysia" location="Sarawak" pageId="6" pageNumber="7" specimenCount="7" specimenCount-juvenile="7" stateProvince="Province">
<emphasis id="B920EA965834FFC3FE76FCBA82F14331" bold="true" box="[439,661,798,825]" italics="true" pageId="6" pageNumber="7">
<collectingCountry id="F34376145834FFC3FE76FCBA82784331" box="[439,540,798,825]" name="Malaysia" pageId="6" pageNumber="7">Sarawak</collectingCountry>
<collectingRegion id="4990F8665834FFC3FDE4FCBB82EA4331" box="[549,654,799,825]" pageId="6" pageNumber="7">Province</collectingRegion>
:
</emphasis>
<specimenCount id="9D52FD0D5834FFC3FD5FFCBB837E4331" box="[670,794,799,825]" pageId="6" pageNumber="7" type="juvenile">7 juveniles</specimenCount>
with same data as for holotype (
<collectionCode id="ED45AE415834FFC3FB52FCBB848A4331" box="[1171,1262,799,825]" country="Netherlands" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="7" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3C3CD95834FFC3FAC2FCBB83174355" ID-GBIF-Occurrence="3866517337" collectingDate="1984-01-04" collectingDateMax="1984-01-10" collectingDateMin="1984-01-04" collectionCode="RMNH" collectorName="C. L. &amp; P. R. Deeleman" country="Malaysia" location="Sarawak" pageId="6" pageNumber="7" specimenCount="4" specimenCount-juvenile="4" stateProvince="Province">
<specimenCount id="9D52FD0D5834FFC3FAC2FCBB85E54331" box="[1283,1409,799,825]" pageId="6" pageNumber="7" type="juvenile">4 juveniles</specimenCount>
, 1 egg-sac with 11 nymphs with same data as for holotype (
<collectionCode id="ED45AE415834FFC3FCD0FCE783084355" box="[785,876,835,861]" country="Netherlands" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="7" type="Museum">RMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B3C3CD95834FFC3FCBFFCE784234355" ID-GBIF-Occurrence="3866517328" box="[894,1095,835,861]" collectionCode="SD" country="Malaysia" pageId="6" pageNumber="7" specimenCode="SD 1370" specimenCount="1" specimenCount-male="1" stateProvince="Province">
<specimenCount id="9D52FD0D5834FFC3FCBFFCE783AD4355" box="[894,969,835,861]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
(
<specimenCode id="DBF29EFF5834FFC3FC19FCE784244355" box="[984,1088,835,861]" collectionCode="SD" pageId="6" pageNumber="7">SD 1370</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B3C3CD95834FFC3FB92FCE785FC4355" ID-GBIF-Occurrence="3866517334" box="[1107,1432,834,861]" collectionCode="SD" country="Malaysia" pageId="6" pageNumber="7" specimenCode="SD 1372" specimenCount="3" specimenCount-female="3" stateProvince="Province">
<specimenCount id="9D52FD0D5834FFC3FB92FCE784DA4355" box="[1107,1214,835,861]" pageId="6" pageNumber="7" type="female">3 females</specimenCount>
(
<specimenCode id="DBF29EFF5834FFC3FB0CFCE785554355" box="[1229,1329,835,861]" collectionCode="SD" pageId="6" pageNumber="7">SD 1372</specimenCode>
&amp; 1560)
</materialsCitation>
,
<materialsCitation id="3B3C3CD95834FFC3FF56FCC381A74389" ID-GBIF-Occurrence="3866517343" box="[151,451,871,897]" collectionCode="SD" country="Malaysia" pageId="6" pageNumber="7" specimenCode="SD 1371" specimenCount="1" specimenCount-subadult="1" stateProvince="Province">
<specimenCount id="9D52FD0D5834FFC3FF56FCC3816E4389" box="[151,266,871,897]" pageId="6" pageNumber="7" type="subadult">1 subadult</specimenCount>
male (
<specimenCode id="DBF29EFF5834FFC3FE94FCC381DF4389" box="[341,443,871,897]" collectionCode="SD" pageId="6" pageNumber="7">SD 1371</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B3C3CD95834FFC3FE0FFCC382A04389" ID-GBIF-Occurrence="3866517302" box="[462,708,871,897]" collectionCode="SD" country="Malaysia" pageId="6" pageNumber="7" specimenCode="SD 1373" specimenCount="3" specimenCount-juvenile="3" stateProvince="Province">
<specimenCount id="9D52FD0D5834FFC3FE0FFCC3822C4389" box="[462,584,871,897]" pageId="6" pageNumber="7" type="juvenile">3 juveniles</specimenCount>
(
<specimenCode id="DBF29EFF5834FFC3FD97FCC382D84389" box="[598,700,871,897]" collectionCode="SD" pageId="6" pageNumber="7">SD 1373</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B3C3CD95834FFC3FD0EFCC3836A43C1" ID-GBIF-Occurrence="3866517332" collectingDate="1985-03-30" collectionCode="RMNH" collectorName="C. L. &amp; P. R. Deeleman" country="Malaysia" county="Borneo" location="Bako National Park" municipality="West" pageId="6" pageNumber="7" specimenCount="4" specimenCount-female="1" specimenCount-juvenile="2" specimenCount-male="1" stateProvince="Sarawak">
1 egg-sac with 4 nymphs,
<collectingCounty id="628A4E085834FFC3FC30FCC384224389" box="[1009,1094,871,897]" pageId="6" pageNumber="7">Borneo</collectingCounty>
,
<collectingMunicipality id="6B8FACFE5834FFC3FB91FCC384E34389" box="[1104,1159,871,897]" pageId="6" pageNumber="7">West</collectingMunicipality>
<collectingRegion id="4990F8665834FFC3FB4CFCC384944389" box="[1165,1264,871,897]" country="Malaysia" name="Sarawak" pageId="6" pageNumber="7">Sarawak</collectingRegion>
,
<location id="8E8B605F5834FFC3FB3AFCC380A943AD" LSID="urn:lsid:plazi:treatment:03FD87925834FFCEFF56FE7E84E34181:8E8B605F5834FFC3FB3AFCC380A943AD" country="Malaysia" county="Borneo" municipality="West" name="Bako National Park" pageId="6" pageNumber="7" stateProvince="Sarawak">Bako National Park</location>
, moeras [= swampy] bog, 29 &amp;
<date id="FFEA10445834FFC3FDF0FC2F82B943AD" box="[561,733,906,933]" pageId="6" pageNumber="7" value="1985-03-30">
<collectingDate id="EFAEE9AC5834FFC3FDF0FC2F82B943AD" box="[561,733,906,933]" pageId="6" pageNumber="7" value="1985-03-30">30 March 1985</collectingDate>
</date>
,
<collectorName id="26A153525834FFC3FD26FC2F838543AD" box="[743,993,907,933]" pageId="6" pageNumber="7">C.L. &amp; P.R. Deeleman</collectorName>
[+
<specimenCount id="9D52FD0D5834FFC3FBC7FC2F841043AD" box="[1030,1140,907,933]" pageId="6" pageNumber="7" type="juvenile">1 juvenile</specimenCount>
<taxonomicName id="4C544D075834FFC3FBBAFC2F856843AD" box="[1147,1292,907,933]" class="Arachnida" family="Sparassidae" genus="Sinopoda" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B920EA965834FFC3FBBAFC2F848143AD" box="[1147,1253,907,933]" italics="true" pageId="6" pageNumber="7">Sinopoda</emphasis>
sp.
</taxonomicName>
, +
<specimenCount id="9D52FD0D5834FFC3FAEFFC2F85F843AD" box="[1326,1436,907,933]" pageId="6" pageNumber="7" type="juvenile">1 juvenile</specimenCount>
<taxonomicName id="4C544D075834FFC3FF56FC0B815843C1" authorityName="Thorell" authorityYear="1873" box="[151,316,943,969]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subFamily" subFamily="Heteropodinae">Heteropodinae</taxonomicName>
indet.] (
<collectionCode id="ED45AE415834FFC3FE5AFC0B819043C1" box="[411,500,943,969]" country="Netherlands" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="6" pageNumber="7" type="Museum">RMNH</collectionCode>
;
<specimenCount id="9D52FD0D5834FFC3FE3EFC0B822A43C1" box="[511,590,943,969]" pageId="6" pageNumber="7" type="male">1 male</specimenCount>
,
<specimenCount id="9D52FD0D5834FFC3FD98FC0B82A443C1" box="[601,704,943,969]" pageId="6" pageNumber="7" type="female">1 female</specimenCount>
: SMF)
</materialsCitation>
;
<materialsCitation id="3B3C3CD95834FFC3FCDBFC0B847643C1" ID-GBIF-Occurrence="3866517331" box="[794,1042,943,969]" collectionCode="SD" country="Malaysia" pageId="6" pageNumber="7" specimenCode="SD 1369" specimenCount="3" specimenCount-juvenile="3" stateProvince="Province">
<specimenCount id="9D52FD0D5834FFC3FCDBFC0B83F043C1" box="[794,916,943,969]" pageId="6" pageNumber="7" type="juvenile">3 juveniles</specimenCount>
(
<specimenCode id="DBF29EFF5834FFC3FC65FC0B846D43C1" box="[932,1033,943,969]" collectionCode="SD" pageId="6" pageNumber="7">SD 1369</specimenCode>
)
</materialsCitation>
<materialsCitation id="3B3C3CD95834FFC3FBD8FC0B827043E5" ID-GBIF-Occurrence="3866517345" collectingDate="1985-03-27" collectingDateMax="1984-01-10" collectingDateMin="1984-01-04" collectionCode="SMF" collectorName="C. L. &amp; P. R. Deeleman" country="Malaysia" location="Sarawak" pageId="6" pageNumber="7" specimenCount="1" stateProvince="Province">
with same data as for holotype except: 24 &amp;
<date id="FFEA10445834FFC3FED5FC7781DB43E5" box="[276,447,978,1005]" pageId="6" pageNumber="7" value="1985-03-27">
<collectingDate id="EFAEE9AC5834FFC3FED5FC7781DB43E5" box="[276,447,978,1005]" pageId="6" pageNumber="7" value="1985-03-27">27 March 1985</collectingDate>
</date>
(
<collectionCode id="ED45AE415834FFC3FE0EFC77826943E5" box="[463,525,979,1005]" country="Germany" lsid="urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="6" pageNumber="7">SMF</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E650F5834FFC3FF06FC5382A9443D" pageId="6" pageNumber="7" type="etymology">
<paragraph id="8BEB36845834FFC3FF06FC5382A9443D" blockId="6.[151,1437,582,2013]" pageId="6" pageNumber="7">
<emphasis id="B920EA965834FFC3FF06FC53812A4419" bold="true" box="[199,334,1015,1041]" pageId="6" pageNumber="7">Etymology.</emphasis>
The species name is derived from the Malayan word “kecil”, meaning “little”, and refers to the small body size of this species; term in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C34E650F5834FFC3FF06FB9A80A04539" pageId="6" pageNumber="7" type="diagnosis">
<paragraph id="8BEB36845834FFC3FF06FB9A80A04539" blockId="6.[151,1437,582,2013]" pageId="6" pageNumber="7">
<emphasis id="B920EA965834FFC3FF06FB9A81264450" bold="true" box="[199,322,1086,1112]" pageId="6" pageNumber="7">Diagnosis.</emphasis>
Small
<taxonomicName id="4C544D075834FFC3FE4EFB9B82774451" authorityName="Bertkau" authorityYear="1872" box="[399,531,1087,1113]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="family">Sparassidae</taxonomicName>
with TL &lt;5.0 (
<figureCitation id="136F2A015834FFC3FD79FB9B835F4451" box="[696,827,1087,1113]" captionStart="FIGURES 82–97" captionStartId="19.[151,264,1899,1923]" captionTargetBox="[152,1435,182,1873]" captionTargetId="figure-19@19.[151,1436,181,1875]" captionTargetPageId="19" captionText="FIGURES 82–97. Heteropodinae gen. &amp; spp. nov., habitus (dorsal and ventral, respectively). 82–85 Borniella parva gen. &amp; spec. nov.; 86–89 Menarik kecil gen. &amp; spec. nov.; 90–93 Micropoda davisae gen. &amp; spec. nov.; 94–97 Tiomaniella ladam gen. &amp; spec. nov. (82–83, 86–87, 90–91, 94–95 holotype male, 84–85, 88–89, 92–93, 96–97 female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6911203" httpUri="https://zenodo.org/record/6911203/files/figure.png" pageId="6" pageNumber="7">Figs 86–89</figureCitation>
). Males of
<taxonomicName id="4C544D075834FFC3FC74FB9B842B4451" authority="Grall &amp; Jäger, 2022" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[949,1103,1087,1113]" class="Arachnida" family="Sparassidae" genus="Menarik" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="kecil" status="sp. nov.">
<emphasis id="B920EA965834FFC3FC74FB9B842B4451" box="[949,1103,1087,1113]" italics="true" pageId="6" pageNumber="7">Menarik kecil</emphasis>
</taxonomicName>
<emphasis id="B920EA965834FFC3FB94FB9B84A24451" bold="true" box="[1109,1222,1087,1113]" pageId="6" pageNumber="7">
<taxonomicNameLabel id="A21357ED5834FFC3FB94FB9B84A24451" box="[1109,1222,1087,1113]" pageId="6" pageNumber="7" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
may be recognised by the following combination of characters (
<figureCitation id="136F2A015834FFC3FD54FBC7837F4475" box="[661,795,1123,1149]" captionStart="FIGURES 17–22" captionStartId="7.[151,264,1857,1881]" captionTargetBox="[151,1436,453,1834]" captionTargetId="figure-168@7.[151,1436,453,1834]" captionTargetPageId="7" captionText="FIGURES 17–22. Menarik kecil gen. &amp; spec. nov., males from Sarawak (17–20 holotype; 21–22 male, SD 1370). 17–19 Left palp (17 prolateral, 18 ventral, 19 retrolateral); 20–22 Palpal tibia (20, 22 dorsal, 21 retrolateral). aRTA—apical part of RTA, bRTA—basal part of RTA, dB—dorsal branch of RTA, ddA—disto-dorsal apophysis, E—embolus, S—spermophor, ST—subtegulum, TA—tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6911167" httpUri="https://zenodo.org/record/6911167/files/figure.png" pageId="6" pageNumber="7">Figs 17–22</figureCitation>
): 1. Apical part of embolus forming almost a circle in a plane oblique to cymbial length axis, 2. Tegular apophysis arising from a 7- to 8-o’clock-position, curved, its tip distad and 3. RTA with basal part bulgy and with various lobes, apical branch acutely tapering. Females are unique in the subfamily
<taxonomicName id="4C544D075834FFC3FE90FB6B819244E1" authorityName="Thorell" authorityYear="1873" box="[337,502,1231,1257]" class="Arachnida" family="Sparassidae" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subFamily" subFamily="Heteropodinae">Heteropodinae</taxonomicName>
having an oval to slit-like copulatory atrium at anterior half of epigyne (
<figureCitation id="136F2A015834FFC3FAD6FB6B85FC44E1" box="[1303,1432,1231,1257]" captionStart="FIGURES 23–27" captionStartId="8.[151,264,1883,1907]" captionTargetBox="[222,1366,190,1859]" captionTargetId="figure-21@8.[222,1366,181,1859]" captionTargetPageId="8" captionText="FIGURES 23–27. Menarik kecil gen. &amp; spec. nov. female paratypes (23–26) and male paratype (27) from Sarawak. 23–24 Epigyne, ventral; 25 Vulva, dorsal; 26 Schematic course of internal duct system, ventral; 27 Trilobate membrane, distal metatarsus of right leg II, dorsal.At—atrium, EF—epigynal field, FB—“fusion bubbles”, FD—fertilization duct, FW—first winding of internal duct system, GA—glandular appendages, MA—muscle attachment point, Sc—scar, SP—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6911169" httpUri="https://zenodo.org/record/6911169/files/figure.png" pageId="6" pageNumber="7">Figs 23–24</figureCitation>
,
<figureCitation id="136F2A015834FFC3FF56FB5780854505" box="[151,225,1267,1293]" captionStart="FIGURES 28–39" captionStartId="9.[151,264,1944,1968]" captionTargetBox="[152,1436,345,1921]" captionTargetId="figure-91@9.[151,1436,345,1921]" captionTargetPageId="9" captionText="FIGURES 28–39. Menarik kecil gen. &amp; spec. nov. female paratypes (28–37) and egg-sac (38–39) from Sarawak. 28–36 Epigyne, ventral (28–35 not dissected, 36 dissected); 37 Vulva, dorsal; 38–39 two views of the same egg-sac. Arrows indicating scars at the anterior epigyne. Note the different shapes of the epigynal atrium." figureDoi="http://doi.org/10.5281/zenodo.6911173" httpUri="https://zenodo.org/record/6911173/files/figure.png" pageId="6" pageNumber="7">28–36</figureCitation>
). Moreover, posterior part with large spherical spermathecae and long antero-laterad fertilisation ducts (
<figureCitation id="136F2A015834FFC3FAAEFB5780D34539" captionStart="FIGURES 23–27" captionStartId="8.[151,264,1883,1907]" captionTargetBox="[222,1366,190,1859]" captionTargetId="figure-21@8.[222,1366,181,1859]" captionTargetPageId="8" captionText="FIGURES 23–27. Menarik kecil gen. &amp; spec. nov. female paratypes (23–26) and male paratype (27) from Sarawak. 23–24 Epigyne, ventral; 25 Vulva, dorsal; 26 Schematic course of internal duct system, ventral; 27 Trilobate membrane, distal metatarsus of right leg II, dorsal.At—atrium, EF—epigynal field, FB—“fusion bubbles”, FD—fertilization duct, FW—first winding of internal duct system, GA—glandular appendages, MA—muscle attachment point, Sc—scar, SP—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6911169" httpUri="https://zenodo.org/record/6911169/files/figure.png" pageId="6" pageNumber="7">Fig. 25</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34E650F5834FFCEFF06FA9E821D416D" lastPageId="11" lastPageNumber="12" pageId="6" pageNumber="7" type="description">
<paragraph id="8BEB36845834FFC3FF06FA9E829A467D" blockId="6.[151,1437,582,2013]" pageId="6" pageNumber="7">
<emphasis id="B920EA965834FFC3FF06FA9E8241455C" bold="true" box="[199,549,1338,1365]" pageId="6" pageNumber="7">
Description. Male (
<typeStatus id="54EF88265834FFC3FE6AFA9E827D455C" box="[427,537,1338,1364]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
):
</emphasis>
TL 3.4; PL 1.8, PW 1.8, AW 1.1; OL 1.6, OL 1.1. Eye measurements (
<figureCitation id="136F2A015834FFC3FAFAFA9F85F4455D" box="[1339,1424,1339,1365]" captionStart="FIGURES 40–46" captionStartId="10.[151,264,1585,1609]" captionTargetBox="[151,1413,356,1562]" captionTargetId="figure-90@10.[151,1436,345,1562]" captionTargetPageId="10" captionText="FIGURES 40–46. Menarik kecil gen. &amp; spec. nov. holotype male (40, 42, 45) and female paratype (41, 43–44, 46) from Sarawak. 40–41 Eyes, dorsal; 42–44 Claws (42, 44 right leg I, retrolateral claw, retrolateral; 43 left palp, prolateral); 45–46 Left chelicera, ventral." figureDoi="http://doi.org/10.5281/zenodo.6911177" httpUri="https://zenodo.org/record/6911177/files/figure.png" pageId="6" pageNumber="7">Fig. 40</figureCitation>
): AME 0.10; ALE 0.18; PME 0.10; PLE 0.20; AME–AME 0.10; AME–ALE 0.02; PME–PME 0.15; PME–PLE 0.21; AME–PME 0.18; ALE–PLE 0.19; clypeus AME 0.14; clypeus ALE 0.12. Leg formula: 2431. Measurements of palp and legs: palp: 2.80 (0.80, 0.45, 0.55, 1.00); I: 6.60 (1.80, 0.80, 1.80, 1.60, 0.60); II: 8.60 (2.40, 0.90, 2.30, 2.20, 0.80); III: 7.20 (2.10, 0.80, 1.90, 1.80, 0.60); IV: 7.40 (2.10, 0.80, 1.85, 1.95, 0.70). Spination: palp: 131, 101, 2001; legs: Fe I 323, Fe II &amp; III 222, Fe IV 221; Pa I–IV 000; Ti I &amp;
<date id="FFEA10445834FFC3FC82FA4B83F34601" box="[835,919,1519,1545]" pageId="6" pageNumber="7" value="1026-02">II 1026</date>
, Ti III &amp;
<date id="FFEA10445834FFC3FC3DFA4B843F4601" box="[1020,1115,1519,1545]" pageId="6" pageNumber="7" value="2226-04">IV 2226</date>
; Mt I &amp; II 0006, Mt
<date id="FFEA10445834FFC3FAFAFA4B85FC4601" box="[1339,1432,1519,1545]" pageId="6" pageNumber="7" value="2036-03">III 2036</date>
, Mt IV 3036. Chelicerae with 3 promarginal, 6 retromarginal teeth, 17 intermarginal denticles and 1 escort seta (
<figureCitation id="136F2A015834FFC3FAAEF9B780D34659" captionStart="FIGURES 40–46" captionStartId="10.[151,264,1585,1609]" captionTargetBox="[151,1413,356,1562]" captionTargetId="figure-90@10.[151,1436,345,1562]" captionTargetPageId="10" captionText="FIGURES 40–46. Menarik kecil gen. &amp; spec. nov. holotype male (40, 42, 45) and female paratype (41, 43–44, 46) from Sarawak. 40–41 Eyes, dorsal; 42–44 Claws (42, 44 right leg I, retrolateral claw, retrolateral; 43 left palp, prolateral); 45–46 Left chelicera, ventral." figureDoi="http://doi.org/10.5281/zenodo.6911177" httpUri="https://zenodo.org/record/6911177/files/figure.png" pageId="6" pageNumber="7">Fig. 45</figureCitation>
). Retrolateral claw of right leg I with 16 teeth (
<figureCitation id="136F2A015834FFC3FD0BF993837A4659" box="[714,798,1591,1617]" captionStart="FIGURES 40–46" captionStartId="10.[151,264,1585,1609]" captionTargetBox="[151,1413,356,1562]" captionTargetId="figure-90@10.[151,1436,345,1562]" captionTargetPageId="10" captionText="FIGURES 40–46. Menarik kecil gen. &amp; spec. nov. holotype male (40, 42, 45) and female paratype (41, 43–44, 46) from Sarawak. 40–41 Eyes, dorsal; 42–44 Claws (42, 44 right leg I, retrolateral claw, retrolateral; 43 left palp, prolateral); 45–46 Left chelicera, ventral." figureDoi="http://doi.org/10.5281/zenodo.6911177" httpUri="https://zenodo.org/record/6911177/files/figure.png" pageId="6" pageNumber="7">Fig. 42</figureCitation>
). Trilobate membrane of right leg II with narrow median hook, slightly shorter than lateral projections (
<figureCitation id="136F2A015834FFC3FD5CF9FF8295467D" box="[669,753,1627,1653]" captionStart="FIGURES 23–27" captionStartId="8.[151,264,1883,1907]" captionTargetBox="[222,1366,190,1859]" captionTargetId="figure-21@8.[222,1366,181,1859]" captionTargetPageId="8" captionText="FIGURES 23–27. Menarik kecil gen. &amp; spec. nov. female paratypes (23–26) and male paratype (27) from Sarawak. 23–24 Epigyne, ventral; 25 Vulva, dorsal; 26 Schematic course of internal duct system, ventral; 27 Trilobate membrane, distal metatarsus of right leg II, dorsal.At—atrium, EF—epigynal field, FB—“fusion bubbles”, FD—fertilization duct, FW—first winding of internal duct system, GA—glandular appendages, MA—muscle attachment point, Sc—scar, SP—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6911169" httpUri="https://zenodo.org/record/6911169/files/figure.png" pageId="6" pageNumber="7">Fig. 27</figureCitation>
).
</paragraph>
<paragraph id="8BEB36845834FFC3FF06F9DB843E470D" blockId="6.[151,1437,582,2013]" pageId="6" pageNumber="7">
Palp as in diagnosis (
<figureCitation id="136F2A015834FFC3FE74F9DB825C4691" box="[437,568,1663,1689]" captionStart="FIGURES 17–22" captionStartId="7.[151,264,1857,1881]" captionTargetBox="[151,1436,453,1834]" captionTargetId="figure-168@7.[151,1436,453,1834]" captionTargetPageId="7" captionText="FIGURES 17–22. Menarik kecil gen. &amp; spec. nov., males from Sarawak (17–20 holotype; 21–22 male, SD 1370). 17–19 Left palp (17 prolateral, 18 ventral, 19 retrolateral); 20–22 Palpal tibia (20, 22 dorsal, 21 retrolateral). aRTA—apical part of RTA, bRTA—basal part of RTA, dB—dorsal branch of RTA, ddA—disto-dorsal apophysis, E—embolus, S—spermophor, ST—subtegulum, TA—tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6911167" httpUri="https://zenodo.org/record/6911167/files/figure.png" pageId="6" pageNumber="7">Figs 17–22</figureCitation>
). Embolus basally wide, apical part filiform. Spermophor curved distinctly at distal part of tegulum, slightly in proximal half (best seen in ventral view). Tegular apophysis wide, curved and tapering. Distal part of tegulum bulging ventrally. Apical part of RTA strongly acute, beak-shaped in retrolateral view with dorsal branch; basal part broad and wide, roughly kidney-shaped in ventral view.
</paragraph>
<paragraph id="8BEB36845834FFC3FF06F8AB831A47B1" blockId="6.[151,1437,582,2013]" pageId="6" pageNumber="7">
Colouration in ethanol (
<figureCitation id="136F2A015834FFC3FE10F8AB82304721" box="[465,596,1807,1833]" captionStart="FIGURES 82–97" captionStartId="19.[151,264,1899,1923]" captionTargetBox="[152,1435,182,1873]" captionTargetId="figure-19@19.[151,1436,181,1875]" captionTargetPageId="19" captionText="FIGURES 82–97. Heteropodinae gen. &amp; spp. nov., habitus (dorsal and ventral, respectively). 82–85 Borniella parva gen. &amp; spec. nov.; 86–89 Menarik kecil gen. &amp; spec. nov.; 90–93 Micropoda davisae gen. &amp; spec. nov.; 94–97 Tiomaniella ladam gen. &amp; spec. nov. (82–83, 86–87, 90–91, 94–95 holotype male, 84–85, 88–89, 92–93, 96–97 female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6911203" httpUri="https://zenodo.org/record/6911203/files/figure.png" pageId="6" pageNumber="7">Figs 86–87</figureCitation>
): Prosoma yellowish-brown, median part with brown semi-circular pattern, posterior and lateral margin dark brown. Sternum yellowish-white. Opisthosoma brown with yellow spots, posteriorly dark brown, ventrally yellow, laterally with brown stripes. Chelicerae yellowish-brown. Coxa I–III medio-dorsally with brown stripe. Palps yellowish-brown. Fe and Pa distally darker brown. Legs yellowish-brown; Fe lateral and distal part darker brown; Pa and Ti laterally with brown spots.
</paragraph>
<paragraph id="8BEB36845834FFC2FF06F867810541A5" blockId="6.[151,1437,582,2013]" lastBlockId="7.[151,1436,150,429]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<emphasis id="B920EA965834FFC3FF06F86781CF47D4" bold="true" box="[199,427,1986,2013]" pageId="6" pageNumber="7">
Female (
<typeStatus id="54EF88265834FFC3FEEDF86681FA47D4" box="[300,414,1986,2012]" pageId="6" pageNumber="7" type="paratype">paratype</typeStatus>
):
</emphasis>
TL 4.4; PL 2.0, PW 1.9, AW 1.2; OL 2.4, OW 1.7. Eye measurements (
<figureCitation id="136F2A015834FFC3FB7AF867856A47D5" box="[1211,1294,1987,2013]" captionStart="FIGURES 40–46" captionStartId="10.[151,264,1585,1609]" captionTargetBox="[151,1413,356,1562]" captionTargetId="figure-90@10.[151,1436,345,1562]" captionTargetPageId="10" captionText="FIGURES 40–46. Menarik kecil gen. &amp; spec. nov. holotype male (40, 42, 45) and female paratype (41, 43–44, 46) from Sarawak. 40–41 Eyes, dorsal; 42–44 Claws (42, 44 right leg I, retrolateral claw, retrolateral; 43 left palp, prolateral); 45–46 Left chelicera, ventral." figureDoi="http://doi.org/10.5281/zenodo.6911177" httpUri="https://zenodo.org/record/6911177/files/figure.png" pageId="6" pageNumber="7">Fig. 41</figureCitation>
): AME 0.10; ALE 0.18; PME 0.11; PLE 0.25; AME–AME 0.12; AME–ALE 0.04; PME–PME 0.16; PME–PLE 0.22; AME– PME 0.21; ALE–PLE 0.18; clypeus AME 0.19; clypeus ALE 0.17. Leg formula: 2431; measurements of palp and legs: palp: 3.00 (0.80, 0.50, 0.70, 1.00); I: 6.80 (1.80, 0.90, 1.80, 1.70, 0.60); II: 8.50 (2.40, 1.00, 2.30, 2.10, 0.70); III: 7.30 (2.10, 0.90, 1.90, 1.80, 0.60); IV: 7.60 (2.10, 0.80, 1.90, 2.10, 0.70). Spination: palp: 131, 101, 2221, 1004; legs: Fe I 222/322, Fe II &amp; III 222, Fe IV 221; Pa I–IV 000; Ti I &amp;
<date id="FFEA10445835FFC2FC43FE8383B24149" box="[898,982,295,321]" pageId="7" pageNumber="8" value="1016-02">II 1016</date>
, Ti
<date id="FFEA10445835FFC2FC3EFE8384394149" box="[1023,1117,295,321]" pageId="7" pageNumber="8" value="2126-03">III 2126</date>
, Ti
<date id="FFEA10445835FFC2FB47FE8384824149" box="[1158,1254,295,321]" pageId="7" pageNumber="8" value="2226-04">IV 2226</date>
; Mt I &amp; II 0006, Mt
<date id="FFEA10445835FFC2FF01FEEF817B416D" box="[192,287,331,357]" pageId="7" pageNumber="8" value="2026-03">III 2026</date>
, MT IV 3036. Chelicerae with 3 promarginal, 6 retromarginal teeth, 16 intermarginal denticles and 1 escort seta (
<figureCitation id="136F2A015835FFC2FEDFFECB81174181" box="[286,371,367,393]" captionStart="FIGURES 40–46" captionStartId="10.[151,264,1585,1609]" captionTargetBox="[151,1413,356,1562]" captionTargetId="figure-90@10.[151,1436,345,1562]" captionTargetPageId="10" captionText="FIGURES 40–46. Menarik kecil gen. &amp; spec. nov. holotype male (40, 42, 45) and female paratype (41, 43–44, 46) from Sarawak. 40–41 Eyes, dorsal; 42–44 Claws (42, 44 right leg I, retrolateral claw, retrolateral; 43 left palp, prolateral); 45–46 Left chelicera, ventral." figureDoi="http://doi.org/10.5281/zenodo.6911177" httpUri="https://zenodo.org/record/6911177/files/figure.png" pageId="7" pageNumber="8">Fig. 46</figureCitation>
). Palpal claw with 6 long teeth and 1 short tooth (
<figureCitation id="136F2A015835FFC2FC6BFECB839A4181" box="[938,1022,367,393]" captionStart="FIGURES 40–46" captionStartId="10.[151,264,1585,1609]" captionTargetBox="[151,1413,356,1562]" captionTargetId="figure-90@10.[151,1436,345,1562]" captionTargetPageId="10" captionText="FIGURES 40–46. Menarik kecil gen. &amp; spec. nov. holotype male (40, 42, 45) and female paratype (41, 43–44, 46) from Sarawak. 40–41 Eyes, dorsal; 42–44 Claws (42, 44 right leg I, retrolateral claw, retrolateral; 43 left palp, prolateral); 45–46 Left chelicera, ventral." figureDoi="http://doi.org/10.5281/zenodo.6911177" httpUri="https://zenodo.org/record/6911177/files/figure.png" pageId="7" pageNumber="8">Fig. 43</figureCitation>
), retrolateral claw of right leg I with 16 teeth (
<figureCitation id="136F2A015835FFC2FEC1FE37813041A5" box="[256,340,403,429]" captionStart="FIGURES 40–46" captionStartId="10.[151,264,1585,1609]" captionTargetBox="[151,1413,356,1562]" captionTargetId="figure-90@10.[151,1436,345,1562]" captionTargetPageId="10" captionText="FIGURES 40–46. Menarik kecil gen. &amp; spec. nov. holotype male (40, 42, 45) and female paratype (41, 43–44, 46) from Sarawak. 40–41 Eyes, dorsal; 42–44 Claws (42, 44 right leg I, retrolateral claw, retrolateral; 43 left palp, prolateral); 45–46 Left chelicera, ventral." figureDoi="http://doi.org/10.5281/zenodo.6911177" httpUri="https://zenodo.org/record/6911177/files/figure.png" pageId="7" pageNumber="8">Fig. 44</figureCitation>
).
</paragraph>
<caption id="DF2B660C5835FFC2FF56F8E5818D47CD" ID-DOI="http://doi.org/10.5281/zenodo.6911167" ID-Zenodo-Dep="6911167" httpUri="https://zenodo.org/record/6911167/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,264,1857,1881]" targetBox="[151,1436,453,1834]" targetPageId="7">
<paragraph id="8BEB36845835FFC2FF56F8E5818D47CD" blockId="7.[151,1437,1857,1989]" pageId="7" pageNumber="8">
<emphasis id="B920EA965835FFC2FF56F8E581304751" bold="true" box="[151,340,1857,1881]" pageId="7" pageNumber="8">FIGURES 17–22.</emphasis>
<taxonomicName id="4C544D075835FFC2FE9BF8E581824751" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[346,486,1857,1881]" class="Arachnida" family="Sparassidae" genus="Menarik" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kecil">
<emphasis id="B920EA965835FFC2FE9BF8E581824751" box="[346,486,1857,1881]" italics="true" pageId="7" pageNumber="8">Menarik kecil</emphasis>
</taxonomicName>
<emphasis id="B920EA965835FFC2FE2CF8E582C64751" bold="true" box="[493,674,1857,1881]" pageId="7" pageNumber="8">gen. &amp; spec. nov.</emphasis>
, males from Sarawak (17–20 holotype; 21–22 male, SD 1370). 17–19 Left palp (17 prolateral, 18 ventral, 19 retrolateral); 20–22 Palpal tibia (20, 22 dorsal, 21 retrolateral). aRTA—apical part of RTA, bRTA—basal part of RTA, dB—dorsal branch of RTA, ddA—disto-dorsal apophysis, E—embolus, S—spermophor, ST—subtegulum, TA—tegular apophysis.
</paragraph>
</caption>
<caption id="DF2B660C583AFFCDFF56F8FF856447D7" ID-DOI="http://doi.org/10.5281/zenodo.6911169" ID-Zenodo-Dep="6911169" httpUri="https://zenodo.org/record/6911169/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,264,1883,1907]" targetBox="[222,1366,190,1859]" targetPageId="8">
<paragraph id="8BEB3684583AFFCDFF56F8FF856447D7" blockId="8.[151,1436,1883,2015]" pageId="8" pageNumber="9">
<emphasis id="B920EA96583AFFCDFF56F8FF8132477B" bold="true" box="[151,342,1883,1907]" pageId="8" pageNumber="9">FIGURES 23–27.</emphasis>
<taxonomicName id="4C544D07583AFFCDFE9FF8FF8188477B" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[350,492,1883,1907]" class="Arachnida" family="Sparassidae" genus="Menarik" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="kecil">
<emphasis id="B920EA96583AFFCDFE9FF8FF8188477B" box="[350,492,1883,1907]" italics="true" pageId="8" pageNumber="9">Menarik kecil</emphasis>
</taxonomicName>
<emphasis id="B920EA96583AFFCDFE35F8FF82CA477B" bold="true" box="[500,686,1883,1907]" pageId="8" pageNumber="9">gen. &amp; spec. nov.</emphasis>
female paratypes (23–26) and male paratype (27) from Sarawak. 23–24 Epigyne, ventral; 25 Vulva, dorsal; 26 Schematic course of internal duct system, ventral; 27 Trilobate membrane, distal metatarsus of right leg II, dorsal. At—atrium, EF—epigynal field, FB—“fusion bubbles”, FD—fertilization duct, FW—first winding of internal duct system, GA—glandular appendages, MA—muscle attachment point, Sc—scar, SP—spermatheca.
</paragraph>
</caption>
<paragraph id="8BEB3684583BFFCCFF06FF3382FC4149" blockId="9.[151,1436,151,321]" pageId="9" pageNumber="10">
Copulatory organ as in diagnosis (
<figureCitation id="136F2A01583BFFCCFD8CFF3382AB40B9" box="[589,719,151,177]" captionStart="FIGURES 23–27" captionStartId="8.[151,264,1883,1907]" captionTargetBox="[222,1366,190,1859]" captionTargetId="figure-21@8.[222,1366,181,1859]" captionTargetPageId="8" captionText="FIGURES 23–27. Menarik kecil gen. &amp; spec. nov. female paratypes (23–26) and male paratype (27) from Sarawak. 23–24 Epigyne, ventral; 25 Vulva, dorsal; 26 Schematic course of internal duct system, ventral; 27 Trilobate membrane, distal metatarsus of right leg II, dorsal.At—atrium, EF—epigynal field, FB—“fusion bubbles”, FD—fertilization duct, FW—first winding of internal duct system, GA—glandular appendages, MA—muscle attachment point, Sc—scar, SP—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6911169" httpUri="https://zenodo.org/record/6911169/files/figure.png" pageId="9" pageNumber="10">Figs 23–26</figureCitation>
,
<figureCitation id="136F2A01583BFFCCFD1DFF33834340B9" box="[732,807,151,177]" captionStart="FIGURES 28–39" captionStartId="9.[151,264,1944,1968]" captionTargetBox="[152,1436,345,1921]" captionTargetId="figure-91@9.[151,1436,345,1921]" captionTargetPageId="9" captionText="FIGURES 28–39. Menarik kecil gen. &amp; spec. nov. female paratypes (28–37) and egg-sac (38–39) from Sarawak. 28–36 Epigyne, ventral (28–35 not dissected, 36 dissected); 37 Vulva, dorsal; 38–39 two views of the same egg-sac. Arrows indicating scars at the anterior epigyne. Note the different shapes of the epigynal atrium." figureDoi="http://doi.org/10.5281/zenodo.6911173" httpUri="https://zenodo.org/record/6911173/files/figure.png" pageId="9" pageNumber="10">28–37</figureCitation>
). Epigynal field wider than long with two long anterior “horns” and two indistinct lateral bulges. Lateral lobes fused, with few “fusion bubbles” in posterior half and with slight median indentation at posterior margin, extending very slightly beyond epigastric furrow. Internal duct system wider than long. Glandular appendages short and mainly visible in ventral view. Fertilization ducts arising posterolaterally, long and slender, tips antero-laterad.
</paragraph>
<caption id="DF2B660C583BFFCCFF56F83C83C847F0" ID-DOI="http://doi.org/10.5281/zenodo.6911173" ID-Zenodo-Dep="6911173" httpUri="https://zenodo.org/record/6911173/files/figure.png" pageId="9" pageNumber="10" startId="9.[151,264,1944,1968]" targetBox="[152,1436,345,1921]" targetPageId="9">
<paragraph id="8BEB3684583BFFCCFF56F83C83C847F0" blockId="9.[151,1437,1944,2040]" pageId="9" pageNumber="10">
<emphasis id="B920EA96583BFFCCFF56F83C813147B8" bold="true" box="[151,341,1944,1968]" pageId="9" pageNumber="10">FIGURES 28–39.</emphasis>
<taxonomicName id="4C544D07583BFFCCFE9AF83C818C47B8" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[347,488,1944,1968]" class="Arachnida" family="Sparassidae" genus="Menarik" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="kecil">
<emphasis id="B920EA96583BFFCCFE9AF83C818C47B8" box="[347,488,1944,1968]" italics="true" pageId="9" pageNumber="10">Menarik kecil</emphasis>
</taxonomicName>
<emphasis id="B920EA96583BFFCCFE2EF83C82C147B8" bold="true" box="[495,677,1944,1968]" pageId="9" pageNumber="10">gen. &amp; spec. nov.</emphasis>
female paratypes (28–37) and egg-sac (38–39) from Sarawak. 28–36 Epigyne, ventral (28–35 not dissected, 36 dissected); 37 Vulva, dorsal; 38–39 two views of the same egg-sac. Arrows indicating scars at the anterior epigyne. Note the different shapes of the epigynal atrium.
</paragraph>
</caption>
<paragraph id="8BEB36845838FFCFFF06FF3384144149" blockId="10.[151,1437,151,321]" pageId="10" pageNumber="11">
Colouration in ethanol (
<figureCitation id="136F2A015838FFCFFE1BFF33820540B9" box="[474,609,151,177]" captionStart="FIGURES 82–97" captionStartId="19.[151,264,1899,1923]" captionTargetBox="[152,1435,182,1873]" captionTargetId="figure-19@19.[151,1436,181,1875]" captionTargetPageId="19" captionText="FIGURES 82–97. Heteropodinae gen. &amp; spp. nov., habitus (dorsal and ventral, respectively). 82–85 Borniella parva gen. &amp; spec. nov.; 86–89 Menarik kecil gen. &amp; spec. nov.; 90–93 Micropoda davisae gen. &amp; spec. nov.; 94–97 Tiomaniella ladam gen. &amp; spec. nov. (82–83, 86–87, 90–91, 94–95 holotype male, 84–85, 88–89, 92–93, 96–97 female paratype)." figureDoi="http://doi.org/10.5281/zenodo.6911203" httpUri="https://zenodo.org/record/6911203/files/figure.png" pageId="10" pageNumber="11">Figs 88–89</figureCitation>
): Prosoma yellowish-brown, medially with semi-circular brown pattern, posterior and lateral margin dark brown. Sternum yellowish-white. Opisthosoma brown with yellow spots, medially with yellow transversal stripes, posterior part darker brown, ventral part medially yellow, laterally with brown stripes. Chelicerae yellowish-brown. Coxa I–III medio-dorsally with brown stripe. Palps yellowish-brown; Fe distally brown. Legs yellowish-brown; Fe distally and medially brown; Pa laterally brown.
</paragraph>
<caption id="DF2B660C5838FFCFFF56F99581294699" ID-DOI="http://doi.org/10.5281/zenodo.6911177" ID-Zenodo-Dep="6911177" httpUri="https://zenodo.org/record/6911177/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,264,1585,1609]" targetBox="[151,1413,356,1562]" targetPageId="10">
<paragraph id="8BEB36845838FFCFFF56F99581294699" blockId="10.[151,1436,1585,1681]" pageId="10" pageNumber="11">
<emphasis id="B920EA965838FFCFFF56F99581334641" bold="true" box="[151,343,1585,1609]" pageId="10" pageNumber="11">FIGURES 40–46.</emphasis>
<taxonomicName id="4C544D075838FFCFFEA1F995818B4641" authorityName="Grall &amp; Jäger" authorityYear="2022" box="[352,495,1585,1609]" class="Arachnida" family="Sparassidae" genus="Menarik" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="kecil">
<emphasis id="B920EA965838FFCFFEA1F995818B4641" box="[352,495,1585,1609]" italics="true" pageId="10" pageNumber="11">Menarik kecil</emphasis>
</taxonomicName>
<emphasis id="B920EA965838FFCFFE39F99582D34641" bold="true" box="[504,695,1585,1609]" pageId="10" pageNumber="11">gen. &amp; spec. nov.</emphasis>
holotype male (40, 42, 45) and female paratype (41, 43–44, 46) from Sarawak. 40–41 Eyes, dorsal; 42–44 Claws (42, 44 right leg I, retrolateral claw, retrolateral; 43 left palp, prolateral); 45–46 Left chelicera, ventral.
</paragraph>
</caption>
<paragraph id="8BEB36845838FFCEFF06F91B85E540DD" blockId="10.[151,1437,1727,2042]" lastBlockId="11.[151,1436,151,393]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="B920EA965838FFCFFF06F91B826046D1" bold="true" box="[199,516,1727,1753]" pageId="10" pageNumber="11">Variation. Male (SD 1370).</emphasis>
TL 4.45; PL 2.05, PW 1.95; OL 2.4, OW 1.65. Measurements of leg II: 9.60 (2.70, 1.00, 2.70, 2.40, 0.80); Spination: palp: Mt 1110; legs: Fe II 223, Fe III 222/322; Ti
<date id="FFEA10445838FFCFFBFDF94084F546F5" box="[1084,1169,1763,1790]" pageId="10" pageNumber="11" value="2026-02">II 2026</date>
; Mt II 006, Mt
<date id="FFEA10445838FFCFFAFBF94085FD46F5" box="[1338,1433,1763,1790]" pageId="10" pageNumber="11" value="2026-03">III 2026</date>
. Chelicerae denticles anterior 3, posterior 5 teeth. In the male from Bako National Park the disto-dorsal apophysis of the male palpal tibia and the retrolateral outline of the basal RTA is differently shaped (
<figureCitation id="136F2A015838FFCFFBBDF88F84B4474D" box="[1148,1232,1835,1861]" captionStart="FIGURES 17–22" captionStartId="7.[151,264,1857,1881]" captionTargetBox="[151,1436,453,1834]" captionTargetId="figure-168@7.[151,1436,453,1834]" captionTargetPageId="7" captionText="FIGURES 17–22. Menarik kecil gen. &amp; spec. nov., males from Sarawak (17–20 holotype; 21–22 male, SD 1370). 17–19 Left palp (17 prolateral, 18 ventral, 19 retrolateral); 20–22 Palpal tibia (20, 22 dorsal, 21 retrolateral). aRTA—apical part of RTA, bRTA—basal part of RTA, dB—dorsal branch of RTA, ddA—disto-dorsal apophysis, E—embolus, S—spermophor, ST—subtegulum, TA—tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6911167" httpUri="https://zenodo.org/record/6911167/files/figure.png" pageId="10" pageNumber="11">Fig. 22</figureCitation>
). Additionally, the dorsal branch of the RTA was longer and more pronounced (
<figureCitation id="136F2A015838FFCFFCFAF8EB83EA4761" box="[827,910,1871,1897]" captionStart="FIGURES 17–22" captionStartId="7.[151,264,1857,1881]" captionTargetBox="[151,1436,453,1834]" captionTargetId="figure-168@7.[151,1436,453,1834]" captionTargetPageId="7" captionText="FIGURES 17–22. Menarik kecil gen. &amp; spec. nov., males from Sarawak (17–20 holotype; 21–22 male, SD 1370). 17–19 Left palp (17 prolateral, 18 ventral, 19 retrolateral); 20–22 Palpal tibia (20, 22 dorsal, 21 retrolateral). aRTA—apical part of RTA, bRTA—basal part of RTA, dB—dorsal branch of RTA, ddA—disto-dorsal apophysis, E—embolus, S—spermophor, ST—subtegulum, TA—tegular apophysis." figureDoi="http://doi.org/10.5281/zenodo.6911167" httpUri="https://zenodo.org/record/6911167/files/figure.png" pageId="10" pageNumber="11">Fig. 21</figureCitation>
).
<emphasis id="B920EA965838FFCFFC62F8EB840A4761" bold="true" box="[931,1134,1871,1897]" pageId="10" pageNumber="11">Females (n = 11).</emphasis>
TL 3.7–4.75; PL 1.8–2.15, PW 1.75–2.05, AW 1.1–1.15; OL 1.8–2.6, OW 1.0–1.9. Measurements leg VI: total length 6.90–7.90, Fe 1.95–2.20, Pa 0.70–0.80, Ti 1.80–2.00, Mt 1.80–2.10, Ta 0.65–0.80. Spination: palp: Ti 2121; legs: Fe I 323, Fe II 322, 232, Fe III 232, Fe IV 321, 231; Ti
<date id="FFEA10445838FFCFFE2EF818822947DD" box="[495,589,1979,2006]" pageId="10" pageNumber="11" value="1006-03">III 1006</date>
, 2026, 2226, Ti
<date id="FFEA10445838FFCFFCC4F818830147DD" box="[773,869,1979,2006]" pageId="10" pageNumber="11" value="2126-04">IV 2126</date>
; Mt
<date id="FFEA10445838FFCFFC59F818839347DD" box="[920,1015,1979,2006]" pageId="10" pageNumber="11" value="1016-03">III 1016</date>
, 2006, 2016, 3036. Chelicerae with 3 prolateral, and 5 retrolateral teeth. One female with 6 (left) and 7 (right) retromarginal teeth. Female epigynes with various shapes of atrium (
<figureCitation id="136F2A015839FFCEFE74FF33825340B9" box="[437,567,151,177]" captionStart="FIGURES 28–39" captionStartId="9.[151,264,1944,1968]" captionTargetBox="[152,1436,345,1921]" captionTargetId="figure-91@9.[151,1436,345,1921]" captionTargetPageId="9" captionText="FIGURES 28–39. Menarik kecil gen. &amp; spec. nov. female paratypes (28–37) and egg-sac (38–39) from Sarawak. 28–36 Epigyne, ventral (28–35 not dissected, 36 dissected); 37 Vulva, dorsal; 38–39 two views of the same egg-sac. Arrows indicating scars at the anterior epigyne. Note the different shapes of the epigynal atrium." figureDoi="http://doi.org/10.5281/zenodo.6911173" httpUri="https://zenodo.org/record/6911173/files/figure.png" pageId="11" pageNumber="12">Figs 28–36</figureCitation>
). Spermathecae in some females antero-laterad, ducts leading to spermathecae were narrower (
<figureCitation id="136F2A015839FFCEFE89FF1F819040DD" box="[328,500,187,213]" captionStart="FIGURES 28–39" captionStartId="9.[151,264,1944,1968]" captionTargetBox="[152,1436,345,1921]" captionTargetId="figure-91@9.[151,1436,345,1921]" captionTargetPageId="9" captionText="FIGURES 28–39. Menarik kecil gen. &amp; spec. nov. female paratypes (28–37) and egg-sac (38–39) from Sarawak. 28–36 Epigyne, ventral (28–35 not dissected, 36 dissected); 37 Vulva, dorsal; 38–39 two views of the same egg-sac. Arrows indicating scars at the anterior epigyne. Note the different shapes of the epigynal atrium." figureDoi="http://doi.org/10.5281/zenodo.6911173" httpUri="https://zenodo.org/record/6911173/files/figure.png" pageId="11" pageNumber="12">Figs 29, 31, 33</figureCitation>
), in some female laterad (
<figureCitation id="136F2A015839FFCEFCD7FF1F83F340DD" box="[790,919,187,213]" captionStart="FIGURES 28–39" captionStartId="9.[151,264,1944,1968]" captionTargetBox="[152,1436,345,1921]" captionTargetId="figure-91@9.[151,1436,345,1921]" captionTargetPageId="9" captionText="FIGURES 28–39. Menarik kecil gen. &amp; spec. nov. female paratypes (28–37) and egg-sac (38–39) from Sarawak. 28–36 Epigyne, ventral (28–35 not dissected, 36 dissected); 37 Vulva, dorsal; 38–39 two views of the same egg-sac. Arrows indicating scars at the anterior epigyne. Note the different shapes of the epigynal atrium." figureDoi="http://doi.org/10.5281/zenodo.6911173" httpUri="https://zenodo.org/record/6911173/files/figure.png" pageId="11" pageNumber="12">Figs 28, 30</figureCitation>
), in some females in between (
<figureCitation id="136F2A015839FFCEFB33FF1F851040DD" box="[1266,1396,187,213]" captionStart="FIGURES 28–39" captionStartId="9.[151,264,1944,1968]" captionTargetBox="[152,1436,345,1921]" captionTargetId="figure-91@9.[151,1436,345,1921]" captionTargetPageId="9" captionText="FIGURES 28–39. Menarik kecil gen. &amp; spec. nov. female paratypes (28–37) and egg-sac (38–39) from Sarawak. 28–36 Epigyne, ventral (28–35 not dissected, 36 dissected); 37 Vulva, dorsal; 38–39 two views of the same egg-sac. Arrows indicating scars at the anterior epigyne. Note the different shapes of the epigynal atrium." figureDoi="http://doi.org/10.5281/zenodo.6911173" httpUri="https://zenodo.org/record/6911173/files/figure.png" pageId="11" pageNumber="12">Figs 32, 34</figureCitation>
).
</paragraph>
<paragraph id="8BEB36845839FFCEFF06FF7B821D416D" blockId="11.[151,1436,151,393]" pageId="11" pageNumber="12">
<emphasis id="B920EA965839FFCEFF06FF7B817540F1" bold="true" box="[199,273,223,249]" pageId="11" pageNumber="12">Notes.</emphasis>
The majority of females examined showed scars on the surface of the epigyne. In most females with scars present they are most distinct in the right half (
<figureCitation id="136F2A015839FFCEFD23FEA7835C4115" box="[738,824,259,285]" captionStart="FIGURES 23–27" captionStartId="8.[151,264,1883,1907]" captionTargetBox="[222,1366,190,1859]" captionTargetId="figure-21@8.[222,1366,181,1859]" captionTargetPageId="8" captionText="FIGURES 23–27. Menarik kecil gen. &amp; spec. nov. female paratypes (23–26) and male paratype (27) from Sarawak. 23–24 Epigyne, ventral; 25 Vulva, dorsal; 26 Schematic course of internal duct system, ventral; 27 Trilobate membrane, distal metatarsus of right leg II, dorsal.At—atrium, EF—epigynal field, FB—“fusion bubbles”, FD—fertilization duct, FW—first winding of internal duct system, GA—glandular appendages, MA—muscle attachment point, Sc—scar, SP—spermatheca." figureDoi="http://doi.org/10.5281/zenodo.6911169" httpUri="https://zenodo.org/record/6911169/files/figure.png" pageId="11" pageNumber="12">Figs 23</figureCitation>
,
<figureCitation id="136F2A015839FFCEFC85FEA783874115" box="[836,995,259,285]" captionStart="FIGURES 28–39" captionStartId="9.[151,264,1944,1968]" captionTargetBox="[152,1436,345,1921]" captionTargetId="figure-91@9.[151,1436,345,1921]" captionTargetPageId="9" captionText="FIGURES 28–39. Menarik kecil gen. &amp; spec. nov. female paratypes (28–37) and egg-sac (38–39) from Sarawak. 28–36 Epigyne, ventral (28–35 not dissected, 36 dissected); 37 Vulva, dorsal; 38–39 two views of the same egg-sac. Arrows indicating scars at the anterior epigyne. Note the different shapes of the epigynal atrium." figureDoi="http://doi.org/10.5281/zenodo.6911173" httpUri="https://zenodo.org/record/6911173/files/figure.png" pageId="11" pageNumber="12">28–29, 32–33</figureCitation>
). Only
<specimenCount id="9D52FD0D5839FFCEFBF4FEA784D44115" box="[1077,1200,259,285]" pageId="11" pageNumber="12" type="female">one female</specimenCount>
showed a larger scar on the left side (
<figureCitation id="136F2A015839FFCEFE8BFE8381FA4149" box="[330,414,295,321]" captionStart="FIGURES 28–39" captionStartId="9.[151,264,1944,1968]" captionTargetBox="[152,1436,345,1921]" captionTargetId="figure-91@9.[151,1436,345,1921]" captionTargetPageId="9" captionText="FIGURES 28–39. Menarik kecil gen. &amp; spec. nov. female paratypes (28–37) and egg-sac (38–39) from Sarawak. 28–36 Epigyne, ventral (28–35 not dissected, 36 dissected); 37 Vulva, dorsal; 38–39 two views of the same egg-sac. Arrows indicating scars at the anterior epigyne. Note the different shapes of the epigynal atrium." figureDoi="http://doi.org/10.5281/zenodo.6911173" httpUri="https://zenodo.org/record/6911173/files/figure.png" pageId="11" pageNumber="12">Fig. 30</figureCitation>
). Egg-sacs contained nymphs with TL 1.2, PL 0.65–0.75, the egg-sac with 11 nymphs had a diameter of
<quantity id="4CAC9B615839FFCEFEDFFEEE8110416D" box="[286,372,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="12" unit="mm" value="3.5">3.5 mm</quantity>
and was 2.0 mm thick.
</paragraph>
</subSubSection>
<subSubSection id="C34E650F5839FFCEFF06FECA84E34181" box="[199,1159,366,393]" pageId="11" pageNumber="12" type="distribution">
<paragraph id="8BEB36845839FFCEFF06FECA84E34181" blockId="11.[151,1436,151,393]" box="[199,1159,366,393]" pageId="11" pageNumber="12">
<emphasis id="B920EA965839FFCEFF06FECA81954180" bold="true" box="[199,497,366,393]" pageId="11" pageNumber="12">Distribution and habitat.</emphasis>
Known only from the
<typeStatus id="54EF88265839FFCEFD30FECB83454181" box="[753,801,367,393]" pageId="11" pageNumber="12">type</typeStatus>
locality (
<figureCitation id="136F2A015839FFCEFC4DFECB83844181" box="[908,992,367,393]" captionStart="FIGURE 98" captionStartId="20.[151,250,1436,1460]" captionTargetBox="[151,1436,956,1413]" captionTargetId="figure-372@20.[151,1436,956,1413]" captionTargetPageId="20" captionText="FIGURE 98. Distribution of the new genera and species in Malaysia, Brunei and Papua New Guinea." figureDoi="http://doi.org/10.5281/zenodo.6911205" httpUri="https://zenodo.org/record/6911205/files/figure.png" pageId="11" pageNumber="12">Fig. 98</figureCitation>
: blue squares).
</paragraph>
</subSubSection>
</treatment>
</document>
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