496 lines
78 KiB
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496 lines
78 KiB
XML
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<mods:title id="A155E2D2CE144F2BCA8E9383F4E06971">What are Halomachilis akkesiensis and Halomachilis kojimai described from Hokkaido, Japan? (Insecta: Archaeognatha: Machilidae)</mods:title>
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<mods:namePart id="F10647A591F77EFBE82D01A64FF2092B">Mtow, Shodo</mods:namePart>
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<mods:affiliation id="0D4EEE207AFEE7D04FB80639DDD58997">Department of Agrobiological Resources, Faculty of Agriculture, Meijo University, 1 - 501 Shiogamaguchi, Tenpaku, Nagoya, Aichi 468 - 8502, Japan. & Faculty of Symbiotic Systems Science, Fukushima University, Kanayagawa 1, Fukushima, Fukushima 960 - 1296, Japan.</mods:affiliation>
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<mods:affiliation id="74994E37C8C9913B0F899FE04EAB19A8">Sugadaira Research Station, Mountain Science Center, University of Tsukuba, Sugadaira Kogen 1278 - 294, Ueda, Nagano 386 - 2204, Japan.</mods:affiliation>
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<mods:date id="9B28476D3DC48571E8EA11E9663026F1">2024</mods:date>
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<taxonomicName id="4C544D1A4F0EFF94C7B8FF67FD9FFF49" authority="(Uchida, 1949)" authorityName="Mtow & Machida" authorityYear="2024" baseAuthorityName="Uchida" baseAuthorityYear="1949" box="[151,630,151,178]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="akkesiensis" status="comb. nov.">
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<emphasis id="B920EA8B4F0EFF94C7B8FF67FE50FF4A" bold="true" box="[151,441,151,178]" italics="true" pageId="2" pageNumber="447">Petrobiellus akkesiensis</emphasis>
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<paragraph id="8BEB36994F0EFF94C7B8FF4DFDEEFF2D" blockId="2.[151,1437,151,629]" box="[151,519,189,213]" pageId="2" pageNumber="447">[Japanese name: kojima-ishinomi]</paragraph>
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(
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,
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<emphasis id="B920EA8B4F0EFF94C7B8FEF0FE71FEE0" box="[151,408,256,280]" italics="true" pageId="2" pageNumber="447">Halomachilis akkesiensis</emphasis>
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<bibRefCitation id="EFC54B684F0EFF94C54FFEF0FCEFFEEF" author="Uchida, H." box="[608,774,255,280]" pageId="2" pageNumber="447" pagination="28 - 34" refId="ref3894" refString="Uchida, H. (1955) Synopsis of the Apterygota of Japan and its vicinity (II). Science Reports of the Faculty of Literature and Science, Hirosaki University, 2 (2), 28 - 34." type="journal article" year="1955">Uchida 1955:32</bibRefCitation>
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<treatmentCitation id="0AF510884F0EFF94C4FCFEF0FB61FEEF" author="Machida, R." box="[979,1160,255,280]" page="58" pageId="2" pageNumber="447" year="1996">
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<bibRefCitation id="EFC54B684F0EFF94C4FCFEF0FB61FEEF" author="Machida, R." box="[979,1160,255,280]" pageId="2" pageNumber="447" refId="ref3214" refString="Machida, R. (1996) 4. Microcoryphia. In: Hidaka, T., Ishii, M., Ohtani, T. & Johki, Y. (Eds.), The Encyclopedia of Animals in Japan, Volume 8, Insects I. Heibonsha, Tokyo, pp. 58. [in Japanese]" type="book" year="1996">Machida 1996:58</bibRefCitation>
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<treatmentCitation id="0AF510884F0EFF94C3BDFEF0FAACFEE0" author="Machida, R." box="[1170,1349,256,280]" page="77" pageId="2" pageNumber="447" year="2020">
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<bibRefCitation id="EFC54B684F0EFF94C3BDFEF0FAACFEE0" author="Machida, R." box="[1170,1349,256,280]" pageId="2" pageNumber="447" pagination="76 - 79" refId="ref3430" refString="Machida, R. (2020) Order Archaeognatha. In: The editional committee of catalogue of the insects of Japan (Ed.), Catalogue of the Insects of Japan, Volume 1 Apterygote Orders. Touka Shobo, Fukuoka, pp. 76 - 79. [in Japanese]" type="book chapter" year="2020">Machida 2020:77</bibRefCitation>
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<emphasis id="B920EA8B4F0EFF94C7B8FEEEFE9BFECE" box="[151,370,286,310]" italics="true" pageId="2" pageNumber="447">Halomachilis kojimai</emphasis>
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<bibRefCitation id="EFC54B684F0EFF94C657FEEEFDF1FECE" author="Uchida, H." box="[376,536,286,310]" pageId="2" pageNumber="447" pagination="2 - 6" refId="ref3783" refString="Uchida, H. (1950) Thysanura. In: Ishii, T., Uchida, S., Esaki, T., Kawamura, T., Kinoshita, S., Kuwayama, S., Shiraki, T. & Yuasa, H. (Eds.), Iconographia Insectorum Japonicorum. Hokuryukan, Tokyo, pp. 2 - 6. [in Japanese]" type="book chapter" year="1950">Uchida, 1950:4</bibRefCitation>
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<treatmentCitation id="0AF510884F0EFF94C50DFEEEFD53FECE" author="Uchida, H." box="[546,698,286,310]" page="1" pageId="2" pageNumber="447" year="1965">
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<bibRefCitation id="EFC54B684F0EFF94C50DFEEEFD53FECE" author="Uchida, H." box="[546,698,286,310]" pageId="2" pageNumber="447" pagination="1 - 2" refId="ref3978" refString="Uchida, H. (1965) Thysanura. In: Asahina, S., Ishihara, T. & Yasumatsu, K. (Eds.), Iconographia Insectorum Japonicorum Colore Naturali Edita, Volumen III. Hokuryukan, Tokyo, pp. 1 - 2, pl. 1. [in Japanese]" type="journal article" year="1965">Uchida 1965:1</bibRefCitation>
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<taxonomicName id="4C544D1A4F0EFF94C7B8FECDFEA2FEAD" box="[151,331,317,341]" class="Insecta" family="Machilidae" genus="Petrobius" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="kojimai">
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<emphasis id="B920EA8B4F0EFF94C7B8FECDFEA2FEAD" box="[151,331,317,341]" italics="true" pageId="2" pageNumber="447">Petrobius kojimai</emphasis>
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:
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<treatmentCitation id="0AF510884F0EFF94C677FECDFDF3FEAD" author="Machida, R." box="[344,538,317,341]" page="799" pageId="2" pageNumber="447" year="1999">
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<bibRefCitation id="EFC54B684F0EFF94C677FECDFDF3FEAD" author="Machida, R." box="[344,538,317,341]" pageId="2" pageNumber="447" pagination="792 - 800" refId="ref3276" refString="Machida, R. (1999) Thysanura. In: Aoki, J. (Ed.), Pictorial Keys to Soil Animals of Japan. Tokai University Press, Hadano, Kanagawa, pp. 792 - 800. [in Japanese]" type="book chapter" year="1999">Machida 1999:799</bibRefCitation>
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<treatmentCitation id="0AF510884F0EFF94C50AFECDFD1AFEAD" author="Machida, R." box="[549,755,317,341]" page="1549" pageId="2" pageNumber="447" year="2015">
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<bibRefCitation id="EFC54B684F0EFF94C50AFECDFD1AFEAD" author="Machida, R." box="[549,755,317,341]" pageId="2" pageNumber="447" pagination="1541 - 1551" refId="ref3379" refString="Machida, R. (2015) Thysanura. In: Aoki, J. (Ed.), Pictorial Keys to Soil Animals of Japan (the Second Edition). Tokai University Press, Hadano, Kanagawa, pp. 1541 - 1551. [in Japanese]" type="book chapter" year="2015">Machida 2015:1549</bibRefCitation>
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.
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<paragraph id="8BEB36994F0EFF94C7B8FE91FDFEFE81" blockId="2.[151,1437,151,629]" box="[151,535,353,377]" pageId="2" pageNumber="447">
|
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<treatmentCitationGroup id="AB4411B74F0EFF94C7B8FE91FDFEFE81" box="[151,535,353,377]" pageId="2" pageNumber="447">
|
||
<taxonomicName id="4C544D1A4F0EFF94C7B8FE91FE8DFE81" box="[151,356,353,377]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="kojimai">
|
||
<emphasis id="B920EA8B4F0EFF94C7B8FE91FE8DFE81" box="[151,356,353,377]" italics="true" pageId="2" pageNumber="447">Petrobiellus kojimai</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<treatmentCitation id="0AF510884F0EFF94C65DFE91FDFEFE81" author="Machida, R." box="[370,535,353,377]" page="6" pageId="2" pageNumber="447" year="2008">
|
||
<bibRefCitation id="EFC54B684F0EFF94C65DFE91FDFEFE81" author="Machida, R." box="[370,535,353,377]" pageId="2" pageNumber="447" pagination="4 - 6" refId="ref3322" refString="Machida, R. (2008) Archaeognatha. In: Hirashima, Y. & Morimoto, K. (Eds.), Iconographia Insectorum Japonicorum Colore Naturali Edita, Volumen III (New Edition). Hokuryukan, Tokyo, pp. 4 - 6, pl. 1. [in Japanese]" type="journal article" year="2008">Machida 2008:6</bibRefCitation>
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E65124F0EFF94C7B8FE75FD7CFD8D" pageId="2" pageNumber="447" type="description">
|
||
<paragraph id="8BEB36994F0EFF94C7B8FE75FD7CFD8D" blockId="2.[151,1437,151,629]" pageId="2" pageNumber="447">
|
||
General features similar to those described by
|
||
<bibRefCitation id="EFC54B684F0EFF94C5BBFE75FCC3FE65" author="Uchida, H." box="[660,810,389,413]" pageId="2" pageNumber="447" refId="ref3756" refString="Uchida, H. (1949) Revision on the Japanese Machilidae. Zoological Magazine, 58 (6), 110. [in Japanese]" type="journal volume" year="1949">Uchida (1949</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC54B684F0EFF94C418FE75FC98FE65" author="Uchida, H." box="[823,881,389,413]" pageId="2" pageNumber="447" pagination="2 - 6" refId="ref3783" refString="Uchida, H. (1950) Thysanura. In: Ishii, T., Uchida, S., Esaki, T., Kawamura, T., Kinoshita, S., Kuwayama, S., Shiraki, T. & Yuasa, H. (Eds.), Iconographia Insectorum Japonicorum. Hokuryukan, Tokyo, pp. 2 - 6. [in Japanese]" type="book chapter" year="1950">1950</bibRefCitation>
|
||
). Body length of males
|
||
<quantity id="4CAC9B7C4F0EFF94C358FE75FB04FE64" box="[1143,1261,389,413]" metricMagnitude="-2" metricUnit="m" metricValue="1.15" metricValueMax="1.2" metricValueMin="1.1" pageId="2" pageNumber="447" unit="mm" value="11.5" valueMax="12.0" valueMin="11.0">11–12 mm</quantity>
|
||
, that of females
|
||
<quantity id="4CAC9B7C4F0EFF94C7B8FE59FEE7FE38" box="[151,270,425,449]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="2" pageNumber="447" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
|
||
. Antennal length approximately
|
||
<quantity id="4CAC9B7C4F0EFF94C542FE59FD55FE38" box="[621,700,425,449]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="447" unit="mm" value="15.0">15 mm</quantity>
|
||
. Cerci strongly curving outwards (
|
||
<figureCitation id="136F2A1C4F0EFF94C31EFE59FB46FE39" box="[1073,1199,425,449]" captionStart="FIGURE 2" captionStartId="3.[151,250,1765,1789]" captionTargetBox="[154,1433,181,1741]" captionTargetId="figure-17@3.[154,1433,181,1741]" captionTargetPageId="3" captionText="FIGURE 2. External features of Petrobiellus akkesiensis comb. nov. (A) Male habitus. (B) Female habitus. (C–H) Female morphology. (C) Right antenna, anterior (medial) view. (D) Left maxilla, lateral view. (E) Labium, posterior view. (F) Left foreleg, posterior (lateral) view. (G) Left middle leg, posterior view. (H) Left hind leg, posterior (medial) view. (I) Male genitalic region, ventral view. (J) Post-abdomen of female, ventrolateral view. Abbreviations: Cx9: coxite IX; Ov: ovipositor; Pa: paramera; Pe: penis; St9: stylus IX. Scale bars: A, B, 5 mm; C–H, J, 1 mm; I, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14386205" httpUri="https://zenodo.org/record/14386205/files/figure.png" pageId="2" pageNumber="447">Figs. 2A, B</figureCitation>
|
||
) and approximately
|
||
<quantity id="4CAC9B7C4F0EFF94C2A0FE59FF29FE1C" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="447" unit="mm" value="5.0">5 mm</quantity>
|
||
long; caudal filament approximately
|
||
<quantity id="4CAC9B7C4F0EFF94C573FE3DFD41FE1C" box="[604,680,461,485]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="447" unit="mm" value="15.0">15 mm</quantity>
|
||
long. The scale pattern on the dorsal body surface as shown in
|
||
<figureCitation id="136F2A1C4F0EFF94C247FE3DFF32FDF0" captionStart="FIGURE 2" captionStartId="3.[151,250,1765,1789]" captionTargetBox="[154,1433,181,1741]" captionTargetId="figure-17@3.[154,1433,181,1741]" captionTargetPageId="3" captionText="FIGURE 2. External features of Petrobiellus akkesiensis comb. nov. (A) Male habitus. (B) Female habitus. (C–H) Female morphology. (C) Right antenna, anterior (medial) view. (D) Left maxilla, lateral view. (E) Labium, posterior view. (F) Left foreleg, posterior (lateral) view. (G) Left middle leg, posterior view. (H) Left hind leg, posterior (medial) view. (I) Male genitalic region, ventral view. (J) Post-abdomen of female, ventrolateral view. Abbreviations: Cx9: coxite IX; Ov: ovipositor; Pa: paramera; Pe: penis; St9: stylus IX. Scale bars: A, B, 5 mm; C–H, J, 1 mm; I, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14386205" httpUri="https://zenodo.org/record/14386205/files/figure.png" pageId="2" pageNumber="447">Figs. 2A, B</figureCitation>
|
||
. Scales absent from the antennae (
|
||
<figureCitation id="136F2A1C4F0EFF94C57BFE01FD40FDF1" box="[596,681,497,521]" captionStart="FIGURE 2" captionStartId="3.[151,250,1765,1789]" captionTargetBox="[154,1433,181,1741]" captionTargetId="figure-17@3.[154,1433,181,1741]" captionTargetPageId="3" captionText="FIGURE 2. External features of Petrobiellus akkesiensis comb. nov. (A) Male habitus. (B) Female habitus. (C–H) Female morphology. (C) Right antenna, anterior (medial) view. (D) Left maxilla, lateral view. (E) Labium, posterior view. (F) Left foreleg, posterior (lateral) view. (G) Left middle leg, posterior view. (H) Left hind leg, posterior (medial) view. (I) Male genitalic region, ventral view. (J) Post-abdomen of female, ventrolateral view. Abbreviations: Cx9: coxite IX; Ov: ovipositor; Pa: paramera; Pe: penis; St9: stylus IX. Scale bars: A, B, 5 mm; C–H, J, 1 mm; I, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14386205" httpUri="https://zenodo.org/record/14386205/files/figure.png" pageId="2" pageNumber="447">Fig. 2C</figureCitation>
|
||
), maxillary and labial palpi (
|
||
<figureCitation id="136F2A1C4F0EFF94C4CBFE01FB8DFDF1" box="[996,1124,497,521]" captionStart="FIGURE 2" captionStartId="3.[151,250,1765,1789]" captionTargetBox="[154,1433,181,1741]" captionTargetId="figure-17@3.[154,1433,181,1741]" captionTargetPageId="3" captionText="FIGURE 2. External features of Petrobiellus akkesiensis comb. nov. (A) Male habitus. (B) Female habitus. (C–H) Female morphology. (C) Right antenna, anterior (medial) view. (D) Left maxilla, lateral view. (E) Labium, posterior view. (F) Left foreleg, posterior (lateral) view. (G) Left middle leg, posterior view. (H) Left hind leg, posterior (medial) view. (I) Male genitalic region, ventral view. (J) Post-abdomen of female, ventrolateral view. Abbreviations: Cx9: coxite IX; Ov: ovipositor; Pa: paramera; Pe: penis; St9: stylus IX. Scale bars: A, B, 5 mm; C–H, J, 1 mm; I, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14386205" httpUri="https://zenodo.org/record/14386205/files/figure.png" pageId="2" pageNumber="447">Figs. 2D, E</figureCitation>
|
||
), thoracic legs (
|
||
<figureCitation id="136F2A1C4F0EFF94C23DFE01FA79FDF1" box="[1298,1424,497,521]" captionStart="FIGURE 2" captionStartId="3.[151,250,1765,1789]" captionTargetBox="[154,1433,181,1741]" captionTargetId="figure-17@3.[154,1433,181,1741]" captionTargetPageId="3" captionText="FIGURE 2. External features of Petrobiellus akkesiensis comb. nov. (A) Male habitus. (B) Female habitus. (C–H) Female morphology. (C) Right antenna, anterior (medial) view. (D) Left maxilla, lateral view. (E) Labium, posterior view. (F) Left foreleg, posterior (lateral) view. (G) Left middle leg, posterior view. (H) Left hind leg, posterior (medial) view. (I) Male genitalic region, ventral view. (J) Post-abdomen of female, ventrolateral view. Abbreviations: Cx9: coxite IX; Ov: ovipositor; Pa: paramera; Pe: penis; St9: stylus IX. Scale bars: A, B, 5 mm; C–H, J, 1 mm; I, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14386205" httpUri="https://zenodo.org/record/14386205/files/figure.png" pageId="2" pageNumber="447">Figs. 2F–H</figureCitation>
|
||
), and coxal and abdominal styli (
|
||
<figureCitation id="136F2A1C4F0EFF94C6CAFDE5FDB7FDD5" box="[485,606,533,557]" captionStart="FIGURE 2" captionStartId="3.[151,250,1765,1789]" captionTargetBox="[154,1433,181,1741]" captionTargetId="figure-17@3.[154,1433,181,1741]" captionTargetPageId="3" captionText="FIGURE 2. External features of Petrobiellus akkesiensis comb. nov. (A) Male habitus. (B) Female habitus. (C–H) Female morphology. (C) Right antenna, anterior (medial) view. (D) Left maxilla, lateral view. (E) Labium, posterior view. (F) Left foreleg, posterior (lateral) view. (G) Left middle leg, posterior view. (H) Left hind leg, posterior (medial) view. (I) Male genitalic region, ventral view. (J) Post-abdomen of female, ventrolateral view. Abbreviations: Cx9: coxite IX; Ov: ovipositor; Pa: paramera; Pe: penis; St9: stylus IX. Scale bars: A, B, 5 mm; C–H, J, 1 mm; I, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14386205" httpUri="https://zenodo.org/record/14386205/files/figure.png" pageId="2" pageNumber="447">Figs. 2G–J</figureCitation>
|
||
). Compound eyes in living specimens army-green in color; dark pigmentation present on the scapus (
|
||
<figureCitation id="136F2A1C4F0EFF94C6BFFDC9FE0FFDA9" box="[400,486,569,593]" captionStart="FIGURE 2" captionStartId="3.[151,250,1765,1789]" captionTargetBox="[154,1433,181,1741]" captionTargetId="figure-17@3.[154,1433,181,1741]" captionTargetPageId="3" captionText="FIGURE 2. External features of Petrobiellus akkesiensis comb. nov. (A) Male habitus. (B) Female habitus. (C–H) Female morphology. (C) Right antenna, anterior (medial) view. (D) Left maxilla, lateral view. (E) Labium, posterior view. (F) Left foreleg, posterior (lateral) view. (G) Left middle leg, posterior view. (H) Left hind leg, posterior (medial) view. (I) Male genitalic region, ventral view. (J) Post-abdomen of female, ventrolateral view. Abbreviations: Cx9: coxite IX; Ov: ovipositor; Pa: paramera; Pe: penis; St9: stylus IX. Scale bars: A, B, 5 mm; C–H, J, 1 mm; I, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14386205" httpUri="https://zenodo.org/record/14386205/files/figure.png" pageId="2" pageNumber="447">Fig. 2C</figureCitation>
|
||
), pedicellus (
|
||
<figureCitation id="136F2A1C4F0EFF94C558FDC9FD24FDA9" box="[631,717,569,593]" captionStart="FIGURE 2" captionStartId="3.[151,250,1765,1789]" captionTargetBox="[154,1433,181,1741]" captionTargetId="figure-17@3.[154,1433,181,1741]" captionTargetPageId="3" captionText="FIGURE 2. External features of Petrobiellus akkesiensis comb. nov. (A) Male habitus. (B) Female habitus. (C–H) Female morphology. (C) Right antenna, anterior (medial) view. (D) Left maxilla, lateral view. (E) Labium, posterior view. (F) Left foreleg, posterior (lateral) view. (G) Left middle leg, posterior view. (H) Left hind leg, posterior (medial) view. (I) Male genitalic region, ventral view. (J) Post-abdomen of female, ventrolateral view. Abbreviations: Cx9: coxite IX; Ov: ovipositor; Pa: paramera; Pe: penis; St9: stylus IX. Scale bars: A, B, 5 mm; C–H, J, 1 mm; I, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14386205" httpUri="https://zenodo.org/record/14386205/files/figure.png" pageId="2" pageNumber="447">Fig. 2C</figureCitation>
|
||
), maxillary and labial palpi (
|
||
<figureCitation id="136F2A1C4F0EFF94C327FDC9FB60FDA9" box="[1032,1161,569,593]" captionStart="FIGURE 2" captionStartId="3.[151,250,1765,1789]" captionTargetBox="[154,1433,181,1741]" captionTargetId="figure-17@3.[154,1433,181,1741]" captionTargetPageId="3" captionText="FIGURE 2. External features of Petrobiellus akkesiensis comb. nov. (A) Male habitus. (B) Female habitus. (C–H) Female morphology. (C) Right antenna, anterior (medial) view. (D) Left maxilla, lateral view. (E) Labium, posterior view. (F) Left foreleg, posterior (lateral) view. (G) Left middle leg, posterior view. (H) Left hind leg, posterior (medial) view. (I) Male genitalic region, ventral view. (J) Post-abdomen of female, ventrolateral view. Abbreviations: Cx9: coxite IX; Ov: ovipositor; Pa: paramera; Pe: penis; St9: stylus IX. Scale bars: A, B, 5 mm; C–H, J, 1 mm; I, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14386205" httpUri="https://zenodo.org/record/14386205/files/figure.png" pageId="2" pageNumber="447">Figs. 2D, E</figureCitation>
|
||
), and thoracic legs (
|
||
<figureCitation id="136F2A1C4F0EFF94C248FDC9FF30FD8D" captionStart="FIGURE 2" captionStartId="3.[151,250,1765,1789]" captionTargetBox="[154,1433,181,1741]" captionTargetId="figure-17@3.[154,1433,181,1741]" captionTargetPageId="3" captionText="FIGURE 2. External features of Petrobiellus akkesiensis comb. nov. (A) Male habitus. (B) Female habitus. (C–H) Female morphology. (C) Right antenna, anterior (medial) view. (D) Left maxilla, lateral view. (E) Labium, posterior view. (F) Left foreleg, posterior (lateral) view. (G) Left middle leg, posterior view. (H) Left hind leg, posterior (medial) view. (I) Male genitalic region, ventral view. (J) Post-abdomen of female, ventrolateral view. Abbreviations: Cx9: coxite IX; Ov: ovipositor; Pa: paramera; Pe: penis; St9: stylus IX. Scale bars: A, B, 5 mm; C–H, J, 1 mm; I, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14386205" httpUri="https://zenodo.org/record/14386205/files/figure.png" pageId="2" pageNumber="447">Figs. 2F–H</figureCitation>
|
||
). Ovipositor extremely long (
|
||
<figureCitation id="136F2A1C4F0EFF94C53DFDADFD61FD8D" box="[530,648,605,629]" captionStart="FIGURE 2" captionStartId="3.[151,250,1765,1789]" captionTargetBox="[154,1433,181,1741]" captionTargetId="figure-17@3.[154,1433,181,1741]" captionTargetPageId="3" captionText="FIGURE 2. External features of Petrobiellus akkesiensis comb. nov. (A) Male habitus. (B) Female habitus. (C–H) Female morphology. (C) Right antenna, anterior (medial) view. (D) Left maxilla, lateral view. (E) Labium, posterior view. (F) Left foreleg, posterior (lateral) view. (G) Left middle leg, posterior view. (H) Left hind leg, posterior (medial) view. (I) Male genitalic region, ventral view. (J) Post-abdomen of female, ventrolateral view. Abbreviations: Cx9: coxite IX; Ov: ovipositor; Pa: paramera; Pe: penis; St9: stylus IX. Scale bars: A, B, 5 mm; C–H, J, 1 mm; I, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14386205" httpUri="https://zenodo.org/record/14386205/files/figure.png" pageId="2" pageNumber="447">Figs. 2B, J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E65124F0EFF94C7B8FD37FD9BF955" pageId="2" pageNumber="447" type="discussion">
|
||
<paragraph id="8BEB36994F0EFF94C7B8FD37FEE0FD19" blockId="2.[151,1437,711,1709]" box="[151,265,711,737]" pageId="2" pageNumber="447">
|
||
<emphasis id="B920EA8B4F0EFF94C7B8FD37FEE0FD19" bold="true" box="[151,265,711,737]" pageId="2" pageNumber="447">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BEB36994F0EFF94C7B8FD1DFE3FFC88" blockId="2.[151,1437,711,1709]" pageId="2" pageNumber="447">
|
||
The ovipositor is much longer than indicated in
|
||
<bibRefCitation id="EFC54B684F0EFF94C5B6FD1DFCDFFCFD" author="Uchida, H." box="[665,822,749,773]" pageId="2" pageNumber="447" pagination="2 - 6" refId="ref3783" refString="Uchida, H. (1950) Thysanura. In: Ishii, T., Uchida, S., Esaki, T., Kawamura, T., Kinoshita, S., Kuwayama, S., Shiraki, T. & Yuasa, H. (Eds.), Iconographia Insectorum Japonicorum. Hokuryukan, Tokyo, pp. 2 - 6. [in Japanese]" type="book chapter" year="1950">Uchida (1950)</bibRefCitation>
|
||
, who had described the ovipositor of the species as short, with its apex slightly surpassing the end of coxite IX. This difference could probably be attributed to the specimens Uchida examined being younger females: the body length in Uchida’s description was
|
||
<quantity id="4CAC9B7C4F0EFF94C300FCC5FB7EFCB4" box="[1071,1175,821,845]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="2" pageNumber="447" unit="mm" value="9.5" valueMax="10.0" valueMin="9.0">9–10 mm</quantity>
|
||
, whereas that of females we examined was
|
||
<quantity id="4CAC9B7C4F0EFF94C674FCA9FE3BFC88" box="[347,466,857,881]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.3" metricValueMin="1.2" pageId="2" pageNumber="447" unit="mm" value="12.5" valueMax="13.0" valueMin="12.0">12–13 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BEB36994F0EFF94C7E8FC8DFDC7FC25" blockId="2.[151,1437,711,1709]" pageId="2" pageNumber="447">
|
||
<taxonomicName id="4C544D1A4F0EFF94C7E8FC8DFEA1FC6D" authorityName="Silvestri" authorityYear="1943" box="[199,328,893,917]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B920EA8B4F0EFF94C7E8FC8DFEA1FC6D" box="[199,328,893,917]" italics="true" pageId="2" pageNumber="447">Petrobiellus</emphasis>
|
||
</taxonomicName>
|
||
is an enigmatic, halophilic genus of
|
||
<taxonomicName id="4C544D1A4F0EFF94C5C5FC8DFC62FC6D" authorityName="Borner" authorityYear="1904" box="[746,907,893,917]" class="Insecta" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="order">Archaeognatha</taxonomicName>
|
||
found only in
|
||
<collectingCountry id="F34376094F0EFF94C317FC8DFB9CFC6C" box="[1080,1141,893,916]" name="Japan" pageId="2" pageNumber="447">Japan</collectingCountry>
|
||
and the Russian Far East. Absence of scales on the appendages is one of the most significant diagnostic features of
|
||
<taxonomicName id="4C544D1A4F0EFF94C344FC51FB05FC41" authorityName="Silvestri" authorityYear="1943" box="[1131,1260,929,953]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B920EA8B4F0EFF94C344FC51FB05FC41" box="[1131,1260,929,953]" italics="true" pageId="2" pageNumber="447">Petrobiellus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFC54B684F0EFF94C3D3FC51FA7EFC41" author="Silvestri, F." box="[1276,1431,929,953]" pageId="2" pageNumber="447" pagination="283 - 306" refId="ref3591" refString="Silvestri, F. (1943) Contributo alla conoscenza dei Machilidae (Insecta, Thysanura) del Giappone. Bollettino del Laboratorio di Zoologia, Generale e Agraria della Facolta Agraria in Portici, 32, 283 - 306." type="journal article" year="1943">Silvestri 1943</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC54B684F0EFF94C7B8FC35FE7EFC25" author="Sturm, H. & Machida, R." box="[151,407,965,989]" pageId="2" pageNumber="447" refId="ref3632" refString="Sturm, H. & Machida, R. (2001) Archaeognatha. In: Kristensen, N. P. & Beutel, R. G. (Eds.), Handbook of Zoology, Volume IV Arthropoda: Insecta Part 37. Walter de Gruyter, Berlin, New York, 213 pp." type="book" year="2001">Sturm & Machida 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC54B684F0EFF94C68DFC35FDCBFC25" author="Mtow, S." box="[418,546,965,989]" pageId="2" pageNumber="447" pagination="108 - 110" refId="ref3484" refString="Mtow, S. (2021) A new record of Petrobiellus takunagae Silvestri, 1943 (Archaeognatha, Machilidae, Petrobiellinae) from Sado Island, Niigata Prefecture, Japan. Japanese Journal of Entomology (New Series), 24 (4), 108 - 110. https: // doi. org / 10.20848 / kontyu. 24.4 _ 108" type="book chapter" year="2021">Mtow 2021</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BEB36994F0EFF94C7E8FC19FACEFABD" blockId="2.[151,1437,711,1709]" pageId="2" pageNumber="447">
|
||
The genus contains of four validly described species:
|
||
<taxonomicName id="4C544D1A4F0EFF94C406FC19FB39FBF9" authority="Silvestri, 1943" authorityName="Silvestri" authorityYear="1943" box="[809,1232,1001,1025]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="takunagae">
|
||
<emphasis id="B920EA8B4F0EFF94C406FC19FBCCFBF9" box="[809,1061,1001,1025]" italics="true" pageId="2" pageNumber="447">Petrobiellus takunagae</emphasis>
|
||
<bibRefCitation id="EFC54B684F0EFF94C31FFC19FB39FBF9" author="Silvestri, F." box="[1072,1232,1001,1025]" pageId="2" pageNumber="447" pagination="283 - 306" refId="ref3591" refString="Silvestri, F. (1943) Contributo alla conoscenza dei Machilidae (Insecta, Thysanura) del Giappone. Bollettino del Laboratorio di Zoologia, Generale e Agraria della Facolta Agraria in Portici, 32, 283 - 306." type="journal article" year="1943">Silvestri, 1943</bibRefCitation>
|
||
</taxonomicName>
|
||
[from Shirahama,
|
||
<collectingRegion id="4990F87B4F0EFF94C7B8FBFDFEF9FBDD" box="[151,272,1037,1061]" country="Japan" name="Wakayama" pageId="2" pageNumber="447">Wakayama</collectingRegion>
|
||
,
|
||
<collectingCountry id="F34376094F0EFF94C632FBFDFEB3FBDC" box="[285,346,1037,1060]" name="Japan" pageId="2" pageNumber="447">Japan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFC54B684F0EFF94C645FBFDFDECFBDD" author="Silvestri, F." box="[362,517,1037,1061]" pageId="2" pageNumber="447" pagination="283 - 306" refId="ref3591" refString="Silvestri, F. (1943) Contributo alla conoscenza dei Machilidae (Insecta, Thysanura) del Giappone. Bollettino del Laboratorio di Zoologia, Generale e Agraria della Facolta Agraria in Portici, 32, 283 - 306." type="journal article" year="1943">Silvestri 1943</bibRefCitation>
|
||
) and from Shimoda,
|
||
<collectingRegion id="4990F87B4F0EFF94C5DEFBFDFCBEFBDD" box="[753,855,1037,1061]" country="Japan" name="Shizuoka" pageId="2" pageNumber="447">Shizuoka</collectingRegion>
|
||
,
|
||
<collectingCountry id="F34376094F0EFF94C44BFBFDFC48FBDC" box="[868,929,1037,1060]" name="Japan" pageId="2" pageNumber="447">Japan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFC54B684F0EFF94C49EFBFDFBB8FBDD" author="Machida, R." box="[945,1105,1037,1061]" pageId="2" pageNumber="447" pagination="792 - 800" refId="ref3276" refString="Machida, R. (1999) Thysanura. In: Aoki, J. (Ed.), Pictorial Keys to Soil Animals of Japan. Tokai University Press, Hadano, Kanagawa, pp. 792 - 800. [in Japanese]" type="book chapter" year="1999">Machida 1999</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC54B684F0EFF94C371FBFDFB70FBDD" author="Machida, R." box="[1118,1177,1037,1061]" pageId="2" pageNumber="447" pagination="76 - 79" refId="ref3430" refString="Machida, R. (2020) Order Archaeognatha. In: The editional committee of catalogue of the insects of Japan (Ed.), Catalogue of the Insects of Japan, Volume 1 Apterygote Orders. Touka Shobo, Fukuoka, pp. 76 - 79. [in Japanese]" type="book chapter" year="2020">2020</bibRefCitation>
|
||
)],
|
||
<taxonomicName id="4C544D1A4F0EFF94C39AFBFEFF25FBB1" authority="Uchida, 1954" authorityName="Uchida" authorityYear="1954" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="curvistylis">
|
||
<emphasis id="B920EA8B4F0EFF94C39AFBFEFAA9FBDD" box="[1205,1344,1037,1061]" italics="true" pageId="2" pageNumber="447">P. curvistylis</emphasis>
|
||
<bibRefCitation id="EFC54B684F0EFF94C266FBFDFF25FBB1" author="Uchida, H." pageId="2" pageNumber="447" pagination="1 - 17" refId="ref3856" refString="Uchida, H. (1954) Apterygota of the Hachijo-Jima and its adjacent islands. Science Reports of the Faculty of Literature and Science, Hirosaki University, 1 (1), 1 - 17." type="journal article" year="1954">Uchida, 1954</bibRefCitation>
|
||
</taxonomicName>
|
||
[from Hachijo Island,
|
||
<collectingRegion id="4990F87B4F0EFF94C6E3FBC1FDFAFBB1" box="[460,531,1073,1097]" country="Japan" name="Tokyo" pageId="2" pageNumber="447">Tokyo</collectingRegion>
|
||
,
|
||
<collectingCountry id="F34376094F0EFF94C50FFBC1FDB4FBB0" box="[544,605,1073,1096]" name="Japan" pageId="2" pageNumber="447">Japan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFC54B684F0EFF94C542FBC1FCE9FBB1" author="Uchida, H." box="[621,768,1073,1097]" pageId="2" pageNumber="447" pagination="1 - 17" refId="ref3856" refString="Uchida, H. (1954) Apterygota of the Hachijo-Jima and its adjacent islands. Science Reports of the Faculty of Literature and Science, Hirosaki University, 1 (1), 1 - 17." type="journal article" year="1954">Uchida 1954</bibRefCitation>
|
||
)],
|
||
<taxonomicName id="4C544D1A4F0EFF94C432FBC2FBC3FBB1" authority="Kaplin, 1980" authorityName="Kaplin" authorityYear="1980" box="[797,1066,1073,1097]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="kusakini">
|
||
<emphasis id="B920EA8B4F0EFF94C432FBC2FC7AFBB1" box="[797,915,1073,1097]" italics="true" pageId="2" pageNumber="447">P. kusakini</emphasis>
|
||
<bibRefCitation id="EFC54B684F0EFF94C4B3FBC1FBC3FBB1" author="Kaplin, V. G." box="[924,1066,1073,1097]" pageId="2" pageNumber="447" pagination="3 - 9" refId="ref2848" refString="Kaplin, V. G. (1980) New species of bristletails (Microcoryphia, Machilidae) from the Kuril Islands and Primorsky Krai. In: Lehr, P. A. (Ed.), Taxonomy of Insects of the Far East. Far Eastern Scientific Center, Academy of Sciences of the Union of Soviet Socialist Republics, Vladivostok, pp. 3 - 9. [in Russian]" type="book chapter" year="1980">Kaplin, 1980</bibRefCitation>
|
||
</taxonomicName>
|
||
[from Simushir Island, Chishima Islands,
|
||
<collectingCountry id="F34376094F0EFF94C7DDFBA5FEC6FB94" box="[242,303,1109,1132]" name="Japan" pageId="2" pageNumber="447">Japan</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFC54B684F0EFF94C611FBA5FE25FB95" author="Kaplin, V. G." box="[318,460,1109,1133]" pageId="2" pageNumber="447" pagination="3 - 9" refId="ref2848" refString="Kaplin, V. G. (1980) New species of bristletails (Microcoryphia, Machilidae) from the Kuril Islands and Primorsky Krai. In: Lehr, P. A. (Ed.), Taxonomy of Insects of the Far East. Far Eastern Scientific Center, Academy of Sciences of the Union of Soviet Socialist Republics, Vladivostok, pp. 3 - 9. [in Russian]" type="book chapter" year="1980">Kaplin 1980</bibRefCitation>
|
||
)], and
|
||
<taxonomicName id="4C544D1A4F0EFF94C53AFBA6FCBFFB95" authority="Kaplin, 2020" authorityName="Kaplin" authorityYear="2020" box="[533,854,1109,1133]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="sachalinensis">
|
||
<emphasis id="B920EA8B4F0EFF94C53AFBA6FD28FB95" box="[533,705,1109,1133]" italics="true" pageId="2" pageNumber="447">P. sachalinensis</emphasis>
|
||
<bibRefCitation id="EFC54B684F0EFF94C5E6FBA5FCBFFB95" author="Kaplin, V. G." box="[713,854,1109,1133]" pageId="2" pageNumber="447" pagination="17 - 22" refId="ref2967" refString="Kaplin, V. G. (2020) A new species of bristletails of the genus Petrobiellus (Microcoryphia: Machilidae) from Sakhalin. Zoosystematica Rossica, 29 (1), 17 - 22. https: // doi. org / 10.31610 / zsr / 2020.29.1.17" type="journal article" year="2020">Kaplin, 2020</bibRefCitation>
|
||
</taxonomicName>
|
||
[from
|
||
<collectingRegion id="4990F87B4F0EFF94C48BFBA5FBE8FB95" box="[932,1025,1109,1133]" country="Russia" name="Sakhalin" pageId="2" pageNumber="447">Sakhalin</collectingRegion>
|
||
Island,
|
||
<collectingCountry id="F34376094F0EFF94C376FBA5FB49FB95" box="[1113,1184,1109,1133]" name="Russia" pageId="2" pageNumber="447">Russia</collectingCountry>
|
||
<bibRefCitation id="EFC54B684F0EFF94C386FBA5FAADFB95" author="Kaplin, V. G." box="[1193,1348,1109,1133]" pageId="2" pageNumber="447" pagination="17 - 22" refId="ref2967" refString="Kaplin, V. G. (2020) A new species of bristletails of the genus Petrobiellus (Microcoryphia: Machilidae) from Sakhalin. Zoosystematica Rossica, 29 (1), 17 - 22. https: // doi. org / 10.31610 / zsr / 2020.29.1.17" type="journal article" year="2020">Kaplin (2020)</bibRefCitation>
|
||
] (
|
||
<bibRefCitation id="EFC54B684F0EFF94C276FBA5FEB8FB69" author="Sturm, H. & Machida, R." pageId="2" pageNumber="447" refId="ref3632" refString="Sturm, H. & Machida, R. (2001) Archaeognatha. In: Kristensen, N. P. & Beutel, R. G. (Eds.), Handbook of Zoology, Volume IV Arthropoda: Insecta Part 37. Walter de Gruyter, Berlin, New York, 213 pp." type="book" year="2001">Sturm & Machida 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC54B684F0EFF94C673FB89FE35FB69" author="Mtow, S." box="[348,476,1145,1169]" pageId="2" pageNumber="447" pagination="108 - 110" refId="ref3484" refString="Mtow, S. (2021) A new record of Petrobiellus takunagae Silvestri, 1943 (Archaeognatha, Machilidae, Petrobiellinae) from Sado Island, Niigata Prefecture, Japan. Japanese Journal of Entomology (New Series), 24 (4), 108 - 110. https: // doi. org / 10.20848 / kontyu. 24.4 _ 108" type="book chapter" year="2021">Mtow 2021</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="EFC54B684F0EFF94C6C1FB89FD77FB69" author="Ma, Y. & He, K. & Yu, P. & Yu, D. & Cheng, X. & Zhang, J." box="[494,670,1145,1169]" pageId="2" pageNumber="447" refId="ref3134" refString="Ma, Y., He, K., Yu, P., Yu, D., Cheng, X. & Zhang, J. (2015) The complete mitochondrial genomes of three bristletails (Insecta: Archaeognatha): the paraphyly of Machilidae and insights into archaeognathan phylogeny. Plos ONE, 10 (1), e 0117669. https: // doi. org / 10.1371 / journal. pone. 0117669" type="journal volume" year="2015">
|
||
Ma
|
||
<emphasis id="B920EA8B4F0EFF94C536FB8AFDA0FB69" box="[537,585,1145,1169]" italics="true" pageId="2" pageNumber="447">et al</emphasis>
|
||
. (2015)
|
||
</bibRefCitation>
|
||
detected two “
|
||
<taxonomicName id="4C544D1A4F0EFF94C46FFB89FC28FB69" authorityName="Silvestri" authorityYear="1943" box="[832,961,1145,1169]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B920EA8B4F0EFF94C46FFB89FC28FB69" box="[832,961,1145,1169]" italics="true" pageId="2" pageNumber="447">Petrobiellus</emphasis>
|
||
</taxonomicName>
|
||
” species from
|
||
<collectingRegion id="4990F87B4F0EFF94C34DFB89FAF4FB69" box="[1122,1309,1145,1169]" country="China" name="Yunnan" pageId="2" pageNumber="447">Yunnan Province</collectingRegion>
|
||
,
|
||
<collectingCountry id="F34376094F0EFF94C208FB89FA80FB69" box="[1319,1385,1145,1169]" name="China" pageId="2" pageNumber="447">China</collectingCountry>
|
||
, i.e.,
|
||
<taxonomicName id="4C544D1A4F0EFF94C7B8FB6EFC0CFB4D" authority="(Mengla County, Xishuangbanna Dai Autonomous Prefecture)" baseAuthorityName="Mengla County, Xishuangbanna Dai Autonomous Prefecture" box="[151,997,1181,1205]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="bannaensis">
|
||
<emphasis id="B920EA8B4F0EFF94C7B8FB6EFF4EFB4D" box="[151,167,1182,1205]" italics="true" pageId="2" pageNumber="447">P</emphasis>
|
||
.
|
||
<emphasis id="B920EA8B4F0EFF94C797FB6DFED9FB4D" box="[184,304,1181,1205]" italics="true" pageId="2" pageNumber="447">bannaensis</emphasis>
|
||
(Mengla County, Xishuangbanna Dai Autonomous Prefecture)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C544D1A4F0EFF94C30FFB6EFB4FFB4D" box="[1056,1190,1181,1205]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="puerensis">
|
||
<emphasis id="B920EA8B4F0EFF94C30FFB6EFBD9FB4D" box="[1056,1072,1182,1205]" italics="true" pageId="2" pageNumber="447">P</emphasis>
|
||
.
|
||
<emphasis id="B920EA8B4F0EFF94C36EFB6EFB4FFB4D" box="[1089,1190,1182,1205]" italics="true" pageId="2" pageNumber="447">puerensis</emphasis>
|
||
</taxonomicName>
|
||
(Ning’er Hani and Yi Autonomous County), based only on molecular data. Their descriptions (and proposed names) are apparently unavailable as
|
||
<bibRefCitation id="EFC54B684F0EFF94C79BFB15FEA0FB05" author="Kaplin, V. G." box="[180,329,1253,1277]" pageId="2" pageNumber="447" pagination="17 - 22" refId="ref2967" refString="Kaplin, V. G. (2020) A new species of bristletails of the genus Petrobiellus (Microcoryphia: Machilidae) from Sakhalin. Zoosystematica Rossica, 29 (1), 17 - 22. https: // doi. org / 10.31610 / zsr / 2020.29.1.17" type="journal article" year="2020">Kaplin (2020)</bibRefCitation>
|
||
mentioned, for the following reasons: 1) no morphological descriptions were included; 2) no
|
||
<typeStatus id="54EF883B4F0EFF94C205FB15FA8BFB05" box="[1322,1378,1253,1277]" pageId="2" pageNumber="447">types</typeStatus>
|
||
were designated; 3) the descriptions were made without designating
|
||
<typeStatus id="54EF883B4F0EFF94C411FAF9FC90FAD9" box="[830,889,1289,1313]" pageId="2" pageNumber="447">types</typeStatus>
|
||
, and 4) the concerned species were not declared to be new species [cf. International Commission of
|
||
<bibRefCitation id="EFC54B684F0EFF94C58DFADDFBEDFABD" author="International Commission on Zoological Nomenclature" box="[674,1028,1325,1349]" pageId="2" pageNumber="447" refId="ref2790" refString="International Commission on Zoological Nomenclature (2000) Kokusai Dobutsu Meimei Kiyaku Dai 4 Han Nihongoyaku [International Code of Zoological Nomenclature Fourth Edition Japanese Translation]. Union of Japanese Societies for Systematic Zoology, Sapporo, XVIII + 133 pp. [Noda, H. & Nishikawa, T. (Eds.)]" type="book" year="2000">Zoological Nomenclature (2000)</bibRefCitation>
|
||
, Arts. 13.1.1., 16.1., 16.4.].
|
||
</paragraph>
|
||
<paragraph id="8BEB36994F0EFF94C7E8FAA1FD9BF955" blockId="2.[151,1437,711,1709]" pageId="2" pageNumber="447">
|
||
<taxonomicName id="4C544D1A4F0EFF94C7E8FAA1FEA1FA91" authorityName="Silvestri" authorityYear="1943" box="[199,328,1361,1385]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B920EA8B4F0EFF94C7E8FAA1FEA1FA91" box="[199,328,1361,1385]" italics="true" pageId="2" pageNumber="447">Petrobiellus</emphasis>
|
||
</taxonomicName>
|
||
was originally described by
|
||
<bibRefCitation id="EFC54B684F0EFF94C551FAA1FCC1FA91" author="Silvestri, F." box="[638,808,1361,1385]" pageId="2" pageNumber="447" pagination="283 - 306" refId="ref3591" refString="Silvestri, F. (1943) Contributo alla conoscenza dei Machilidae (Insecta, Thysanura) del Giappone. Bollettino del Laboratorio di Zoologia, Generale e Agraria della Facolta Agraria in Portici, 32, 283 - 306." type="journal article" year="1943">Silvestri (1943)</bibRefCitation>
|
||
, based only on females. Later,
|
||
<bibRefCitation id="EFC54B684F0EFF94C35AFAA1FAF8FA91" author="Uchida, H." box="[1141,1297,1361,1385]" pageId="2" pageNumber="447" pagination="1 - 17" refId="ref3856" refString="Uchida, H. (1954) Apterygota of the Hachijo-Jima and its adjacent islands. Science Reports of the Faculty of Literature and Science, Hirosaki University, 1 (1), 1 - 17." type="journal article" year="1954">Uchida (1954)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C544D1A4F0EFF94C2AAFAA2FEECFA75" authorityName="Uchida" authorityYear="1954" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="curvistylis">
|
||
<emphasis id="B920EA8B4F0EFF94C2AAFAA2FA7CFA91" box="[1413,1429,1362,1385]" italics="true" pageId="2" pageNumber="447">P</emphasis>
|
||
.
|
||
<emphasis id="B920EA8B4F0EFF94C7B8FA85FEECFA75" box="[151,261,1397,1421]" italics="true" pageId="2" pageNumber="447">curvistylis</emphasis>
|
||
</taxonomicName>
|
||
, including information on males, and the junior author (RM) succeeded in collecting the males of
|
||
<taxonomicName id="4C544D1A4F0EFF94C225FA86FA7CFA75" authorityName="Silvestri" authorityYear="1943" box="[1290,1429,1397,1421]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="takunagae">
|
||
<emphasis id="B920EA8B4F0EFF94C225FA86FAF3FA75" box="[1290,1306,1398,1421]" italics="true" pageId="2" pageNumber="447">P</emphasis>
|
||
.
|
||
<emphasis id="B920EA8B4F0EFF94C209FA85FA7CFA75" box="[1318,1429,1397,1421]" italics="true" pageId="2" pageNumber="447">takunagae</emphasis>
|
||
</taxonomicName>
|
||
, which is the
|
||
<typeStatus id="54EF883B4F0EFF94C60FFA69FEA7FA49" box="[288,334,1433,1457]" pageId="2" pageNumber="447">type</typeStatus>
|
||
species of
|
||
<taxonomicName id="4C544D1A4F0EFF94C6E8FA69FDA1FA49" authorityName="Silvestri" authorityYear="1943" box="[455,584,1433,1457]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B920EA8B4F0EFF94C6E8FA69FDA1FA49" box="[455,584,1433,1457]" italics="true" pageId="2" pageNumber="447">Petrobiellus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFC54B684F0EFF94C578FA69FCB2FA49" author="Sturm, H. & Machida, R." box="[599,859,1433,1457]" pageId="2" pageNumber="447" refId="ref3632" refString="Sturm, H. & Machida, R. (2001) Archaeognatha. In: Kristensen, N. P. & Beutel, R. G. (Eds.), Handbook of Zoology, Volume IV Arthropoda: Insecta Part 37. Walter de Gruyter, Berlin, New York, 213 pp." type="book" year="2001">Sturm & Machida 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC54B684F0EFF94C449FA69FBEAFA49" author="Machida, R." box="[870,1027,1433,1457]" pageId="2" pageNumber="447" pagination="4 - 6" refId="ref3322" refString="Machida, R. (2008) Archaeognatha. In: Hirashima, Y. & Morimoto, K. (Eds.), Iconographia Insectorum Japonicorum Colore Naturali Edita, Volumen III (New Edition). Hokuryukan, Tokyo, pp. 4 - 6, pl. 1. [in Japanese]" type="journal article" year="2008">Machida 2008</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC54B684F0EFF94C321FA69FAC5FA49" author="Klass, K. - D. & Matushkina, N. A." box="[1038,1324,1433,1457]" pageId="2" pageNumber="447" pagination="235 - 294" refId="ref3017" refString="Klass, K. - D. & Matushkina, N. A. (2018) The exoskeleton of the male genitalic region in Archaeognatha, with hypotheses on the early evolution and the morphological interpretation of genitalia in insects. Arthropod Systematics & Phylogeny, 76 (2), 235 - 294. https: // doi. org / 10.3897 / asp. 76. e 31928" type="journal article" year="2018">Klass & Matushkina 2018</bibRefCitation>
|
||
):
|
||
<bibRefCitation id="EFC54B684F0EFF94C210FA69FF37FA2D" author="Machida, R." pageId="2" pageNumber="447" pagination="76 - 79" refId="ref3430" refString="Machida, R. (2020) Order Archaeognatha. In: The editional committee of catalogue of the insects of Japan (Ed.), Catalogue of the Insects of Japan, Volume 1 Apterygote Orders. Touka Shobo, Fukuoka, pp. 76 - 79. [in Japanese]" type="book chapter" year="2020">Machida (2020)</bibRefCitation>
|
||
had suggested
|
||
<taxonomicName id="4C544D1A4F0EFF94C6A7FA4EFDFCFA2D" authorityName="Uchida" authorityYear="1954" box="[392,533,1469,1493]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="curvistylis">
|
||
<emphasis id="B920EA8B4F0EFF94C6A7FA4EFE71FA2D" box="[392,408,1470,1493]" italics="true" pageId="2" pageNumber="447">P</emphasis>
|
||
.
|
||
<emphasis id="B920EA8B4F0EFF94C688FA4DFDFCFA2D" box="[423,533,1469,1493]" italics="true" pageId="2" pageNumber="447">curvistylis</emphasis>
|
||
</taxonomicName>
|
||
could possibly be a synonym of
|
||
<taxonomicName id="4C544D1A4F0EFF94C4ADFA4EFBF8FA2D" authorityName="Silvestri" authorityYear="1943" box="[898,1041,1469,1493]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="takunagae">
|
||
<emphasis id="B920EA8B4F0EFF94C4ADFA4EFC7BFA2D" box="[898,914,1470,1493]" italics="true" pageId="2" pageNumber="447">P</emphasis>
|
||
.
|
||
<emphasis id="B920EA8B4F0EFF94C48DFA4DFBF8FA2D" box="[930,1041,1469,1493]" italics="true" pageId="2" pageNumber="447">takunagae</emphasis>
|
||
</taxonomicName>
|
||
. As for
|
||
<taxonomicName id="4C544D1A4F0EFF94C343FA4EFAEDFA2D" authorityName="Uchida" authorityYear="1949" box="[1132,1284,1469,1493]" class="Insecta" family="Machilidae" genus="Halomachilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="akkesiensis">
|
||
<emphasis id="B920EA8B4F0EFF94C343FA4EFB95FA2D" box="[1132,1148,1470,1493]" italics="true" pageId="2" pageNumber="447">P</emphasis>
|
||
.
|
||
<emphasis id="B920EA8B4F0EFF94C3A3FA4DFAEDFA2D" box="[1164,1284,1469,1493]" italics="true" pageId="2" pageNumber="447">akkesiensis</emphasis>
|
||
</taxonomicName>
|
||
, no male was identified (
|
||
<bibRefCitation id="EFC54B684F0EFF94C624FA11FE70FA01" author="Uchida, H." box="[267,409,1505,1529]" pageId="2" pageNumber="447" pagination="2 - 6" refId="ref3783" refString="Uchida, H. (1950) Thysanura. In: Ishii, T., Uchida, S., Esaki, T., Kawamura, T., Kinoshita, S., Kuwayama, S., Shiraki, T. & Yuasa, H. (Eds.), Iconographia Insectorum Japonicorum. Hokuryukan, Tokyo, pp. 2 - 6. [in Japanese]" type="book chapter" year="1950">Uchida 1950</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFC54B684F0EFF94C68BFA11FE35FA01" author="Uchida, H." box="[420,476,1505,1529]" pageId="2" pageNumber="447" pagination="1 - 2" refId="ref3978" refString="Uchida, H. (1965) Thysanura. In: Asahina, S., Ishihara, T. & Yasumatsu, K. (Eds.), Iconographia Insectorum Japonicorum Colore Naturali Edita, Volumen III. Hokuryukan, Tokyo, pp. 1 - 2, pl. 1. [in Japanese]" type="journal article" year="1965">1965</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC54B684F0EFF94C6C7FA11FD6CFA01" author="Machida, R." box="[488,645,1505,1529]" pageId="2" pageNumber="447" pagination="4 - 6" refId="ref3322" refString="Machida, R. (2008) Archaeognatha. In: Hirashima, Y. & Morimoto, K. (Eds.), Iconographia Insectorum Japonicorum Colore Naturali Edita, Volumen III (New Edition). Hokuryukan, Tokyo, pp. 4 - 6, pl. 1. [in Japanese]" type="journal article" year="2008">Machida 2008</bibRefCitation>
|
||
), but the senior author (SM) recently successfully collected and identified males (
|
||
<figureCitation id="136F2A1C4F0EFF94C7CBF9F5FEB3F9E5" box="[228,346,1541,1565]" captionStart="FIGURE 2" captionStartId="3.[151,250,1765,1789]" captionTargetBox="[154,1433,181,1741]" captionTargetId="figure-17@3.[154,1433,181,1741]" captionTargetPageId="3" captionText="FIGURE 2. External features of Petrobiellus akkesiensis comb. nov. (A) Male habitus. (B) Female habitus. (C–H) Female morphology. (C) Right antenna, anterior (medial) view. (D) Left maxilla, lateral view. (E) Labium, posterior view. (F) Left foreleg, posterior (lateral) view. (G) Left middle leg, posterior view. (H) Left hind leg, posterior (medial) view. (I) Male genitalic region, ventral view. (J) Post-abdomen of female, ventrolateral view. Abbreviations: Cx9: coxite IX; Ov: ovipositor; Pa: paramera; Pe: penis; St9: stylus IX. Scale bars: A, B, 5 mm; C–H, J, 1 mm; I, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14386205" httpUri="https://zenodo.org/record/14386205/files/figure.png" pageId="2" pageNumber="447">Figs. 2A, I</figureCitation>
|
||
). The males of
|
||
<taxonomicName id="4C544D1A4F0EFF94C6D0F9F6FD7CF9E5" authorityName="Uchida" authorityYear="1949" box="[511,661,1541,1565]" class="Insecta" family="Machilidae" genus="Halomachilis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="akkesiensis">
|
||
<emphasis id="B920EA8B4F0EFF94C6D0F9F6FDE6F9E5" box="[511,527,1542,1565]" italics="true" pageId="2" pageNumber="447">P</emphasis>
|
||
.
|
||
<emphasis id="B920EA8B4F0EFF94C532F9F5FD7CF9E5" box="[541,661,1541,1565]" italics="true" pageId="2" pageNumber="447">akkesiensis</emphasis>
|
||
</taxonomicName>
|
||
share several remarkably similar features of the genitalia with the males of
|
||
<taxonomicName id="4C544D1A4F0EFF94C79AF9DAFEA8F9B9" authorityName="Uchida" authorityYear="1954" box="[181,321,1577,1601]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="curvistylis">
|
||
<emphasis id="B920EA8B4F0EFF94C79AF9DAFF2CF9B9" box="[181,197,1578,1601]" italics="true" pageId="2" pageNumber="447">P</emphasis>
|
||
.
|
||
<emphasis id="B920EA8B4F0EFF94C7FCF9D9FEA8F9B9" box="[211,321,1577,1601]" italics="true" pageId="2" pageNumber="447">curvistylis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C544D1A4F0EFF94C659F9DAFDEDF9B9" authorityName="Silvestri" authorityYear="1943" box="[374,516,1577,1601]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="takunagae">
|
||
<emphasis id="B920EA8B4F0EFF94C659F9DAFE6FF9B9" box="[374,390,1578,1601]" italics="true" pageId="2" pageNumber="447">P</emphasis>
|
||
.
|
||
<emphasis id="B920EA8B4F0EFF94C6BAF9D9FDEDF9B9" box="[405,516,1577,1601]" italics="true" pageId="2" pageNumber="447">takunagae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFC54B684F0EFF94C53DF9D9FD4BF9B9" author="Uchida, H." box="[530,674,1577,1601]" pageId="2" pageNumber="447" pagination="1 - 17" refId="ref3856" refString="Uchida, H. (1954) Apterygota of the Hachijo-Jima and its adjacent islands. Science Reports of the Faculty of Literature and Science, Hirosaki University, 1 (1), 1 - 17." type="journal article" year="1954">Uchida 1954</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC54B684F0EFF94C582F9D9FC46F9B9" author="Sturm, H. & Machida, R." box="[685,943,1577,1601]" pageId="2" pageNumber="447" refId="ref3632" refString="Sturm, H. & Machida, R. (2001) Archaeognatha. In: Kristensen, N. P. & Beutel, R. G. (Eds.), Handbook of Zoology, Volume IV Arthropoda: Insecta Part 37. Walter de Gruyter, Berlin, New York, 213 pp." type="book" year="2001">Sturm & Machida 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFC54B684F0EFF94C495F9D9FB30F9B9" author="Klass, K. - D. & Matushkina, N. A." box="[954,1241,1577,1601]" pageId="2" pageNumber="447" pagination="235 - 294" refId="ref3017" refString="Klass, K. - D. & Matushkina, N. A. (2018) The exoskeleton of the male genitalic region in Archaeognatha, with hypotheses on the early evolution and the morphological interpretation of genitalia in insects. Arthropod Systematics & Phylogeny, 76 (2), 235 - 294. https: // doi. org / 10.3897 / asp. 76. e 31928" type="journal article" year="2018">Klass & Matushkina 2018</bibRefCitation>
|
||
): 1) the distal part of the penis is sclerotized and hook-shaped, 2) coxites IX are medially strongly concave, 3) styli IX are robust with the concaved medial side, on which dark strong setae are densely distributed, and 4) the paramerae, which are present only in abdominal segment IX, are slender (
|
||
<figureCitation id="136F2A1C4F0EFF94C535F965FD8CF955" box="[538,613,1685,1709]" captionStart="FIGURE 2" captionStartId="3.[151,250,1765,1789]" captionTargetBox="[154,1433,181,1741]" captionTargetId="figure-17@3.[154,1433,181,1741]" captionTargetPageId="3" captionText="FIGURE 2. External features of Petrobiellus akkesiensis comb. nov. (A) Male habitus. (B) Female habitus. (C–H) Female morphology. (C) Right antenna, anterior (medial) view. (D) Left maxilla, lateral view. (E) Labium, posterior view. (F) Left foreleg, posterior (lateral) view. (G) Left middle leg, posterior view. (H) Left hind leg, posterior (medial) view. (I) Male genitalic region, ventral view. (J) Post-abdomen of female, ventrolateral view. Abbreviations: Cx9: coxite IX; Ov: ovipositor; Pa: paramera; Pe: penis; St9: stylus IX. Scale bars: A, B, 5 mm; C–H, J, 1 mm; I, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.14386205" httpUri="https://zenodo.org/record/14386205/files/figure.png" pageId="2" pageNumber="447">Fig. 2I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E65124F0EFF92C7B8F90FFC0EFC8C" lastPageId="4" lastPageNumber="449" pageId="2" pageNumber="447" type="biology_ecology">
|
||
<paragraph id="8BEB36994F0EFF94C7B8F90FFEB2F8E1" blockId="2.[151,1437,1791,2033]" box="[151,347,1791,1817]" pageId="2" pageNumber="447">
|
||
<heading id="D0A381F54F0EFF94C7B8F90FFEB2F8E1" bold="true" box="[151,347,1791,1817]" fontSize="11" level="1" pageId="2" pageNumber="447" reason="1">
|
||
<emphasis id="B920EA8B4F0EFF94C7B8F90FFEB2F8E1" bold="true" box="[151,347,1791,1817]" pageId="2" pageNumber="447">Biological notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BEB36994F0EFF94C7B8F8D5FE74F809" blockId="2.[151,1437,1791,2033]" pageId="2" pageNumber="447">
|
||
<taxonomicName id="4C544D1A4F0EFF94C7B8F8D5FE70F8C5" authorityName="Mtow & Machida" authorityYear="2024" baseAuthorityName="Uchida" baseAuthorityYear="1949" box="[151,409,1829,1853]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="akkesiensis">
|
||
<emphasis id="B920EA8B4F0EFF94C7B8F8D5FE70F8C5" box="[151,409,1829,1853]" italics="true" pageId="2" pageNumber="447">Petrobiellus akkesiensis</emphasis>
|
||
</taxonomicName>
|
||
was collected by us on
|
||
<date id="FFEA10594F0EFF94C5B0F8D5FC84F8C5" box="[671,877,1829,1853]" pageId="2" pageNumber="447" value="1990-09-01">September 1, 1990</date>
|
||
, and
|
||
<date id="FFEA10594F0EFF94C488F8D5FBA6F8C5" box="[935,1103,1829,1853]" pageId="2" pageNumber="447" value="2022-06-06" valueMax="2022-06-07" valueMin="2022-06-06">June 6–7, 2022</date>
|
||
, from the supralittoral cliffs or from under rolling stones along the rocky coasts, around the Akkeshi Marine Station, Field Science Centre for Northern Biosphere,
|
||
<collectingRegion id="4990F87B4F0EFF94C63BF89DFE94F87D" box="[276,381,1901,1925]" country="Japan" name="Hokkaido" pageId="2" pageNumber="447">Hokkaido</collectingRegion>
|
||
University, Aikappu, Akkeshi Town, Akkeshi County, Kushiro Subprefecture,
|
||
<collectingRegion id="4990F87B4F0EFF94C3C9F89DFABBF87D" box="[1254,1362,1901,1925]" country="Japan" name="Hokkaido" pageId="2" pageNumber="447">Hokkaido</collectingRegion>
|
||
,
|
||
<collectingCountry id="F34376094F0EFF94C270F89DFA75F87C" box="[1375,1436,1901,1924]" name="Japan" pageId="2" pageNumber="447">Japan</collectingCountry>
|
||
(
|
||
<figureCitation id="136F2A1C4F0EFF94C7B0F861FEC9F851" box="[159,288,1937,1961]" captionStart="FIGURE 1" captionStartId="1.[151,250,1346,1370]" captionTargetBox="[292,1296,487,1322]" captionTargetId="figure-168@1.[292,1296,487,1322]" captionTargetPageId="1" captionText="FIGURE 1. Collection locality of Petrobiellus akkesiensis comb. nov. (A–C) Maps of the collection sites. (B) and (C) were constructed using the GSI Map Vector (https://maps.gsi.go.jp/vector/). Dotted line in (C) indicates submerged island, see text. (D) Collection sites around the Akkeshi Marine Station, Akkeshi Town, Hokkaido, Japan." figureDoi="http://doi.org/10.5281/zenodo.14386201" httpUri="https://zenodo.org/record/14386201/files/figure.png" pageId="2" pageNumber="447">Figs. 1A–D</figureCitation>
|
||
).
|
||
<bibRefCitation id="EFC54B684F0EFF94C61EF861FE25F851" author="Uchida, H." box="[305,460,1937,1961]" pageId="2" pageNumber="447" pagination="1 - 2" refId="ref3978" refString="Uchida, H. (1965) Thysanura. In: Asahina, S., Ishihara, T. & Yasumatsu, K. (Eds.), Iconographia Insectorum Japonicorum Colore Naturali Edita, Volumen III. Hokuryukan, Tokyo, pp. 1 - 2, pl. 1. [in Japanese]" type="journal article" year="1965">Uchida (1965)</bibRefCitation>
|
||
had previously reported that
|
||
<taxonomicName id="4C544D1A4F0EFF94C42EF862FC7FF851" authorityName="Mtow & Machida" authorityYear="2024" baseAuthorityName="Uchida" baseAuthorityYear="1949" box="[769,918,1937,1961]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="2" pageNumber="447" phylum="Arthropoda" rank="species" species="akkesiensis">
|
||
<emphasis id="B920EA8B4F0EFF94C42EF862FCF8F851" box="[769,785,1938,1961]" italics="true" pageId="2" pageNumber="447">P</emphasis>
|
||
.
|
||
<emphasis id="B920EA8B4F0EFF94C431F861FC7FF851" box="[798,918,1937,1961]" italics="true" pageId="2" pageNumber="447">akkesiensis</emphasis>
|
||
</taxonomicName>
|
||
also inhabited the Kaki Islands in Lake Akkeshi, but the Kaki Islands now no longer exist due to subsidence (the approximate area where the islands were is depicted with a dotted line in
|
||
<figureCitation id="136F2A1C4F0EFF94C612F829FE79F809" box="[317,400,2009,2033]" captionStart="FIGURE 1" captionStartId="1.[151,250,1346,1370]" captionTargetBox="[292,1296,487,1322]" captionTargetId="figure-168@1.[292,1296,487,1322]" captionTargetPageId="1" captionText="FIGURE 1. Collection locality of Petrobiellus akkesiensis comb. nov. (A–C) Maps of the collection sites. (B) and (C) were constructed using the GSI Map Vector (https://maps.gsi.go.jp/vector/). Dotted line in (C) indicates submerged island, see text. (D) Collection sites around the Akkeshi Marine Station, Akkeshi Town, Hokkaido, Japan." figureDoi="http://doi.org/10.5281/zenodo.14386201" httpUri="https://zenodo.org/record/14386201/files/figure.png" pageId="2" pageNumber="447">Fig. 1C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF2B66114F0FFF95C7B8F915FBCEF875" ID-DOI="http://doi.org/10.5281/zenodo.14386205" ID-Zenodo-Dep="14386205" httpUri="https://zenodo.org/record/14386205/files/figure.png" pageId="3" pageNumber="448" startId="3.[151,250,1765,1789]" targetBox="[154,1433,181,1741]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BEB36994F0FFF95C7B8F915FBCEF875" blockId="3.[151,1437,1765,1933]" pageId="3" pageNumber="448">
|
||
<emphasis id="B920EA8B4F0FFF95C7B8F915FEFCF905" bold="true" box="[151,277,1765,1789]" pageId="3" pageNumber="448">FIGURE 2.</emphasis>
|
||
External features of
|
||
<taxonomicName id="4C544D1A4F0FFF95C6DBF915FD00F905" authorityName="Mtow & Machida" authorityYear="2024" baseAuthorityName="Uchida" baseAuthorityYear="1949" box="[500,745,1765,1789]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="3" pageNumber="448" phylum="Arthropoda" rank="species" species="akkesiensis" status="comb. nov.">
|
||
<emphasis id="B920EA8B4F0FFF95C6DBF915FD00F905" box="[500,745,1765,1789]" italics="true" pageId="3" pageNumber="448">Petrobiellus akkesiensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B920EA8B4F0FFF95C5DDF915FC81F905" bold="true" box="[754,872,1765,1789]" pageId="3" pageNumber="448">
|
||
<taxonomicNameLabel id="A21357F04F0FFF95C5DDF915FC81F905" box="[754,872,1765,1789]" pageId="3" pageNumber="448" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A) Male habitus. (B) Female habitus. (C–H) Female morphology. (C) Right antenna, anterior (medial) view. (D) Left maxilla, lateral view. (E) Labium, posterior view. (F) Left foreleg, posterior (lateral) view. (G) Left middle leg, posterior view. (H) Left hind leg, posterior (medial) view. (I) Male genitalic region, ventral view. (J) Post-abdomen of female, ventrolateral view. Abbreviations: Cx9: coxite IX; Ov: ovipositor; Pa: paramera; Pe: penis; St9: stylus IX. Scale bars: A, B, 5 mm; C–H, J, 1 mm; I, 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF2B66114F08FF92C7B8FD90FF08FD64" ID-DOI="http://doi.org/10.5281/zenodo.14386207" ID-Zenodo-Dep="14386207" httpUri="https://zenodo.org/record/14386207/files/figure.png" pageId="4" pageNumber="449" startId="4.[151,250,608,632]" targetBox="[302,1286,181,584]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BEB36994F08FF92C7B8FD90FF08FD64" blockId="4.[151,1437,608,668]" pageId="4" pageNumber="449">
|
||
<emphasis id="B920EA8B4F08FF92C7B8FD90FEFFFD80" bold="true" box="[151,278,608,632]" pageId="4" pageNumber="449">FIGURE 3.</emphasis>
|
||
Field observations of
|
||
<taxonomicName id="4C544D1A4F08FF92C529FD90FD15FD80" authorityName="Mtow & Machida" authorityYear="2024" baseAuthorityName="Uchida" baseAuthorityYear="1949" box="[518,764,608,632]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="4" pageNumber="449" phylum="Arthropoda" rank="species" species="akkesiensis" status="comb. nov.">
|
||
<emphasis id="B920EA8B4F08FF92C529FD90FD15FD80" box="[518,764,608,632]" italics="true" pageId="4" pageNumber="449">Petrobiellus akkesiensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B920EA8B4F08FF92C429FD90FC94FD80" bold="true" box="[774,893,608,632]" pageId="4" pageNumber="449">
|
||
<taxonomicNameLabel id="A21357F04F08FF92C429FD90FC94FD80" box="[774,893,608,632]" pageId="4" pageNumber="449" rank="species">comb. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A) Adults gathered on a rock. (B) Exuviae left in cluster.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEB36994F08FF92C7E8FD00FC0EFC8C" blockId="4.[151,1437,752,884]" pageId="4" pageNumber="449">
|
||
<taxonomicName id="4C544D1A4F08FF92C7E8FD00FE27FCF0" authorityName="Mtow & Machida" authorityYear="2024" baseAuthorityName="Uchida" baseAuthorityYear="1949" box="[199,462,752,776]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="4" pageNumber="449" phylum="Arthropoda" rank="species" species="akkesiensis">
|
||
<emphasis id="B920EA8B4F08FF92C7E8FD00FE27FCF0" box="[199,462,752,776]" italics="true" pageId="4" pageNumber="449">Petrobiellus akkesiensis</emphasis>
|
||
</taxonomicName>
|
||
individuals live close to each other in high density (
|
||
<figureCitation id="136F2A1C4F08FF92C36EFD00FB74FCF0" box="[1089,1181,752,776]" captionStart="FIGURE 3" captionStartId="4.[151,250,608,632]" captionTargetBox="[302,1286,181,584]" captionTargetId="figure-21@4.[302,1286,181,584]" captionTargetPageId="4" captionText="FIGURE 3. Field observations of Petrobiellus akkesiensis comb. nov. (A) Adults gathered on a rock. (B) Exuviae left in cluster." figureDoi="http://doi.org/10.5281/zenodo.14386207" httpUri="https://zenodo.org/record/14386207/files/figure.png" pageId="4" pageNumber="449">Fig. 3A</figureCitation>
|
||
), similar to halophilic archaeognathan
|
||
<taxonomicName id="4C544D1A4F08FF92C66AFCE4FE36FCD4" box="[325,479,788,812]" class="Insecta" family="Machilidae" genus="Petrobius" kingdom="Animalia" order="Archaeognatha" pageId="4" pageNumber="449" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B920EA8B4F08FF92C66AFCE4FE45FCD4" box="[325,428,788,812]" italics="true" pageId="4" pageNumber="449">Petrobius</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EFC54B684F08FF92C6C3FCE4FD9DFCD4" author="Larink, O." box="[492,628,788,812]" pageId="4" pageNumber="449" pagination="124 - 129" refId="ref3088" refString="Larink, O. (1968) Zur Biologie des kustenbewohnenden Machiliden Petrobius brevistylis (Thysanura, Insecta). Helgolander Wissenschaftliche Meeresuntersuchungen, 18 (1 - 2), 124 - 129. https: // doi. org / 10.1007 / BF 01611670" type="journal article" year="1968">Larink 1968</bibRefCitation>
|
||
). The sex ratio of
|
||
<taxonomicName id="4C544D1A4F08FF92C41BFCE5FC23FCD4" authorityName="Mtow & Machida" authorityYear="2024" baseAuthorityName="Uchida" baseAuthorityYear="1949" box="[820,970,788,812]" class="Insecta" family="Machilidae" genus="Petrobiellus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Archaeognatha" pageId="4" pageNumber="449" phylum="Arthropoda" rank="species" species="akkesiensis">
|
||
<emphasis id="B920EA8B4F08FF92C41BFCE5FCADFCD4" box="[820,836,789,812]" italics="true" pageId="4" pageNumber="449">P</emphasis>
|
||
.
|
||
<emphasis id="B920EA8B4F08FF92C47DFCE4FC23FCD4" box="[850,970,788,812]" italics="true" pageId="4" pageNumber="449">akkesiensis</emphasis>
|
||
</taxonomicName>
|
||
seems strongly biased towards females. On
|
||
<date id="FFEA10594F08FF92C7B8FCC9FEF6FCA8" box="[151,287,824,848]" pageId="4" pageNumber="449" value="2022-06-06">June 6, 2022</date>
|
||
, 75 individuals were collected, among which seven were males and 68 were females; and among 122 exuviae found in a crevice of the cliff eight were males and 114 were females (
|
||
<figureCitation id="136F2A1C4F08FF92C4A8FCACFC33FC8C" box="[903,986,860,884]" captionStart="FIGURE 3" captionStartId="4.[151,250,608,632]" captionTargetBox="[302,1286,181,584]" captionTargetId="figure-21@4.[302,1286,181,584]" captionTargetPageId="4" captionText="FIGURE 3. Field observations of Petrobiellus akkesiensis comb. nov. (A) Adults gathered on a rock. (B) Exuviae left in cluster." figureDoi="http://doi.org/10.5281/zenodo.14386207" httpUri="https://zenodo.org/record/14386207/files/figure.png" pageId="4" pageNumber="449">Fig. 3B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |