215 lines
26 KiB
XML
215 lines
26 KiB
XML
<document id="AFD3E84620AC455C92641FFC698001EF" ID-DOI="10.11646/zootaxa.5367.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10145106" ID-ZooBank="5A1E5321-D2BA-4B92-BA23-A7C1CDBA5723" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700167363318" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2023" docId="03FD878BFFD9FFFAFCBEFBBE58983DE6" docLanguage="en" docName="zootaxa.5367.1.1.pdf" docOrigin="Zootaxa 5367 (1)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5367.1.1/52225" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydraena (Hydraenopsis) belilymba Perkins 2023, new species" docType="treatment" docVersion="2" lastPageNumber="12" masterDocId="FFC4FFF3FFD0FFF1FC29FFA5596F3C37" masterDocTitle="New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)" masterLastPageNumber="86" masterPageNumber="1" pageNumber="10" updateTime="1700168056660" updateUser="ExternalLinkService">
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<mods:title id="77C18C71243CB693AF3E733D843005E6">New Neotropical and Nearctic species of water beetles in the genera Hydraena Kugelann and Ochthebius Leach, a key to North American genera and subgenera of the family, new distribution records, and a synopsis of ecology, behavior and morphology related to aquatic life (Coleoptera: Hydraenidae)</mods:title>
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<mods:namePart id="8F0921F53574F6C162D42AEC4878AC14">Perkins, Philip D.</mods:namePart>
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<mods:title id="B949EAE01F70F02C7942208DA8EECE37">Zootaxa</mods:title>
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<mods:part id="8D2AA14BBFB5854C0834095822E4B8E6">
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<mods:date id="8ACA7CF269E3A0100B189CFC7C6BCAE1">2023</mods:date>
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<mods:number id="8D939C0FBD58270F66622A5DB737CF1F">2023-11-07</mods:number>
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<mods:number id="8486137C3E7762EDF621C9D65667E0C3">5367</mods:number>
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<mods:url id="CED23D81AE3D5ED6C8DA87DE42994E81">https://www.mapress.com/zt/article/download/zootaxa.5367.1.1/52225</mods:url>
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<mods:classification id="68FEEAD1E44CFD105863D9BFB348325C">journal article</mods:classification>
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<mods:identifier id="856A017E603321815744077C87AE6E9D" type="DOI">10.11646/zootaxa.5367.1.1</mods:identifier>
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<mods:identifier id="C722D3F1C292B734010CB851B82E485F" type="ISSN">1175-5334</mods:identifier>
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</mods:mods>
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<treatment id="03FD878BFFD9FFFAFCBEFBBE58983DE6" LSID="urn:lsid:plazi:treatment:03FD878BFFD9FFFAFCBEFBBE58983DE6" httpUri="http://treatment.plazi.org/id/03FD878BFFD9FFFAFCBEFBBE58983DE6" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
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<subSubSection id="C34E6516FFD9FFF8FCBEFBBE5B833802" box="[151,748,1051,1078]" pageId="9" pageNumber="10" type="nomenclature">
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<paragraph id="8BEB369DFFD9FFF8FCBEFBBE5B833802" blockId="9.[151,764,1051,1114]" box="[151,748,1051,1078]" pageId="9" pageNumber="10">
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<heading id="D0A381F1FFD9FFF8FCBEFBBE5B833802" bold="true" box="[151,748,1051,1078]" fontSize="11" level="2" pageId="9" pageNumber="10" reason="4">
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<emphasis id="B920EA8FFFD9FFF8FCBEFBBE5B833802" bold="true" box="[151,748,1051,1078]" pageId="9" pageNumber="10">
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<taxonomicName id="4C544D1EFFD9FFF8FCBEFBBE5B223801" authority="Perkins, 2023" authorityName="Perkins" authorityYear="2023" box="[151,589,1051,1078]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="belilymba" status="sp. nov." subGenus="Hydraenopsis">
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<emphasis id="B920EA8FFFD9FFF8FCBEFBBE5B223801" bold="true" box="[151,589,1051,1078]" italics="true" pageId="9" pageNumber="10">Hydraena (Hydraenopsis) belilymba</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A21357F4FFD9FFF8FE75FBBE5B833802" box="[604,748,1051,1077]" pageId="9" pageNumber="10" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E6516FFD9FFF8FCBEFB9A5B93386E" box="[151,764,1087,1114]" pageId="9" pageNumber="10" type="description">
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<paragraph id="8BEB369DFFD9FFF8FCBEFB9A5B93386E" blockId="9.[151,764,1051,1114]" box="[151,764,1087,1114]" pageId="9" pageNumber="10">
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<figureCitation id="136F2A18FFD9FFF8FCBEFB9A598C386E" box="[151,227,1087,1113]" captionStart="FIGURE 2" captionStartId="6.[151,250,1867,1891]" captionTargetBox="[228,1360,181,1844]" captionTargetId="figure-35@6.[228,1360,181,1844]" captionTargetPageId="6" captionText="FIGURE 2. Dorsal habitus of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.10145110" httpUri="https://zenodo.org/record/10145110/files/figure.png" pageId="9" pageNumber="10">Figs. 2</figureCitation>
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(habitus comparison), 5 (habitus and aedeagus)
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E6516FFD9FFF8FCBEFB225AD338DE" pageId="9" pageNumber="10" type="materials_examined">
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||
<paragraph id="8BEB369DFFD9FFF8FCBEFB225AD338DE" blockId="9.[151,1437,1159,1942]" pageId="9" pageNumber="10">
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||
<emphasis id="B920EA8FFFD9FFF8FCBEFB22582A3896" bold="true" box="[151,325,1159,1185]" pageId="9" pageNumber="10">Type Material.</emphasis>
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<materialsCitation id="3B3C3CC0FFD9FFF8FD61FB225A8138F2" collectingDate="2006-08-11" collectingDateMax="2006-08-15" collectingDateMin="2006-08-11" collectionCode="MCZ" collectorName="P. W. Kovarik" country="Belize" latitude="16.39825" location="Union Camp" longLatPrecision="1" longitude="-89.14281" municipality="Columbia Forest Reserve" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Toledo" typeStatus="holotype">
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<emphasis id="B920EA8FFFD9FFF8FD61FB2258DA3896" bold="true" box="[328,437,1159,1185]" pageId="9" pageNumber="10">
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<typeStatus id="54EF883FFFD9FFF8FD61FB2258DA3896" box="[328,437,1159,1185]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
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</emphasis>
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(male):“
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<collectingCountry id="F343760DFFD9FFF8FE3EFB225B113896" box="[535,638,1159,1185]" name="Belize" pageId="9" pageNumber="10">BELIZE</collectingCountry>
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:
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<collectingRegion id="4990F87FFFD9FFF8FEADFB225A433896" box="[644,812,1159,1185]" country="Belize" name="Toledo" pageId="9" pageNumber="10">Toledo District</collectingRegion>
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,
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<collectingMunicipality id="6B8FACE7FFD9FFF8FF1AFB225D243895" box="[819,1099,1159,1186]" pageId="9" pageNumber="10">Columbia Forest Reserve</collectingMunicipality>
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||
,
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||
<location id="8E8B6046FFD9FFF8F87AFB225D8D3896" LSID="urn:lsid:plazi:treatment:03FD878BFFD9FFFAFCBEFBBE58983DE6:8E8B6046FFD9FFF8F87AFB225D8D3896" box="[1107,1250,1159,1185]" country="Belize" latitude="16.39825" longLatPrecision="1" longitude="-89.14281" municipality="Columbia Forest Reserve" name="Union Camp" pageId="9" pageNumber="10" stateProvince="Toledo">Union Camp</location>
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, (
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<geoCoordinate id="EE60505AFFD9FFF8F8DBFB225CF73895" box="[1266,1432,1159,1186]" degrees="16" direction="north" minutes="23" orientation="latitude" pageId="9" pageNumber="10" precision="1" seconds="53.7" value="16.39825">16°23’53.7’’N</geoCoordinate>
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,
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<geoCoordinate id="EE60505AFFD9FFF8FCBEFB0E585E38F1" box="[151,305,1195,1222]" degrees="89" direction="west" minutes="8" orientation="longitude" pageId="9" pageNumber="10" precision="1" seconds="34.1" value="-89.14281">89°8’34.1’’W</geoCoordinate>
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),
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<date id="FFEA105DFFD9FFF8FD6EFB0E589B38F2" box="[327,500,1195,1222]" pageId="9" pageNumber="10" value="2006-08-11" valueMax="2006-08-15" valueMin="2006-08-11">
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<collectingDate id="EFAEE9B5FFD9FFF8FD6EFB0E589B38F2" box="[327,500,1195,1222]" pageId="9" pageNumber="10" value="2006-08-11" valueMax="2006-08-15" valueMin="2006-08-11">11–15 viii 2006</collectingDate>
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</date>
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//
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<collectingMethod id="52154E8AFFD9FFF8FE26FB0E5B8138F1" box="[527,750,1195,1222]" pageId="9" pageNumber="10">flight intercept traps</collectingMethod>
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<collectorName id="26A1534BFFD9FFF8FEDDFB095AE138F2" box="[756,910,1195,1222]" pageId="9" pageNumber="10">P. W. Kovarik</collectorName>
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” (
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<collectionCode id="ED45AE58FFD9FFF8FF8CFB0E5A8B38F2" box="[933,996,1195,1221]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="9" pageNumber="10" type="Museum">MCZ</collectionCode>
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<emphasis id="B920EA8FFFD9FFF8FFCDFB0E5A8138F2" bold="true" box="[996,1006,1195,1221]" pageId="9" pageNumber="10">)</emphasis>
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||
</materialsCitation>
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||
<emphasis id="B920EA8FFFD9FFF8FFC7FB0E5A9B38F2" bold="true" box="[1006,1012,1195,1221]" pageId="9" pageNumber="10">.</emphasis>
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<materialsCitation id="3B3C3CC0FFD9FFF8FFD3FB0E5AD838DE" collectingDate="1981-06-30" collectingDateMax="2006-08-15" collectingDateMin="2006-08-11" collectionCode="MCZ, NMNH" collectorName="W. E. Steiner" country="Belize" latitude="16.39825" location="Same" longLatPrecision="1" longitude="-89.14281" municipality="Blue Creek" pageId="9" pageNumber="10" specimenCount="2" specimenCount-male="2" stateProvince="Toledo" typeStatus="paratype">
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||
<emphasis id="B920EA8FFFD9FFF8FFD3FB0E5DCF38F2" bold="true" box="[1018,1184,1195,1221]" pageId="9" pageNumber="10">
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||
<typeStatus id="54EF883FFFD9FFF8FFD3FB0E5D1D38F2" box="[1018,1138,1195,1221]" pageId="9" pageNumber="10" type="paratype">Paratypes</typeStatus>
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(4):
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</emphasis>
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||
<location id="8E8B6046FFD9FFF8F88FFB0E5D8B38F2" LSID="urn:lsid:plazi:treatment:03FD878BFFD9FFFAFCBEFBBE58983DE6:8E8B6046FFD9FFF8F88FFB0E5D8B38F2" box="[1190,1252,1195,1221]" municipality="Blue Creek" name="Same" pageId="9" pageNumber="10">Same</location>
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data as holotype (
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<specimenCount id="9D52FD14FFD9FFF8FC89FB6A599838DE" box="[160,247,1231,1257]" count="2" pageId="9" pageNumber="10" type="male">2 males</specimenCount>
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<collectionCode id="ED45AE58FFD9FFF8FCD7FB6A582F38DE" box="[254,320,1231,1257]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="9" pageNumber="10" type="Museum">MCZ</collectionCode>
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);
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<collectingMunicipality id="6B8FACE7FFD9FFF8FD7AFB6A58BA38DE" box="[339,469,1231,1257]" pageId="9" pageNumber="10">Blue Creek</collectingMunicipality>
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,
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<date id="FFEA105DFFD9FFF8FDF6FB6A5B3638DE" box="[479,601,1231,1257]" pageId="9" pageNumber="10" value="1981-06-30">
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<collectingDate id="EFAEE9B5FFD9FFF8FDF6FB6A5B3638DE" box="[479,601,1231,1257]" pageId="9" pageNumber="10" value="1981-06-30">30 vi 1981</collectingDate>
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</date>
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, coll.
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<collectorName id="26A1534BFFD9FFF8FEB4FB755A5E38DE" box="[669,817,1231,1258]" pageId="9" pageNumber="10">W. E. Steiner</collectorName>
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(2
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<collectionCode id="ED45AE58FFD9FFF8FF7FFB755AC338DD" box="[854,940,1232,1258]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10" type="Museum">NMNH</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C34E6516FFD9FFF8FCEEFB565B2C39F5" pageId="9" pageNumber="10" type="diagnosis">
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<paragraph id="8BEB369DFFD9FFF8FCEEFB565B2C39F5" blockId="9.[151,1437,1159,1942]" pageId="9" pageNumber="10">
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<emphasis id="B920EA8FFFD9FFF8FCEEFB5658B9393A" bold="true" box="[199,470,1267,1293]" pageId="9" pageNumber="10">Differential Diagnosis.</emphasis>
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This new species is a member of the
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<taxonomicName id="4C544D1EFFD9FFF8FFBFFB515D75393A" box="[918,1050,1267,1293]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="colymba">
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<emphasis id="B920EA8FFFD9FFF8FFBFFB515D75393A" box="[918,1050,1267,1293]" italics="true" pageId="9" pageNumber="10">H. colymba</emphasis>
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||
</taxonomicName>
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Complex of the
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<taxonomicName id="4C544D1EFFD9FFF8F8CAFB515CF3393A" box="[1251,1436,1267,1293]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="marginicollis">
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<emphasis id="B920EA8FFFD9FFF8F8CAFB515CF3393A" box="[1251,1436,1267,1293]" italics="true" pageId="9" pageNumber="10">H. marginicollis</emphasis>
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</taxonomicName>
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Group (
|
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<bibRefCitation id="EFC54B6CFFD9FFF8FCC7FAB258DD3906" author="Perkins, P. D." box="[238,434,1303,1329]" pageId="9" pageNumber="10" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins 1980: 37</bibRefCitation>
|
||
). Other members of the complex include
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||
<emphasis id="B920EA8FFFD9FFF8FFAFFABD5C263906" box="[902,1353,1303,1329]" italics="true" pageId="9" pageNumber="10">
|
||
<taxonomicName id="4C544D1EFFD9FFF8FFAFFABD5D643906" box="[902,1035,1303,1329]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="colymba">H. colymba</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C544D1EFFD9FFF8F83FFABD5DFF3906" authorityName="Perkins" authorityYear="1980" box="[1046,1168,1303,1329]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="malkini">H. malkini</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C544D1EFFD9FFF8F8B5FABD5C2A3906" authorityName="Perkins" authorityYear="1980" box="[1180,1349,1304,1329]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="nevermanni">H. nevermanni</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C544D1EFFD9FFF8F9A8FABD587A3962" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="pontequula">
|
||
<emphasis id="B920EA8FFFD9FFF8F9A8FABD587A3962" italics="true" pageId="9" pageNumber="10">H. pontequula</emphasis>
|
||
</taxonomicName>
|
||
(all
|
||
<bibRefCitation id="EFC54B6CFFD9FFF8FD63FA9E588D3962" author="Perkins, P. D." box="[330,482,1339,1365]" pageId="9" pageNumber="10" pagination="3 - 554" refId="ref49723" refString="Perkins, P. D. (1980) Aquatic beetles of the family Hydraenidae in the Western Hemisphere: classification, biogeography and inferred phylogeny (Insecta: Coleoptera). Quaestiones Entomologicae, 16, 3 - 554. [Available free from the Biodiversity Heritage Library website]" type="journal article" year="1980">Perkins 1980</bibRefCitation>
|
||
), plus
|
||
<taxonomicName id="4C544D1EFFD9FFF8FE05FA995BDD3962" authorityName="Brojer & Jach" authorityYear="2011" box="[556,690,1339,1365]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lagamba">
|
||
<emphasis id="B920EA8FFFD9FFF8FE05FA995BDD3962" box="[556,690,1339,1365]" italics="true" pageId="9" pageNumber="10">H. lagamba</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C544D1EFFD9FFF8FEC3FA995DFD3962" authority="(Brojer & Jach 2011)" baseAuthorityName="Brojer & Jach" baseAuthorityYear="2011" box="[746,1170,1339,1365]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="lascrucensis">
|
||
<emphasis id="B920EA8FFFD9FFF8FEC3FA995AF83962" box="[746,919,1339,1365]" italics="true" pageId="9" pageNumber="10">H. lascrucensis</emphasis>
|
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(
|
||
<bibRefCitation id="EFC54B6CFFD9FFF8FF8FFA9E5DE43962" author="Brojer, M. & Jach, M. A." box="[934,1163,1339,1365]" pageId="9" pageNumber="10" pagination="69 - 92" refId="ref49588" refString="Brojer, M. & Jach, M. A. (2011) Update on the hydraenid fauna of Costa Rica, with descriptions of six new species. Koleopterologische Rundschau, 81, 69 - 92." type="journal article" year="2011">Brojer & Jäch 2011</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. These species are very similar in dorsal and most ventral characters; reliable determinations require dissection of males.
|
||
<taxonomicName id="4C544D1EFFD9FFF8F8C2FAC55C12394E" authority="Perkins, 2023" authorityName="Perkins" authorityYear="2023" box="[1259,1405,1375,1401]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="belilymba" status="n. sp.">
|
||
<emphasis id="B920EA8FFFD9FFF8F8C2FAC55C12394E" box="[1259,1405,1375,1401]" italics="true" pageId="9" pageNumber="10">H. belilymba</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B920EA8FFFD9FFF8F9AFFAFA59D739AA" bold="true" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="A21357F4FFD9FFF8F9AFFAFA59D739AA" pageId="9" pageNumber="10" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be differentiated from other members of the complex by the combination of the lack of metaventral plaques, and the aedeagal morphology (
|
||
<figureCitation id="136F2A18FFD9FFF8FDD8FA025B5A39F5" box="[497,565,1447,1474]" captionStart="FIGURE 5" captionStartId="10.[151,250,1792,1816]" captionTargetBox="[313,1275,181,1768]" captionTargetId="figure-35@10.[313,1275,181,1768]" captionTargetPageId="10" captionText="FIGURE 5. Hydraena (Hydraenopsis) belilymba new species, holotype habitus and aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10145116" httpUri="https://zenodo.org/record/10145116/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E6516FFD9FFFAFCEEFA6E5A3D3D52" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10" type="description">
|
||
<paragraph id="8BEB369DFFD9FFF8FCEEFA6E5D293B32" blockId="9.[151,1437,1159,1942]" pageId="9" pageNumber="10">
|
||
<emphasis id="B920EA8FFFD9FFF8FCEEFA6E583E39D2" bold="true" box="[199,337,1483,1509]" pageId="9" pageNumber="10">Description</emphasis>
|
||
. Size:
|
||
<typeStatus id="54EF883FFFD9FFF8FD8DFA6E5B6939D2" box="[420,518,1483,1509]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
|
||
(length/width, mm): body (length to elytral apices) 1.47/0.63; head width 0.36; pronotum 0.37/0.48, PA 0.40, PB 0.43; elytra 0.90/0.63. Habitus as illustrated (
|
||
<figureCitation id="136F2A18FFD9FFF8F805FA4A5D1C3A3D" box="[1068,1139,1519,1546]" captionStart="FIGURE 5" captionStartId="10.[151,250,1792,1816]" captionTargetBox="[313,1275,181,1768]" captionTargetId="figure-35@10.[313,1275,181,1768]" captionTargetPageId="10" captionText="FIGURE 5. Hydraena (Hydraenopsis) belilymba new species, holotype habitus and aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10145116" httpUri="https://zenodo.org/record/10145116/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
|
||
). Dorsum with head dark brown to black, pronotum with distinctive dark brown macula surrounded by wide testaceous margins, elytra brown, palpi testaceous, tip of last palpomere not darker. Dorsum of head moderately finely moderately sparsely punctulate; pronotum very finely very sparsely punctulate, more sparsely punctulate on macula than on surrounding areas, interstices strongly shining; punctures of frons ca. 1xef, those of pronotal disc slightly smaller, interstices on pronotal disc ca. 2–5xpd or slightly larger; punctures of clypeus very fine, lateral areas microreticulate. Labrum apicomedially excised, free margins weakly upturned. Pronotum with anterior margin weakly emarginate over middle 1/2; anterolateral pronotal fovea moderately deep, other usual foveae absent.
|
||
</paragraph>
|
||
<paragraph id="8BEB369DFFD9FFF8FCEEF8AA5C633BA2" blockId="9.[151,1437,1159,1942]" pageId="9" pageNumber="10">Elytra with summit of posterior declivity slightly beyond midlength; lateral explanate margins narrow; on basal 1/3 punctures slightly larger than largest pronotal punctures, punctures becoming gradually smaller toward posterior. Intervals not raised, shining, width ca. 1–2xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
|
||
<caption id="DF2B6615FFDAFFFBFCBEF8A55D383B2F" ID-DOI="http://doi.org/10.5281/zenodo.10145116" ID-Zenodo-Dep="10145116" box="[151,1111,1792,1816]" httpUri="https://zenodo.org/record/10145116/files/figure.png" pageId="10" pageNumber="11" startId="10.[151,250,1792,1816]" targetBox="[313,1275,181,1768]" targetPageId="10" targetType="figure">
|
||
<paragraph id="8BEB369DFFDAFFFBFCBEF8A55D383B2F" blockId="10.[151,1111,1792,1816]" box="[151,1111,1792,1816]" pageId="10" pageNumber="11">
|
||
<emphasis id="B920EA8FFFDAFFFBFCBEF8A5587C3B2F" bold="true" box="[151,275,1792,1816]" pageId="10" pageNumber="11">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C544D1EFFDAFFFBFD30F8A55BE23B2F" authority="Perkins, 2023" authorityName="Perkins" authorityYear="2023" box="[281,653,1792,1816]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="belilymba" status="sp. nov." subGenus="Hydraenopsis">
|
||
<emphasis id="B920EA8FFFDAFFFBFD30F8A558113B2F" box="[281,382,1792,1816]" italics="true" pageId="10" pageNumber="11">Hydraena</emphasis>
|
||
(
|
||
<emphasis id="B920EA8FFFDAFFFBFDA4F8A55B753B2F" box="[397,538,1792,1816]" italics="true" pageId="10" pageNumber="11">Hydraenopsis</emphasis>
|
||
)
|
||
<emphasis id="B920EA8FFFDAFFFBFE00F8A55BE23B2F" box="[553,653,1792,1816]" italics="true" pageId="10" pageNumber="11">belilymba</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A21357F4FFDAFFFBFEBAF8A45A7F3B2F" box="[659,784,1792,1816]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
|
||
, holotype habitus and aedeagus.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BEB369DFFDBFFFAFCEEFF325A3D3D52" blockId="11.[151,1437,151,465]" pageId="11" pageNumber="12">Venter: Mentum moderately densely very finely punctulate; postmentum very finely moderately densely micropunctulate. Genae raised, shining, without posterior ridge. P1 laminate; median carina sinuate in profile. P2 moderately narrow, l/w ca. 4/1, sides converging to blunt apex, apex raised slightly above mesoventral intercoxal process. Plaques absent, metaventrite with deep median depression.AIS width at straight posterior margin ca. 3x P2. Protibiae slightly arcuate, inner margin widened inb distal 1/3; meso- and metatibiae slender, straight. Abdominal apex symmetrical; last tergite with apicomedian emargination.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E6516FFDBFFFAFCEEFECB5B9D3DBE" box="[199,754,366,393]" pageId="11" pageNumber="12" type="distribution">
|
||
<paragraph id="8BEB369DFFDBFFFAFCEEFECB5B9D3DBE" blockId="11.[151,1437,151,465]" box="[199,754,366,393]" pageId="11" pageNumber="12">
|
||
<emphasis id="B920EA8FFFDBFFFAFCEEFECB580E3DBF" bold="true" box="[199,353,366,392]" pageId="11" pageNumber="12">Distribution.</emphasis>
|
||
Currently known only from
|
||
<collectingCountry id="F343760DFFDBFFFAFE8AFECA5B813DBE" box="[675,750,367,393]" name="Belize" pageId="11" pageNumber="12">Belize</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C34E6516FFDBFFFAFCEEFE3658983DE6" pageId="11" pageNumber="12" type="etymology">
|
||
<paragraph id="8BEB369DFFDBFFFAFCEEFE3658983DE6" blockId="11.[151,1437,151,465]" pageId="11" pageNumber="12">
|
||
<emphasis id="B920EA8FFFDBFFFAFCEEFE3658213D9A" bold="true" box="[199,334,403,429]" pageId="11" pageNumber="12">Etymology.</emphasis>
|
||
Named in reference to the geographical distribution and the relationship to
|
||
<taxonomicName id="4C544D1EFFDBFFFAF8B5FE365C153D9A" authority="Perkins" authorityName="Perkins" box="[1180,1402,403,429]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="colymba">
|
||
<emphasis id="B920EA8FFFDBFFFAF8B5FE365C723D9A" box="[1180,1309,403,429]" italics="true" pageId="11" pageNumber="12">H. colymba</emphasis>
|
||
Perkins
|
||
</taxonomicName>
|
||
, as indicated by the male genitalia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |