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<document id="E8230F9784E31E694B2C74E90B96D86C" ID-DOI="10.11646/zootaxa.3941.4.1" ID-GBIF-Dataset="4a423dfe-e6dc-48fc-8149-d17562f026b3" ID-ISSN="1175-5326" ID-Zenodo-Dep="242977" ID-ZooBank="0282A923-893A-45BC-941A-6CDECBE36618" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461027129719" checkinUser="plazi" docAuthor="Hartop, Emily A., Brown, Brian V. &amp; Disney, R. Henry L." docDate="2015" docId="03FD8784FFA4FFFBC5FAFF10FA95FCCA" docLanguage="en" docName="zt03941p484.pdf" docOrigin="Zootaxa 3941 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Megaselia steptoeae Hartop, Brown &amp; Disney, 2015, new species" docType="treatment" docVersion="9" lastPageNumber="480" masterDocId="FFC4FFFCFFB9FFE6C56DFF87FFB2FFFD" masterDocTitle="Opportunity in our Ignorance: Urban Biodiversity Study Reveals 30 New Species and One New Nearctic Record for Megaselia (Diptera: Phoridae) in Los Angeles (California, USA)" masterLastPageNumber="484" masterPageNumber="451" pageNumber="480" updateTime="1698572556843" updateUser="plazi">
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<mods:title id="402CBA3E20CEE8183D7C0A0684AF0D8C">Opportunity in our Ignorance: Urban Biodiversity Study Reveals 30 New Species and One New Nearctic Record for Megaselia (Diptera: Phoridae) in Los Angeles (California, USA)</mods:title>
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<mods:namePart id="7B09550C1DF7291011AD711B0175ECE3">Hartop, Emily A.</mods:namePart>
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<mods:namePart id="1768D6A0974D3ED50B4AD84832EF0B57">Brown, Brian V.</mods:namePart>
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<mods:namePart id="FE9E4DD0C62845FC7B7EA6755A166956">Disney, R. Henry L.</mods:namePart>
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<mods:date id="C7E8232E16642096BB97C41CBBC70F8B">2015</mods:date>
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<treatment id="03FD8784FFA4FFFBC5FAFF10FA95FCCA" ID-DOI="http://doi.org/10.5281/zenodo.6093918" ID-GBIF-Taxon="119606912" ID-Zenodo-Dep="6093918" LSID="urn:lsid:plazi:treatment:03FD8784FFA4FFFBC5FAFF10FA95FCCA" httpUri="http://treatment.plazi.org/id/03FD8784FFA4FFFBC5FAFF10FA95FCCA" lastPageNumber="480" pageId="29" pageNumber="480">
<subSubSection id="C34E6519FFA4FFFBC5FAFF10FC9FFF4F" box="[151,813,151,178]" pageId="29" pageNumber="480" type="nomenclature">
<paragraph id="8BEB3692FFA4FFFBC5FAFF10FC9FFF4F" blockId="29.[151,813,151,178]" box="[151,813,151,178]" pageId="29" pageNumber="480">
<heading id="D0A381FEFFA4FFFBC5FAFF10FC9FFF4F" bold="true" box="[151,813,151,178]" fontSize="11" level="1" pageId="29" pageNumber="480" reason="1">
<emphasis id="B920EA80FFA4FFFBC5FAFF10FC9FFF4F" bold="true" box="[151,813,151,178]" pageId="29" pageNumber="480">
<taxonomicName id="4C544D11FFA4FFFBC5FAFF10FE3AFF4C" ID-CoL="8CS8F" box="[151,392,151,177]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="steptoeae" status="sp. nov.">
<emphasis id="B920EA80FFA4FFFBC5FAFF10FE3AFF4C" bold="true" box="[151,392,151,177]" italics="true" pageId="29" pageNumber="480">Megaselia steptoeae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21357FBFFA4FFFBC4FDFF1FFD92FF4F" box="[400,544,152,178]" pageId="29" pageNumber="480" rank="species">new species</taxonomicNameLabel>
(
<figureCitation id="136F2A17FFA4FFFBC742FF1FFD21FF4F" box="[559,659,152,178]" captionStart="FIGURES 28 30" captionStartId="16.[151,264,1919,1942]" captionTargetBox="[164,1415,207,1883]" captionTargetId="figure@16.[151,1436,195,1898]" captionTargetPageId="16" captionText="FIGURES 28 30. Habitus images (lateral). 28. Megaselia sidneyae. 29. Megaselia steptoeae. 30. Megaselia wiegmanae. FIGURE 31. Midfemur. Megaselia pisanoi." httpUri="https://zenodo.org/record/242980/files/figure.png" pageId="29" pageNumber="480">Figs. 29</figureCitation>
,
<figureCitation id="136F2A17FFA4FFFBC7CDFF1FFD0CFF4F" box="[672,702,152,178]" captionStart="FIGURES 53 62" captionStartId="22.[151,264,1872,1895]" captionTargetBox="[169,1416,273,1795]" captionTargetId="figure@22.[151,1436,265,1811]" captionTargetPageId="22" captionText="FIGURES 53 62. Right wing, dorsal. 53. Megaselia mikejohnsoni. 54. Megaselia oxboroughae. 55. Megaselia pisanoi. 56. Megaselia renwickorum. 57. Megaselia rodriguezorum. 58. Megaselia sacatelensis. 59. Megaselia seaverorum. 60. Megaselia sidneyae. 61. Megaselia steptoeae. 62. Megaselia wiegmanae." httpUri="https://zenodo.org/record/242983/files/figure.png" pageId="29" pageNumber="480">61</figureCitation>
,
<figureCitation id="136F2A17FFA4FFFBC7A1FF1FFD58FF4F" box="[716,746,152,178]" captionStart="FIGURES 87 92" captionStartId="28.[151,264,1880,1903]" captionTargetBox="[167,1419,255,1809]" captionTargetId="figure@28.[150,1436,241,1829]" captionTargetPageId="28" captionText="FIGURES 87 92. Male genitalia, left lateral. 87. Megaselia rodriguezorum. 88. Megaselia sacatelensis. 89. Megaselia seaverorum. 90. Megaselia sidneyae. 91. Megaselia steptoeae. 92. Megaselia wiegmanae." httpUri="https://zenodo.org/record/242988/files/figure.png" pageId="29" pageNumber="480">91</figureCitation>
,
<figureCitation id="136F2A17FFA4FFFBC79AFF1FFC96FF4F" box="[759,804,152,178]" captionStart="FIGURES 109 124" captionStartId="31.[151,264,1876,1899]" captionTargetBox="[168,1420,392,1840]" captionTargetId="figure@31.[151,1436,375,1854]" captionTargetPageId="31" captionText="FIGURES 109 124. Hypandria (ventral unless noted otherwise) 109. Megaselia isaacmajorum. 110. Megaselia kelleri. 111. Megaselia lombardorum. 112. Megaselia marquezi. 113. Megaselia mikejohnsoni. 114. Megaselia oxboroughae. 115. Megaselia pisanoi. 116. Megaselia renwickorum. 117. Megaselia rodriguezorum. 118. Megaselia sacatelensis. 119. Megaselia seaverorum. 120. Megaselia sidneyae. 121. Megaselia sidneyae (detail left process). 122. Megaselia steptoeae. 123. Megaselia wiegmanae." httpUri="https://zenodo.org/record/242990/files/figure.png" pageId="29" pageNumber="480">123</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34E6519FFA4FFFBC5FAFF59FCAFFE56" pageId="29" pageNumber="480" type="diagnosis">
<paragraph id="8BEB3692FFA4FFFBC5FAFF59FCAFFE56" blockId="29.[151,1436,222,823]" pageId="29" pageNumber="480">
<emphasis id="B920EA80FFA4FFFBC5FAFF59FEA3FF0A" bold="true" box="[151,273,222,247]" pageId="29" pageNumber="480">Diagnosis.</emphasis>
Male. In the group III key of
<bibRefCitation id="EFC54B63FFA4FFFBC732FF59FC98FF0A" author="Borgmeier" box="[607,810,222,247]" pageId="29" pageNumber="480" refString="Borgmeier, T. (1964) Revision of the North American phorid flies. Part II. The species of the genus Megaselia, subgenus Aphiochaeta. Studia Entomologica, 7, 257 - 416." type="journal article" year="1964">Borgmeier (1964)</bibRefCitation>
,
<taxonomicName id="4C544D11FFA4FFFBC658FF58FC76FF0B" box="[821,964,223,246]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="steptoeae">
<emphasis id="B920EA80FFA4FFFBC658FF58FC76FF0B" box="[821,964,223,246]" italics="true" pageId="29" pageNumber="480">M. steptoeae</emphasis>
</taxonomicName>
keys to couplet 13 where it can readily be distinguished from both
<taxonomicName id="4C544D11FFA4FFFBC4C6FE84FDC7FEE7" authority="Brues" authorityName="Brues" box="[427,629,259,283]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="scopalis">
<emphasis id="B920EA80FFA4FFFBC4C6FE84FD9EFEE6" box="[427,556,259,283]" italics="true" pageId="29" pageNumber="480">M. scopalis</emphasis>
Brues
</taxonomicName>
and
<taxonomicName id="4C544D11FFA4FFFBC7C1FE84FC2DFEE6" authority="Borgmeier." authorityName="Borgmeier." box="[684,927,258,283]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="renata">
<emphasis id="B920EA80FFA4FFFBC7C1FE84FCA9FEE7" box="[684,795,259,282]" italics="true" pageId="29" pageNumber="480">M. renata</emphasis>
Borgmeier.
</taxonomicName>
<taxonomicName id="4C544D11FFA4FFFBC6C8FE84FB87FEE7" box="[933,1077,259,282]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="steptoeae">
<emphasis id="B920EA80FFA4FFFBC6C8FE84FB87FEE7" box="[933,1077,259,282]" italics="true" pageId="29" pageNumber="480">M. steptoeae</emphasis>
</taxonomicName>
lacks the forceps-like epandrial process and strong anipisternal bristle of
<taxonomicName id="4C544D11FFA4FFFBC700FEA0FD42FEC3" box="[621,752,294,318]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="scopalis">
<emphasis id="B920EA80FFA4FFFBC700FEA0FD42FEC3" box="[621,752,294,318]" italics="true" pageId="29" pageNumber="480">M. scopalis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C544D11FFA4FFFBC66DFEA0FC23FEC3" box="[768,913,295,318]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="steptoeae">
<emphasis id="B920EA80FFA4FFFBC66DFEA0FC23FEC3" box="[768,913,295,318]" italics="true" pageId="29" pageNumber="480">M. steptoeae</emphasis>
</taxonomicName>
is differentiated easily from
<taxonomicName id="4C544D11FFA4FFFBC1B0FEA0FAFEFEC3" box="[1245,1356,295,318]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="renata">
<emphasis id="B920EA80FFA4FFFBC1B0FEA0FAFEFEC3" box="[1245,1356,295,318]" italics="true" pageId="29" pageNumber="480">M. renata</emphasis>
</taxonomicName>
by the structure of the genitalia. In
<taxonomicName id="4C544D11FFA4FFFBC4B3FECCFDFCFE9F" box="[478,590,331,354]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="renata">
<emphasis id="B920EA80FFA4FFFBC4B3FECCFDFCFE9F" box="[478,590,331,354]" italics="true" pageId="29" pageNumber="480">M. renata</emphasis>
</taxonomicName>
, the epandrium has a triangular notch near the rear margin and numerous stout bristles near the lower margin of the epandrium;
<taxonomicName id="4C544D11FFA4FFFBC79FFEE8FC32FE7B" box="[754,896,367,390]" class="Insecta" family="Phoridae" genus="Megaselia" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="480" phylum="Arthropoda" rank="species" species="steptoeae">
<emphasis id="B920EA80FFA4FFFBC79FFEE8FC32FE7B" box="[754,896,367,390]" italics="true" pageId="29" pageNumber="480">M. steptoeae</emphasis>
</taxonomicName>
lacks such a notch and has few bristles arranged in a more-or-less vertical row on the epandrium (
<figureCitation id="136F2A17FFA4FFFBC7D1FE15FCBDFE56" box="[700,783,402,427]" captionStart="FIGURES 87 92" captionStartId="28.[151,264,1880,1903]" captionTargetBox="[167,1419,255,1809]" captionTargetId="figure@28.[150,1436,241,1829]" captionTargetPageId="28" captionText="FIGURES 87 92. Male genitalia, left lateral. 87. Megaselia rodriguezorum. 88. Megaselia sacatelensis. 89. Megaselia seaverorum. 90. Megaselia sidneyae. 91. Megaselia steptoeae. 92. Megaselia wiegmanae." httpUri="https://zenodo.org/record/242988/files/figure.png" pageId="29" pageNumber="480">Fig. 91</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34E6519FFA4FFFBC5AAFE31FE5AFE33" box="[199,488,438,463]" pageId="29" pageNumber="480" type="description">
<paragraph id="8BEB3692FFA4FFFBC5AAFE31FE5AFE33" blockId="29.[151,1436,222,823]" box="[199,488,438,463]" pageId="29" pageNumber="480">
<emphasis id="B920EA80FFA4FFFBC5AAFE31FEEBFE32" bold="true" box="[199,345,438,463]" pageId="29" pageNumber="480">Description.</emphasis>
See
<tableCitation id="C6D60329FFA4FFFBC4E3FE31FE56FE33" box="[398,484,438,463]" captionStart="TABLE 7" captionStartId="21.[151,239,440,463]" captionText="TABLE 7. Species descriptions, M. sacatelensis M. wiegmanae. Character remarks in parentheses, general remarks in last row." pageId="29" pageNumber="480">Table 7</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34E6519FFA4FFFBC5AAFE5DFD7EFE0E" box="[199,716,474,499]" pageId="29" pageNumber="480" type="distribution">
<paragraph id="8BEB3692FFA4FFFBC5AAFE5DFD7EFE0E" blockId="29.[151,1436,222,823]" box="[199,716,474,499]" pageId="29" pageNumber="480">
<emphasis id="B920EA80FFA4FFFBC5AAFE5DFED2FE0E" bold="true" box="[199,352,474,499]" pageId="29" pageNumber="480">Distribution.</emphasis>
Los Angeles, California (
<collectingCountry id="F3437602FFA4FFFBC7EFFE5CFD72FE0E" box="[642,704,475,499]" name="United States of America" pageId="29" pageNumber="480">USA</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="C34E6519FFA4FFFBC5AAFE79FB89FDEA" box="[199,1083,510,535]" pageId="29" pageNumber="480" type="etymology">
<paragraph id="8BEB3692FFA4FFFBC5AAFE79FB89FDEA" blockId="29.[151,1436,222,823]" box="[199,1083,510,535]" pageId="29" pageNumber="480">
<emphasis id="B920EA80FFA4FFFBC5AAFE79FEFCFDEA" bold="true" box="[199,334,510,535]" pageId="29" pageNumber="480">Etymology.</emphasis>
Named in honor of LaChristian Steptoe, host of BioSCAN site 10.
</paragraph>
</subSubSection>
<subSubSection id="C34E6519FFA4FFFBC5AAFDA5FE16FDC6" box="[199,420,546,571]" pageId="29" pageNumber="480" type="biology_ecology">
<paragraph id="8BEB3692FFA4FFFBC5AAFDA5FE16FDC6" blockId="29.[151,1436,222,823]" box="[199,420,546,571]" pageId="29" pageNumber="480">
<emphasis id="B920EA80FFA4FFFBC5AAFDA5FE95FDC6" bold="true" box="[199,295,546,571]" pageId="29" pageNumber="480">Biology.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34E6519FFA4FFFBC5AAFDC1FA95FCCA" pageId="29" pageNumber="480" type="materials_examined">
<paragraph id="8BEB3692FFA4FFFBC5AAFDC1FF4EFD7E" blockId="29.[151,1436,222,823]" pageId="29" pageNumber="480">
<emphasis id="B920EA80FFA4FFFBC5AAFDC1FE89FDA2" bold="true" box="[199,315,582,607]" pageId="29" pageNumber="480">
<typeStatus id="54EF8830FFA4FFFBC5AAFDC1FE84FDA2" box="[199,310,582,607]" pageId="29" pageNumber="480" type="holotype">Holotype</typeStatus>
.
</emphasis>
♂,
<collectingCountry id="F3437602FFA4FFFBC41DFDC0FE1EFDA2" box="[368,428,583,607]" name="United States of America" pageId="29" pageNumber="480">USA</collectingCountry>
: CALIFORNIA: Los Angeles, Los Feliz,
<date id="FFEA1052FFA4FFFBC6C9FDC0FB8CFDA2" box="[932,1086,583,607]" pageId="29" pageNumber="480" value="2014-04-04" valueMax="2014-04-11" valueMin="2014-04-04">411.IV.2014</date>
, Louie, Malaise trap (
<collectionCode id="ED45AE57FFA4FFFBC024FDC0FA2EFDA2" LSID="urn:lsid:biocol.org:col:34802" box="[1353,1436,583,607]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="29" pageNumber="480">LACM</collectionCode>
329834).
</paragraph>
<paragraph id="8BEB3692FFA4FFFBC5AAFD09FD90FD36" blockId="29.[151,1436,222,823]" pageId="29" pageNumber="480">
<emphasis id="B920EA80FFA4FFFBC5AAFD09FEF4FD5A" bold="true" box="[199,326,654,679]" pageId="29" pageNumber="480">
<typeStatus id="54EF8830FFA4FFFBC5AAFD09FEF3FD5A" box="[199,321,654,679]" pageId="29" pageNumber="480" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="9D52FD1BFFA4FFFBC423FD08FEC9FD5B" box="[334,379,655,679]" pageId="29" pageNumber="480" type="generic">4 ♂</specimenCount>
,
<collectingCountry id="F3437602FFA4FFFBC4E4FD08FE77FD5A" box="[393,453,655,679]" name="United States of America" pageId="29" pageNumber="480">USA</collectingCountry>
: CALIFORNIA: Los Angeles, Koreatown,
<date id="FFEA1052FFA4FFFBC6D9FD09FBFCFD5A" box="[948,1102,654,679]" pageId="29" pageNumber="480" value="2014-04-05" valueMax="2014-04-12" valueMin="2014-04-05">512.IV.2014</date>
, Ralph, Malaise trap (
<collectionCode id="ED45AE57FFA4FFFBC024FD08FA2EFD5A" LSID="urn:lsid:biocol.org:col:34802" box="[1353,1436,655,679]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="29" pageNumber="480">LACM</collectionCode>
329835,
<collectionCode id="ED45AE57FFA4FFFBC594FD34FEF8FD36" LSID="urn:lsid:biocol.org:col:34802" box="[249,330,691,715]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="29" pageNumber="480">LACM</collectionCode>
329836, 2
<collectionCode id="ED45AE57FFA4FFFBC4A5FD34FDA9FD36" box="[456,539,691,715]" httpUri="http://grbio.org/cool/da58-p0xp" name="Chulalongkorn University Museum of Natural History" pageId="29" pageNumber="480">CUMZ</collectionCode>
)
</paragraph>
<paragraph id="8BEB3692FFA4FFFBC5AAFD51FA95FCCA" blockId="29.[151,1436,222,823]" pageId="29" pageNumber="480">
<emphasis id="B920EA80FFA4FFFBC5AAFD51FD8DFD12" bold="true" box="[199,575,726,751]" pageId="29" pageNumber="480">Additional Material Examined.</emphasis>
<collectingCountry id="F3437602FFA4FFFBC72AFD50FD31FD12" box="[583,643,727,751]" name="United States of America" pageId="29" pageNumber="480">USA</collectingCountry>
: CALIFORNIA: Los Angeles: Exposition Park, Los Feliz, Silverlake, Leimert Park, Koreatown, Mid-City, Elysian Park, Gardena, Hollywood, Glendale, Larchmont, Carthay, Echo Park, Eagle Rock, Pico-Union, Highland Park,
<specimenCount id="9D52FD1BFFA4FFFBC7C5FC99FD50FCCB" box="[680,738,798,823]" pageId="29" pageNumber="480" type="generic">45 ♂</specimenCount>
,
<date id="FFEA1052FFA4FFFBC783FC98FCE5FCCA" box="[750,855,799,823]" pageId="29" pageNumber="480" value="2014-01" valueMax="2014-05-31" valueMin="2014-01-00">IV.2014</date>
, Malaise traps (
<collectionCode id="ED45AE57FFA4FFFBC16AFC98FBEEFCCA" LSID="urn:lsid:biocol.org:col:34802" box="[1031,1116,799,823]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34802" name="Natural History Museum of Los Angeles County" pageId="29" pageNumber="480">LACM</collectionCode>
,
<collectionCode id="ED45AE57FFA4FFFBC105FC98FB08FCCA" box="[1128,1210,799,823]" pageId="29" pageNumber="480">MCZC</collectionCode>
,
<collectionCode id="ED45AE57FFA4FFFBC1A8FC98FAA9FCCA" LSID="urn:lsid:biocol.org:col:34871" box="[1221,1307,799,823]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="29" pageNumber="480">USNM</collectionCode>
).
</paragraph>
</subSubSection>
</treatment>
</document>