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<mods:title id="6C28A3FFF37C6144C5560B1398259C7F">Revision of the genus Sadoletus Distant, with description of new species from China and Australia (Hemiptera: Heteroptera: Heterogastridae)</mods:title>
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Genus
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(
<figureCitation id="136FCFA684263154FF48FCE5FEC7FC64" box="[159,280,873,900]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 13" captionStart-11="FIGURE 14" captionStart-12="FIGURE 15" captionStart-13="FIGURE 16" captionStart-14="FIGURE 18" captionStart-15="FIGURE 19" captionStart-16="FIGURE 20" captionStart-17="FIGURE 21" captionStart-18="FIGURE 22" captionStart-19="FIGURE 23" captionStart-2="FIGURE 3" captionStart-20="FIGURE 25" captionStart-21="FIGURE 26" captionStart-3="FIGURE 4" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 9" captionStart-7="FIGURE 10" captionStart-8="FIGURE 11" captionStart-9="FIGURE 12" captionStartId-0="4.[151,250,1910,1935]" captionStartId-1="5.[151,250,1972,1997]" captionStartId-10="19.[151,250,1767,1792]" captionStartId-11="20.[151,250,1975,2000]" captionStartId-12="21.[151,250,1618,1643]" captionStartId-13="23.[151,250,1974,1999]" captionStartId-14="26.[151,250,1904,1929]" captionStartId-15="27.[151,250,1662,1687]" captionStartId-16="28.[151,250,917,942]" captionStartId-17="29.[151,250,1980,2005]" captionStartId-18="30.[151,250,1476,1501]" captionStartId-19="31.[151,250,1980,2005]" captionStartId-2="6.[151,250,1918,1943]" captionStartId-20="34.[151,250,1980,2005]" captionStartId-21="35.[151,250,1637,1662]" captionStartId-3="7.[151,250,1742,1767]" captionStartId-4="9.[151,250,1958,1983]" captionStartId-5="11.[151,250,1942,1967]" captionStartId-6="13.[151,250,1898,1923]" captionStartId-7="15.[151,250,1691,1716]" captionStartId-8="16.[151,250,1651,1676]" captionStartId-9="18.[151,250,1956,1981]" captionTargetBox-0="[163,1436,181,1870]" captionTargetBox-1="[246,1321,180,1942]" captionTargetBox-10="[158,1436,217,1729]" captionTargetBox-11="[286,1312,193,1924]" captionTargetBox-12="[170,1417,177,1578]" captionTargetBox-13="[173,1415,523,1938]" captionTargetBox-14="[151,1436,460,1870]" captionTargetBox-15="[199,1386,190,1641]" captionTargetBox-16="[151,1436,177,897]" captionTargetBox-17="[165,1427,185,1956]" captionTargetBox-18="[151,1436,177,1457]" captionTargetBox-19="[294,1304,210,1933]" captionTargetBox-2="[168,1427,384,1870]" captionTargetBox-20="[263,1343,193,1915]" captionTargetBox-21="[217,1386,293,1597]" captionTargetBox-3="[151,1436,177,1712]" captionTargetBox-4="[151,1436,416,1924]" captionTargetBox-5="[193,1392,561,1860]" captionTargetBox-6="[151,1436,200,1856]" captionTargetBox-7="[151,1436,304,1671]" captionTargetBox-8="[151,1436,268,1631]" captionTargetBox-9="[234,1376,180,1911]" captionTargetId-0="figure@4.[151,1436,177,1890]" captionTargetId-1="figure@5.[169,1418,177,1952]" captionTargetId-10="figure@19.[151,1436,177,1747]" captionTargetId-11="figure@20.[163,1425,177,1955]" captionTargetId-12="figure@21.[170,1417,177,1598]" captionTargetId-13="figure@23.[173,1415,517,1955]" captionTargetId-14="figure@26.[151,1436,460,1870]" captionTargetId-15="figure@27.[189,1398,177,1643]" captionTargetId-16="figure@28.[151,1436,177,897]" captionTargetId-17="figure@29.[151,1436,177,1960]" captionTargetId-18="figure@30.[151,1436,177,1457]" captionTargetId-19="figure@31.[195,1392,195,1961]" captionTargetId-2="figure@6.[151,1436,365,1906]" captionTargetId-20="figure@34.[186,1402,177,1960]" captionTargetId-21="figure@35.[189,1398,269,1618]" captionTargetId-3="figure@7.[151,1436,177,1722]" captionTargetId-4="figure@9.[151,1436,416,1924]" captionTargetId-5="figure@11.[193,1395,553,1923]" captionTargetId-6="figure@13.[151,1436,200,1856]" captionTargetId-7="figure@15.[151,1436,304,1671]" captionTargetId-8="figure@16.[151,1436,268,1631]" captionTargetId-9="figure@18.[151,1436,177,1937]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-10="19" captionTargetPageId-11="20" captionTargetPageId-12="21" captionTargetPageId-13="23" captionTargetPageId-14="26" captionTargetPageId-15="27" captionTargetPageId-16="28" captionTargetPageId-17="29" captionTargetPageId-18="30" captionTargetPageId-19="31" captionTargetPageId-2="6" captionTargetPageId-20="34" captionTargetPageId-21="35" captionTargetPageId-3="7" captionTargetPageId-4="9" captionTargetPageId-5="11" captionTargetPageId-6="13" captionTargetPageId-7="15" captionTargetPageId-8="18" captionTargetPageId-9="18" captionText-0="FIGURE 1. Types of Sadoletus validus Distant, 1903 and their labels. a, lectotype, female, dorsal view; b, same, lateral view; c, paralectotype, male, dorsal view; d, same, lateral view; e, paralectotype, pygophore, posterodorsal view; fi, paralectotype, left paramere, four different aspects; j, lectotype, labels; k, paralectotype, labels. Abbreviations: mp, median projection." captionText-1="FIGURE 2. Sadoletus validus Distant, 1903. a, male, dorsal view; b, same, ventral view; c, female, dorsal view; d, same, ventral view." captionText-10="FIGURE 13. Sadoletus melasmus Ban, 2018. a, female abdominal dorsum, dorsal view; b, female abdominal venter, ventral view; c, ovipositor, posteroventral view; d, spermathecal bulb; e, pygophore, posterodorsal view; fi, left paramere, four different aspects." captionText-11="FIGURE 14. Sadoletus biprotuberans sp. nov. a, holotype, male, dorsal view; b, same, lateral view; c, paratype, female, dorsal view; d, same, lateral view." captionText-12="FIGURE 15. Types of Sadoletus biprotuberans sp. nov. a, holotype, male abdominal dorsum, dorsal view; b, same, male abdominal venter, ventral view; c, paratype, female abdominal dorsum, dorsal view; d, same, female abdominal venter, ventral view; ej, holotype; e, pygophore, posterodorsal view; fi, left paramere, four different aspects; j, phallus, lateral view; kn, paratype; k, ovipositor, posteroventral view; l, spermathecal bulb; m, n, ovipositor.Abbreviations: 1 gpo, 2 gpo, first and second gonapophysis; og = outgrowth; ph, phallotheca; pt VIII, pt IX, eighth and ninth paratergites; sm, spermatheca; sp, spiracle; t VIII, eighth tergite." captionText-13="FIGURE 16. Sadoletus variabilis sp. nov. paratypes. a, male, dorsal view; b, same, ventral view; c, male, dorsal view; d, same, ventral view; e, female, dorsal view; f, same, ventral view." captionText-14="FIGURE 18. Lectotype of Sadoletus voluptarius Bergroth, 1918 and its labels. a, dorsal view; b, ventral view; c, pygophore, posterodorsal view; d, labels (Photographed by Pekka Malinen, MZH)." captionText-15="FIGURE 19. Sadoletus voluptarius Bergroth, 1918. a, male habitus, dorsal view; b, same, ventral view; c, male abdominal dorsum, dorsal view; d, male abdominal venter, ventral view; e, pygophore, posterodorsal view; f, pygophore, dorsal view; gj, left paramere, four different aspects. Abbreviations: cp, cuplike sclerite; mp, median projection; og, outgrowth." captionText-16="FIGURE 20. Type of Sadoletus corvus Distant, 1908 and its labels. a, dorsal view (photograph from Előd Kondorosy, Hun- gary); b, labels (photograph from Mick Webb, NHM)." captionText-17="FIGURE 21. Lectotype of Sadoletus bakeri Bergroth, 1918 and its labels. a, dorsal view; b, ventral view; c, ovipositor, ventral view; d, labels (Photographed by Pekka Malinen, MZH)." captionText-18="FIGURE 22. Type of Sadoletus vulvatus Bergroth, 1918 and its labels. a, dorsal view; b, same, ventral view; c, ovipositor, ventral view; d, labels (Photographed by Pekka Malinen, MZH)." captionText-19="FIGURE 23. Sadoletus abathonotus sp. nov. paratypes. a, male, dorsal view; b, same, ventral view; c, female, dorsal view; f, same, ventral view." captionText-2="FIGURE 3. Sadoletus validus Distant, 1903. a, male abdominal dorsum, dorsal view; b, male abdominal venter, ventral view; c, female abdominal dorsum, dorsal view; d, female abdominal venter, ventral view; e, pygophore, posterodorsal view; f, phallotheca, lateral view; gj, left paramere, four different aspects. Abbreviations: IVII, abdominal segments; ho, horn-like outgrowth; sgs, scent gland scar." captionText-20="FIGURE 25. Sadoletus alphus sp. nov. a, holotype, male, dorsal view; b, same, ventral view; c, paratype, female, dorsal view; d, same, ventral view." captionText-21="FIGURE 26. Sadoletus alphus sp. nov. ab, paratype. a, female abdominal dorsum, dorsal view; b, female abdominal venter, inner view; ch, holotype. c, pygophore, posterodorsal view; d, phallus, lateral view; eh, left paramere, four different aspects; i, paratype, spermatheca. Abbreviations: sm, spermatheca." captionText-3="FIGURE 4. Sadoletus validus Distant, 1903. a, head, ventral view; b, meso- and metapleura; c, thoracic scent gland; d, sternite V; e, anterior trichobothria on sternite V; f, apex of abdomen. Abbreviations: VII, seventh sternite; sg, scent gland; sp, spiracle; tr, trichobothria." captionText-4="FIGURE 6. Paralectotype of Sadoletus valdezi Bergroth, 1918 and its labels. a, dorsal view; b, ventral view; c, ovipositor, ventral view; d, labels (Photographed by Pekka Malinen, MZH)." captionText-5="FIGURE 7. Types of Sadoletus pallescens Distant, 1909 and their labels. a, lectotype, female, dorsal view; b, same, lateral view; c, same, abdominal venter, ventral view; d, paralectotype, dorsal view; e, same, lateral view; f, lectotype, labels; g, paralectotype, labels." captionText-6="FIGURE 9. Sadoletus planus sp. nov. paratypes. a, male abdominal dorsum, dorsal view; b, male abdominal venter, ventral view; c, female abdominal dorsum, dorsal view; d, female abdominal venter, ventral view; e, pygophore, posterodorsal view; f, phallotheca, lateral view; gj, left paramere, four different aspects; k, female, tergite VIII, dorsal view; l, ovipositor, inner view. Abbreviations: ap, apodeme; sm, spermatheca." captionText-7="FIGURE 10. Lectotype of Sadoletus montanellus Bergroth, 1918 and its labels. a, dorsal view; b, ventral view; c, pygophore, posterodorsal view; d, labels (Photographed by Pekka Malinen, MZH)." captionText-8="FIGURE 11. Lectotype of Sadoletus montivagus Bergroth, 1918 and its labels. a, dorsal view; b, ventral view; c, pygophore, posterodorsal view; d, labels (Photographed by Pekka Malinen, MZH)." captionText-9="FIGURE 12. Sadoletus melasmus Ban, 2018. a, male, dorsal view; b, same, ventral view; c, female, dorsal view; d, same, ventral view." httpUri-0="https://zenodo.org/record/3238904/files/figure.png" httpUri-1="https://zenodo.org/record/3238906/files/figure.png" httpUri-10="https://zenodo.org/record/3238920/files/figure.png" httpUri-11="https://zenodo.org/record/3238922/files/figure.png" httpUri-12="https://zenodo.org/record/3238924/files/figure.png" httpUri-13="https://zenodo.org/record/3238926/files/figure.png" httpUri-14="https://zenodo.org/record/3238928/files/figure.png" httpUri-15="https://zenodo.org/record/3238930/files/figure.png" httpUri-16="https://zenodo.org/record/3238932/files/figure.png" httpUri-17="https://zenodo.org/record/3238934/files/figure.png" httpUri-18="https://zenodo.org/record/3238936/files/figure.png" httpUri-19="https://zenodo.org/record/3238938/files/figure.png" httpUri-2="https://zenodo.org/record/3238908/files/figure.png" httpUri-20="https://zenodo.org/record/3238940/files/figure.png" httpUri-21="https://zenodo.org/record/3238942/files/figure.png" httpUri-3="https://zenodo.org/record/3238910/files/figure.png" httpUri-4="https://zenodo.org/record/3238912/files/figure.png" httpUri-5="https://zenodo.org/record/3238914/files/figure.png" httpUri-6="https://zenodo.org/record/3238916/files/figure.png" httpUri-7="https://zenodo.org/record/3405169/files/figure.png" httpUri-8="https://zenodo.org/record/3405171/files/figure.png" httpUri-9="https://zenodo.org/record/3238918/files/figure.png" pageId="1" pageNumber="252">Figs. 126</figureCitation>
)
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<emphasis id="B9200F3184263154FF40FC22FF25FC26" box="[151,250,942,966]" italics="true" pageId="1" pageNumber="252">Sadoletus</emphasis>
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<bibRefCitation id="EFC5AED284263154FED7FC21FE53FC26" author="Distant, W. L." box="[256,396,941,966]" pageId="1" pageNumber="252" refId="ref15319" refString="Distant, W. L. (1903) The Fauna of British India, including Ceylon and Burma. Rhynchota. - Vol. II. (Heteroptera). Part 1. Taylor and Francis, London, 242 pp." type="book" year="1903">Distant, 1903</bibRefCitation>
: 35
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, 37.
<typeStatus id="54EF6D8184263154FE36FC22FDCCFC26" box="[481,531,942,966]" pageId="1" pageNumber="252">Type</typeStatus>
species by original designation:
<taxonomicName id="4C54A8A084263154FCB6FC22FB7AFC26" ID-CoL="4TXKP" authority="Distant, 1903" authorityName="Distant" authorityYear="1903" box="[865,1189,941,966]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="validus">
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<emphasis id="B9200F3184263154FF40FC40FF25FC04" box="[151,250,972,996]" italics="true" pageId="1" pageNumber="252">Sadoletus</emphasis>
:
<treatmentCitation id="0AF5F53284263154FEDFFC40FE17FC04" author="Distant, W. L." box="[264,456,972,997]" page="128" pageId="1" pageNumber="252" year="1908">
<bibRefCitation id="EFC5AED284263154FEDFFC40FE49FC04" author="Distant, W. L." box="[264,406,972,997]" pageId="1" pageNumber="252" pagination="127 - 151" refId="ref15362" refString="Distant, W. L. (1908) Rhynchota Malayana. Part I. Records of the Indian Museum, 2, 127 - 151, pls. VII-VIII." type="journal article" year="1908">Distant (1908</bibRefCitation>
: 128
</treatmentCitation>
)
</taxonomicName>
(fauna of
<collectingCountry id="F34393B384263154FDEDFC40FD44FC05" box="[570,667,972,997]" name="Malaysia" pageId="1" pageNumber="252">Malaysia</collectingCountry>
),
<treatmentCitation id="0AF5F53284263154FD7BFC40FCA3FC04" author="Bergroth, E." box="[684,892,972,997]" page="112" pageId="1" pageNumber="252" year="1918">
<bibRefCitation id="EFC5AED284263154FD7BFC40FC93FC04" author="Bergroth, E." box="[684,844,972,997]" pageId="1" pageNumber="252" pagination="43 - 73" refId="ref15240" refString="Bergroth, E. (1918) Studies in Philippine Heteroptera, I. Philippine Journal of Science, D, 13 (2 &amp; 3), 43 - 73 &amp; 75 - 126." type="journal article" year="1918">Bergroth (1918</bibRefCitation>
: 112
</treatmentCitation>
) (diagnostic characters, distribution),
<treatmentCitation id="0AF5F53284263154FAD7FC40FF31FBE3" author="Scudder, G. G. E." page="120" pageId="1" pageNumber="252" year="1962">
<bibRefCitation id="EFC5AED284263154FAD7FC40FA46FC05" author="Scudder, G. G. E." box="[1280,1433,972,997]" pageId="1" pageNumber="252" pagination="117 - 127" refId="ref15779" refString="Scudder, G. G. E. (1962) New Heterogastrinae (Hemiptera) with a key to the genera of the world. Opuscula Entomologica, 27, 117 - 127." type="journal article" year="1962">Scudder (1962</bibRefCitation>
: 120
</treatmentCitation>
) (in key),
<treatmentCitation id="0AF5F53284263154FE82FC66FDDBFBE3" author="Slater, J. A." box="[341,516,1002,1027]" page="776" pageId="1" pageNumber="252" year="1964">
<bibRefCitation id="EFC5AED284263154FE82FC66FE0BFBE3" author="Slater, J. A." box="[341,468,1002,1027]" pageId="1" pageNumber="252" pagination="779 - 1668" refId="ref15814" refString="Slater, J. A. (1964) A catalogue of the Lygaeidae of the world. Vols. I-II. University of Connecticut, Storrs, XVIII + 1668 pp. [I: I-XVIII, 1 - 778; II: 779 - 1668]" type="book chapter" year="1964">Slater (1964</bibRefCitation>
: 776
</treatmentCitation>
) (catalogue),
<treatmentCitation id="0AF5F53284263154FD59FC66FC54FBE3" author="Zheng, L. Y. &amp; Zou, H. G." box="[654,907,1002,1027]" page="108" pageId="1" pageNumber="252" year="1981">
<bibRefCitation id="EFC5AED284263154FD59FC66FC84FBE3" author="Zheng, L. Y. &amp; Zou, H. G." box="[654,859,1002,1027]" pageId="1" pageNumber="252" pagination="1 - 215" refId="ref15862" refString="Zheng, L. Y. &amp; Zou, H. G. (1981) Lygaeidae. In: Hsiao, T. Y., Ren, S. Z., Zheng, L. Y., Jing, X. L., Zou, H. G. &amp; Liu, S. L. (Eds.), A handbook for the determination of the Chinese Hemiptera-Heteroptera. Vol. 2. Science Press, Beijing, pp. 1 - 215, plates 1 - 26. [in Chinese, English summary]" type="book chapter" year="1981">Zheng &amp; Zou (1981</bibRefCitation>
: 108
</treatmentCitation>
, 114) (in key, diagnosis, fauna of
<collectingCountry id="F34393B384263154FB0CFC66FAC4FBE3" box="[1243,1307,1002,1027]" name="China" pageId="1" pageNumber="252">China</collectingCountry>
),
<treatmentCitation id="0AF5F53284263154FAFBFC67FF3DFBC2" author="Zou, H. G." page="93" pageId="1" pageNumber="252" year="1985">
<bibRefCitation id="EFC5AED284263154FAFBFC67FA46FBE3" author="Zou, H. G." box="[1324,1433,1002,1027]" pageId="1" pageNumber="252" pagination="93 - 96" refId="ref15965" refString="Zou, H. G. (1985) A new genus and species of Heterogastrinae from China (Hemiptera: Lygaeidae). Acta Scientiarum Naturalium Universitatis Nankaiensis, 1985 (2), 93 - 96. [in Chinese, English summary]" type="journal article" year="1985">Zou (1985</bibRefCitation>
: 93
</treatmentCitation>
, 96) (comparison with
<taxonomicName id="4C54A8A084263154FE07FB86FD8CFBC2" ID-CoL="8QPJ7" authorityName="Zou" authorityYear="1985" box="[464,595,1034,1058]" class="Insecta" family="Heterogastridae" genus="Equatobursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="genus">
<emphasis id="B9200F3184263154FE07FB86FD8CFBC2" box="[464,595,1034,1058]" italics="true" pageId="1" pageNumber="252">Equatobursa</emphasis>
</taxonomicName>
), Péricart (2001: 105) (catalogue, Palaearctic),
<treatmentCitation id="0AF5F53284263154FBEEFB85FA4AFBC2" author="Ishikawa, T. &amp; Tomokuni, M." box="[1081,1429,1033,1058]" page="316" pageId="1" pageNumber="252" year="2012">
<bibRefCitation id="EFC5AED284263154FBEEFB85FABBFBC2" author="Ishikawa, T. &amp; Tomokuni, M." box="[1081,1380,1033,1058]" pageId="1" pageNumber="252" pagination="315 - 317" refId="ref15550" refString="Ishikawa, T. &amp; Tomokuni, M. (2012) Family Heterogastridae Stal, 1872. In: Ishikawa, T., Takai, M. &amp; Yasunaga, T. (Eds.), A field guide to Japanese bugs - Terrestrial heteropterans. Vol. 3. Zenkoku Noson Kyoiku Kyokai Publishing Co., Tokyo, pp. 315 - 317, pl. 67. [in Japanese]" type="journal article" year="2012">
<collectingRegion id="49901DC184263154FBEEFB85FB4BFBC2" box="[1081,1172,1033,1058]" country="Japan" name="Ishikawa" pageId="1" pageNumber="252">Ishikawa</collectingRegion>
&amp; Tomokuni (2012
</bibRefCitation>
: 316
</treatmentCitation>
) (fauna of
<collectingCountry id="F34393B384263154FEFFFBA4FEB9FBA0" box="[296,358,1064,1088]" name="Japan" pageId="1" pageNumber="252">Japan</collectingCountry>
),
<treatmentCitation id="0AF5F53284263154FEA0FBA5FD50FBA1" author="Ban, T. &amp; Ishikawa, T." box="[375,655,1064,1089]" page="196" pageId="1" pageNumber="252" year="2013">
<bibRefCitation id="EFC5AED284263154FEA0FBA5FD81FBA0" author="Ban, T. &amp; Ishikawa, T." box="[375,606,1064,1089]" pageId="1" pageNumber="252" pagination="195 - 203" refId="ref15189" refString="Ban, T. &amp; Ishikawa, T. (2013) The lygaeoid bug genus Sadoletus Distant (Hemiptera: Heteroptera: Heterogastridae) from Japan. Zootaxa, 3609 (2), 195 - 203. https: // doi. org / 10.11646 / zootaxa. 3609.2.5" type="journal article" year="2013">
Ban &amp;
<collectingRegion id="49901DC184263154FE68FBA4FDC5FBA1" box="[447,538,1064,1089]" country="Japan" name="Ishikawa" pageId="1" pageNumber="252">Ishikawa</collectingRegion>
(2013
</bibRefCitation>
: 196
</treatmentCitation>
) (diagnosis, fauna of
<collectingCountry id="F34393B384263154FCBEFBA4FC78FBA0" box="[873,935,1064,1088]" name="Japan" pageId="1" pageNumber="252">Japan</collectingCountry>
);
<treatmentCitation id="0AF5F53284263154FC6FFBA5FB97FBA1" ID-DOI="http://doi.org/10.5281/zenodo.5974605" ID-Zenodo-Dep="5974605" author="Ban, T." box="[952,1096,1064,1089]" httpUri="http://treatment.plazi.org/id/9B474B53FFEA3D6BFF01B46BFADD567F" page="55" pageId="1" pageNumber="252" year="2018">
<bibRefCitation id="EFC5AED284263154FC6FFBA5FBF9FBA0" author="Ban, T." box="[952,1062,1064,1088]" pageId="1" pageNumber="252" pagination="1" refId="ref15135" refString="Ban, T. (2018) The lygaeoid bug family Heterogastridae (Hemiptera: Heteroptera: Lygaeoidea) from Laos and Thailand, with descriptions of three new species. Zootaxa, 4446 (1), 051 - 067. https: // doi. org / 10.11646 / zootaxa. 4446.1.4" type="journal article" year="2018">Ban (2018</bibRefCitation>
: 55
</treatmentCitation>
) (in key, diagnosis, fauna of
<collectingCountry id="F34393B384263154FABCFBA5FA43FBA0" box="[1387,1436,1065,1088]" name="Laos" pageId="1" pageNumber="252">Laos</collectingCountry>
and
<collectingCountry id="F34393B384263154FF25FBCAFE8EFBBF" box="[242,337,1094,1119]" name="Thailand" pageId="1" pageNumber="252">Thailand</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BEBD32384263154FF40FBEAFCB5FB9E" blockId="1.[151,1437,941,1150]" box="[151,874,1125,1150]" pageId="1" pageNumber="252">
<treatmentCitationGroup id="AB44F40D84263154FF40FBEAFCB5FB9E" box="[151,874,1125,1150]" pageId="1" pageNumber="252">
<taxonomicName id="4C54A8A084263154FF40FBEAFE6EFB9E" ID-CoL="8QPJ7" authority="Zou, 1985: 93" authorityName="Zou" authorityPageNumber="93" authorityYear="1985" box="[151,433,1125,1150]" class="Insecta" family="Heterogastridae" genus="Equatobursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="genus">
<emphasis id="B9200F3184263154FF40FBEAFEC5FB9E" box="[151,282,1126,1150]" italics="true" pageId="1" pageNumber="252">Equatobursa</emphasis>
<treatmentCitation id="0AF5F53284263154FEF7FBEAFE6EFB9E" author="Zou, H. G." box="[288,433,1125,1150]" page="93" pageId="1" pageNumber="252" year="1985">
<bibRefCitation id="EFC5AED284263154FEF7FBEAFE53FB9E" author="Zou, H. G." box="[288,396,1125,1150]" pageId="1" pageNumber="252" pagination="93 - 96" refId="ref15965" refString="Zou, H. G. (1985) A new genus and species of Heterogastrinae from China (Hemiptera: Lygaeidae). Acta Scientiarum Naturalium Universitatis Nankaiensis, 1985 (2), 93 - 96. [in Chinese, English summary]" type="journal article" year="1985">Zou, 1985</bibRefCitation>
: 93
</treatmentCitation>
</taxonomicName>
, 96. Synonymized by
<bibRefCitation id="EFC5AED284263154FD44FBEAFCBAFB9E" author="Gao, C. Q. &amp; Redei, D." box="[659,869,1125,1150]" pageId="1" pageNumber="252" pagination="300 - 306" refId="ref15491" refString="Gao, C. Q. &amp; Redei, D. (2017) The identity of Equatobursa, with proposal of new genus and species level synonymies (Hemiptera: Heteroptera: Heterogastridae). Zootaxa, 4237 (2), 300 - 306. https: // doi. org / 10.11646 / zootaxa. 4237.2.4" type="journal article" year="2017">Gao &amp; Rédei (2017)</bibRefCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34E80A884263154FF40FB27FAF5F8CB" pageId="1" pageNumber="252" type="description">
<paragraph id="8BEBD32384263154FF40FB27FCF5FB0A" blockId="1.[151,1437,1195,1835]" pageId="1" pageNumber="252">
<emphasis id="B9200F3184263154FF40FB27FE9AFB26" bold="true" box="[151,325,1195,1222]" pageId="1" pageNumber="252">Redescription.</emphasis>
Following are additions and/ or alterations to the detailed redescription of the genus
<taxonomicName id="4C54A8A084263154FB2DFB20FAB8FB26" authorityName=": Distant" authorityYear="1908" box="[1274,1383,1196,1222]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="genus">
<emphasis id="B9200F3184263154FB2DFB20FAB8FB26" box="[1274,1383,1196,1222]" italics="true" pageId="1" pageNumber="252">Sadoletus</emphasis>
</taxonomicName>
provided by
<bibRefCitation id="EFC5AED284263154FF28FB5DFDD8FB0A" author="Ban, T. &amp; Ishikawa, T." box="[255,519,1232,1259]" pageId="1" pageNumber="252" pagination="195 - 203" refId="ref15189" refString="Ban, T. &amp; Ishikawa, T. (2013) The lygaeoid bug genus Sadoletus Distant (Hemiptera: Heteroptera: Heterogastridae) from Japan. Zootaxa, 3609 (2), 195 - 203. https: // doi. org / 10.11646 / zootaxa. 3609.2.5" type="journal article" year="2013">
Ban &amp;
<collectingRegion id="49901DC184263154FE87FB5CFE6AFB0A" box="[336,437,1232,1258]" country="Japan" name="Ishikawa" pageId="1" pageNumber="252">Ishikawa</collectingRegion>
(2013)
</bibRefCitation>
and
<bibRefCitation id="EFC5AED284263154FDEAFB5CFCF9FB0A" author="Gao, C. Q. &amp; Redei, D." box="[573,806,1232,1258]" pageId="1" pageNumber="252" pagination="300 - 306" refId="ref15491" refString="Gao, C. Q. &amp; Redei, D. (2017) The identity of Equatobursa, with proposal of new genus and species level synonymies (Hemiptera: Heteroptera: Heterogastridae). Zootaxa, 4237 (2), 300 - 306. https: // doi. org / 10.11646 / zootaxa. 4237.2.4" type="journal article" year="2017">Gao &amp; Rédei (2017)</bibRefCitation>
.
</paragraph>
<paragraph id="8BEBD32384263154FF10FB78FDABF9CE" blockId="1.[151,1437,1195,1835]" pageId="1" pageNumber="252">
Buccula as in
<figureCitation id="136FCFA684263154FEB2FB78FE69FAEE" box="[357,438,1268,1294]" captionStart="FIGURE 4" captionStartId="7.[151,250,1742,1767]" captionTargetBox="[151,1436,177,1712]" captionTargetId="figure@7.[151,1436,177,1722]" captionTargetPageId="7" captionText="FIGURE 4. Sadoletus validus Distant, 1903. a, head, ventral view; b, meso- and metapleura; c, thoracic scent gland; d, sternite V; e, anterior trichobothria on sternite V; f, apex of abdomen. Abbreviations: VII, seventh sternite; sg, scent gland; sp, spiracle; tr, trichobothria." httpUri="https://zenodo.org/record/3238910/files/figure.png" pageId="1" pageNumber="252">Fig. 4a</figureCitation>
; metathoracic scent gland as in
<figureCitation id="136FCFA684263154FCCDFB78FC59FAEF" box="[794,902,1268,1295]" captionStart="FIGURE 4" captionStartId="7.[151,250,1742,1767]" captionTargetBox="[151,1436,177,1712]" captionTargetId="figure@7.[151,1436,177,1722]" captionTargetPageId="7" captionText="FIGURE 4. Sadoletus validus Distant, 1903. a, head, ventral view; b, meso- and metapleura; c, thoracic scent gland; d, sternite V; e, anterior trichobothria on sternite V; f, apex of abdomen. Abbreviations: VII, seventh sternite; sg, scent gland; sp, spiracle; tr, trichobothria." httpUri="https://zenodo.org/record/3238910/files/figure.png" pageId="1" pageNumber="252">Fig. 4b, c</figureCitation>
; tergites I and II fused; sutures between tergites IIIIV straight or slightly curved caudad, IVV moderately and VVI strongly or IVV and VVI equally curved caudad in the middle area, VIVII straight or slightly sinuate, and with scent gland scars only between IVV and VVI well developed and subequal in width. Outer laterotergite II present, very small; outer laterotergite III narrowed towards base of abdomen, and IVVI elongated rectangle; distal area of outer laterotergite VII and tergite VII fused. Tergites IVI with regular dense punctures, and tergite VII with sparse smaller irregular punctures. Sternum covered with very fine punctures, almost indistinct. Trichobothria in full trichobothria complement (3:3:3:3:2); detailed structure of trichobothria as in
<figureCitation id="136FCFA684263154FD83FA7CFD1EF9EB" box="[596,705,1520,1547]" captionStart="FIGURE 4" captionStartId="7.[151,250,1742,1767]" captionTargetBox="[151,1436,177,1712]" captionTargetId="figure@7.[151,1436,177,1722]" captionTargetPageId="7" captionText="FIGURE 4. Sadoletus validus Distant, 1903. a, head, ventral view; b, meso- and metapleura; c, thoracic scent gland; d, sternite V; e, anterior trichobothria on sternite V; f, apex of abdomen. Abbreviations: VII, seventh sternite; sg, scent gland; sp, spiracle; tr, trichobothria." httpUri="https://zenodo.org/record/3238910/files/figure.png" pageId="1" pageNumber="252">Fig. 4d, e</figureCitation>
; bothrium not sunken below surface of the cuticle and trichomes present, with short spine-like microtrichia.
</paragraph>
<paragraph id="8BEBD32384263154FF10F9B4FAF5F8CB" blockId="1.[151,1437,1195,1835]" pageId="1" pageNumber="252">
Ovipositor short, only reaching posterior margin of abdominal sternite VI; abdominal sternite VII short and medially divided fully to its base, with the two halves gradually broadly rounded and flap-like, either leaving a narrow gap in the middle (as in
<taxonomicName id="4C54A8A084263154FE75F90CFDD9F97A" authorityName="Bergroth" authorityYear="1918" box="[418,518,1664,1690]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="bakeri">
<emphasis id="B9200F3184263154FE75F90CFDD9F97A" box="[418,518,1664,1690]" italics="true" pageId="1" pageNumber="252">S. bakeri</emphasis>
</taxonomicName>
) (
<figureCitation id="136FCFA684263154FDC9F90CFDA4F97A" box="[542,635,1664,1690]" captionStart="FIGURE 21" captionStartId="29.[151,250,1980,2005]" captionTargetBox="[165,1427,185,1956]" captionTargetId="figure@29.[151,1436,177,1960]" captionTargetPageId="29" captionText="FIGURE 21. Lectotype of Sadoletus bakeri Bergroth, 1918 and its labels. a, dorsal view; b, ventral view; c, ovipositor, ventral view; d, labels (Photographed by Pekka Malinen, MZH)." httpUri="https://zenodo.org/record/3238934/files/figure.png" pageId="1" pageNumber="252">Fig. 21c</figureCitation>
) or the flaps are slightly (for example in
<taxonomicName id="4C54A8A084263154FB95F90CFB73F97A" authorityName="Bergroth" authorityYear="1918" box="[1090,1196,1664,1690]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="valdezi">
<emphasis id="B9200F3184263154FB95F90CFB73F97A" box="[1090,1196,1664,1690]" italics="true" pageId="1" pageNumber="252">S. valdezi</emphasis>
</taxonomicName>
) (
<figureCitation id="136FCFA684263154FB12F90CFACBF97B" box="[1221,1300,1664,1691]" captionStart="FIGURE 6" captionStartId="9.[151,250,1958,1983]" captionTargetBox="[151,1436,416,1924]" captionTargetId="figure@9.[151,1436,416,1924]" captionTargetPageId="9" captionText="FIGURE 6. Paralectotype of Sadoletus valdezi Bergroth, 1918 and its labels. a, dorsal view; b, ventral view; c, ovipositor, ventral view; d, labels (Photographed by Pekka Malinen, MZH)." httpUri="https://zenodo.org/record/3238912/files/figure.png" pageId="1" pageNumber="252">Fig. 6c</figureCitation>
) or strongly (for example in
<taxonomicName id="4C54A8A084263154FE9DF928FE0BF95E" authorityName="Ban" authorityYear="2018" box="[330,468,1700,1726]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="melasmus">
<emphasis id="B9200F3184263154FE9DF928FE0BF95E" box="[330,468,1700,1726]" italics="true" pageId="1" pageNumber="252">S. melasmus</emphasis>
</taxonomicName>
) (
<figureCitation id="136FCFA684263154FE3AF928FD94F95F" box="[493,587,1700,1727]" captionStart="FIGURE 13" captionStartId="19.[151,250,1767,1792]" captionTargetBox="[158,1436,217,1729]" captionTargetId="figure@19.[151,1436,177,1747]" captionTargetPageId="19" captionText="FIGURE 13. Sadoletus melasmus Ban, 2018. a, female abdominal dorsum, dorsal view; b, female abdominal venter, ventral view; c, ovipositor, posteroventral view; d, spermathecal bulb; e, pygophore, posterodorsal view; fi, left paramere, four different aspects." httpUri="https://zenodo.org/record/3238920/files/figure.png" pageId="1" pageNumber="252">Fig. 13b</figureCitation>
) overlapping in the middle, and the sternites IIIVI are complete, with their posterior margins entire, almost straight, not divided or narrowed in the middle (
<figureCitation id="136FCFA684263154FBDAF944FBB5F902" box="[1037,1130,1736,1762]" captionStart="FIGURE 3" captionStartId="6.[151,250,1918,1943]" captionTargetBox="[168,1427,384,1870]" captionTargetId="figure@6.[151,1436,365,1906]" captionTargetPageId="6" captionText="FIGURE 3. Sadoletus validus Distant, 1903. a, male abdominal dorsum, dorsal view; b, male abdominal venter, ventral view; c, female abdominal dorsum, dorsal view; d, female abdominal venter, ventral view; e, pygophore, posterodorsal view; f, phallotheca, lateral view; gj, left paramere, four different aspects. Abbreviations: IVII, abdominal segments; ho, horn-like outgrowth; sgs, scent gland scar." httpUri="https://zenodo.org/record/3238908/files/figure.png" pageId="1" pageNumber="252">Figs. 3d</figureCitation>
;
<figureCitation id="136FCFA684263154FBA2F944FB73F903" box="[1141,1196,1736,1763]" captionStart="FIGURE 6" captionStartId="9.[151,250,1958,1983]" captionTargetBox="[151,1436,416,1924]" captionTargetId="figure@9.[151,1436,416,1924]" captionTargetPageId="9" captionText="FIGURE 6. Paralectotype of Sadoletus valdezi Bergroth, 1918 and its labels. a, dorsal view; b, ventral view; c, ovipositor, ventral view; d, labels (Photographed by Pekka Malinen, MZH)." httpUri="https://zenodo.org/record/3238912/files/figure.png" pageId="1" pageNumber="252">6b, c</figureCitation>
), moderate or weakly concave (Fig. 8e). Pygophore dorsal opening with a well developed median projection on its posterior margin; median projection fused with cuplike sclerite arising from inner side (except in
<taxonomicName id="4C54A8A084263154FC27F89CFBACF8CA" authorityName="Ban" authorityYear="2018" box="[1008,1139,1808,1834]" class="Insecta" family="Heterogastridae" genus="Sadoletus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="252" phylum="Arthropoda" rank="species" species="laosensis">
<emphasis id="B9200F3184263154FC27F89CFBACF8CA" box="[1008,1139,1808,1834]" italics="true" pageId="1" pageNumber="252">S. laosensis</emphasis>
</taxonomicName>
) (
<figureCitation id="136FCFA684263154FB5CF89CFB38F8CA" box="[1163,1255,1808,1834]" captionStart="FIGURE 1" captionStartId="4.[151,250,1910,1935]" captionTargetBox="[163,1436,181,1870]" captionTargetId="figure@4.[151,1436,177,1890]" captionTargetPageId="4" captionText="FIGURE 1. Types of Sadoletus validus Distant, 1903 and their labels. a, lectotype, female, dorsal view; b, same, lateral view; c, paralectotype, male, dorsal view; d, same, lateral view; e, paralectotype, pygophore, posterodorsal view; fi, paralectotype, left paramere, four different aspects; j, lectotype, labels; k, paralectotype, labels. Abbreviations: mp, median projection." httpUri="https://zenodo.org/record/3238904/files/figure.png" pageId="1" pageNumber="252">Figs. 1e</figureCitation>
;
<figureCitation id="136FCFA684263154FB23F89CFAC2F8CB" box="[1268,1309,1808,1835]" captionStart="FIGURE 19" captionStartId="27.[151,250,1662,1687]" captionTargetBox="[199,1386,190,1641]" captionTargetId="figure@27.[189,1398,177,1643]" captionTargetPageId="27" captionText="FIGURE 19. Sadoletus voluptarius Bergroth, 1918. a, male habitus, dorsal view; b, same, ventral view; c, male abdominal dorsum, dorsal view; d, male abdominal venter, ventral view; e, pygophore, posterodorsal view; f, pygophore, dorsal view; gj, left paramere, four different aspects. Abbreviations: cp, cuplike sclerite; mp, median projection; og, outgrowth." httpUri="https://zenodo.org/record/3238930/files/figure.png" pageId="1" pageNumber="252">19f</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>