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<document id="17244A141217BF3041C1F7FCDFB4CA7D" ID-CLB-Dataset="305320" ID-DOI="10.4314/met.v32i1.6" ID-GBIF-Dataset="968b6c66-e35b-4f42-8e73-e380d3c599a8" ID-ISSN="2307-5031" ID-Zenodo-Dep="14199458" ID-ZooBank="92B77F73-1BBE-4B6F-AF9E-FEE578A75420" IM.bibliography_approvedBy="julia" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1731461411424" checkinUser="felipe" docAuthor="Libert, Michel &amp; Collins, Steve C." docDate="2021" docId="03FCBF1CFF837C73FF137CB8EF74F43F" docLanguage="en" docName="Metamorphosis.32.30-35.pdf" docOrigin="Metamorphosis 32" docStyle="DocumentStyle:73CBAE06DE6A380ACC54A94377F5FCBA.1:Metamorphosis.2021-.journal_article" docStyleId="73CBAE06DE6A380ACC54A94377F5FCBA" docStyleName="Metamorphosis.2021-.journal_article" docStyleVersion="1" docTitle="Liptena eketi Bethune-Baker 1926" docType="treatment" docVersion="3" lastPageNumber="32" masterDocId="FFC5C764FF817C71FF977E42EA68F72D" masterDocTitle="Taxonomic notes on Liptena eketi Bethune-Baker, 1926 and related species (Papilionoidea: Lycaenidae: Poritiinae)" masterLastPageNumber="35" masterPageNumber="30" pageNumber="32" updateTime="1732213473018" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
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<mods:title id="ECADE79030A866B09DF0D48F544B7FA9">Taxonomic notes on Liptena eketi Bethune-Baker, 1926 and related species (Papilionoidea: Lycaenidae: Poritiinae)</mods:title>
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<treatment id="03FCBF1CFF837C73FF137CB8EF74F43F" ID-DOI="http://doi.org/10.5281/zenodo.14199520" ID-Zenodo-Dep="14199520" LSID="urn:lsid:plazi:treatment:03FCBF1CFF837C73FF137CB8EF74F43F" httpUri="http://treatment.plazi.org/id/03FCBF1CFF837C73FF137CB8EF74F43F" lastPageNumber="32" pageId="2" pageNumber="32">
<subSubSection id="C34F5D81FF837C73FF137CB8EB97F43F" box="[132,511,762,786]" pageId="2" pageNumber="32" type="nomenclature">
<paragraph id="8BEA0E0AFF837C73FF137CB8EB97F43F" blockId="2.[132,511,762,786]" box="[132,511,762,786]" pageId="2" pageNumber="32">
<heading id="D0A2B966FF837C73FF137CB8EB97F43F" box="[132,511,762,786]" fontSize="10" level="2" pageId="2" pageNumber="32" reason="8">
<taxonomicName id="4C557589FF837C73FF137CB8EB97F43F" ID-CoL="3VC8K" authority="Bethune-Baker, 1926" authorityName="Bethune-Baker" authorityYear="1926" box="[132,511,762,786]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="eketi">
<emphasis id="B921D218FF837C73FF137CB8EB7CF43F" bold="true" box="[132,276,762,786]" italics="true" pageId="2" pageNumber="32">Liptena eketi</emphasis>
<bibRefCitation id="EFC473FBFF837C73FE8D7CB8EB97F43F" author="BETHUNE-BAKER, G. T." box="[282,511,762,786]" pageId="2" pageNumber="32" pagination="384 - 402" part="9" refId="ref3623" refString="BETHUNE-BAKER, G. T. 1926. Annals and Magazine of Natural History (9) 17: 391 (384 - 402)." title="Annals and Magazine of Natural History" type="book chapter" year="1926">Bethune-Baker, 1926</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34F5D81FF837C73FF137D73E8EFF37B" pageId="2" pageNumber="32" type="materials_examined">
<paragraph id="8BEA0E0AFF837C73FF137D73EBA0F464" blockId="2.[132,456,817,841]" box="[132,456,817,841]" pageId="2" pageNumber="32">
<typeStatus id="54EEB0A8FF837C73FF137D73EAD3F464" box="[132,187,817,841]" pageId="2" pageNumber="32">Type</typeStatus>
material and distribution
</paragraph>
<paragraph id="8BEA0E0AFF817C71FCDE796DE9FFF06B" blockId="0.[841,1456,1838,1986]" box="[841,919,1838,1862]" pageId="0" pageNumber="30">
<taxonomicName id="4C557589FF817C71FCDE796DE9FFF06B" authorityName="Bethune-Baker" authorityYear="1926" box="[841,919,1838,1862]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="0" pageNumber="30" phylum="Arthropoda" rank="species" species="eketi">
<emphasis id="B921D218FF817C71FCDE796DE9FFF06B" box="[841,919,1838,1862]" italics="true" pageId="0" pageNumber="30">L. eketi</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BEA0E0AFF817C71FCDE790FE98BF0EC" blockId="0.[841,1456,1838,1986]" pageId="0" pageNumber="30">
<specimenCount id="9D53C583FF817C71FCDE790FE903F048" box="[841,875,1869,1893]" count="3" pageId="0" pageNumber="30" type="male">3♂</specimenCount>
Rhoko, Iko Esai Community Forest,
<collectingRegion id="4991C0E8FF817C71FB66790FEFC3F048" box="[1265,1451,1869,1893]" country="Nigeria" name="Cross River" pageId="0" pageNumber="30">Cross River State</collectingRegion>
,
<collectingCountry id="F3424E9AFF817C71FCDE7929E9F1F0AE" box="[841,921,1899,1923]" name="Nigeria" pageId="0" pageNumber="30">Nigeria</collectingCountry>
,
<date id="FFEB28CAFF817C71FC3E7929EEEDF0AE" box="[937,1157,1899,1923]" pageId="0" pageNumber="30" value="2010-11-28" valueMax="2010-12-20" valueMin="2010-11-28">28.xi. to 20.xii.2010</date>
, leg. J. Brophy,
<collectionCode id="ED4496CFFF817C71FAA97929EF15F0AE" box="[1342,1405,1899,1923]" pageId="0" pageNumber="30">ABRI</collectionCode>
;
<specimenCount id="9D53C583FF817C71FA197929EFC7F0AE" box="[1422,1455,1899,1923]" count="5" pageId="0" pageNumber="30" type="male">5♂</specimenCount>
as above, RWC;
<specimenCount id="9D53C583FF817C71FB9679C8EE4BF08F" box="[1025,1059,1930,1954]" count="2" pageId="0" pageNumber="30" type="female">2♀</specimenCount>
near Umuhaia, Eastern
<collectingCountry id="F3424E9AFF817C71FABE79C8EF14F08F" box="[1321,1404,1930,1954]" name="Nigeria" pageId="0" pageNumber="30">Nigeria</collectingCountry>
, leg. Davey,
<collectionCode id="ED4496CFFF817C71FC0B79EBE9B4F0EC" box="[924,988,1961,1985]" pageId="0" pageNumber="30">ABRI</collectionCode>
.
</paragraph>
<paragraph id="8BEA0E0AFF837C73FF137D2BE8EFF37B" blockId="2.[132,747,872,1110]" pageId="2" pageNumber="32">
<taxonomicName id="4C557589FF837C73FF137D2BEABDF4AD" authorityName="Bethune-Baker" authorityYear="1926" box="[132,213,872,896]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="eketi">
<emphasis id="B921D218FF837C73FF137D2BEABDF4AD" box="[132,213,872,896]" italics="true" pageId="2" pageNumber="32">L. eketi</emphasis>
</taxonomicName>
was described from a male collected at Eket, in south-eastern
<collectingCountry id="F3424E9AFF837C73FE8E7DC4EB04F4B3" box="[281,364,902,926]" name="Nigeria" pageId="2" pageNumber="32">Nigeria</collectingCountry>
.
<taxonomicName id="4C557589FF837C73FEE17DC5EBADF4B3" authorityName="Bethune-Baker" authorityYear="1926" box="[374,453,902,926]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="eketi">
<emphasis id="B921D218FF837C73FEE17DC5EBADF4B3" box="[374,453,902,926]" italics="true" pageId="2" pageNumber="32">L. eketi</emphasis>
</taxonomicName>
was illustrated by
<bibRefCitation id="EFC473FBFF837C73FD197DC4EB6EF490" author="D'ABRERA, B." bookContentInfo="260 pp." journalOrPublisher="Hill House, Melbourne, London" pageId="2" pageNumber="32" refId="ref3722" refString="D'ABRERA, B., 2009. Butterflies of the Afrotropical Region. New and revised edition. Part III, Lycaenidae, Riodinidae. Hill House, Melbourne, London 260 pp." title="Butterflies of the Afrotropical Region. New and revised edition. Part III, Lycaenidae, Riodinidae" type="book" year="2009">d'Abrera (2009: 649)</bibRefCitation>
, with the upper side of the
<typeStatus id="54EEB0A8FF837C73FDA27DE7E8FAF490" box="[565,658,933,957]" pageId="2" pageNumber="32" type="holotype">holotype</typeStatus>
and the underside of another male. A female captured in western
<collectingCountry id="F3424E9AFF837C73FF137DA1EA9BF4D6" box="[132,243,995,1019]" name="Cameroon" pageId="2" pageNumber="32">Cameroon</collectingCountry>
was attributed to
<taxonomicName id="4C557589FF837C73FE207DA6E86EF4D6" authorityName="Bethune-Baker" authorityYear="1926" box="[439,518,995,1019]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="eketi">
<emphasis id="B921D218FF837C73FE207DA6E86EF4D6" box="[439,518,995,1019]" italics="true" pageId="2" pageNumber="32">L. eketi</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC473FBFF837C73FDA67DA1E883F4D6" author="LIBERT, M." box="[561,747,995,1019]" journalOrPublisher="Lambillionea" pageId="2" pageNumber="32" pagination="21 - 34" part="92" refId="ref3756" refString="LIBERT, M., 1992. Notes faunistiques sur les Lepidopteres Rhopaloceres du Cameroun. I - Liste d'especes non connues du Cameroun. Lambillionea 92 (1): 21 - 34." title="Notes faunistiques sur les Lepidopteres Rhopaloceres du Cameroun. I - Liste d'especes non connues du Cameroun" type="journal article" year="1992">Libert (1992: 28)</bibRefCitation>
but is now considered to be
<taxonomicName id="4C557589FF837C73FE3C7A40E84BF334" box="[427,547,1025,1049]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="kiellandi">
<emphasis id="B921D218FF837C73FE3C7A40E84BF334" box="[427,547,1025,1049]" italics="true" pageId="2" pageNumber="32">L. kiellandi</emphasis>
</taxonomicName>
(see below). From the available data, the range of
<taxonomicName id="4C557589FF837C73FE4D7A63E841F315" authorityName="Bethune-Baker" authorityYear="1926" box="[474,553,1056,1080]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="eketi">
<emphasis id="B921D218FF837C73FE4D7A63E841F315" box="[474,553,1056,1080]" italics="true" pageId="2" pageNumber="32">L. eketi</emphasis>
</taxonomicName>
therefore appears to be restricted to south-eastern
<collectingCountry id="F3424E9AFF837C73FE4D7A7CE842F37B" box="[474,554,1086,1110]" name="Nigeria" pageId="2" pageNumber="32">Nigeria</collectingCountry>
(
<figureCitation id="136E128FFF837C73FDAF7A7CE81FF37B" box="[568,631,1086,1110]" captionStart="Figure 1" captionStartId="1.[479,547,1019,1040]" captionTargetBox="[133,1435,361,987]" captionTargetId="figure-267@1.[133,1435,361,987]" captionTargetPageId="1" captionText="Figure 1 Distribution map for L. eketi, L. seyboui, L. kiellandi" figureDoi="http://doi.org/10.5281/zenodo.14199460" httpUri="https://zenodo.org/record/14199460/files/figure.png" pageId="2" pageNumber="32">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34F5D81FF837C73FF137A34EF74F43F" pageId="2" pageNumber="32" type="description">
<paragraph id="8BEA0E0AFF837C73FF137A34EB0CF3A3" blockId="2.[132,356,1142,1166]" box="[132,356,1142,1166]" pageId="2" pageNumber="32">
Facies (
<figureCitation id="136E128FFF837C73FF4F7A34EB33F3A3" box="[216,347,1142,1166]" captionStart="Plate 1" captionStartId="2.[165,217,620,641]" captionTargetBox="[153,1446,162,585]" captionTargetId="figure-918@2.[153,1446,162,585]" captionTargetPageId="2" captionText="Plate 1 A, B: L. seyboui, Alépé (Ivory Coast) A: ♂ Recto, B: ♀ Verso; C, D: ♂ L. eketi, Rhoko forest (Nigeria) C: Recto, D: Verso E, F: ♂ L. kiellandi, Maliva (NE DRC) E: Recto, F: Verso; G, H: ♀ L. kiellandi, Manki (Cameroon) G: Recto, H: Verso." figureDoi="http://doi.org/10.5281/zenodo.14199464" httpUri="https://zenodo.org/record/14199464/files/figure.png" pageId="2" pageNumber="32">Plate 1 C, D</figureCitation>
)
</paragraph>
<paragraph id="8BEA0E0AFF837C73FF137AEFE806F3E8" blockId="2.[132,622,1197,1221]" box="[132,622,1197,1221]" pageId="2" pageNumber="32">
See the description by
<bibRefCitation id="EFC473FBFF837C73FEE07AEFE80FF3E8" author="BETHUNE-BAKER, G. T." box="[375,615,1197,1221]" pageId="2" pageNumber="32" pagination="384 - 402" part="9" refId="ref3623" refString="BETHUNE-BAKER, G. T. 1926. Annals and Magazine of Natural History (9) 17: 391 (384 - 402)." title="Annals and Magazine of Natural History" type="book chapter" year="1926">Bethune-Baker (1926)</bibRefCitation>
.
</paragraph>
<paragraph id="8BEA0E0AFF837C73FF137AA6EBFAF3D1" blockId="2.[132,402,1252,1276]" box="[132,402,1252,1276]" pageId="2" pageNumber="32">
Male genitalia (
<figureCitation id="136E128FFF837C73FEBB7AA6EB07F3D1" box="[300,367,1252,1276]" captionStart="Figures 2" captionStartId="1.[148,225,1948,1969]" captionTargetBox="[133,1440,1070,1913]" captionTargetId="figure-270@1.[133,1440,1070,1913]" captionTargetPageId="1" captionText="Figures 2 to 7 Male genitalia of Liptena eketi (♂ Rhoko Forest, Eastern Nigeria, gen. prep. M. Libert 118-143) 2 left lateral view; 3 dorsal view of uncus; 4 posterior view; 5 ventral view of saccus and penis; 6 penis, left lateral view; 7 penis, dorsal view of the distal part." figureDoi="http://doi.org/10.5281/zenodo.14199462" httpUri="https://zenodo.org/record/14199462/files/figure.png" pageId="2" pageNumber="32">Figs 2</figureCitation>
7)
</paragraph>
<paragraph id="8BEA0E0AFF837C73FF137B59EB75F14B" blockId="2.[132,747,1307,1638]" pageId="2" pageNumber="32">
The saccus of
<taxonomicName id="4C557589FF837C73FEA77B5EEBECF21E" authorityName="Bethune-Baker" authorityYear="1926" box="[304,388,1307,1331]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="eketi">
<emphasis id="B921D218FF837C73FEA77B5EEBECF21E" box="[304,388,1307,1331]" italics="true" pageId="2" pageNumber="32">L. eketi</emphasis>
</taxonomicName>
is so developed that it can be described as hypertrophied (even unique). The lateral view of the genitalia shows that its extremity curls up (
<figureCitation id="136E128FFF837C73FD287B1BEAFAF2A2" captionStart="Figures 2" captionStartId="1.[148,225,1948,1969]" captionTargetBox="[133,1440,1070,1913]" captionTargetId="figure-270@1.[133,1440,1070,1913]" captionTargetPageId="1" captionText="Figures 2 to 7 Male genitalia of Liptena eketi (♂ Rhoko Forest, Eastern Nigeria, gen. prep. M. Libert 118-143) 2 left lateral view; 3 dorsal view of uncus; 4 posterior view; 5 ventral view of saccus and penis; 6 penis, left lateral view; 7 penis, dorsal view of the distal part." figureDoi="http://doi.org/10.5281/zenodo.14199462" httpUri="https://zenodo.org/record/14199462/files/figure.png" pageId="2" pageNumber="32">Fig. 2</figureCitation>
) and the posterior view that the upraised part presents a significant notch (Fig. 4). In the natural position, the distal end of the penis is very close to the saccus (
<figureCitation id="136E128FFF837C73FD347BF7E88DF2E0" box="[675,741,1461,1485]" captionStart="Figures 2" captionStartId="1.[148,225,1948,1969]" captionTargetBox="[133,1440,1070,1913]" captionTargetId="figure-270@1.[133,1440,1070,1913]" captionTargetPageId="1" captionText="Figures 2 to 7 Male genitalia of Liptena eketi (♂ Rhoko Forest, Eastern Nigeria, gen. prep. M. Libert 118-143) 2 left lateral view; 3 dorsal view of uncus; 4 posterior view; 5 ventral view of saccus and penis; 6 penis, left lateral view; 7 penis, dorsal view of the distal part." figureDoi="http://doi.org/10.5281/zenodo.14199462" httpUri="https://zenodo.org/record/14199462/files/figure.png" pageId="2" pageNumber="32">Fig. 2</figureCitation>
, arrow), and it can be hypothesised that the notch constitutes a guide for the penis. While the saccus rarely offers any interesting character, that of
<taxonomicName id="4C557589FF837C73FDFA7850E8AFF104" authorityName="Bethune-Baker" authorityYear="1926" box="[621,711,1553,1577]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="eketi">
<emphasis id="B921D218FF837C73FDFA7850E8AFF104" box="[621,711,1553,1577]" italics="true" pageId="2" pageNumber="32">L. eketi</emphasis>
</taxonomicName>
is genuinely exceptional, both for its size and the role it seems to play.
</paragraph>
<paragraph id="8BEA0E0AFF837C73FF1378C7E8E0F059" blockId="2.[132,747,1669,1908]" pageId="2" pageNumber="32">
The dorsal edge of the valve is adorned with a structure composed of two processes, one distal and the other internal. There is no ventral process comparable to
<taxonomicName id="4C557589FF837C73FF1378A0EA9BF1D4" box="[132,243,1761,1785]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="seyboui">
<emphasis id="B921D218FF837C73FF1378A0EA9BF1D4" box="[132,243,1761,1785]" italics="true" pageId="2" pageNumber="32">L. seyboui</emphasis>
</taxonomicName>
. The distal process, visible on the lateral view (
<figureCitation id="136E128FFF837C73FF1B7942EABDF035" box="[140,213,1792,1816]" captionStart="Figures 2" captionStartId="1.[148,225,1948,1969]" captionTargetBox="[133,1440,1070,1913]" captionTargetId="figure-270@1.[133,1440,1070,1913]" captionTargetPageId="1" captionText="Figures 2 to 7 Male genitalia of Liptena eketi (♂ Rhoko Forest, Eastern Nigeria, gen. prep. M. Libert 118-143) 2 left lateral view; 3 dorsal view of uncus; 4 posterior view; 5 ventral view of saccus and penis; 6 penis, left lateral view; 7 penis, dorsal view of the distal part." figureDoi="http://doi.org/10.5281/zenodo.14199462" httpUri="https://zenodo.org/record/14199462/files/figure.png" pageId="2" pageNumber="32">Fig. 2</figureCitation>
), is massive and well sclerotized; an internal process, much thinner, is approximately perpendicular to the distal process and to the valve and therefore only visible in the dorsal and lateral views (Figs 4-5).
</paragraph>
<paragraph id="8BEA0E0AFF837C73FF1379D1EF74F43F" blockId="2.[132,747,1939,2055]" lastBlockId="2.[841,1456,700,786]" pageId="2" pageNumber="32">
Other differences are less important, but the subunci are shorter in
<taxonomicName id="4C557589FF837C73FE9779F1EB3EF0E7" authorityName="Bethune-Baker" authorityYear="1926" box="[256,342,1970,1994]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="eketi">
<emphasis id="B921D218FF837C73FE9779F1EB3EF0E7" box="[256,342,1970,1994]" italics="true" pageId="2" pageNumber="32">L. eketi</emphasis>
</taxonomicName>
(
<figureCitation id="136E128FFF837C73FEFB79F0EBDEF0E7" box="[364,438,1970,1994]" captionStart="Figures 2" captionStartId="1.[148,225,1948,1969]" captionTargetBox="[133,1440,1070,1913]" captionTargetId="figure-270@1.[133,1440,1070,1913]" captionTargetPageId="1" captionText="Figures 2 to 7 Male genitalia of Liptena eketi (♂ Rhoko Forest, Eastern Nigeria, gen. prep. M. Libert 118-143) 2 left lateral view; 3 dorsal view of uncus; 4 posterior view; 5 ventral view of saccus and penis; 6 penis, left lateral view; 7 penis, dorsal view of the distal part." figureDoi="http://doi.org/10.5281/zenodo.14199462" httpUri="https://zenodo.org/record/14199462/files/figure.png" pageId="2" pageNumber="32">Fig. 2</figureCitation>
). The penis of
<taxonomicName id="4C557589FF837C73FDE379F1E8A2F0E7" authorityName="Bethune-Baker" authorityYear="1926" box="[628,714,1970,1994]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="eketi">
<emphasis id="B921D218FF837C73FDE379F1E8A2F0E7" box="[628,714,1970,1994]" italics="true" pageId="2" pageNumber="32">L. eketi</emphasis>
</taxonomicName>
is distinctly twisted, its distal part turned to the right (Fig. 7 shows a dorsal view of this part, the basal part turned left). It is possible that the penis of
<taxonomicName id="4C557589FF837C73FB017CFFEF6EF5F9" box="[1174,1286,700,724]" class="Insecta" family="Lycaenidae" genus="Liptena" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="32" phylum="Arthropoda" rank="species" species="seyboui">
<emphasis id="B921D218FF837C73FB017CFFEF6EF5F9" box="[1174,1286,700,724]" italics="true" pageId="2" pageNumber="32">L. seyboui</emphasis>
</taxonomicName>
is also twisted, and that the unusual shape of its distal end is an artefact (
<bibRefCitation id="EFC473FBFF837C73FCC67CB8EEDAF43F" author="SAFIAN, Sz. &amp; LORENC-BRUDECKA, J." box="[849,1202,762,786]" journalOrPublisher="Metamorphosis" pageId="2" pageNumber="32" pagination="4 - 6" part="31" refId="ref3869" refString="SAFIAN, Sz. &amp; LORENC-BRUDECKA, J., 2020. Unique genitalic structure in a West African lycaenid butterfly, Liptena seyboui Warren-Gash &amp; Larsen, 2003 (Lepidoptera: Lycaenidae: Poritiinae: Liptenini). Metamorphosis 31: 4 - 6." title="Unique genitalic structure in a West African lycaenid butterfly, Liptena seyboui Warren-Gash &amp; Larsen, 2003 (Lepidoptera: Lycaenidae: Poritiinae: Liptenini)" type="journal article" year="2020">Sáfián &amp; Lorenc-Brudecka 2020</bibRefCitation>
, Fig. 4C).
</paragraph>
</subSubSection>
</treatment>
</document>