treatments-xml/data/03/FB/8C/03FB8C32F81CFFB1FF60FE99FB654D8D.xml
2024-06-21 12:22:17 +02:00

295 lines
37 KiB
XML
Raw Permalink Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="16AEBF9709B085086012FAC8703EABF3" ID-CLB-Dataset="27714" ID-DOI="10.11646/zootaxa.4543.1.1" ID-GBIF-Dataset="22ef92b8-1d9c-4e9f-b01c-ce1f5d3faa7b" ID-ISSN="1175-5326" ID-Zenodo-Dep="2617607" ID-ZooBank="31ABFA6E-6126-4603-B84F-4BEC7632D1E8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553927968043" checkinUser="plazi" docAuthor="Fresneda, Javier, Faille, Arnaud, Fery, Hans &amp; Ribera, Ignacio" docDate="2019" docId="03FB8C32F81CFFB1FF60FE99FB654D8D" docLanguage="en" docName="zootaxa.4543.1.1.pdf" docOrigin="Zootaxa 4543 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Speonemadus brusteli Fresneda &amp; Faille &amp; Fery &amp; Ribera 2019, sp. n." docType="treatment" docVersion="8" lastPageNumber="6" masterDocId="FFC2F44AF819FFB4FFF7FF98FFAD4B19" masterDocTitle="A molecular phylogeny of Speonemadus Jeannel, 1922 with description of two new species from Morocco (Coleoptera: Leiodidae: Cholevinae: Anemadini)" masterLastPageNumber="36" masterPageNumber="1" pageNumber="6" updateTime="1698715325246" updateUser="ExternalLinkService">
<mods:mods id="24866FA8EE2AE10F7A5C73A1DCF7C727" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E104F02BB03274D84775BB1119088CF4">
<mods:title id="9B3793EE9874F84D4AFB610436B1B913">A molecular phylogeny of Speonemadus Jeannel, 1922 with description of two new species from Morocco (Coleoptera: Leiodidae: Cholevinae: Anemadini)</mods:title>
</mods:titleInfo>
<mods:name id="A2B49187AB9D462647188B2C486B83A0" type="personal">
<mods:role id="36A196B5D57FC958E5C4199557E880CC">
<mods:roleTerm id="4E60F39F26A905B216BF559EC9D66FD6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="75E042A009545606D0DB782728F808FD">Fresneda, Javier</mods:namePart>
</mods:name>
<mods:name id="8AA6A36F37FE16A96C4469E24C727A1B" type="personal">
<mods:role id="E207DAD38D11D71BFAF3BF1FAE63AA16">
<mods:roleTerm id="4433EA407866DA28DB72168491278B65">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3309FD00048F1FE5419BB08142D9BE50">Faille, Arnaud</mods:namePart>
</mods:name>
<mods:name id="26C8DAC8DB25713998094B6C67F383A3" type="personal">
<mods:role id="C61E69DEA49D68E76C450EAAB72088FE">
<mods:roleTerm id="A2312D31755142580F05953FF99F680A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AA901503B6C54A7A30FCF6A974CECA1C">Fery, Hans</mods:namePart>
</mods:name>
<mods:name id="60356B07214F2B838C0902F01ABD022D" type="personal">
<mods:role id="CA071D1EE7B3F07A17D0C53D00D59722">
<mods:roleTerm id="50792C032B5989BF6007C66EA501A8CD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F1438D9C2D23CFF951FB77757AE25220">Ribera, Ignacio</mods:namePart>
</mods:name>
<mods:typeOfResource id="AF4F64C1741B7F303E1FA17813ED2B79">text</mods:typeOfResource>
<mods:relatedItem id="BE4A0AA9506D67EED9AEB7FBF694D815" type="host">
<mods:titleInfo id="AC850855B66B20ACC3A56F96E1CC19EA">
<mods:title id="B90DC89E6A9DD5EF6FA109F8CB863F97">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="97217DCADD0773D4606C5E356208E024">
<mods:date id="1B7D6FC9DF2847ED4B48C2A73927C0B1">2019</mods:date>
<mods:detail id="B310AA7A957366B21435A3E515D07061" type="pubDate">
<mods:number id="0E19CDA0A22A93F092C9C285FB709F10">2019-01-04</mods:number>
</mods:detail>
<mods:detail id="1DCE925A8CE729DEB65EC7C0B81473B6" type="volume">
<mods:number id="5DEE7457D7427EE1EF9E1F26AD652F6D">4543</mods:number>
</mods:detail>
<mods:detail id="66FECF170450961DBCBEC9F17DCF3A87" type="issue">
<mods:number id="F105A9DA15FB7B4BD4EC2EB21671513C">1</mods:number>
</mods:detail>
<mods:extent id="C1F95F2CA864868FB6BABE5F47B1E5B7" unit="page">
<mods:start id="7E504726E0CEE92B4C94A600BF90F5A8">1</mods:start>
<mods:end id="F54D8DC8DACF505DF188AA7EC210819D">36</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4E03690E1BB51FBD5FECBA5A6D518CC1">journal article</mods:classification>
<mods:identifier id="6D32F4A5D96FC0E8FAA9B743B0350BC2" type="CLB-Dataset">27714</mods:identifier>
<mods:identifier id="0763135D8F54CA50713A77E15BAC7202" type="DOI">10.11646/zootaxa.4543.1.1</mods:identifier>
<mods:identifier id="15F7D8BF70A64B7FF59D083DDFFC9C5C" type="GBIF-Dataset">22ef92b8-1d9c-4e9f-b01c-ce1f5d3faa7b</mods:identifier>
<mods:identifier id="D10A2762C5B061201AD9BD9C58A9B10C" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E35B62EE25C24F9CF9BFC8CE82C52C94" type="Zenodo-Dep">2617607</mods:identifier>
<mods:identifier id="489D4EDCCD87C72D691690201ADDBBAB" type="ZooBank">31ABFA6E-6126-4603-B84F-4BEC7632D1E8</mods:identifier>
</mods:mods>
<treatment id="03FB8C32F81CFFB1FF60FE99FB654D8D" ID-DOI="http://doi.org/10.5281/zenodo.5927124" ID-GBIF-Taxon="154529607" ID-Zenodo-Dep="5927124" LSID="urn:lsid:plazi:treatment:03FB8C32F81CFFB1FF60FE99FB654D8D" httpUri="http://treatment.plazi.org/id/03FB8C32F81CFFB1FF60FE99FB654D8D" lastPageNumber="6" pageId="5" pageNumber="6">
<subSubSection id="C3486EAFF81CFFB1FF60FE99FE474A02" box="[151,490,257,283]" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="8BED3D24F81CFFB1FF60FE99FE474A02" blockId="5.[151,490,257,317]" box="[151,490,257,283]" pageId="5" pageNumber="6">
<heading id="D0A58A48F81CFFB1FF60FE99FE474A02" bold="true" box="[151,490,257,283]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="B926E136F81CFFB1FF60FE99FE474A02" bold="true" box="[151,490,257,283]" pageId="5" pageNumber="6">
<taxonomicName id="4C5246A7F81CFFB1FF60FE99FE0C4A02" ID-CoL="8Y9NL" authority="Fresneda &amp; Faille &amp; Fery &amp; Ribera, 2019" authorityName="Fresneda &amp; Faille &amp; Fery &amp; Ribera" authorityYear="2019" box="[151,417,257,283]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="brusteli" status="sp. nov.">
<emphasis id="B926E136F81CFFB1FF60FE99FE0C4A02" bold="true" box="[151,417,257,283]" italics="true" pageId="5" pageNumber="6">Speonemadus brusteli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2155C4DF81CFFB1FE5FFE99FE474A02" box="[424,490,257,283]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3486EAFF81CFFB1FF60FEBCFE8F4A25" box="[151,290,292,317]" pageId="5" pageNumber="6" type="description">
<paragraph id="8BED3D24F81CFFB1FF60FEBCFE8F4A25" blockId="5.[151,490,257,317]" box="[151,290,292,317]" pageId="5" pageNumber="6">
<figureCitation id="136921A1F81CFFB1FF60FEBCFF714A25" box="[151,220,292,317]" captionStart="FIGURES 12" captionStartId="6.[151,264,1618,1640]" captionTargetBox="[183,1395,224,1571]" captionTargetId="figure@6.[151,1437,193,1597]" captionTargetPageId="6" captionText="FIGURES 12. Habitus of the holotypes of: (1) Speonemadus brusteli sp. n.; (2) S. comasi sp. n." httpUri="https://zenodo.org/record/2617609/files/figure.png" pageId="5" pageNumber="6">Figs 1</figureCitation>
,
<figureCitation id="136921A1F81CFFB1FF1DFEBCFF574A24" box="[234,250,292,317]" captionStart="FIGURES 56" captionStartId="8.[151,264,1938,1960]" captionTargetBox="[352,1105,457,1799]" captionTargetId="figure@8.[294,1292,411,1916]" captionTargetPageId="8" captionText="FIGURES 56. Aedeagi in ventral view of: (5) Speonemadus. orchesioides (Ammi-Moussa, Algeria); (6) holotype of S. brusteli sp. n. (Talembote, Morocco)." httpUri="https://zenodo.org/record/2617613/files/figure.png" pageId="5" pageNumber="6">6</figureCitation>
,
<figureCitation id="136921A1F81CFFB1FEF1FEBCFE8F4A25" box="[262,290,292,316]" captionStart="FIGURES 1328" captionStartId="12.[151,264,1731,1753]" captionTargetBox="[170,1387,215,1697]" captionTargetId="figure@12.[151,1436,193,1710]" captionTargetPageId="12" captionText="FIGURES 1328. Apex of the parameres of: (13) Speonemadus orchesioides (Ammi-Moussa, Algeria); (14) holotype of S. brusteli sp. n. (Talembote, Morocco); (15) S. gracilis (Hornos, Spain); (16) S. tenuipes (Ifri Semedane, Algeria); (17) S. vandalitiae (Villanúa, Spain); (18) S. pulchellus (Ficuzza, Sicily); (19) holotype of S. algarvensis (Gruta do Vale Telheiro, Portugal); (20) S. angusticollis (Rute, Spain); (21) S. bolivari (Ardales, Spain); (22) S. breuili (Motillas, Spain); (23) S. escalerai (Jumilla, Spain); (24) S. transversostriatus (La Bastida, Spain); (25) S. clathratus (Linarejos, Spain); (26) S. maroccanus (Taza, Morocco); (27) holotype of S. comasi sp. n. (Tagelft, Morocco); (28) Anemadus hajeki (Yunnan, China)." httpUri="https://zenodo.org/record/2617619/files/figure.png" pageId="5" pageNumber="6">14</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3486EAFF81CFFB1FF60FEF4FC754966" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8BED3D24F81CFFB1FF60FEF4FF194ABE" blockId="5.[151,1437,364,1685]" pageId="5" pageNumber="6">
<materialsCitation id="3B3A3779F81CFFB1FF60FEF4FF1D4ABE" ID-GBIF-Occurrence="2147524523" country="Morocco" county="Talassemtane" elevation="1400" latitude="35.083332" location="Talembote" longLatPrecision="1195" longitude="-5.133333" municipality="Moroccan" pageId="5" pageNumber="6" specimenCount="1" typeStatus="holotype">
<emphasis id="B926E136F81CFFB1FF60FEF4FE974A9C" bold="true" box="[151,314,364,389]" pageId="5" pageNumber="6">
<typeStatus id="54E98386F81CFFB1FF60FEF4FF7F4A9C" box="[151,210,364,389]" pageId="5" pageNumber="6">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F3457DB4F81CFFB1FEB4FEF5FE064A9D" box="[323,427,365,388]" name="Morocco" pageId="5" pageNumber="6">Morocco</collectingCountry>
,
<location id="8E8D6BFFF81CFFB1FE40FEF4FD9D4A9C" LSID="urn:lsid:plazi:treatment:03FB8C32F81CFFB1FF60FE99FB654D8D:8E8D6BFFF81CFFB1FE40FEF4FD9D4A9C" box="[439,560,364,389]" country="Morocco" county="Talassemtane" latitude="35.083332" longLatPrecision="1195" longitude="-5.133333" municipality="Moroccan" name="Talembote" pageId="5" pageNumber="6">Talembote</location>
,
<collectingMunicipality id="6B89A75EF81CFFB1FDCAFEF5FD024A9D" box="[573,687,365,388]" pageId="5" pageNumber="6">Moroccan</collectingMunicipality>
fir (
<taxonomicName id="4C5246A7F81CFFB1FD14FEF4FC344A9D" box="[739,921,364,389]" class="Pinopsida" family="Pinaceae" genus="Abies" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="5" pageNumber="6" phylum="Tracheophyta" rank="species" species="marocana">
<emphasis id="B926E136F81CFFB1FD14FEF4FC344A9D" box="[739,921,364,389]" italics="true" pageId="5" pageNumber="6">Abies marocana</emphasis>
</taxonomicName>
) forest of
<collectingCounty id="628C45A8F81CFFB1FBE4FEF4FB014A9C" box="[1043,1196,364,389]" pageId="5" pageNumber="6">Talassemtane</collectingCounty>
,
<geoCoordinate id="EE665BE3F81CFFB1FB4EFEF4FAA54A9C" box="[1209,1288,364,389]" direction="north" orientation="latitude" pageId="5" pageNumber="6" precision="925" value="35.083332">35°5'N</geoCoordinate>
<geoCoordinate id="EE665BE3F81CFFB1FAE7FEF4FAFB4A9C" box="[1296,1366,364,389]" direction="west" orientation="longitude" pageId="5" pageNumber="6" precision="925" value="-5.133333">5°8'W</geoCoordinate>
,
<quantity id="4CAA90C1F81CFFB1FA93FEF4FF1D4ABE" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="6" unit="m" value="1400.0">
<elevation id="007FDA17F81CFFB1FA93FEF4FF1D4ABE" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="6" unit="m" value="1400.0">1400 m</elevation>
</quantity>
</materialsCitation>
.
</paragraph>
<paragraph id="8BED3D24F81CFFB1FF30FE2CFC754966" blockId="5.[151,1437,364,1685]" pageId="5" pageNumber="6">
<emphasis id="B926E136F81CFFB1FF30FE2CFE754AD4" bold="true" box="[199,472,436,461]" pageId="5" pageNumber="6">
Type series:
<materialsCitation id="3B3A3779F81CFFB1FE95FE2CFE754AD4" ID-GBIF-Occurrence="2147524521" box="[354,472,436,461]" collectingDate="2010-11-13" collectionCode="MNHN" collectorName="Herve Brustel" country="Canada" location="Canada" pageId="5" pageNumber="6" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54E98386F81CFFB1FE95FE2CFE7E4AD4" box="[354,467,436,461]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
:
</materialsCitation>
</emphasis>
<specimenCount id="9D54F6ADF81CFFB1FE15FE2DFE574AD5" box="[482,506,437,460]" pageId="5" pageNumber="6" type="male"></specimenCount>
: &quot;
<collectorName id="26A758F2F81CFFB1FDE5FE2CFD194AD4" box="[530,692,436,461]" pageId="5" pageNumber="6">Hervé Brustel</collectorName>
leg. / MO Rif Talembote / (Sapinière) Talassemtane /
<date id="FFEC1BE4F81CFFB1FAC3FE2CFA314AD5" box="[1332,1436,436,460]" pageId="5" pageNumber="6" value="2010-11-13">
<collectingDate id="EFA8E20CF81CFFB1FAC3FE2CFA314AD5" box="[1332,1436,436,460]" pageId="5" pageNumber="6" value="2010-11-13">10/13 XI</collectingDate>
</date>
2015&quot; [printed], &quot;
<typeStatus id="54E98386F81CFFB1FE92FE40FDA24AE9" box="[357,527,472,496]" pageId="5" pageNumber="6" type="holotype">HOLOTYPUS</typeStatus>
/
<taxonomicName id="4C5246A7F81CFFB1FDC6FE4FFD664AE9" authorityName="Jeannel" authorityYear="1922" box="[561,715,471,496]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="B926E136F81CFFB1FDC6FE4FFD664AE9" box="[561,715,471,496]" italics="true" pageId="5" pageNumber="6">Speonemadus</emphasis>
</taxonomicName>
/
<taxonomicName id="4C5246A7F81CFFB1FD1AFE4FFCEF4AE9" authority="Fresneda &amp; Faille &amp; Fery &amp; Ribera, 2019" authorityName="Fresneda &amp; Faille &amp; Fery &amp; Ribera" authorityYear="2019" box="[749,834,471,496]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="brusteli" status="sp. nov.">
<emphasis id="B926E136F81CFFB1FD1AFE4FFCEF4AE9" box="[749,834,471,496]" italics="true" pageId="5" pageNumber="6">brusteli</emphasis>
</taxonomicName>
<emphasis id="B926E136F81CFFB1FCB8FE4FFC384AE9" bold="true" box="[847,917,471,496]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A2155C4DF81CFFB1FCB8FE4FFC384AE9" box="[847,917,471,496]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
/ JF-AF-HF-IR det. 2016&quot; [red label, hw]; aedeagus mounted in
<collectingCountry id="F3457DB4F81CFFB1FE7AFE64FE4C490C" box="[397,481,508,533]" name="Canada" pageId="5" pageNumber="6">Canada</collectingCountry>
balsam on a transparent microslide pinned with the specimen (MNHN).
<materialsCitation id="3B3A3779F81CFFB1FAEBFE64FC794966" ID-GBIF-Occurrence="2147524151" collectionCode="CAF, CFL, CHB, CHF, IBE" country="Canada" pageId="5" pageNumber="6" specimenCount="5" typeStatus="paratype">
<emphasis id="B926E136F81CFFB1FAEBFE64FA3E490C" bold="true" box="[1308,1427,508,533]" pageId="5" pageNumber="6">
<typeStatus id="54E98386F81CFFB1FAEBFE64FA3E490C" box="[1308,1427,508,533]" pageId="5" pageNumber="6" type="paratype">Paratypes</typeStatus>
</emphasis>
: 15 exs, same labels as
<typeStatus id="54E98386F81CFFB1FE64FD87FE554921" box="[403,504,543,568]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
, except &quot;
<typeStatus id="54E98386F81CFFB1FDACFDB8FD524921" box="[603,767,544,568]" pageId="5" pageNumber="6" type="paratype">PARATYPUS</typeStatus>
&quot; instead of &quot;
<typeStatus id="54E98386F81CFFB1FC7CFDB8FB9A4921" box="[907,1079,544,568]" pageId="5" pageNumber="6" type="holotype">HOLOTYPUS</typeStatus>
&quot; (
<collectionCode id="ED43A5E1F81CFFB1FBB9FDB8FB2A4921" box="[1102,1159,544,568]" country="China" lsid="urn:lsid:biocol.org:col:12802" name="Chinese Academy of Forestry" pageId="5" pageNumber="6" type="Herbarium">CAF</collectionCode>
,
<collectionCode id="ED43A5E1F81CFFB1FB66FDB8FB6A4921" box="[1169,1223,544,568]" pageId="5" pageNumber="6">CFL</collectionCode>
,
<collectionCode id="ED43A5E1F81CFFB1FB25FDB8FAA34921" box="[1234,1294,544,568]" pageId="5" pageNumber="6">CHB</collectionCode>
,
<collectionCode id="ED43A5E1F81CFFB1FAEDFDB8FAF94921" box="[1306,1364,544,568]" pageId="5" pageNumber="6">CHF</collectionCode>
;
<collectionCode id="ED43A5E1F81CFFB1FAA8FDB8FA3C4921" box="[1375,1425,544,568]" country="Spain" httpUri="http://grbio.org/cool/qyi3-sxew" name="Institut de Biologia Evolutiva, (CSIC-UPF)" pageId="5" pageNumber="6">IBE</collectionCode>
). Two
<typeStatus id="54E98386F81CFFB1FF38FDDDFE964944" box="[207,315,581,605]" pageId="5" pageNumber="6" type="paratype">paratypes</typeStatus>
were used for DNA extraction and sequencing, with voucher numbers IBE-AN489 and IBE-AN523 respectively. Extracted specimens and DNA aliquots deposited in the
<collectionCode id="ED43A5E1F81CFFB1FC54FDF0FC794966" box="[931,980,616,639]" country="Spain" httpUri="http://grbio.org/cool/qyi3-sxew" name="Institut de Biologia Evolutiva, (CSIC-UPF)" pageId="5" pageNumber="6">IBE</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3486EAFF81CFFB1FF30FD14FADA4F61" pageId="5" pageNumber="6" type="description">
<paragraph id="8BED3D24F81CFFB1FF30FD14FAA84F4C" blockId="5.[151,1437,364,1685]" pageId="5" pageNumber="6">
<emphasis id="B926E136F81CFFB1FF30FD14FDD649BC" bold="true" box="[199,635,652,677]" pageId="5" pageNumber="6">
Description of the
<typeStatus id="54E98386F81CFFB1FE53FD14FDA249BC" box="[420,527,652,677]" pageId="5" pageNumber="6" type="holotype">holotype</typeStatus>
: Habitus
</emphasis>
as in
<figureCitation id="136921A1F81CFFB1FD49FD14FCAC49BD" box="[702,769,652,677]" captionStart="FIGURES 12" captionStartId="6.[151,264,1618,1640]" captionTargetBox="[183,1395,224,1571]" captionTargetId="figure@6.[151,1437,193,1597]" captionTargetPageId="6" captionText="FIGURES 12. Habitus of the holotypes of: (1) Speonemadus brusteli sp. n.; (2) S. comasi sp. n." httpUri="https://zenodo.org/record/2617609/files/figure.png" pageId="5" pageNumber="6">Fig. 1</figureCitation>
. Body length 2.3 mm. Body shape elongate; colouration of body reddish-brownish, somewhat depigmented.
<emphasis id="B926E136F81CFFB1FD4BFD37FCAE49D1" bold="true" box="[700,771,687,712]" pageId="5" pageNumber="6">Head.</emphasis>
Eyes well developed; surface with dense punctation; suture between epistome and frons visible; antennomeres VIIXI forming a lose club, antennomeres VIIIX clearly transverse.
<emphasis id="B926E136F81CFFB1FEEAFD6FFE3B4809" bold="true" box="[285,406,759,784]" pageId="5" pageNumber="6">Pronotum</emphasis>
transverse (maximum width 0.9 mm; maximum length 0.6 mm); sides more arcuate posteriad; base straight, slightly wider than base of elytra; surface with dense, rough punctation.
<emphasis id="B926E136F81CFFB1FB20FC84FA8E482C" bold="true" box="[1239,1315,796,821]" pageId="5" pageNumber="6">Elytra</emphasis>
. Elongate, broadening posteriad in first third of length, then regularly tapering towards apex; with parasutural striae. Transverse striola on elytra broad and widely spaced.
<emphasis id="B926E136F81CFFB1FCE2FCFCFCE04864" bold="true" box="[789,845,868,893]" pageId="5" pageNumber="6">Legs</emphasis>
. All tarsi pentamerous; protibiae simple; pro- and mesotibia slightly dilated; first three protarsomeres and first two mesotarsomeres dilated.
<emphasis id="B926E136F81CFFB1FB7AFC1FFA9548B9" bold="true" box="[1165,1336,903,928]" pageId="5" pageNumber="6">Male genitalia</emphasis>
(
<figureCitation id="136921A1F81CFFB1FABFFC1FFA2048B9" box="[1352,1421,903,928]" captionStart="FIGURES 56" captionStartId="8.[151,264,1938,1960]" captionTargetBox="[352,1105,457,1799]" captionTargetId="figure@8.[294,1292,411,1916]" captionTargetPageId="8" captionText="FIGURES 56. Aedeagi in ventral view of: (5) Speonemadus. orchesioides (Ammi-Moussa, Algeria); (6) holotype of S. brusteli sp. n. (Talembote, Morocco)." httpUri="https://zenodo.org/record/2617613/files/figure.png" pageId="5" pageNumber="6">Fig. 6</figureCitation>
). Aedeagus slightly curved towards ventral side; basal lamina of median lobe longer than apical part; parameres much longer than median lobe, with straight basal part, but apical third bent inwards with strong angle; apical third broader in middle region, tapering towards apex, here provided with a triangular expansion on external side; apex of parameres with three setae, without conical spine (
<figureCitation id="136921A1F81CFFB1FD05FB8FFCEB4F29" box="[754,838,1047,1072]" captionStart="FIGURES 1328" captionStartId="12.[151,264,1731,1753]" captionTargetBox="[170,1387,215,1697]" captionTargetId="figure@12.[151,1436,193,1710]" captionTargetPageId="12" captionText="FIGURES 1328. Apex of the parameres of: (13) Speonemadus orchesioides (Ammi-Moussa, Algeria); (14) holotype of S. brusteli sp. n. (Talembote, Morocco); (15) S. gracilis (Hornos, Spain); (16) S. tenuipes (Ifri Semedane, Algeria); (17) S. vandalitiae (Villanúa, Spain); (18) S. pulchellus (Ficuzza, Sicily); (19) holotype of S. algarvensis (Gruta do Vale Telheiro, Portugal); (20) S. angusticollis (Rute, Spain); (21) S. bolivari (Ardales, Spain); (22) S. breuili (Motillas, Spain); (23) S. escalerai (Jumilla, Spain); (24) S. transversostriatus (La Bastida, Spain); (25) S. clathratus (Linarejos, Spain); (26) S. maroccanus (Taza, Morocco); (27) holotype of S. comasi sp. n. (Tagelft, Morocco); (28) Anemadus hajeki (Yunnan, China)." httpUri="https://zenodo.org/record/2617619/files/figure.png" pageId="5" pageNumber="6">Fig. 14</figureCitation>
). Median lobe in dorsal view with broad, short apex. Endophallus with two series of sclerotised denticles, not joined in apical part, forming an apical dent.
</paragraph>
<paragraph id="8BED3D24F81CFFB1FF30FBC7FADA4F61" blockId="5.[151,1437,364,1685]" box="[199,1399,1119,1144]" pageId="5" pageNumber="6">
<emphasis id="B926E136F81CFFB1FF30FBC7FE844F61" bold="true" box="[199,297,1119,1144]" pageId="5" pageNumber="6">Females</emphasis>
similar to males except for pro- and mesotibiae, which are thinner; pro- and mesotarsi not dilated.
</paragraph>
</subSubSection>
<subSubSection id="C3486EAFF81CFFB1FF30FB1CFBA94FD9" pageId="5" pageNumber="6" type="etymology">
<paragraph id="8BED3D24F81CFFB1FF30FB1CFBA94FD9" blockId="5.[151,1437,364,1685]" pageId="5" pageNumber="6">
<emphasis id="B926E136F81CFFB1FF30FB1CFEE34F84" bold="true" box="[199,334,1156,1181]" pageId="5" pageNumber="6">Etymology.</emphasis>
The species is dedicated to our friend and colleague Hervé Brustel, from l'Ecole d'Ingénieurs de PURPAN (
<collectingCountry id="F3457DB4F81CFFB1FEE6FB30FECF4FD9" box="[273,354,1192,1216]" name="France" pageId="5" pageNumber="6">France</collectingCountry>
, Toulouse). The species name is a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3486EAFF81CFFB1FF30FB54FAF24D69" pageId="5" pageNumber="6" type="discussion">
<paragraph id="8BED3D24F81CFFB1FF30FB54FAF24D69" blockId="5.[151,1437,364,1685]" pageId="5" pageNumber="6">
<emphasis id="B926E136F81CFFB1FF30FB54FE994FFC" bold="true" box="[199,308,1228,1253]" pageId="5" pageNumber="6">Remarks</emphasis>
. According to the morphology and in particular to the aedeagus,
<taxonomicName id="4C5246A7F81CFFB1FBC1FB54FA9D4FFC" authority="Fresneda &amp; Faille &amp; Fery &amp; Ribera, 2019" authorityName="Fresneda &amp; Faille &amp; Fery &amp; Ribera" authorityYear="2019" box="[1078,1328,1228,1253]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="brusteli" status="sp. nov.">
<emphasis id="B926E136F81CFFB1FBC1FB54FA9D4FFC" box="[1078,1328,1228,1253]" italics="true" pageId="5" pageNumber="6">Speonemadus brusteli</emphasis>
</taxonomicName>
<emphasis id="B926E136F81CFFB1FACBFB54FAD24FFC" bold="true" box="[1340,1407,1228,1253]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A2155C4DF81CFFB1FACBFB54FAD24FFC" box="[1340,1407,1228,1253]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is included in the
<taxonomicName id="4C5246A7F81CFFB1FEB3FB68FE734E11" authorityName="" baseAuthorityName="Heyden" baseAuthorityYear="1870" box="[324,478,1263,1288]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="vandalitiae">
<emphasis id="B926E136F81CFFB1FEB3FB68FE734E11" box="[324,478,1263,1288]" italics="true" pageId="5" pageNumber="6">S. vandalitiae</emphasis>
</taxonomicName>
-group of
<bibRefCitation id="EFC340D5F81CFFB1FDBAFB77FCD94E11" author="Giachino, P. M. &amp; Vailati, D." box="[589,884,1263,1288]" pageId="5" pageNumber="6" refId="ref13309" refString="Giachino, P. M. &amp; Vailati, D. (1993) Revisione degli Anemadinae. Monografie di &quot; Natura Bresciana &quot; No. 18. Museo Civico di Scienze Naturali di Brescia, Brescia, 314 pp." type="book" year="1993">Giachino &amp; Vailati (1993)</bibRefCitation>
, near
<taxonomicName id="4C5246A7F81CFFB1FC40FB68FBF24E11" authorityName="Fairmaire" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" box="[951,1119,1263,1288]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F81CFFB1FC40FB68FBF24E11" box="[951,1119,1263,1288]" italics="true" pageId="5" pageNumber="6">S. orchesioides</emphasis>
</taxonomicName>
. This group is characterised by: transverse pronotum, with dense to rugose punctation; transverse striola on the elytra either fine and dense or thick and sparse; protibiae simple; sutural angle of elytra rounded in both sexes; parameres much longer than median lobe, with 35 apical setae and without conical spine (
<figureCitation id="136921A1F81CFFB1FC98FAC4FB8C4E6C" box="[879,1057,1372,1397]" captionStart="FIGURES 1328" captionStartId="12.[151,264,1731,1753]" captionTargetBox="[170,1387,215,1697]" captionTargetId="figure@12.[151,1436,193,1710]" captionTargetPageId="12" captionText="FIGURES 1328. Apex of the parameres of: (13) Speonemadus orchesioides (Ammi-Moussa, Algeria); (14) holotype of S. brusteli sp. n. (Talembote, Morocco); (15) S. gracilis (Hornos, Spain); (16) S. tenuipes (Ifri Semedane, Algeria); (17) S. vandalitiae (Villanúa, Spain); (18) S. pulchellus (Ficuzza, Sicily); (19) holotype of S. algarvensis (Gruta do Vale Telheiro, Portugal); (20) S. angusticollis (Rute, Spain); (21) S. bolivari (Ardales, Spain); (22) S. breuili (Motillas, Spain); (23) S. escalerai (Jumilla, Spain); (24) S. transversostriatus (La Bastida, Spain); (25) S. clathratus (Linarejos, Spain); (26) S. maroccanus (Taza, Morocco); (27) holotype of S. comasi sp. n. (Tagelft, Morocco); (28) Anemadus hajeki (Yunnan, China)." httpUri="https://zenodo.org/record/2617619/files/figure.png" pageId="5" pageNumber="6">Figs 13, 1518</figureCitation>
); endophallus with two series of sclerotised denticles, without apical indentation. The species of the group are externally very homogeneous, and can only be separated by the study of the aedeagus (
<bibRefCitation id="EFC340D5F81CFFB1FCE6FA3CFB944EA5" author="Giachino, P. M. &amp; Vailati, D." box="[785,1081,1444,1469]" pageId="5" pageNumber="6" refId="ref13309" refString="Giachino, P. M. &amp; Vailati, D. (1993) Revisione degli Anemadinae. Monografie di &quot; Natura Bresciana &quot; No. 18. Museo Civico di Scienze Naturali di Brescia, Brescia, 314 pp." type="book" year="1993">Giachino &amp; Vailati 1993</bibRefCitation>
). The general structure of the aedeagus is similar to that of
<taxonomicName id="4C5246A7F81CFFB1FE16FA50FD274EF9" authorityName="Fairmaire" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" box="[481,650,1479,1504]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F81CFFB1FE16FA50FD274EF9" box="[481,650,1479,1504]" italics="true" pageId="5" pageNumber="6">S. orchesioides</emphasis>
</taxonomicName>
(
<figureCitation id="136921A1F81CFFB1FD6EFA5FFD704EF9" box="[665,733,1479,1504]" captionStart="FIGURES 56" captionStartId="8.[151,264,1938,1960]" captionTargetBox="[352,1105,457,1799]" captionTargetId="figure@8.[294,1292,411,1916]" captionTargetPageId="8" captionText="FIGURES 56. Aedeagi in ventral view of: (5) Speonemadus. orchesioides (Ammi-Moussa, Algeria); (6) holotype of S. brusteli sp. n. (Talembote, Morocco)." httpUri="https://zenodo.org/record/2617613/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
). However, the dilated apex of the parameres with a triangular expansion on the external side clearly separates
<taxonomicName id="4C5246A7F81CFFB1FD37FA75FC9E4D1C" authority="Fresneda &amp; Faille &amp; Fery &amp; Ribera, 2019" authorityName="Fresneda &amp; Faille &amp; Fery &amp; Ribera" authorityYear="2019" box="[704,819,1516,1541]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="brusteli" status="sp. nov.">
<emphasis id="B926E136F81CFFB1FD37FA75FC9E4D1C" box="[704,819,1516,1541]" italics="true" pageId="5" pageNumber="6">S. brusteli</emphasis>
</taxonomicName>
<emphasis id="B926E136F81CFFB1FCCAFA74FCD24D1C" bold="true" box="[829,895,1516,1541]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A2155C4DF81CFFB1FCCAFA74FCD24D1C" box="[829,895,1516,1541]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
from this species, which has simple parameres (
<figureCitation id="136921A1F81CFFB1FF68F997FF5E4D31" box="[159,243,1551,1576]" captionStart="FIGURES 1328" captionStartId="12.[151,264,1731,1753]" captionTargetBox="[170,1387,215,1697]" captionTargetId="figure@12.[151,1436,193,1710]" captionTargetPageId="12" captionText="FIGURES 1328. Apex of the parameres of: (13) Speonemadus orchesioides (Ammi-Moussa, Algeria); (14) holotype of S. brusteli sp. n. (Talembote, Morocco); (15) S. gracilis (Hornos, Spain); (16) S. tenuipes (Ifri Semedane, Algeria); (17) S. vandalitiae (Villanúa, Spain); (18) S. pulchellus (Ficuzza, Sicily); (19) holotype of S. algarvensis (Gruta do Vale Telheiro, Portugal); (20) S. angusticollis (Rute, Spain); (21) S. bolivari (Ardales, Spain); (22) S. breuili (Motillas, Spain); (23) S. escalerai (Jumilla, Spain); (24) S. transversostriatus (La Bastida, Spain); (25) S. clathratus (Linarejos, Spain); (26) S. maroccanus (Taza, Morocco); (27) holotype of S. comasi sp. n. (Tagelft, Morocco); (28) Anemadus hajeki (Yunnan, China)." httpUri="https://zenodo.org/record/2617619/files/figure.png" pageId="5" pageNumber="6">Fig. 13</figureCitation>
). The expansion on the apex of the median lobe is also regularly arched in
<taxonomicName id="4C5246A7F81CFFB1FBB8F988FB6C4D31" authority="Fresneda &amp; Faille &amp; Fery &amp; Ribera, 2019" authorityName="Fresneda &amp; Faille &amp; Fery &amp; Ribera" authorityYear="2019" box="[1103,1217,1551,1576]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="brusteli" status="sp. nov.">
<emphasis id="B926E136F81CFFB1FBB8F988FB6C4D31" box="[1103,1217,1551,1576]" italics="true" pageId="5" pageNumber="6">S. brusteli</emphasis>
</taxonomicName>
<emphasis id="B926E136F81CFFB1FB3CF997FAA64D31" bold="true" box="[1227,1291,1551,1576]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A2155C4DF81CFFB1FB3CF997FAA64D31" box="[1227,1291,1551,1576]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="136921A1F81CFFB1FAEBF997FACF4D31" box="[1308,1378,1551,1576]" captionStart="FIGURES 56" captionStartId="8.[151,264,1938,1960]" captionTargetBox="[352,1105,457,1799]" captionTargetId="figure@8.[294,1292,411,1916]" captionTargetPageId="8" captionText="FIGURES 56. Aedeagi in ventral view of: (5) Speonemadus. orchesioides (Ammi-Moussa, Algeria); (6) holotype of S. brusteli sp. n. (Talembote, Morocco)." httpUri="https://zenodo.org/record/2617613/files/figure.png" pageId="5" pageNumber="6">Fig. 6</figureCitation>
), but roughly pentagonal in
<taxonomicName id="4C5246A7F81CFFB1FE6EF9ADFDEE4D54" authorityName="Fairmaire" baseAuthorityName="Fairmaire" baseAuthorityYear="1879" box="[409,579,1588,1613]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="orchesioides">
<emphasis id="B926E136F81CFFB1FE6EF9ADFDEE4D54" box="[409,579,1588,1613]" italics="true" pageId="5" pageNumber="6">S. orchesioides</emphasis>
</taxonomicName>
(
<figureCitation id="136921A1F81CFFB1FDA3F9ACFD374D54" box="[596,666,1588,1613]" captionStart="FIGURES 56" captionStartId="8.[151,264,1938,1960]" captionTargetBox="[352,1105,457,1799]" captionTargetId="figure@8.[294,1292,411,1916]" captionTargetPageId="8" captionText="FIGURES 56. Aedeagi in ventral view of: (5) Speonemadus. orchesioides (Ammi-Moussa, Algeria); (6) holotype of S. brusteli sp. n. (Talembote, Morocco)." httpUri="https://zenodo.org/record/2617613/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
). Molecular data place
<taxonomicName id="4C5246A7F81CFFB1FC5FF9ADFBB74D54" authority="Fresneda &amp; Faille &amp; Fery &amp; Ribera, 2019" authorityName="Fresneda &amp; Faille &amp; Fery &amp; Ribera" authorityYear="2019" box="[936,1050,1588,1613]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="brusteli" status="sp. nov.">
<emphasis id="B926E136F81CFFB1FC5FF9ADFBB74D54" box="[936,1050,1588,1613]" italics="true" pageId="5" pageNumber="6">S. brusteli</emphasis>
</taxonomicName>
<emphasis id="B926E136F81CFFB1FBD3F9ACFBC84D54" bold="true" box="[1060,1125,1588,1613]" pageId="5" pageNumber="6">
<taxonomicNameLabel id="A2155C4DF81CFFB1FBD3F9ACFBC84D54" box="[1060,1125,1588,1613]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
as sister to
<taxonomicName id="4C5246A7F81CFFB1FB18F9ADFA384D55" baseAuthorityName="Jeannel" baseAuthorityYear="1936" box="[1263,1429,1589,1612]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="maroccanus">
<emphasis id="B926E136F81CFFB1FB18F9ADFA384D55" box="[1263,1429,1589,1612]" italics="true" pageId="5" pageNumber="6">S. maroccanus</emphasis>
</taxonomicName>
, with a considerably different aedeagus and included in the
<taxonomicName id="4C5246A7F81CFFB1FCDDF9CFFC364D69" baseAuthorityName="Perris" baseAuthorityYear="1864" box="[810,923,1623,1648]" class="Insecta" family="Leiodidae" genus="Speonemadus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="clathratus">
<emphasis id="B926E136F81CFFB1FCDDF9CFFC364D69" box="[810,923,1623,1648]" italics="true" pageId="5" pageNumber="6">clathratus</emphasis>
</taxonomicName>
-group
<emphasis id="B926E136F81CFFB1FC1AF9C1FB864D69" box="[1005,1067,1625,1648]" italics="true" pageId="5" pageNumber="6">sensu</emphasis>
<bibRefCitation id="EFC340D5F81CFFB1FBC4F9CFFAF14D69" author="Giachino, P. M. &amp; Vailati, D." box="[1075,1372,1623,1648]" pageId="5" pageNumber="6" refId="ref13309" refString="Giachino, P. M. &amp; Vailati, D. (1993) Revisione degli Anemadinae. Monografie di &quot; Natura Bresciana &quot; No. 18. Museo Civico di Scienze Naturali di Brescia, Brescia, 314 pp." type="book" year="1993">Giachino &amp; Vailati (1993)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3486EAFF81CFFB1FF30F9E4FB654D8D" box="[199,1224,1660,1685]" pageId="5" pageNumber="6" type="distribution">
<paragraph id="8BED3D24F81CFFB1FF30F9E4FB654D8D" blockId="5.[151,1437,364,1685]" box="[199,1224,1660,1685]" pageId="5" pageNumber="6">
<emphasis id="B926E136F81CFFB1FF30F9E4FEF44D8C" bold="true" box="[199,345,1660,1685]" pageId="5" pageNumber="6">Distribution</emphasis>
(
<figureCitation id="136921A1F81CFFB1FE9FF9E4FE114D8D" box="[360,444,1660,1685]" captionStart="FIGURE 38" captionStartId="21.[151,250,1234,1256]" captionTargetBox="[259,1364,471,1140]" captionTargetId="figure@21.[205,1395,421,1189]" captionTargetPageId="21" captionText="FIGURE 38. Distribution maps of: (38) Speonemadus brusteli sp. n., S. comasi sp. n., S. orchesioides, S. pulchellus, S. tenuipes (empty circles are dubious records of which no material could be studied; the records from continental Italy, given originally as S. orchesoides, are considered S. pulchellus, P.M. Giachino, personal communication 2018; see main text). See Appendix for detailed locality data." httpUri="https://zenodo.org/record/2617639/files/figure.png" pageId="5" pageNumber="6">Fig. 38</figureCitation>
). So far only known from the
<typeStatus id="54E98386F81CFFB1FCFAF9E5FC904D8C" box="[781,829,1661,1685]" pageId="5" pageNumber="6">type</typeStatus>
locality in the Rif chain,
<collectingCountry id="F3457DB4F81CFFB1FBACF9E5FB694D8D" box="[1115,1220,1661,1684]" name="Morocco" pageId="5" pageNumber="6">Morocco</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>